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<mods:title id="95F2785C907186835679442091C18A9B">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<treatment id="03B787C7FFA47656FEB6D27A4F0DF5C1" ID-DOI="http://doi.org/10.5281/zenodo.6967994" ID-GBIF-Taxon="198618694" ID-Zenodo-Dep="6967994" LSID="urn:lsid:plazi:treatment:03B787C7FFA47656FEB6D27A4F0DF5C1" httpUri="http://treatment.plazi.org/id/03B787C7FFA47656FEB6D27A4F0DF5C1" lastPageId="123" lastPageNumber="540" pageId="121" pageNumber="538">
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<paragraph id="8BA136D1FFA47654FEB6D27A48A4F321" blockId="121.[285,656,1570,1595]" box="[285,656,1570,1595]" pageId="121" pageNumber="538">
<heading id="D0E981BDFFA47654FEB6D27A48A4F321" box="[285,656,1570,1595]" centered="true" fontSize="9" level="2" pageId="121" pageNumber="538" reason="2">
<taxonomicName id="4C1E4D52FFA47654FEB6D27A48A4F321" ID-CoL="992LJ" authority="(OI, 1977)" baseAuthorityName="Oi" baseAuthorityYear="1977" box="[285,656,1570,1595]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FEB6D27A4823F320" box="[285,535,1570,1594]" italics="true" pageId="121" pageNumber="538">ATYPENA FORMOSANA</emphasis>
(
<bibRefCitation id="EF8F4B20FFA47654FD8DD27A48BCF321" author="Oi R" box="[550,648,1570,1595]" pageId="121" pageNumber="538" pagination="23 - 26" refId="ref93859" refString="Oi R. 1977. A new erigonid spider from Formosa. Acta Arachnologica 27: 23 - 26." type="journal article" year="1977">OI, 1977</bibRefCitation>
)
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<subSubSection id="C304655AFFA47654FEF1D2114867F378" box="[346,595,1609,1634]" pageId="121" pageNumber="538" type="description">
<paragraph id="8BA136D1FFA47654FEF1D2114867F378" blockId="121.[346,595,1609,1634]" box="[346,595,1609,1634]" pageId="121" pageNumber="538">
(
<figureCitation id="13252A54FFA47654FEC8D2114B8AF37A" box="[355,446,1609,1634]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, Oedothorax paracymbialis Tanasevitch, 2015 (traced from photograph). D, Oe. kodaikanal Tanasevitch, 2015 (traced from photograph). E, Oe. cunur Tanasevitch, 2015 (traced from photograph). F, Oe. stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, Oe. nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="121" pageNumber="538">FIGS 7H</figureCitation>
,
<figureCitation id="13252A54FFA47654FE60D2114BC4F37A" box="[459,496,1609,1634]" captionStart="Figure 67" captionStartId="110.[144,224,1301,1323]" captionTargetBox="[147,1426,197,1260]" captionTargetId="figure-175@110.[145,1428,195,1261]" captionTargetPageId="110" captionText="Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967988" httpUri="https://zenodo.org/record/6967988/files/figure.png" pageId="121" pageNumber="538">67I</figureCitation>
,
<figureCitation id="13252A54FFA47654FE56D2114815F37A" box="[509,545,1609,1634]" captionStart="Figure 68" captionStartId="111.[163,245,1308,1330]" captionTargetBox="[164,1444,196,1268]" captionTargetId="figure-26@111.[163,1446,195,1269]" captionTargetPageId="111" captionText="Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967990" httpUri="https://zenodo.org/record/6967990/files/figure.png" pageId="121" pageNumber="538">68I</figureCitation>
,
<figureCitation id="13252A54FFA47654FD84D211487FF378" box="[559,587,1609,1634]" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="121" pageNumber="538">73</figureCitation>
)
</paragraph>
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<paragraph id="8BA136D1FFA47654FF08D22948DDF39D" blockId="121.[163,780,1649,1886]" box="[163,745,1649,1671]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FF08D22948DDF39D" box="[163,745,1649,1671]" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FF08D2294801F39C" author="Oi R" box="[163,565,1649,1671]" page="23" pageId="121" pageNumber="538" year="1977">
<taxonomicName id="4C1E4D52FFA47654FF08D2294801F39C" ID-CoL="5X826" authority="Oi, 1977: 23" authorityName="Oi" authorityPageNumber="23" authorityYear="1977" box="[163,565,1649,1671]" class="Arachnida" family="Linyphiidae" genus="Callitrichia" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FF08D2294B92F39C" box="[163,422,1649,1670]" italics="true" pageId="121" pageNumber="538">Callitrichia formosana</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FE06D2294801F39C" author="Oi R" box="[429,565,1649,1671]" pageId="121" pageNumber="538" pagination="23 - 26" refId="ref93859" refString="Oi R. 1977. A new erigonid spider from Formosa. Acta Arachnologica 27: 23 - 26." type="journal article" year="1977">Oi, 1977: 23</bibRefCitation>
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, figs 15 (Dmf).
