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<document id="6E496AF4254D84D36E1AC3ED89D3131E" ID-CLB-Dataset="115951" ID-DOI="10.1093/zoolinnean/zlab033" ID-GBIF-Dataset="75b10efe-a17d-4d10-b46e-094c6a7ebf01" ID-ISSN="0024-4082" ID-Zenodo-Dep="6967774" ID-ZooBank="BE2B3859-8F6A-4543-8A69-91840F82377B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1653991067759" checkinUser="plazi" docAuthor="Lin, Shou-Wang, Lopardo, Lara &amp; Uhl, Gabriele" docDate="2022" docId="03B787C7FFA07650FF6DD1C94E6EF1C8" docLanguage="en" docName="zlab033.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oedothorax bifoveatus TANASEVITCH 2017, INCERTAE" docType="treatment" docVersion="7" lastPageNumber="542" masterDocId="FF8EFFBFFFDD762DFFABD4584A34F51A" masterDocTitle="Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)" masterLastPageNumber="584" masterPageNumber="417" pageNumber="542" updateTime="1699338434129" updateUser="ExternalLinkService">
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<mods:title id="D598F669D9AA4F16A19CEE0BE767AE8C">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<mods:namePart id="135B967D11CE393432424FA6F1D9A8E0">Lin, Shou-Wang</mods:namePart>
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<mods:namePart id="594D057B7D9667D141D77AFDC39AA6AB">Lopardo, Lara</mods:namePart>
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<mods:namePart id="AFFC845C0C7FAB43A514096CF42A0DA3">Uhl, Gabriele</mods:namePart>
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<mods:date id="47123CEA91FCF091AAE342F701198212">2022</mods:date>
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<treatment id="03B787C7FFA07650FF6DD1C94E6EF1C8" ID-DOI="http://doi.org/10.5281/zenodo.6968014" ID-GBIF-Taxon="198618718" ID-Zenodo-Dep="6968014" LSID="urn:lsid:plazi:treatment:03B787C7FFA07650FF6DD1C94E6EF1C8" httpUri="http://treatment.plazi.org/id/03B787C7FFA07650FF6DD1C94E6EF1C8" lastPageNumber="542" pageId="125" pageNumber="542">
<subSubSection id="C304655AFFA07650FF6DD1C94807F0DD" pageId="125" pageNumber="542" type="nomenclature">
<paragraph id="8BA136D1FFA07650FF6DD1C94807F0DD" blockId="125.[198,743,1425,1479]" pageId="125" pageNumber="542">
<heading id="D0E981BDFFA07650FF6DD1C94807F0DD" centered="true" fontSize="9" level="2" pageId="125" pageNumber="542" reason="2">
<taxonomicName id="4C1E4D52FFA07650FF6DD1C94BDEF0DD" ID-CoL="48WTJ" authority="TANASEVITCH, 2017 INCERTAE" authorityName="TANASEVITCH" authorityYear="2017" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="125" pageNumber="542" phylum="Arthropoda" rank="species" species="bifoveatus">
<emphasis id="B96AEAC3FFA07650FF6DD1C94BCAF0B2" box="[198,510,1425,1448]" italics="true" pageId="125" pageNumber="542">OEDOTHORAX BIFOVEATUS</emphasis>
TANASEVITCH, 2017
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<emphasis id="B96AEAC3FFA07650FED1D1EB4BDEF0DD" box="[378,490,1459,1479]" italics="true" pageId="125" pageNumber="542">INCERTAE</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="B96AEAC3FFA07650FE5AD1EB4807F0DD" box="[497,563,1459,1479]" italics="true" pageId="125" pageNumber="542">SEDIS</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA07650FF08D1804B9DF316" pageId="125" pageNumber="542" type="reference_group">
<paragraph id="8BA136D1FFA07650FF08D1804B9DF316" blockId="125.[163,778,1496,1548]" pageId="125" pageNumber="542">
<treatmentCitationGroup id="AB0E11FFFFA07650FF08D1804B9DF316" pageId="125" pageNumber="542">
