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<document id="E81FC95CC69339650ED571CBD81B5041" ID-DOI="10.5281/zenodo.178554" ID-GBIF-Dataset="881da8c7-ff5e-4163-8ab5-625dedd4f460" ID-ISSN="1175-5326" ID-Zenodo-Dep="178554" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459900171483" checkinUser="plazi" docAuthor="Sonnenberg, Rainer" docDate="2007" docId="03B787BF496A8721FCC689D7FB10F6A4" docLanguage="en" docName="zt01591p038.pdf" docOrigin="Zootaxa 1591" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Chromaphyosemion omega Sonnenberg, 2007, sp. nov." docType="treatment" docVersion="10" lastPageNumber="16" masterDocId="FF8EFFC74961872EFC518F6AFFF9F24E" masterDocTitle="Description of three new species of the genus Chromaphyosemion Radda, 1971 (Cyprinodontiformes: Nothobranchiidae) from the coastal plains of Cameroon with a preliminary review of the Chromaphyosemion splendopleure complex" masterLastPageNumber="38" masterPageNumber="1" pageNumber="12" updateTime="1698228761725" updateUser="plazi">
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<mods:title id="EAF89DAABDB4AA843B1AEFF1A3CE565C">Description of three new species of the genus Chromaphyosemion Radda, 1971 (Cyprinodontiformes: Nothobranchiidae) from the coastal plains of Cameroon with a preliminary review of the Chromaphyosemion splendopleure complex</mods:title>
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<mods:name id="FA5A498B738C5CA64E2FB31C08E5F981" type="personal">
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<mods:roleTerm id="6211BCC77FF8176664F0ECE9B6BE942B">Author</mods:roleTerm>
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<mods:namePart id="1871C817C5AA7918E2CEDAF76F8ED314">Sonnenberg, Rainer</mods:namePart>
</mods:name>
<mods:typeOfResource id="A55061DBAD11E454B6F2DB83EB0F245D">text</mods:typeOfResource>
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<mods:date id="D1A8201A165FB644D399B22422099A11">2007</mods:date>
<mods:detail id="B375CB7F94208C6B03ED96AD5522F5F2" type="volume">
<mods:number id="625183C1F0441ED7C669FA6C71A76D5A">1591</mods:number>
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<mods:classification id="B7EDC2E9BF8B1BE6256B2F6C27046F59">journal article</mods:classification>
<mods:identifier id="A74D1CCB145B74555ADA44750E8560A1" type="DOI">10.5281/zenodo.178554</mods:identifier>
<mods:identifier id="2FF125F2C413B55E7820A627BE69E9D3" type="GBIF-Dataset">881da8c7-ff5e-4163-8ab5-625dedd4f460</mods:identifier>
<mods:identifier id="20713844393FF5AE5B2955ABD403B557" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5FCA5C9D2EF33CCF65CBB569B16ED3EE" type="Zenodo-Dep">178554</mods:identifier>
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<treatment id="03B787BF496A8721FCC689D7FB10F6A4" ID-DOI="http://doi.org/10.5281/zenodo.5680201" ID-GBIF-Taxon="119372949" ID-Zenodo-Dep="5680201" LSID="urn:lsid:plazi:treatment:03B787BF496A8721FCC689D7FB10F6A4" httpUri="http://treatment.plazi.org/id/03B787BF496A8721FCC689D7FB10F6A4" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
<subSubSection id="C3046522496A8725FCC689D7FEDCF4B4" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BA136A9496A8725FCC689D7FDBEF499" blockId="11.[151,583,1725,1786]" box="[151,583,1725,1751]" pageId="11" pageNumber="12">
<heading id="D0E981C5496A8725FCC689D7FDBEF499" bold="true" box="[151,583,1725,1751]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B96AEABB496A8725FCC689D7FDBEF499" bold="true" box="[151,583,1725,1751]" pageId="11" pageNumber="12">
<taxonomicName id="4C1E4D2A496A8725FCC689D7FE1BF499" ID-CoL="5YGL9" box="[151,482,1725,1751]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="omega" status="sp. nov.">
<emphasis id="B96AEABB496A8725FCC689D7FE1BF499" bold="true" box="[151,482,1725,1751]" italics="true" pageId="11" pageNumber="12">Chromaphyosemion omega</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957C0496A8725FDB889D7FDBEF499" box="[489,583,1725,1751]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA136A9496A8725FCC6898AFEDCF4B4" blockId="11.[151,583,1725,1786]" box="[151,293,1760,1786]" pageId="11" pageNumber="12">
(
<figureCitation id="13252A2C496A8725FCCE898AFEE5F4B4" box="[159,284,1760,1786]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="12.[151,255,1817,1841]" captionStartId-1="13.[151,255,991,1015]" captionStartId-2="13.[151,255,1779,1803]" captionTargetBox-0="[248,1338,1189,1790]" captionTargetBox-1="[246,1341,277,964]" captionTargetBox-2="[246,1341,1068,1755]" captionTargetId-0="figure@12.[248,1339,1187,1791]" captionTargetId-1="figure@13.[246,1342,277,964]" captionTargetId-2="figure@13.[246,1342,1068,1755]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="13" captionText-0="FIGURE 9. Chromaphyosemion omega, males from the type locality, not preserved. Photographed directly after collection." captionText-1="FIGURE 10. Chromaphyosemion omega, male from the strain C 89 / 35, not preserved. Photo by W. Eigelshofen" captionText-2="FIGURE 11. Chromaphyosemion omega, female from the type locality, not preserved. Photographed after some months in the aquarium. Photo by W. Eigelshofen" httpUri-0="https://zenodo.org/record/178563/files/figure.png" httpUri-1="https://zenodo.org/record/178564/files/figure.png" httpUri-2="https://zenodo.org/record/178565/files/figure.png" pageId="11" pageNumber="12">Figs. 9–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3046522496A8722FCC6885AFEB6F3E1" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8BA136A9496A8725FCC6885AFED8F529" blockId="11.[151,1436,1840,2023]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB0E1187496A8725FCC6885AFED8F529" pageId="11" pageNumber="12">