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</paragraph>
<paragraph id="8BA136D1FFA47654FF08D2C848A1F3BF" blockId="121.[163,780,1649,1886]" box="[163,661,1680,1702]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FF08D2C848A1F3BF" box="[163,661,1680,1702]" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FF08D2C848A5F3BF" author="Brignoli PM" box="[163,657,1680,1702]" page="349" pageId="121" pageNumber="538" year="1983">
<taxonomicName id="4C1E4D52FFA47654FF08D2C848A5F3BF" authority="Brignoli, 1983: 349" authorityName="Brignoli" authorityPageNumber="349" authorityYear="1983" box="[163,657,1680,1702]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B96AEAC3FFA47654FF08D2C84B85F3BF" box="[163,433,1680,1701]" italics="true" pageId="121" pageNumber="538">Oedothorax formosanus</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FE1CD2C848A5F3BF" author="Brignoli PM" box="[439,657,1680,1702]" pageId="121" pageNumber="538" refId="ref89503" refString="Brignoli PM. 1983. A catalogue of the Araneae described between 1940 and 1981. Manchester: Manchester University Press." type="book" year="1983">Brignoli, 1983: 349</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BA136D1FFA47654FF08D2F64BA6F3F9" blockId="121.[163,780,1649,1886]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FF08D2F64BA6F3F9" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FF08D2F6485EF3DE" author="Song DX" box="[163,618,1710,1732]" page="144" pageId="121" pageNumber="538" year="1987">
<taxonomicName id="4C1E4D52FFA47654FF08D2F6485EF3DE" ID-CoL="5X826" authority="Song, 1987: 144" authorityName="Song" authorityPageNumber="144" authorityYear="1987" box="[163,618,1710,1732]" class="Arachnida" family="Linyphiidae" genus="Callitrichia" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FF08D2F64B98F3DE" box="[163,428,1710,1732]" italics="true" pageId="121" pageNumber="538">Callitrichia formosana</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FE18D2F7485EF3DE" author="Song DX" box="[435,618,1711,1732]" pageId="121" pageNumber="538" refId="ref95430" refString="Song DX. 1987. Spiders from agricultural regions of China (Arachnida: Araneae). Beijing: Agriculture Publishing House." type="book" year="1987">Song, 1987: 144</bibRefCitation>
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, fig. 104 (Tmf from
<taxonomicName id="4C1E4D52FFA47654FF54D2954BB6F3F8" ID-CoL="6792" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[255,386,1741,1762]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B96AEAC3FFA47654FF54D2954BB6F3F8" box="[255,386,1741,1762]" italics="true" pageId="121" pageNumber="538">Oedothorax</emphasis>
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).