<treatmentCitation id="0ABF10C0FFA07650FF08D18048E7F0F7" ID-DOI="http://doi.org/10.5281/zenodo.5698138" ID-Zenodo-Dep="5698138" author="Tanasevitch AV" box="[163,723,1496,1518]" httpUri="http://treatment.plazi.org/id/03C487DDFFB9FFC0885CD94FFDE3F40F" page="148" pageId="125" pageNumber="542" year="2017">
<taxonomicName id="4C1E4D52FFA07650FF08D18048E7F0F7" ID-CoL="48WTJ" authority="Tanasevitch, 2017 a: 148" authorityName="TANASEVITCH" authorityPageNumber="148" authorityYear="2017" box="[163,723,1496,1518]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="125" pageNumber="542" phylum="Arthropoda" rank="species" species="bifoveatus">
<emphasis id="B96AEAC3FFA07650FF08D1804B9FF0F7" box="[163,427,1496,1517]" italics="true" pageId="125" pageNumber="542">Oedothorax bifoveatus</emphasis>
<bibRefCitation id="EF8F4B20FFA07650FE18D18048E7F0F7" author="Tanasevitch AV" box="[435,723,1496,1518]" pageId="125" pageNumber="542" pagination="141 - 155" refId="ref95919" refString="Tanasevitch AV. 2017 a. New genera and new species of the family Linyphiidae from Borneo, Sumatra and Java (Arachnida, Araneae). Revue Suisse de Zoologie 124: 141 - 155." type="journal article" year="2017">Tanasevitch, 2017a: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 3134, 4651 (Dmf).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA07650FF08D26E4F2FF42D" pageId="125" pageNumber="542" type="materials_examined">
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<emphasis id="B96AEAC3FFA07650FF08D26E4B7FF350" box="[163,331,1589,1611]" italics="true" pageId="125" pageNumber="542">Type material:</emphasis>
<typeStatus id="54A58873FFA07650FECAD26D4BF8F351" box="[353,460,1589,1611]" pageId="125" pageNumber="542" type="holotype">Holotype</typeStatus>
:
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<collectingCountry id="F3097641FFA07650FE73D26D486FF351" box="[472,603,1589,1611]" name="Indonesia" pageId="125" pageNumber="542">Indonesia</collectingCountry>
:
</emphasis>
<collectingRegion id="49DAF833FFA07650FDC2D26E4895F351" box="[617,673,1590,1611]" country="Indonesia" name="Jawa Timur" pageId="125" pageNumber="542">Java</collectingRegion>
, Cibodas Botanical Garden, near Cipanas,
<emphasis id="B96AEAC3FFA07650FDBFD20D4810F370" box="[532,548,1621,1642]" italics="true" pageId="125" pageNumber="542">c.</emphasis>
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east of Bogor,
<quantity id="4CE69B34FFA07650FF08D22B4ACFF392" box="[163,251,1651,1672]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="125" pageNumber="542" unit="m" value="1400.0">1400 m</quantity>
, vegetational debris in montane
<taxonomicName id="4C1E4D52FFA07650FDD4D22A493FF39D" box="[639,779,1650,1671]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="125" pageNumber="542" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAC3FFA07650FDD4D22A493FF39D" box="[639,779,1650,1671]" italics="true" pageId="125" pageNumber="542">Lithocarpus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D52FFA07650FF6BD2C94B7AF3BC" box="[192,334,1681,1702]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="125" pageNumber="542" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAC3FFA07650FF6BD2C94B7AF3BC" box="[192,334,1681,1702]" italics="true" pageId="125" pageNumber="542">Castanopsis</emphasis>
</taxonomicName>
forest, sifting;
<specimenCount id="9D18FD58FFA07650FDACD2CC4823F3BC" box="[519,535,1684,1702]" pageId="125" pageNumber="542" type="male"></specimenCount>
<date id="FFA01011FFA07650FD8AD2C94886F3BD" box="[545,690,1681,1703]" pageId="125" pageNumber="542" value="1989-11-03" valueMax="1989-11-06" valueMin="1989-11-03">3.6.xi.1989</date>
, leg. D. Burckhardt, I. LöbI &amp; D.Agosti (not examined).