<taxonomicName id="4C1E4D2A496A8725FCC6885AFE60F509" ID-CoL="FL59" box="[151,409,1840,1863]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="bivittatum">
<emphasis id="B96AEABB496A8725FCC6885AFE60F509" box="[151,409,1840,1863]" italics="true" pageId="11" pageNumber="12">Aphyosemion bivittatum</emphasis>
</taxonomicName>
(non
<bibRefCitation id="EF8F4B58496A8725FD87885AFD7AF509" author="Lonnberg" box="[470,643,1840,1863]" pageId="11" pageNumber="36" refString="Lonnberg, E. (1895) Notes on fishes collected in the Cameroons by Mr. Y. Sjostedt. Ofversigt af Kongliga Vetenskaps- Akademiens Forhandlingar, 3, 179 - 195." type="journal article" year="1895">Lönnberg, 1895</bibRefCitation>
),
<treatmentCitation id="0ABF10B8496A8725FEC4885AFCB4F509" author="Scheel" box="[661,845,1840,1863]" page="117" pageId="11" pageNumber="12" year="1968">
<bibRefCitation id="EF8F4B58496A8725FEC4885AFCE5F509" author="Scheel" box="[661,796,1840,1863]" pageId="11" pageNumber="37" refString="Scheel, J. J. (1968) Rivulins of the Old World. TFH Publications, New Jersey, 480 pp." type="book" year="1968">Scheel 1968</bibRefCitation>
: 117
</treatmentCitation>
–127 (in part, distribution, morphology, taxonomy, synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136A9496A8725FCC6881AFE71F5E9" blockId="11.[151,1436,1840,2023]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB0E1187496A8725FCC6881AFE71F5E9" pageId="11" pageNumber="12">
<taxonomicName id="4C1E4D2A496A8725FCC6881AFD59F5C9" ID-CoL="FL4Z" box="[151,672,1904,1927]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="bitaeniatum" subGenus="Chromaphyosemion">
<emphasis id="B96AEABB496A8725FCC6881AFD59F5C9" box="[151,672,1904,1927]" italics="true" pageId="11" pageNumber="12">Aphyosemion (Chromaphyosemion) bitaeniatum</emphasis>
</taxonomicName>
(non
<bibRefCitation id="EF8F4B58496A8725FEB4881AFCAFF5C9" author="Ahl" box="[741,854,1904,1927]" pageId="11" pageNumber="35" refString="Ahl, E. (1924) Neue afrikanische Zahnkarpfen aus dem zoologischen Museum Berlin. Zoologischer Anzeiger, 61, 135 - 145." type="journal article" year="1924">Ahl, 1924</bibRefCitation>
),
<treatmentCitation id="0ABF10B8496A8725FF3D881AFBD3F5C9" author="Scheel" box="[876,1066,1904,1927]" page="168" pageId="11" pageNumber="12" year="1990">
<bibRefCitation id="EF8F4B58496A8725FF3D881AFC0FF5C9" author="Scheel" box="[876,1014,1904,1927]" pageId="11" pageNumber="37" refString="Scheel, J. J. (1990) Atlas of the Killifishes of the Old World. TFH Publication, Neptune City, New Jersey, 448 pp." type="book" year="1990">Scheel 1990</bibRefCitation>
: 168
</treatmentCitation>
–173 (in part, distribution, karyotypes, taxonomy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136A9496A8722FCC688DAFC9FF341" blockId="11.[151,1436,1840,2023]" lastBlockId="12.[151,1436,152,431]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<treatmentCitationGroup id="AB0E1187496A8722FCC688DAFC9FF341" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<taxonomicName id="4C1E4D2A496A8725FCC688DAFD4BF589" ID-CoL="FLDB" box="[151,690,1968,1991]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="splendopleure" subGenus="Chromaphyosemion">
<emphasis id="B96AEABB496A8725FCC688DAFD4BF589" box="[151,690,1968,1991]" italics="true" pageId="11" pageNumber="12">Aphyosemion (Chromaphyosemion) splendopleure</emphasis>
</taxonomicName>
(non
<bibRefCitation id="EF8F4B58496A8725FEA188DAFC60F589" author="Bruning" box="[752,921,1968,1991]" pageId="11" pageNumber="35" refString="Bruning, C. (1929 a) Der grune Leuchtaugenfisch und der afrikanische Glanzkarpfling. Wochenschrift fur Aquarien- und Terrarienkunde, 26, 356." type="journal article" year="1929" yearSuffix="a">Brüning, 1929a</bibRefCitation>
),
<bibRefCitation id="EF8F4B58496A8725FFFC88DAFB8DF589" author="Agnese" box="[941,1140,1968,1991]" pageId="11" pageNumber="35" refString="Agnese, J. - F., Zentz, F., Legros, O. &amp; Sellos, D. (2006) Phylogenetic relationships and phylogeography of the killifish species of the subgenus Chromaphyosemion (Radda, 1971) in West Africa, inferred from mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 40, 332 - 346." type="journal article" year="2006">Agnèse et al. 2006</bibRefCitation>
(in part, phylogeny, collection localities, taxonomy),
<treatmentCitation id="0ABF10B8496A8725FDB788BAFD68F5A9" author="Amiet" box="[486,657,2000,2023]" page="49" pageId="11" pageNumber="12" year="1987">
<bibRefCitation id="EF8F4B58496A8725FDB788BAFD90F5A9" author="Amiet" box="[486,617,2000,2023]" pageId="11" pageNumber="35" refString="Amiet, J. L. (1987) Le Genre Aphyosemion Myers (Pisces, Teleostei, Cyprinodontiformes). Faune du Cameroun, Volume 2. Sciences Nat, Compiegne, France, 262 pp." type="book" year="1987">Amiet 1987</bibRefCitation>
: 49
</treatmentCitation>
, 176, plate 36, fig. 15 (in part, collection localities, color pattern),
<treatmentCitation id="0ABF10B8496A8722F90B88BAFECBF2E1" author="Huber" lastPageId="12" lastPageNumber="13" page="20" pageId="11" pageNumber="12" year="1998">
<bibRefCitation id="EF8F4B58496A8722F90B88BAFEF5F2E1" author="Huber" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="36" refString="Huber, J. H. (1998 a) Aphyosemion alpha n. sp. - Eine neue Art der Untergattung Chromaphyosemion mit einem ausgepragtem Farbungsmuster und einer besonders sudlichen Verbreitung. Das Aquarium, 350, 15 - 23." type="journal article" year="1998" yearSuffix="a">Huber 1998a</bibRefCitation>
: 20
</treatmentCitation>
–21 (in part, taxonomy, distribution), 2000: 471 (in part, collection localities),