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</paragraph>
<paragraph id="8BA136D1FFA47654FF08D2B448EFF23A" blockId="121.[163,780,1649,1886]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FF08D2B448EFF23A" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FF08D2B4486EF21B" author="Tazoe S" box="[163,602,1772,1793]" page="212" pageId="121" pageNumber="538" year="1992">
<taxonomicName id="4C1E4D52FFA47654FF08D2B4486EF21B" ID-CoL="992LJ" authority="Tazoe, 1992: 212" authorityName="Tazoe" authorityPageNumber="212" authorityYear="1992" box="[163,602,1772,1793]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FF08D2B44BBDF21B" box="[163,393,1772,1793]" italics="true" pageId="121" pageNumber="538">Atypena formosana</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FE3AD2B4486EF21B" author="Tazoe S" box="[401,602,1772,1793]" pageId="121" pageNumber="538" pagination="211 - 214" refId="ref96185" refString="Tazoe S. 1992. A newly recorded spider from Japan, Atypena formosana (Oi, 1977), comb. nov. (Linyphiidae). Acta Arachnologica 41: 211 - 214." type="journal article" year="1992">Tazoe, 1992: 212</bibRefCitation>
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, figs 110 (mf; NB: generic placement following
<bibRefCitation id="EF8F4B20FFA47654FD91D35348FFF23A" author="Jocque R" box="[570,715,1803,1825]" pageId="121" pageNumber="538" pagination="235 - 245" refId="ref92149" refString="Jocque R. 1983. Sur la synonymie de Callitrichia Fage et Atypena Simon avec la redescription de quelques especes paleotropicales (Araneae, Linyphiidae). Bulletin du Museum National d'Histoire Naturelle de Paris 5: 235 - 245." type="journal article" year="1983">Jocqué, 1983</bibRefCitation>
).
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</paragraph>
<paragraph id="8BA136D1FFA47654FF08D3714B02F244" blockId="121.[163,780,1649,1886]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FF08D3714B02F244" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FF08D37148EDF224" author="Okuma C &amp; Kamal NQ &amp; Hirashima Y &amp; Alam MZ &amp; Ogata K" box="[163,729,1833,1855]" page="13" pageId="121" pageNumber="538" year="1993">
<taxonomicName id="4C1E4D52FFA47654FF08D37148EDF224" authority="Okuma et al., 1993: 13" authorityName="Okuma" authorityPageNumber="13" authorityYear="1993" box="[163,729,1833,1855]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B96AEAC3FFA47654FF08D3714B89F224" box="[163,445,1833,1854]" italics="true" pageId="121" pageNumber="538">Oedothorax formosanus</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FE6CD37148EDF224" author="Okuma C &amp; Kamal NQ &amp; Hirashima Y &amp; Alam MZ &amp; Ogata K" box="[455,729,1833,1855]" pageId="121" pageNumber="538" refId="ref93879" refString="Okuma C, Kamal NQ, Hirashima Y, Alam MZ, Ogata K. 1993. Illustrated Monograph of the Rice Field Spiders of Bangladesh. Institute of Postgraduate Studies in Agriculture (Salna, Gazipur, Bangladesh) (Vol. 1). Japan International Cooperation Agency Project Publication." type="book" year="1993">
Okuma
<emphasis id="B96AEAC3FFA47654FD8CD3724857F224" box="[551,611,1833,1855]" italics="true" pageId="121" pageNumber="538">et al.</emphasis>
, 1993: 13
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</taxonomicName>
</treatmentCitation>
, fig. 11AB (f).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D49D4E76F5E3" blockId="121.[827,1442,197,372]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FC90D49D4E76F5E3" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FC90D49D4FAAF5C0" author="Barrion AT &amp; Litsinger JA" box="[827,1438,197,219]" page="319" pageId="121" pageNumber="538" year="1994">