<typeStatus id="54A58873FFA07650FD3DD2E84933F3DF" box="[662,775,1712,1733]" pageId="125" pageNumber="542" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B96AEAC3FFA07650FF08D2964B11F3FE" bold="true" box="[163,293,1742,1764]" pageId="125" pageNumber="542">
<collectingCountry id="F3097641FFA07650FF08D2964B15F3FE" box="[163,289,1742,1764]" name="Indonesia" pageId="125" pageNumber="542">Indonesia</collectingCountry>
:
</emphasis>
<specimenCount id="9D18FD58FFA07650FE87D2974B7DF3F9" box="[300,329,1743,1764]" pageId="125" pageNumber="542" type="male">2♂</specimenCount>
<specimenCount id="9D18FD58FFA07650FEE2D2974B47F3F9" box="[329,371,1743,1764]" pageId="125" pageNumber="542" type="female">17♀</specimenCount>
, collected together with the
<typeStatus id="54A58873FFA07650FD07D296493FF3FE" box="[684,779,1742,1764]" pageId="125" pageNumber="542" type="holotype">holotype</typeStatus>
(not examined);
<collectingRegion id="49DAF833FFA07650FEFAD2B64BB2F219" box="[337,390,1774,1795]" country="Indonesia" name="Jawa Timur" pageId="125" pageNumber="542">Java</collectingRegion>
, Gunung Gede-Pangrango National Park, near Cibodas (
<geoCoordinate id="EE2A5016FFA07650FE21D3544BDBF23B" box="[394,495,1804,1826]" degrees="6" direction="south" minutes="47" orientation="latitude" pageId="125" pageNumber="542" precision="15" seconds="0" value="-6.7833333">6°470” S</geoCoordinate>
,
<geoCoordinate id="EE2A5016FFA07650FE51D3544849F23B" box="[506,637,1804,1825]" degrees="107" direction="east" minutes="01" orientation="longitude" pageId="125" pageNumber="542" precision="15" seconds="0" value="107.01667">107°010” E</geoCoordinate>
),
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,
<specimenCount id="9D18FD58FFA07650FF6CD3734AD1F225" box="[199,229,1835,1856]" pageId="125" pageNumber="542" type="male">1♂</specimenCount>
<specimenCount id="9D18FD58FFA07650FF4ED3734B24F225" box="[229,272,1835,1856]" pageId="125" pageNumber="542" type="female">21♀</specimenCount>
<date id="FFA01011FFA07650FEB3D3724B9CF25A" box="[280,424,1834,1856]" pageId="125" pageNumber="542" value="2005-05-04" valueMax="2005-05-11" valueMin="2005-05-04">4.11.v.2005</date>
, leg. A. Schulz (not examined).
<emphasis id="B96AEAC3FFA07650FF08D3114B55F245" bold="true" box="[163,353,1865,1887]" pageId="125" pageNumber="542">
East
<collectingCountry id="F3097641FFA07650FF42D3114B55F245" box="[233,353,1865,1887]" name="Malaysia" pageId="125" pageNumber="542">Malaysia</collectingCountry>
</emphasis>
: Borneo Island,
<collectingRegion id="49DAF833FFA07650FD84D3114848F245" box="[559,636,1865,1887]" country="Malaysia" name="Sabah" pageId="125" pageNumber="542">Sabah</collectingRegion>
, Tambunan District, Crocker Range, near pass,
<quantity id="4CE69B34FFA07650FB60D49D4F50F5C1" box="[1227,1380,197,219]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.65" metricValueMin="1.55" pageId="125" pageNumber="542" unit="m" value="1600.0" valueMax="1650.0" valueMin="1550.0">15501650 m</quantity>
, road Kota Kinabalu to Tambunan,
<taxonomicName id="4C1E4D52FFA07650FBD7D4BC4F34F5E3" box="[1148,1280,228,249]" class="Magnoliopsida" family="Fagaceae" genus="Lithocarpus" kingdom="Plantae" order="Fagales" pageId="125" pageNumber="542" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAC3FFA07650FBD7D4BC4F34F5E3" box="[1148,1280,228,249]" italics="true" pageId="125" pageNumber="542">Lithocarpus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D52FFA07650FAB5D4BC4F97F5E3" box="[1310,1443,228,249]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="125" pageNumber="542" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEAC3FFA07650FAB5D4BC4F97F5E3" box="[1310,1443,228,249]" italics="true" pageId="125" pageNumber="542">Castanopsis</emphasis>