<treatmentCitation id="0ABF10B8496D8722F82F8FF2FA77F2E1" author="Legros" box="[1150,1422,152,175]" page="20" pageId="12" pageNumber="13" year="2007">
<bibRefCitation id="EF8F4B58496D8722F82F8FF2FA90F2E1" author="Legros" box="[1150,1385,152,175]" pageId="12" pageNumber="36" refString="Legros, O. &amp; Zentz, F. (2007) Aphyosemion malumbresi n. sp., un nouveau Chromaphyosemion (Teleostei: Aplocheilidae) originaire de Guinee Equatoriale continentale. AKFB Killi-Contact, no. 6, decembre 2006, 1 - 28." type="journal article" year="2007">Legros &amp; Zentz 2007</bibRefCitation>
: 20
</treatmentCitation>
– 21 (phylogeny),
<treatmentCitation id="0ABF10B8496D8722FD278FD2FD81F281" author="Radda" box="[374,632,184,207]" page="24" pageId="12" pageNumber="13" year="1982">
<bibRefCitation id="EF8F4B58496D8722FD278FD2FDAAF281" author="Radda" box="[374,595,184,207]" pageId="12" pageNumber="37" refString="Radda, A. C. &amp; Purzl, E. (1982) Killifische aus aller Welt. Feldfuhrer der Cyprinodontiformes der Lander der Bucht von Biafra (SO-Nigeria, West-Kamerun). Band 2. Verlag Otto Hofmann, Wien, 72 pp." type="book" year="1982">Radda &amp; Pürzl 1982</bibRefCitation>
: 24
</treatmentCitation>
–25 (in part, distribution),
<treatmentCitation id="0ABF10B8496D8722FFC18FD2FB68F281" author="Radda" box="[912,1169,184,207]" page="20" pageId="12" pageNumber="13" year="1983">
<bibRefCitation id="EF8F4B58496D8722FFC18FD2FB95F281" author="Radda" box="[912,1132,184,207]" pageId="12" pageNumber="37" refString="Radda, A. C. &amp; Purzl, E. (1983) Killifische aus aller Welt. Feldfuhrer der Cyprinodontiformes der Lander der Kustenlander Zentralafrikas I Ostkamerun. Band 3. Verlag Otto Hofmann, Wien, 80 pp." type="book" year="1983">Radda &amp; Pürzl 1983</bibRefCitation>
: 20
</treatmentCitation>
–21 (in part, distribution) 1987: 70 (in part, distribution),
<treatmentCitation id="0ABF10B8496D8722FE728FB2FD16F2A1" author="Seegers" box="[547,751,216,239]" page="159" pageId="12" pageNumber="13" year="1981">
<bibRefCitation id="EF8F4B58496D8722FE728FB2FD41F2A1" author="Seegers" box="[547,696,216,239]" pageId="12" pageNumber="37" refString="Seegers, L. (1981) Chromaphyosemion - kleine Farbwunder aus Westafrika. Die Aquarien- und Terrarienzeitschrift, 34, 155 - 163." type="journal article" year="1981">Seegers 1981</bibRefCitation>
: 159
</treatmentCitation>
, 161 (in part, distribution), 1986: 52–54 (in part, distribution),
<treatmentCitation id="0ABF10B8496D8722FC968F92FE4AF341" author="Wildekamp" box="[199,435,248,271]" page="277" pageId="12" pageNumber="13" year="1993">
<bibRefCitation id="EF8F4B58496D8722FC968F92FE7BF341" author="Wildekamp" box="[199,386,248,271]" pageId="12" pageNumber="38" refString="Wildekamp, R. H. (1993) A World of Killies. Atlas of the Oviparous Cyprinodontiform Fishes of the World. Vol um e 1. The American Killifish Association, Mishawaka, Indiana, 311 pp." type="book" year="1993">Wildekamp 1993</bibRefCitation>
: 277
</treatmentCitation>
–279 (in part, distribution, morphology).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136A9496D8722FCC68E72FD9BF321" blockId="12.[151,1436,152,431]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB0E1187496D8722FCC68E72FD9BF321" bridged="true" pageId="12" pageNumber="13">
<taxonomicName id="4C1E4D2A496D8722FCC68E72FD4EF361" ID-CoL="FLDB" box="[151,695,280,303]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="splendopleure" subGenus="Chromaphyosemion">
<emphasis id="B96AEABB496D8722FCC68E72FD4EF361" box="[151,695,280,303]" italics="true" pageId="12" pageNumber="13">Aphyosemion (Chromaphyosemion) splendopleure</emphasis>
</taxonomicName>
'Kopongo' (non
<bibRefCitation id="EF8F4B58496D8722FF218E72FBE5F361" author="Bruning" box="[880,1052,280,303]" pageId="12" pageNumber="35" refString="Bruning, C. (1929 a) Der grune Leuchtaugenfisch und der afrikanische Glanzkarpfling. Wochenschrift fur Aquarien- und Terrarienkunde, 26, 356." type="journal article" year="1929" yearSuffix="a">Brüning, 1929a</bibRefCitation>
),
<treatmentCitation id="0ABF10B8496D8722F8608E72FB2FF361" author="Eberl" box="[1073,1238,280,303]" page="48" pageId="12" pageNumber="13" year="1996">
<bibRefCitation id="EF8F4B58496D8722F8608E72FB57F361" author="Eberl" box="[1073,1198,280,303]" pageId="12" pageNumber="36" refString="Eberl, W. (1996) Die Untergattung Chromaphyosemion. Deutsche Killifisch Gemeinschaft Journal, Supplementheft 4, 88 pp." type="book" year="1996">Eberl 1996</bibRefCitation>
: 48
</treatmentCitation>
-50 (color pattern, collection localities, color photo p. 49, lower picture, erroneously exchanged with the above picture),
<treatmentCitation id="0ABF10B8496D8722F94C8E52FEEEF321" author="Eigelshofen" page="9" pageId="12" pageNumber="13" year="1998">
<bibRefCitation id="EF8F4B58496D8722F94C8E52FEF9F321" author="Eigelshofen" pageId="12" pageNumber="36" refString="Eigelshofen, W. (1998) Chromaphyosemion. Beeindruckend in Form und Farbe. Aquaristik Aktuell, 9 - 10, 8 - 13." type="book chapter" year="1998">Eigelshofen 1998</bibRefCitation>
: 9
</treatmentCitation>
–12 (distribution, color photo).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136A9496D8722FCC68E12FEB6F3E1" blockId="12.[151,1436,152,431]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB0E1187496D8722FCC68E12FEB6F3E1" pageId="12" pageNumber="13">
<taxonomicName id="4C1E4D2A496D8722FCC68E12FE94F3C1" box="[151,365,376,399]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B96AEABB496D8722FCC68E12FE94F3C1" box="[151,365,376,399]" italics="true" pageId="12" pageNumber="13">Chromaphyosemion</emphasis>
</taxonomicName>
sp.