<taxonomicName id="4C1E4D52FFA47654FC90D49D4FAAF5C0" ID-CoL="992LJ" authority="Barrion &amp; Litsinger, 1994: 319" authorityName="Barrion &amp; Litsinger" authorityPageNumber="319" authorityYear="1994" box="[827,1438,197,219]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FC90D49D4E15F5C0" box="[827,1057,197,218]" italics="true" pageId="121" pageNumber="538">Atypena formosana</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FB80D49D4FAAF5C0" author="Barrion AT &amp; Litsinger JA" box="[1067,1438,197,219]" pageId="121" pageNumber="538" pagination="283 - 359" refId="ref88774" refString="Barrion AT, Litsinger JA. 1994. Taxonomy of rice insect pests and their arthropod parasites and predators. In: Heinrichs EA, ed. Biology and management of rice insects. New Delhi: Wiley Eastern, 13 - 15, 283 - 359." type="journal article" year="1994">Barrion &amp; Litsinger, 1994: 319</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 16731676 (mf).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D55A49D0F42D" blockId="121.[827,1442,197,372]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FC90D55A49D0F42D" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FC90D55A4F59F402" author="Song DX &amp; Zhu MS &amp; Chen J" box="[827,1389,258,280]" page="160" pageId="121" pageNumber="538" year="1999">
<taxonomicName id="4C1E4D52FFA47654FC90D55A4F59F402" ID-CoL="5X826" authority="Song et al., 1999: 160" authorityName="Song" authorityPageNumber="160" authorityYear="1999" box="[827,1389,258,280]" class="Arachnida" family="Linyphiidae" genus="Callitrichia" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FC90D55A4E61F40D" box="[827,1109,258,279]" italics="true" pageId="121" pageNumber="538">Callitrichia formosana</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FBF4D55B4F59F402" author="Song DX &amp; Zhu MS &amp; Chen J" box="[1119,1389,258,280]" pageId="121" pageNumber="538" refId="ref95453" refString="Song DX, Zhu MS, Chen J. 1999. The spiders of China. Shijiazhuang: Hebei University of Science and Techology Publishing House." type="book" year="1999">
Song
<emphasis id="B96AEAC3FFA47654FB0ED55B4ED0F40D" box="[1189,1252,258,280]" italics="true" pageId="121" pageNumber="538">et al.</emphasis>
, 1999: 160
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 88KL (mf).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D51849A5F46E" blockId="121.[827,1442,197,372]" pageId="121" pageNumber="538">
<treatmentCitationGroup id="AB0E11FFFFA47654FC90D51849A5F46E" pageId="121" pageNumber="538">
<treatmentCitation id="0ABF10C0FFA47654FC90D5184F12F44C" author="Ono H &amp; Matsuda M &amp; Saito H" box="[827,1318,320,342]" page="267" pageId="121" pageNumber="538" year="2009">
<taxonomicName id="4C1E4D52FFA47654FC90D5184F12F44C" ID-CoL="5X826" authority="Ono et al., 2009: 267" authorityName="Ono" authorityPageNumber="267" authorityYear="2009" box="[827,1318,320,342]" class="Arachnida" family="Linyphiidae" genus="Callitrichia" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA47654FC90D5184E09F44F" box="[827,1085,320,341]" italics="true" pageId="121" pageNumber="538">Callitrichia formosana</emphasis>
<bibRefCitation id="EF8F4B20FFA47654FBE9D5184F12F44C" author="Ono H &amp; Matsuda M &amp; Saito H" box="[1090,1318,320,342]" pageId="121" pageNumber="538" pagination="253 - 344" refId="ref93932" refString="Ono H, Matsuda M, Saito H. 2009. Linyphiidae, Pimoidae. In: Ono H, ed. The spiders of Japan with keys to the families and genera and illustrations of the species. Kanagawa: Tokai University Press, 253 - 344." type="book chapter" year="2009">
Ono
<emphasis id="B96AEAC3FFA47654FBDCD5184E99F44F" box="[1143,1197,320,341]" italics="true" pageId="121" pageNumber="538">et al.</emphasis>
, 2009: 267
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 9599 (mf).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA47656FC90D5C54F0DF5C1" lastPageId="123" lastPageNumber="540" pageId="121" pageNumber="538" type="materials_examined">
<paragraph id="8BA136D1FFA47654FC90D5C54EB0F4EA" blockId="121.[827,1443,413,496]" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D5C54E13F4A8" box="[827,1063,413,434]" italics="true" pageId="121" pageNumber="538">Examined material:</emphasis>