</taxonomicName>
forest, sifting dead wood, leaves and moss,
<specimenCount id="9D18FD58FFA07650FB51D55B4F2CF40D" box="[1274,1304,259,280]" pageId="125" pageNumber="542" type="male">3♂</specimenCount>
<specimenCount id="9D18FD58FFA07650FABCD55B4F00F40D" box="[1303,1332,259,280]" pageId="125" pageNumber="542" type="female">1♀</specimenCount>
<date id="FFA01011FFA07650FA9CD55A4FABF402" box="[1335,1439,258,280]" pageId="125" pageNumber="542" value="1987-05-16">16.v.1987</date>
, leg. D. Burckhardt &amp; I. Löbl (not examined).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA07650FC90D53B49F3F482" pageId="125" pageNumber="542" type="distribution">
<paragraph id="8BA136D1FFA07650FC90D53B49F3F482" blockId="125.[827,1443,355,408]" pageId="125" pageNumber="542">
<emphasis id="B96AEAC3FFA07650FC90D53B49F8F462" box="[827,972,355,376]" italics="true" pageId="125" pageNumber="542">Distribution:</emphasis>
East
<collectingCountry id="F3097641FFA07650FBB9D53B4E49F463" box="[1042,1149,355,377]" name="Malaysia" pageId="125" pageNumber="542">Malaysia</collectingCountry>
: Borneo Island;
<collectingCountry id="F3097641FFA07650FA86D53B4F94F463" box="[1325,1440,355,377]" name="Indonesia" pageId="125" pageNumber="542">Indonesia</collectingCountry>
:
<collectingRegion id="49DAF833FFA07650FC90D5DB4945F482" box="[827,881,387,408]" country="Indonesia" name="Jawa Timur" pageId="125" pageNumber="542">Java</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA07650FC90D59D4E50F636" pageId="125" pageNumber="542" type="diagnosis">
<paragraph id="8BA136D1FFA07650FC90D59D4986F4C0" blockId="125.[827,946,453,474]" box="[827,946,453,474]" pageId="125" pageNumber="542">
<emphasis id="B96AEAC3FFA07650FC90D59D4986F4C0" box="[827,946,453,474]" italics="true" pageId="125" pageNumber="542">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FFA07650FC90D65F49C5F7AC" blockId="125.[827,1443,519,694]" pageId="125" pageNumber="542">
<emphasis id="B96AEAC3FFA07650FC90D65F49B2F706" box="[827,902,519,540]" italics="true" pageId="125" pageNumber="542">Males:</emphasis>
This species can be identified by the shape of palpal tibia, which has a largely elevated margin on the dorsal side and a slender, setae-bearing retrolateral apophysis (fig.
<quantity id="4CE69B34FFA07650FC5DD63B4E01F763" box="[1014,1077,611,633]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" pageId="125" pageNumber="542" unit="in" value="49.0">49 in</quantity>
<bibRefCitation id="EF8F4B20FFA07650FB95D63B4F1FF762" author="Tanasevitch AV" box="[1086,1323,611,633]" pageId="125" pageNumber="542" pagination="141 - 155" refId="ref95919" refString="Tanasevitch AV. 2017 a. New genera and new species of the family Linyphiidae from Borneo, Sumatra and Java (Arachnida, Araneae). Revue Suisse de Zoologie 124: 141 - 155." type="journal article" year="2017">Tanasevitch, 2017a</bibRefCitation>
), and the curved shape of the radix [convector in
<bibRefCitation id="EF8F4B20FFA07650FABDD6DA49A4F7AC" author="Tanasevitch AV" pageId="125" pageNumber="542" pagination="141 - 155" refId="ref95919" refString="Tanasevitch AV. 2017 a. New genera and new species of the family Linyphiidae from Borneo, Sumatra and Java (Arachnida, Araneae). Revue Suisse de Zoologie 124: 141 - 155." type="journal article" year="2017">Tanasevitch (2017a)</bibRefCitation>
, fig. 50].