<taxonomicName id="4C1E4D2A496D8722FDCE8E12FD9AF3C1" ID-CoL="FLDB" box="[415,611,376,399]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="splendopleure">
aff.
<emphasis id="B96AEABB496D8722FD9C8E12FD9AF3C1" box="[461,611,376,399]" italics="true" pageId="12" pageNumber="13">splendopleure</emphasis>
</taxonomicName>
'Kopongo',
<treatmentCitation id="0ABF10B8496D8722FEBA8E12FC20F3C1" author="Sonnenberg" box="[747,985,376,399]" page="81" pageId="12" pageNumber="13" year="2000">
<bibRefCitation id="EF8F4B58496D8722FEBA8E12FC57F3C1" author="Sonnenberg" box="[747,942,376,399]" pageId="12" pageNumber="38" refString="Sonnenberg, R. (2000) The Distribution of Chromaphyosemion Radda, 1971 (Teleostei: Cyprinodontiformes) on the Coastal Plains of West and Central Africa. In Rheinwald, G. (Ed.) Isolated Vertebrate Communities in the Tropics. Proceedings of the 4 th International Symposium, Bonn. Bonner zoologische Monographien 46, pp. 79 - 94." type="journal article" year="2000">Sonnenberg 2000</bibRefCitation>
: 81
</treatmentCitation>
, 85–86 (color pattern, distribution, color photo Nr. 4).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3046522496D8722FCC68EB5FA64F03C" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8BA136A9496D8722FCC68EB5FD79F004" blockId="12.[151,1437,479,1106]" pageId="12" pageNumber="13">
<emphasis id="B96AEABB496D8722FCC68EB5FEE9F3B7" bold="true" box="[151,272,479,505]" pageId="12" pageNumber="13">
<typeStatus id="54A5880B496D8722FCC68EB5FEF2F3B7" box="[151,267,479,505]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED0FAE6C496D8722FD4A8E8AFE95F3B4" LSID="urn:lsid:biocol.org:col:34613" box="[283,364,480,506]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="12" pageNumber="13">ZFMK</collectionCode>
39970, male (
<quantity id="4CE69B4C496D8722FE4F8E8AFD72F3B4" box="[542,651,480,506]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="12" pageNumber="13" unit="mm" value="31.3">31.3 mm</quantity>
SL),
<collectingCountry id="F3097639496D8722FE828E8AFCA9F3B4" box="[723,848,480,506]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
, old road from Edea to Douala, small river Ngo Njock near the village Ndog Nyang, (
<geoCoordinate id="EE2A506E496D8722FE208D62FCD8F06C" box="[625,801,520,546]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="3.9867778">03°59'12.4'' N</geoCoordinate>
,
<geoCoordinate id="EE2A506E496D8722FF638D62FC24F06C" box="[818,989,520,546]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="10.097305">10°05'50.3'' E</geoCoordinate>
), U. Kämpf, R. Sonnenberg and A. Tränkner leg., collection locality
<collectionCode id="ED0FAE6C496D8722FE4C8D5AFD9DF004" box="[541,612,560,586]" httpUri="http://grbio.org/cool/e1dy-v5iv" name="Culture Collection of Phytopathogenic Fungi (Colecao de Culturas de Fungos Fitopatogenicos Prof. Maria Menezes)" pageId="12" pageNumber="13">CMM</collectionCode>
7.
</paragraph>
<paragraph id="8BA136A9496D8722FC978D3DFA64F03C" blockId="12.[151,1437,479,1106]" box="[198,1437,599,626]" pageId="12" pageNumber="13">
<emphasis id="B96AEABB496D8722FC978D3DFEB2F03F" bold="true" box="[198,331,599,625]" pageId="12" pageNumber="13">
<typeStatus id="54A5880B496D8722FC978D3DFEBFF03F" box="[198,326,599,625]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED0FAE6C496D8722FD038D32FE5DF03C" LSID="urn:lsid:biocol.org:col:34613" box="[338,420,600,626]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="12" pageNumber="13">ZFMK</collectionCode>
39971–39980 (
<specimenCount id="9D18FD20496D8722FE0D8D32FD40F03C" box="[604,697,600,626]" pageId="12" pageNumber="13" type="male">7 males</specimenCount>
,
<specimenCount id="9D18FD20496D8722FE948D32FCC1F03C" box="[709,824,600,626]" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
,
<specimenCount id="9D18FD20496D8722FF158D32FC45F03C" box="[836,956,600,626]" pageId="12" pageNumber="13" type="juvenile">1 juvenile</specimenCount>
), paratopotypes, collected with
<typeStatus id="54A5880B496D8722F97F8D32FA61F03C" box="[1326,1432,600,626]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3046522496D8722FC978DEAFB14F174" pageId="12" pageNumber="13" type="description">
<paragraph id="8BA136A9496D8722FC978DEAFE4BF0A4" blockId="12.[151,1437,479,1106]" pageId="12" pageNumber="13">
ZFMK 39981–39986 (
<specimenCount id="9D18FD20496D8722FD808DEAFDD4F0D4" box="[465,557,640,666]" pageId="12" pageNumber="13" type="male">5 males</specimenCount>
,
<specimenCount id="9D18FD20496D8722FE688DEAFD5BF0D4" box="[569,674,640,666]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
,
<specimenCount id="9D18FD20496D8722FEFC8DEAFCDFF0D4" box="[685,806,640,666]" pageId="12" pageNumber="13" type="juvenile">1 juvenile</specimenCount>
),
<collectingCountry id="F3097639496D8722FF688DEAFC4FF0D4" box="[825,950,640,666]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
, creek crossing the old road from Edea to Douala, near Bonepoupa (
<geoCoordinate id="EE2A506E496D8722FD998DC2FD8BF08C" box="[456,626,680,706]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="4.1083612">04°06'30.1'' N</geoCoordinate>
,
<geoCoordinate id="EE2A506E496D8722FE2F8DC2FCDAF08C" box="[638,803,680,706]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="10.020166">10°01'12.6'' E</geoCoordinate>
), H. Kullmann, B. Misof and R. Sonnenberg leg., collection locality C 01/62.