<materialsCitation id="3B763C8CFFA47654FB9ED5C54EB4F4EA" ID-GBIF-Occurrence="3880945427" accessionNumber="SMF 56485" collectingDate="1995-04-02" collectionCode="SMF" collectorName="A. Tanikawa &amp; Tanasevitch" country="Japan" determinerName="A. Tanikawa" location="Urauchi" municipality="Iromotejima Island" pageId="121" pageNumber="538" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Okinawa">
<emphasis id="B96AEAC3FFA47654FB9ED5C54EBBF4A8" bold="true" box="[1077,1167,413,434]" pageId="121" pageNumber="538">
<collectingCountry id="F3097641FFA47654FB9ED5C54EBCF4A8" box="[1077,1160,413,434]" name="Japan" pageId="121" pageNumber="538">Japan</collectingCountry>
:
</emphasis>
<collectingRegion id="49DAF833FFA47654FB3CD5C54F37F4A9" box="[1175,1283,413,435]" country="Japan" name="Okinawa" pageId="121" pageNumber="538">Okinawa</collectingRegion>
,
<collectingMunicipality id="6BC5ACABFFA47654FABAD5C549B1F4C8" pageId="121" pageNumber="538">Iromotejima Island</collectingMunicipality>
,
<location id="8EC1600AFFA47654FC25D5E449D9F4C8" LSID="urn:lsid:plazi:treatment:03B787C7FFA47656FEB6D27A4F0DF5C1:8EC1600AFFA47654FC25D5E449D9F4C8" box="[910,1005,444,466]" country="Japan" municipality="Iromotejima Island" name="Urauchi" pageId="121" pageNumber="538" stateProvince="Okinawa">Urauchi</location>
,
<specimenCount id="9D18FD58FFA47654FC5CD5E44E21F4CB" box="[1015,1045,444,465]" pageId="121" pageNumber="538" type="male">2♂</specimenCount>
<specimenCount id="9D18FD58FFA47654FBBED5E44E06F4CB" box="[1045,1074,444,465]" pageId="121" pageNumber="538" type="female">3♀</specimenCount>
<date id="FFA01011FFA47654FB93D5E44E98F4C8" box="[1080,1196,444,466]" pageId="121" pageNumber="538" value="1995-04-02">
<collectingDate id="EFE4E9F9FFA47654FB93D5E44E98F4C8" box="[1080,1196,444,466]" pageId="121" pageNumber="538" value="1995-04-02">02.iv.1995</collectingDate>
</date>
, leg.
<collectorName id="26EB5307FFA47654FB4BD5E44F5BF4C8" box="[1248,1391,444,466]" pageId="121" pageNumber="538">
<determinerName id="7E7DDA15FFA47654FB4BD5E44F5BF4C8" box="[1248,1391,444,466]" pageId="121" pageNumber="538">A. Tanikawa</determinerName>
</collectorName>
, det. A,
<collectorName id="26EB5307FFA47654FCF3D58249D6F4EA" box="[856,994,474,496]" pageId="121" pageNumber="538">Tanasevitch</collectorName>
(
<collectionCode id="ED0FAE14FFA47654FC5AD5834E1DF4EA" box="[1009,1065,475,496]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="121" pageNumber="538">SMF</collectionCode>
56485)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D6424986F735" blockId="121.[827,946,538,559]" box="[827,946,538,559]" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D6424986F735" box="[827,946,538,559]" italics="true" pageId="121" pageNumber="538">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D6004FBDF7B1" blockId="121.[827,1441,600,683]" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D60049B9F777" box="[827,909,600,621]" italics="true" pageId="121" pageNumber="538">Males:</emphasis>
This species can be distinguished from
<taxonomicName id="4C1E4D52FFA47654FC90D62F49E1F791" baseAuthorityName="Heimer" baseAuthorityYear="1984" box="[827,981,630,652]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="species" species="cirrifrons">
<emphasis id="B96AEAC3FFA47654FC90D62F49E1F791" box="[827,981,630,652]" italics="true" pageId="121" pageNumber="538">A. cirrifrons</emphasis>
</taxonomicName>
by the proportionally shorter distal suprategular apophysis and ventral radical process.
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D68C4EDDF63D" blockId="121.[827,1443,724,807]" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D68C4996F7F3" box="[827,930,724,745]" italics="true" pageId="121" pageNumber="538">Females:</emphasis>
No study of
<taxonomicName id="4C1E4D52FFA47654FBE6D68C4E9AF7F3" baseAuthorityName="Jocque" baseAuthorityYear="1983" box="[1101,1198,724,745]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="121" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FFA47654FBE6D68C4E9AF7F3" box="[1101,1198,724,745]" italics="true" pageId="121" pageNumber="538">Atypena</emphasis>
</taxonomicName>
comparative female morphology exists to date; such diagnosis is, therefore, beyond the scope of the current study.