</paragraph>
<paragraph id="8BA136D1FFA07650FC90D6824E50F636" blockId="125.[827,1443,730,813]" pageId="125" pageNumber="542">
<emphasis id="B96AEAC3FFA07650FC90D6824996F7F5" box="[827,930,730,751]" italics="true" pageId="125" pageNumber="542">Females:</emphasis>
Characterized by the presence of a large socket on each side of the dorsal plate of epigyne (fig.
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<bibRefCitation id="EF8F4B20FFA07650FCD1D74F4E61F636" author="Tanasevitch AV" box="[890,1109,791,813]" pageId="125" pageNumber="542" pagination="141 - 155" refId="ref95919" refString="Tanasevitch AV. 2017 a. New genera and new species of the family Linyphiidae from Borneo, Sumatra and Java (Arachnida, Araneae). Revue Suisse de Zoologie 124: 141 - 155." type="journal article" year="2017">Tanasevitch, 2017a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA07650FC90D7084E0EF67C" box="[827,1082,848,870]" pageId="125" pageNumber="542" type="biology_ecology">
<paragraph id="8BA136D1FFA07650FC90D7084E0EF67C" blockId="125.[827,1082,848,870]" box="[827,1082,848,870]" pageId="125" pageNumber="542">
<emphasis id="B96AEAC3FFA07650FC90D70849ADF67F" box="[827,921,848,869]" italics="true" pageId="125" pageNumber="542">Habitat:</emphasis>
Forest litter.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFFA07650FC90D7D24E6EF1C8" pageId="125" pageNumber="542" type="discussion">
<paragraph id="8BA136D1FFA07650FC90D7D24E6EF1C8" blockId="125.[827,1443,906,1234]" pageId="125" pageNumber="542">
<emphasis id="B96AEAC3FFA07650FC90D7D2499DF685" box="[827,937,906,927]" italics="true" pageId="125" pageNumber="542">Remarks:</emphasis>
Features in the male palpal configuration (e.g. paracymbium distal part massive, distal setae group distally situated; anterior radical process ending in several short, pointed processes; embolus retrolaterally spiral), as well as the epigyne (e.g. two large, rounded sockets on either side of the dorsal plate), differ significantly from
<emphasis id="B96AEAC3FFA07650FAB3D019496AF16C" italics="true" pageId="125" pageNumber="542">
<taxonomicName id="4C1E4D52FFA07650FAB3D0194F96F14C" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[1304,1442,1089,1110]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="125" pageNumber="542" phylum="Arthropoda" rank="genus" sensu="stricto">Oedothorax</taxonomicName>
<taxonomicNameLabel id="A25957B8FFA07650FC90D039496AF16C" box="[827,862,1121,1142]" pageId="125" pageNumber="542" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
. Furthermore, its Indomalayan distribution in constrast to the Holarctic distribution of
<taxonomicName id="4C1E4D52FFA07650FAB6D0274F97F18E" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[1309,1443,1151,1172]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="125" pageNumber="542" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAC3FFA07650FAB6D0274F97F18E" box="[1309,1443,1151,1172]" italics="true" pageId="125" pageNumber="542">Oedothorax</emphasis>
</taxonomicName>
also suggests that this species presumably does not belong to
<emphasis id="B96AEAC3FFA07650FC02D0E44E6EF1C8" box="[937,1114,1212,1234]" italics="true" pageId="125" pageNumber="542">
<taxonomicName id="4C1E4D52FFA07650FC02D0E44E18F1CB" authorityName="Lin &amp; Lopardo &amp; Uhl" authorityYear="2022" box="[937,1068,1212,1233]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="125" pageNumber="542" phylum="Arthropoda" rank="genus" sensu="stricto">Oedothorax</taxonomicName>
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..
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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