</paragraph>
<paragraph id="8BA136A9496D8722FC978D92FBBAF15C" blockId="12.[151,1437,479,1106]" box="[198,1091,760,786]" pageId="12" pageNumber="13">
CENAREST, Batoke,
<collectingCountry id="F3097639496D8722FD9D8D92FDBFF15C" box="[460,582,760,786]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
(
<specimenCount id="9D18FD20496D8722FE088D92FD53F15C" box="[601,682,760,786]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
), same collection data as
<typeStatus id="54A5880B496D8722FF858D92FBC7F15C" box="[980,1086,760,786]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BA136A9496D8722FC978C4AFB14F174" blockId="12.[151,1437,479,1106]" box="[198,1261,800,826]" pageId="12" pageNumber="13">
CENAREST, Batoke,
<collectingCountry id="F3097639496D8722FD9D8C4AFDBFF174" box="[460,582,800,826]" name="Cameroon" pageId="12" pageNumber="13">Cameroon</collectingCountry>
(
<specimenCount id="9D18FD20496D8722FE088C4AFD38F174" box="[601,705,800,826]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
), same collection data as ZFMK 39981–39986.
</paragraph>
</subSubSection>
<subSubSection id="C3046522496D8722FC978C2DFABAF61C" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BA136A9496D8722FC978C2DFABAF61C" blockId="12.[151,1437,479,1106]" pageId="12" pageNumber="13">
<emphasis id="B96AEABB496D8722FC978C2DFEBFF12F" bold="true" box="[198,326,839,865]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
<taxonomicName id="4C1E4D2A496D8722FD018C22FD6BF12F" box="[336,658,840,865]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496D8722FD018C22FD6BF12F" box="[336,658,840,865]" italics="true" pageId="12" pageNumber="13">Chromaphyosemion omega</emphasis>
</taxonomicName>
is distinguished from all surrounding and closely related congeners by completely orange anal and pelvic fins without red pattern, a red submarginal and a blue marginal band, in some specimens a small greenish basal part with a row of basal dark red dots in anal fin versus incomplete orange anal fin with red markings and always blue in lower half, but above submarginal red band in
<taxonomicName id="4C1E4D2A496D8722FCE48C82FEA7F64F" box="[181,350,1000,1025]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="loennbergii">
<emphasis id="B96AEABB496D8722FCE48C82FEA7F64F" box="[181,350,1000,1025]" italics="true" pageId="12" pageNumber="13">C. loennbergii</emphasis>
</taxonomicName>
, greenish in
<taxonomicName id="4C1E4D2A496D8722FDA78C82FD49F64F" box="[502,688,1000,1026]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="koungueense">
<emphasis id="B96AEABB496D8722FDA78C82FDF0F64F" box="[502,521,1000,1025]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B96AEABB496D8722FE488C82FD49F64F" box="[537,688,1000,1025]" italics="true" pageId="12" pageNumber="13">koungueense</emphasis>
</taxonomicName>
, only partially orange with yellow or greenish anal in
<taxonomicName id="4C1E4D2A496D8722F9638C82FF07F667" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" sensu="lato" species="splendopleure">
<emphasis id="B96AEABB496D8722F9638C82FF07F667" italics="true" pageId="12" pageNumber="13">C. splendopleure</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957C0496D8722FD548B7AFEDEF664" box="[261,295,1040,1066]" pageId="12" pageNumber="13" sensu="lato">s.l.</taxonomicNameLabel>
and
<taxonomicName id="4C1E4D2A496D8722FD308B7AFDEFF667" box="[353,534,1040,1066]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="punctulatum">
<emphasis id="B96AEABB496D8722FD308B7AFE8DF667" box="[353,372,1040,1065]" italics="true" pageId="12" pageNumber="13">C</emphasis>
.
<emphasis id="B96AEABB496D8722FDD28B7AFDEFF667" box="[387,534,1040,1065]" italics="true" pageId="12" pageNumber="13">punctulatum</emphasis>
</taxonomicName>
, or with a yellow or greenish marginal band in
<taxonomicName id="4C1E4D2A496D8722F8128B7AFAF2F667" box="[1091,1291,1040,1065]" class="Actinopterygii" family="Nothobranchiidae" genus="Aphyosemion" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" sensu="lato" species="splendopleure">
<emphasis id="B96AEABB496D8722F8128B7AFAF2F667" box="[1091,1291,1040,1065]" italics="true" pageId="12" pageNumber="13">C. splendopleure</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957C0496D8722F9438B7AFACDF664" box="[1298,1332,1040,1066]" pageId="12" pageNumber="13" sensu="lato">s.l.</taxonomicNameLabel>
It is distinguished from all remaining
<emphasis id="B96AEABB496D8722FDAA8B52FD2EF61F" box="[507,727,1080,1105]" italics="true" pageId="12" pageNumber="13">Chromphyosemion</emphasis>
species by the unique combination of color patterns.
</paragraph>
</subSubSection>
<caption id="DF616621496D8722FCC68873FF31F51D" httpUri="https://zenodo.org/record/178563/files/figure.png" pageId="12" pageNumber="13" targetBox="[248,1338,1189,1790]" targetPageId="12">
<paragraph id="8BA136A9496D8722FCC68873FF31F51D" blockId="12.[151,1436,1817,1875]" pageId="12" pageNumber="13">
<emphasis id="B96AEABB496D8722FCC68873FEE3F57F" bold="true" box="[151,282,1817,1841]" pageId="12" pageNumber="13">FIGURE 9.</emphasis>
<taxonomicName id="4C1E4D2A496D8722FD718873FDBDF57E" box="[288,580,1817,1840]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496D8722FD718873FDBDF57E" box="[288,580,1817,1840]" italics="true" pageId="12" pageNumber="13">Chromaphyosemion omega</emphasis>
</taxonomicName>
, males from the type locality, not preserved. Photographed directly after collection.