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D70849F7F67F" blockId="121.[827,963,848,869]" box="[827,963,848,869]" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D70849F7F67F" box="[827,963,848,869]" italics="true" pageId="121" pageNumber="538">Description:</emphasis>
</paragraph>
<paragraph id="8BA136D1FFA47654FC90D7D64F1DF3E4" blockId="121.[827,1444,910,1791]" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D7D649FBF6BE" box="[827,975,910,932]" italics="true" pageId="121" pageNumber="538">
Male (
<collectionCode id="ED0FAE14FFA47654FC2DD7D749F5F6BE" box="[902,961,911,932]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="121" pageNumber="538">SMF</collectionCode>
):
</emphasis>
Total length: 1.76. Prosoma: 0.78 long, 0.62 wide, PME region largely elevated, interocular region with translucent setae laterally curved (
<figureCitation id="13252A54FFA47654FADDD7944969F11A" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, Oedothorax paracymbialis Tanasevitch, 2015 (traced from photograph). D, Oe. kodaikanal Tanasevitch, 2015 (traced from photograph). E, Oe. cunur Tanasevitch, 2015 (traced from photograph). F, Oe. stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, Oe. nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="121" pageNumber="538">Fig. 7H</figureCitation>
). Eyes: AME-AME: 0.03, AME width: 0.06, AME- ALE: 0.05, ALE width: 0.08, ALE-PLE: 0.01, PLE width: 0.08, PLE-PME: 0.14, PME width: 0.07, PME- PME: 0.17. Clypeus: upper-half hirsute, one sub- AME seta. Sternum: 0.46 long, 0.46 wide. Chelicerae: mastidia absent; stridulatory striae imbricated, rows compressed proximally (
<figureCitation id="13252A54FFA47654FB02D0FA4F3CF1A2" box="[1193,1288,1186,1208]" captionStart="Figure 67" captionStartId="110.[144,224,1301,1323]" captionTargetBox="[147,1426,197,1260]" captionTargetId="figure-175@110.[145,1428,195,1261]" captionTargetPageId="110" captionText="Figure 67. Male chelicerae, stridulatory striae on lateral side. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967988" httpUri="https://zenodo.org/record/6967988/files/figure.png" pageId="121" pageNumber="538">Fig. 67I</figureCitation>
). Legs: tibia chaetotaxy 2-2-1-1, dorsal proximal macroseta on tibia I, II, III and IV 0.65, 0.82, 1.45 and 1.87 times diameter of tibia, respectively; Tm I: 0.78. All metatarsi with trichobothrium. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, two retrolateral trichobothria; TPA short, pointed and sclerotized at tip; PC base not visible from dorsal view, distal setae close to distal clasp, terminal part slightly enlarged, distal clasp without striae, clasp extended apically (
<figureCitation id="13252A54FFA47654FC04D18D4E23F0F0" box="[943,1047,1493,1515]" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="121" pageNumber="538">Fig. 73A</figureCitation>
); T without papillae; PT small, without papillae; TS long and slender, without papillae (
<figureCitation id="13252A54FFA47654FCE8D24A4993F33D" box="[835,935,1554,1576]" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="121" pageNumber="538">Fig. 73D</figureCitation>
); MSA absent; DSA wide, truncated at tip, ventral corner more extended than dorsal corner (
<figureCitation id="13252A54FFA47654FAD3D269495DF37F" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="121" pageNumber="538">Fig. 73A</figureCitation>
); EM broad and flat, without papillae, exceeding ARP; ARP flat, unsclerotized; LER retrolateral to E; VRP long, slender; R without papillae; TP long, slender; E short and stout, slightly retrolaterally curved (
<figureCitation id="13252A54FFA47654FAD3D2F4495DF3C5" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="121" pageNumber="538">Fig. 73B</figureCitation>
). Opisthosoma: dorsal pattern see
<figureCitation id="13252A54FFA47654FAA8D2924F68F3FA" box="[1283,1372,1738,1760]" captionStart="Figure 68" captionStartId="111.[163,245,1308,1330]" captionTargetBox="[164,1444,196,1268]" captionTargetId="figure-26@111.[163,1446,195,1269]" captionTargetPageId="111" captionText="Figure 68. Male opisthosoma, dorsal view. A, Jilinus hulongensis (Zhu &amp; Wen, 1980). B, Cornitibia simplicithorax (Tanasevitch, 1998). C, Emertongone montifera (Emerton, 1882). D, Oedothorax paracymbialis Tanasevitch, 2015. E, Oe. kodaikanal Tanasevitch, 2015. F, Oe. cunur Tanasevitch, 2015. G, Oe. stylus Tanasevitch, 2015. H, Atypena cirrifrons (Heimer, 1984). I, A. formosana (Oi, 1977). J, Oe. nazareti Scharff, 1989. K, Gongylidium rufipes (Linnaeus, 1758). L, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906). M, U. esyunini (Zhang, Zhang &amp; Yu, 2003). N, Hylyphantes graminicola (Sundevall, 1830). O, Tmeticus tolli Kulczyński, 1908. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967990" httpUri="https://zenodo.org/record/6967990/files/figure.png" pageId="121" pageNumber="538">Fig. 68I</figureCitation>
; PMS with mAP, three AC; PLS with triad, 3+ AC.