</paragraph>
</caption>
<subSubSection id="C3046522496D8721FC9788C9FB77F0F4" lastPageId="15" lastPageNumber="16" pageId="12" pageNumber="13" type="description">
<paragraph id="8BA136A9496D8723FC9788C9FB12F294" blockId="12.[151,1436,1955,2022]" lastBlockId="13.[151,1436,152,218]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="B96AEABB496D8722FC9788C9FEA6F5F3" bold="true" box="[198,351,1955,1981]" pageId="12" pageNumber="13">Description.</emphasis>
See
<figureCitation id="13252A2C496D8722FDCE88CEFDC2F5F0" box="[415,571,1956,1982]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="12.[151,255,1817,1841]" captionStartId-1="13.[151,255,991,1015]" captionStartId-2="13.[151,255,1779,1803]" captionTargetBox-0="[248,1338,1189,1790]" captionTargetBox-1="[246,1341,277,964]" captionTargetBox-2="[246,1341,1068,1755]" captionTargetId-0="figure@12.[248,1339,1187,1791]" captionTargetId-1="figure@13.[246,1342,277,964]" captionTargetId-2="figure@13.[246,1342,1068,1755]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="13" captionText-0="FIGURE 9. Chromaphyosemion omega, males from the type locality, not preserved. Photographed directly after collection." captionText-1="FIGURE 10. Chromaphyosemion omega, male from the strain C 89 / 35, not preserved. Photo by W. Eigelshofen" captionText-2="FIGURE 11. Chromaphyosemion omega, female from the type locality, not preserved. Photographed after some months in the aquarium. Photo by W. Eigelshofen" httpUri-0="https://zenodo.org/record/178563/files/figure.png" httpUri-1="https://zenodo.org/record/178564/files/figure.png" httpUri-2="https://zenodo.org/record/178565/files/figure.png" pageId="12" pageNumber="13">Figures 9–11</figureCitation>
for general appearance and
<tableCitation id="C69C0312496D8722FFC988CEFC0AF5F0" box="[920,1011,1956,1982]" captionStart="TABLE 3" captionStartId="14.[151,242,157,181]" captionTargetBox="[165,1420,249,1186]" captionTargetPageId="14" captionText="TABLE 3. Morphometrics of Chromaphyosemion omega sp. nov .. All measurements in percents of standard length (SL), except standard length in mm. For abbreviations see Table 2." httpUri="http://table.plazi.org/id/DF616621496F8720FCC68FF7FC99F299" pageId="12" pageNumber="13" tableUuid="DF616621496F8720FCC68FF7FC99F299">Table 3</tableCitation>
for morphometric data of the
<typeStatus id="54A5880B496D8722F93888CEFA65F5F0" box="[1385,1436,1956,1982]" pageId="12" pageNumber="13">type</typeStatus>
series and for general description under
<taxonomicName id="4C1E4D2A496D8722FE2888A6FCCDF5AB" box="[633,820,1996,2021]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="koungueense">
<emphasis id="B96AEABB496D8722FE2888A6FCCDF5AB" box="[633,820,1996,2021]" italics="true" pageId="12" pageNumber="13">C. koungueense</emphasis>
</taxonomicName>
. First ray of dorsal fin in front or above 1st or 2nd anal fin ray. Dorsal fin with 11–13 rays, anal fin with 13–15 rays. Pectoral fins extending over origin of pelvic fins, pelvic fins not or nearly reaching anal fin. Pectoral fin rays dorsally longer then ventrally.
</paragraph>
<caption id="DF616621496C8723FCC68CB5FABCF1B8" box="[151,1349,991,1015]" httpUri="https://zenodo.org/record/178564/files/figure.png" pageId="13" pageNumber="14" targetBox="[246,1341,277,964]" targetPageId="13">
<paragraph id="8BA136A9496C8723FCC68CB5FABCF1B8" blockId="13.[151,1349,991,1015]" box="[151,1349,991,1015]" pageId="13" pageNumber="14">
<emphasis id="B96AEABB496C8723FCC68CB5FEDFF1B9" bold="true" box="[151,294,991,1015]" pageId="13" pageNumber="14">FIGURE 10.</emphasis>
<taxonomicName id="4C1E4D2A496C8723FD7C8CB5FDA8F1B8" box="[301,593,991,1014]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496C8723FD7C8CB5FDA8F1B8" box="[301,593,991,1014]" italics="true" pageId="13" pageNumber="14">Chromaphyosemion omega</emphasis>
</taxonomicName>
, male from the strain C 89/35, not preserved. Photo by W. Eigelshofen
</paragraph>
</caption>
<caption id="DF616621496C8723FCC68999FDACF563" httpUri="https://zenodo.org/record/178565/files/figure.png" pageId="13" pageNumber="14" targetBox="[246,1341,1068,1755]" targetPageId="13">
<paragraph id="8BA136A9496C8723FCC68999FDACF563" blockId="13.[151,1437,1779,1837]" pageId="13" pageNumber="14">
<emphasis id="B96AEABB496C8723FCC68999FEDCF545" bold="true" box="[151,293,1779,1803]" pageId="13" pageNumber="14">FIGURE 11.</emphasis>
<taxonomicName id="4C1E4D2A496C8723FD7D899EFDB6F545" box="[300,591,1780,1803]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496C8723FD7D899EFDB6F545" box="[300,591,1780,1803]" italics="true" pageId="13" pageNumber="14">Chromaphyosemion omega</emphasis>
</taxonomicName>
, female from the type locality, not preserved. Photographed after some months in the aquarium. Photo by W. Eigelshofen
</paragraph>
</caption>
<paragraph id="8BA136A9496C8723FC97880BFD86F5BD" blockId="13.[151,1436,1889,2035]" pageId="13" pageNumber="14">Scales cycloid, body and head entirely scaled, except ventral surface of head. Frontal squamation of Gtype, scales on mid-longitudinal series 23–26 + 2–3 on caudal fin base. Transverse row of scales above pelvic fins 8–9, circum-peduncular scale row 12–13. Open frontal neuromast system with two separate grooves, preopercular neuromast system with 6 pores.</paragraph>
<caption id="DF616621496F8720FCC68FF7FC99F299" ID-Table-UUID="DF616621496F8720FCC68FF7FC99F299" httpUri="http://table.plazi.org/id/DF616621496F8720FCC68FF7FC99F299" pageId="14" pageNumber="15" targetBox="[165,1420,249,1186]" targetIsTable="true" targetPageId="14">
<paragraph id="8BA136A9496F8720FCC68FF7FC99F299" blockId="14.[151,1435,157,215]" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720FCC68FF7FEF6F2FB" bold="true" box="[151,271,157,181]" pageId="14" pageNumber="15">TABLE 3.</emphasis>
Morphometrics of
<taxonomicName id="4C1E4D2A496F8720FDB88FF4FCE8F2FB" box="[489,785,158,181]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="omega" status="sp. nov.">
<emphasis id="B96AEABB496F8720FDB88FF4FCE8F2FB" box="[489,785,158,181]" italics="true" pageId="14" pageNumber="15">Chromaphyosemion omega</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957C0496F8720FF4A8FF7FC97F2FA" box="[795,878,157,180]" pageId="14" pageNumber="15" rank="species">sp. nov</taxonomicNameLabel>
.. All measurements in percents of standard length (SL), except standard length in mm. For abbreviations see Table 2.
</paragraph>
</caption>
<paragraph id="8BA136A9496F8720FCE18E41FAACF6EC" pageId="14" pageNumber="15">
<table id="F91EC409496F78D1FCF48F93FA75F6EC" box="[165,1420,249,1186]" gridcols="7" gridrows="19" pageId="14" pageNumber="15">
<tr id="352E34EB496F78D1FCF48F93FA75F30C" box="[165,1420,249,322]" gridrow="0" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48F93FF20F30C" box="[165,217,249,322]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">SL</th>
<th id="76FF5D97496F78D1FCAA8F93FE9AF30C" box="[251,355,249,322]" gridcol="1" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720FCAA8F91FE9AF35D" bold="true" box="[251,355,251,275]" pageId="14" pageNumber="15">Holotype</emphasis>
31.3
</th>
<th id="76FF5D97496F78D1FDD78F93FDA1F30C" box="[390,600,249,322]" gridcol="2" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720FDD78F97FE01F35B" bold="true" box="[390,504,253,277]" pageId="14" pageNumber="15">Paratypes</emphasis>
ɗ
<emphasis id="B96AEABB496F8720FE4B8F97FDA1F35B" bold="true" box="[538,600,253,277]" pageId="14" pageNumber="15">mean</emphasis>
23.3
</th>
<th id="76FF5D97496F78D1FE2B8F93FCB3F30C" box="[634,842,249,322]" gridcol="3" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720FE2B8F91FD15F35D" bold="true" box="[634,748,251,275]" pageId="14" pageNumber="15">Paratypes</emphasis>
Ψ
<emphasis id="B96AEABB496F8720FF5D8F91FCB3F35D" bold="true" box="[780,842,251,275]" pageId="14" pageNumber="15">mean</emphasis>
21.6
</th>
<th id="76FF5D97496F78D1FF3B8F93FBF6F30C" box="[874,1039,249,322]" gridcol="4" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720FF3B8F91FBF6F35D" bold="true" box="[874,1039,251,275]" pageId="14" pageNumber="15">Paratypes juv.</emphasis>
18.0
</th>
<th id="76FF5D97496F78D1F8618F93FB25F30C" box="[1072,1244,249,322]" gridcol="5" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720F8618F91FB25F35D" bold="true" box="[1072,1244,251,275]" pageId="14" pageNumber="15">all Types mean</emphasis>
22.9
</th>
<th id="76FF5D97496F78D1F8AD8F93FA75F30C" box="[1276,1420,249,322]" gridcol="6" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720F8AD8F91FA75F35D" bold="true" box="[1276,1420,251,275]" pageId="14" pageNumber="15">all Types SD</emphasis>
4.6
</th>
</tr>
<tr id="352E34EB496F78D1FCF48E31FA75F33C" box="[165,1420,347,370]" gridrow="1" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48E31FF20F33C" box="[165,217,347,370]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">TL</th>
<td id="76FF5D97496F78D1FCAA8E31FE9AF33C" box="[251,355,347,370]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">132.7</td>
<td id="76FF5D97496F78D1FDD78E31FDA1F33C" box="[390,600,347,370]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">126.4</td>
<td id="76FF5D97496F78D1FE2B8E31FCB3F33C" box="[634,842,347,370]" gridcol="3" gridrow="1" pageId="14" pageNumber="15">128.4</td>
<td id="76FF5D97496F78D1FF3B8E31FBF6F33C" box="[874,1039,347,370]" gridcol="4" gridrow="1" pageId="14" pageNumber="15">126.6</td>
<td id="76FF5D97496F78D1F8618E31FB25F33C" box="[1072,1244,347,370]" gridcol="5" gridrow="1" pageId="14" pageNumber="15">127.1</td>
<td id="76FF5D97496F78D1F8AD8E31FA75F33C" box="[1276,1420,347,370]" gridcol="6" gridrow="1" pageId="14" pageNumber="15">2.5</td>
</tr>
<tr id="352E34EB496F78D1FCF48EE1FA75F3EC" box="[165,1420,395,418]" gridrow="2" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48EE1FF20F3EC" box="[165,217,395,418]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">PD</th>
<td id="76FF5D97496F78D1FCAA8EE1FE9AF3EC" box="[251,355,395,418]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">59.3</td>
<td id="76FF5D97496F78D1FDD78EE1FDA1F3EC" box="[390,600,395,418]" gridcol="2" gridrow="2" pageId="14" pageNumber="15">59.9</td>
<td id="76FF5D97496F78D1FE2B8EE1FCB3F3EC" box="[634,842,395,418]" gridcol="3" gridrow="2" pageId="14" pageNumber="15">62.9</td>
<td id="76FF5D97496F78D1FF3B8EE1FBF6F3EC" box="[874,1039,395,418]" gridcol="4" gridrow="2" pageId="14" pageNumber="15">63.9</td>
<td id="76FF5D97496F78D1F8618EE1FB25F3EC" box="[1072,1244,395,418]" gridcol="5" gridrow="2" pageId="14" pageNumber="15">60.7</td>
<td id="76FF5D97496F78D1F8AD8EE1FA75F3EC" box="[1276,1420,395,418]" gridcol="6" gridrow="2" pageId="14" pageNumber="15">2.7</td>
</tr>
<tr id="352E34EB496F78D1FCF48ED1FA75F39C" box="[165,1420,443,466]" gridrow="3" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48ED1FF20F39C" box="[165,217,443,466]" gridcol="0" gridrow="3" pageId="14" pageNumber="15">HL</th>