</paragraph>
<paragraph id="8BA136D1FFA47656FC90D3704B33F44F" blockId="121.[827,1442,1832,1884]" lastBlockId="123.[163,780,197,342]" lastPageId="123" lastPageNumber="540" pageId="121" pageNumber="538">
<emphasis id="B96AEAC3FFA47654FC90D37049DCF227" box="[827,1000,1832,1853]" italics="true" pageId="121" pageNumber="538">
Female (
<collectionCode id="ED0FAE14FFA47654FC34D37049EDF227" box="[927,985,1832,1853]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="121" pageNumber="538">SMF</collectionCode>
):
</emphasis>
See also
<bibRefCitation id="EF8F4B20FFA47654FBCCD3704EE1F224" author="Oi R" box="[1127,1237,1832,1854]" pageId="121" pageNumber="538" pagination="23 - 26" refId="ref93859" refString="Oi R. 1977. A new erigonid spider from Formosa. Acta Arachnologica 27: 23 - 26." type="journal article" year="1977">Oi (1977)</bibRefCitation>
; epigyne without prominent external modification, copulatory ducts slightly curved, trojactory simple, entrances to spermathecae mesal to exits of fertilixation ducts. (
<figureCitation id="13252A54FFA67656FD4BD4BC4A8BF402" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="123" pageNumber="540">Fig. 73</figureCitation>
). Opisthosoma: dorsal pattern same as male; PMS with CY, mAP, 24 AC; PLS with triad, 2 CY, 3+ AC (
<figureCitation id="13252A54FFA67656FF00D5184AC3F44F" box="[171,247,320,342]" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="123" pageNumber="540">Fig. 73</figureCitation>
).
</paragraph>
<caption id="DF616659FFA77657FF3AD23748B6F3C7" ID-DOI="http://doi.org/10.5281/zenodo.6968006" ID-Zenodo-Dep="6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="122" pageNumber="539" startId="122.[145,225,1647,1669]" targetBox="[158,1428,198,1592]" targetPageId="122">
<paragraph id="8BA136D1FFA77657FF3AD23748B6F3C7" blockId="122.[145,1425,1647,1757]" pageId="122" pageNumber="539">
<emphasis id="B96AEAC3FFA77657FF3AD2374B3DF39E" bold="true" box="[145,265,1647,1669]" pageId="122" pageNumber="539">Figure 73.</emphasis>
<taxonomicName id="4C1E4D52FFA77657FEBFD2374867F39F" authority="(Oi, 1977)" baseAuthorityName="Oi" baseAuthorityYear="1977" box="[276,595,1647,1669]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="122" pageNumber="539" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B96AEAC3FFA77657FEBFD2374BD5F39F" box="[276,481,1647,1669]" italics="true" pageId="122" pageNumber="539">Atypena formosana</emphasis>
(
<bibRefCitation id="EF8F4B20FFA77657FE5BD2374878F39F" author="Oi R" box="[496,588,1647,1669]" pageId="122" pageNumber="539" pagination="23 - 26" refId="ref93859" refString="Oi R. 1977. A new erigonid spider from Formosa. Acta Arachnologica 27: 23 - 26." type="journal article" year="1977">Oi, 1977</bibRefCitation>
)
</taxonomicName>
. AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA136D1FFA67656FF08D5D1480BF485" blockId="123.[163,575,393,415]" box="[163,575,393,415]" pageId="123" pageNumber="540">
<emphasis id="B96AEAC3FFA67656FF08D5D14B00F484" box="[163,308,393,414]" italics="true" pageId="123" pageNumber="540">Distribution:</emphasis>
<collectingCountry id="F3097641FFA67656FEE3D5D14BFAF485" box="[328,462,393,415]" name="Bangladesh" pageId="123" pageNumber="540">Bangladesh</collectingCountry>
to
<collectingCountry id="F3097641FFA67656FE59D5D2480FF485" box="[498,571,394,415]" name="Japan" pageId="123" pageNumber="540">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8BA136D1FFA67656FF08D58D4B59F79E" blockId="123.[163,780,469,644]" pageId="123" pageNumber="540">
<emphasis id="B96AEAC3FFA67656FF08D58D4B3AF4F0" box="[163,270,469,490]" italics="true" pageId="123" pageNumber="540">Remarks:</emphasis>
As in
<taxonomicName id="4C1E4D52FFA67656FEF0D58D4BD7F4F0" baseAuthorityName="Heimer" baseAuthorityYear="1984" box="[347,483,469,490]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="123" pageNumber="540" phylum="Arthropoda" rank="species" species="cirrifrons">
<emphasis id="B96AEAC3FFA67656FEF0D58D4BD7F4F0" box="[347,483,469,490]" italics="true" pageId="123" pageNumber="540">A. cirrifrons</emphasis>
</taxonomicName>
, a scaly sac-like structure is observed at the paracymbial base (
<figureCitation id="13252A54FFA67656FDDED5AC4931F713" box="[629,773,500,522]" captionStart="Figure 73" captionStartId="122.[145,225,1647,1669]" captionTargetBox="[158,1428,198,1592]" captionTargetId="figure-25@122.[145,1431,195,1607]" captionTargetPageId="122" captionText="Figure 73. Atypena formosana (Oi, 1977). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. E, F, epigyne. E, ventral view. F, external morphology. G, female spinnerets, first individual, dorsal view. H, female posterior median spinnerets, second individual, dorsal view. I, female posterior median spinnerets, first individual, ventral view. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968006" httpUri="https://zenodo.org/record/6968006/files/figure.png" pageId="123" pageNumber="540">Figs 73A, C</figureCitation>
,
<figureCitation id="13252A54FFA67656FF08D64B4AC5F732" box="[163,241,531,552]" captionStart="Figure 74" captionStartId="123.[163,243,1819,1841]" captionTargetBox="[166,1446,730,1777]" captionTargetId="figure-215@123.[163,1449,718,1780]" captionTargetPageId="123" captionText="Figure 74. Atypena cirrifrons (Heimer, 1984). AE, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, male right posterior lateral spinneret and posterior median spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6968010" httpUri="https://zenodo.org/record/6968010/files/figure.png" pageId="123" pageNumber="540">74A, C</figureCitation>
). According to the results of our phylogenetic analysis, and the similarities in male genital organs and prosomal modifications, this species is congeneric with
<taxonomicName id="4C1E4D52FFA67656FF76D6374B52F799" baseAuthorityName="Heimer" baseAuthorityYear="1984" box="[221,358,622,644]" class="Arachnida" family="Linyphiidae" genus="Atypena" kingdom="Animalia" order="Araneae" pageId="123" pageNumber="540" phylum="Arthropoda" rank="species" species="cirrifrons">
<emphasis id="B96AEAC3FFA67656FF76D6374B52F799" box="[221,358,622,644]" italics="true" pageId="123" pageNumber="540">A. cirrifrons</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA136D1FFA67656FC0ED49C4F0DF5C1" blockId="123.[933,1337,196,221]" box="[933,1337,196,221]" pageId="123" pageNumber="540">ADDITIONAL MORPHOLOGICAL DATA</paragraph>
</subSubSection>
</treatment>
</document>