<td id="76FF5D97496F78D1FCAA8ED1FE9AF39C" box="[251,355,443,466]" gridcol="1" gridrow="3" pageId="14" pageNumber="15">28.1</td>
<td id="76FF5D97496F78D1FDD78ED1FDA1F39C" box="[390,600,443,466]" gridcol="2" gridrow="3" pageId="14" pageNumber="15">28.5</td>
<td id="76FF5D97496F78D1FE2B8ED1FCB3F39C" box="[634,842,443,466]" gridcol="3" gridrow="3" pageId="14" pageNumber="15">28.7</td>
<td id="76FF5D97496F78D1FF3B8ED1FBF6F39C" box="[874,1039,443,466]" gridcol="4" gridrow="3" pageId="14" pageNumber="15">30.1</td>
<td id="76FF5D97496F78D1F8618ED1FB25F39C" box="[1072,1244,443,466]" gridcol="5" gridrow="3" pageId="14" pageNumber="15">28.7</td>
<td id="76FF5D97496F78D1F8AD8ED1FA75F39C" box="[1276,1420,443,466]" gridcol="6" gridrow="3" pageId="14" pageNumber="15">1.6</td>
</tr>
<tr id="352E34EB496F78D1FCF48E81FA75F04C" box="[165,1420,491,514]" gridrow="4" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48E81FF20F04C" box="[165,217,491,514]" gridcol="0" gridrow="4" pageId="14" pageNumber="15">pPD</th>
<td id="76FF5D97496F78D1FCAA8E81FE9AF04C" box="[251,355,491,514]" gridcol="1" gridrow="4" pageId="14" pageNumber="15">46.9</td>
<td id="76FF5D97496F78D1FDD78E81FDA1F04C" box="[390,600,491,514]" gridcol="2" gridrow="4" pageId="14" pageNumber="15">47.1</td>
<td id="76FF5D97496F78D1FE2B8E81FCB3F04C" box="[634,842,491,514]" gridcol="3" gridrow="4" pageId="14" pageNumber="15">48.1</td>
<td id="76FF5D97496F78D1FF3B8E81FBF6F04C" box="[874,1039,491,514]" gridcol="4" gridrow="4" pageId="14" pageNumber="15">48.2</td>
<td id="76FF5D97496F78D1F8618E81FB25F04C" box="[1072,1244,491,514]" gridcol="5" gridrow="4" pageId="14" pageNumber="15">47.4</td>
<td id="76FF5D97496F78D1F8AD8E81FA75F04C" box="[1276,1420,491,514]" gridcol="6" gridrow="4" pageId="14" pageNumber="15">1.6</td>
</tr>
<tr id="352E34EB496F78D1FCF48D71FA75F07C" box="[165,1420,539,562]" gridrow="5" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48D71FF20F07C" box="[165,217,539,562]" gridcol="0" gridrow="5" pageId="14" pageNumber="15">pAD</th>
<td id="76FF5D97496F78D1FCAA8D71FE9AF07C" box="[251,355,539,562]" gridcol="1" gridrow="5" pageId="14" pageNumber="15">59.7</td>
<td id="76FF5D97496F78D1FDD78D71FDA1F07C" box="[390,600,539,562]" gridcol="2" gridrow="5" pageId="14" pageNumber="15">59.6</td>
<td id="76FF5D97496F78D1FE2B8D71FCB3F07C" box="[634,842,539,562]" gridcol="3" gridrow="5" pageId="14" pageNumber="15">59.8</td>
<td id="76FF5D97496F78D1FF3B8D71FBF6F07C" box="[874,1039,539,562]" gridcol="4" gridrow="5" pageId="14" pageNumber="15">59.9</td>
<td id="76FF5D97496F78D1F8618D71FB25F07C" box="[1072,1244,539,562]" gridcol="5" gridrow="5" pageId="14" pageNumber="15">59.7</td>
<td id="76FF5D97496F78D1F8AD8D71FA75F07C" box="[1276,1420,539,562]" gridcol="6" gridrow="5" pageId="14" pageNumber="15">2.2</td>
</tr>
<tr id="352E34EB496F78D1FCF48D21FA75F02C" box="[165,1420,587,610]" gridrow="6" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48D21FF20F02C" box="[165,217,587,610]" gridcol="0" gridrow="6" pageId="14" pageNumber="15">HB</th>
<td id="76FF5D97496F78D1FCAA8D21FE9AF02C" box="[251,355,587,610]" gridcol="1" gridrow="6" pageId="14" pageNumber="15">20.8</td>
<td id="76FF5D97496F78D1FDD78D21FDA1F02C" box="[390,600,587,610]" gridcol="2" gridrow="6" pageId="14" pageNumber="15">19.6</td>
<td id="76FF5D97496F78D1FE2B8D21FCB3F02C" box="[634,842,587,610]" gridcol="3" gridrow="6" pageId="14" pageNumber="15">20.0</td>
<td id="76FF5D97496F78D1FF3B8D21FBF6F02C" box="[874,1039,587,610]" gridcol="4" gridrow="6" pageId="14" pageNumber="15">19.2</td>
<td id="76FF5D97496F78D1F8618D21FB25F02C" box="[1072,1244,587,610]" gridcol="5" gridrow="6" pageId="14" pageNumber="15">19.7</td>
<td id="76FF5D97496F78D1F8AD8D21FA75F02C" box="[1276,1420,587,610]" gridcol="6" gridrow="6" pageId="14" pageNumber="15">1.2</td>
</tr>
<tr id="352E34EB496F78D1FCF48D11FA75F0DC" box="[165,1420,635,658]" gridrow="7" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48D11FF20F0DC" box="[165,217,635,658]" gridcol="0" gridrow="7" pageId="14" pageNumber="15">HC</th>
<td id="76FF5D97496F78D1FCAA8D11FE9AF0DC" box="[251,355,635,658]" gridcol="1" gridrow="7" pageId="14" pageNumber="15">12.3</td>
<td id="76FF5D97496F78D1FDD78D11FDA1F0DC" box="[390,600,635,658]" gridcol="2" gridrow="7" pageId="14" pageNumber="15">12.0</td>
<td id="76FF5D97496F78D1FE2B8D11FCB3F0DC" box="[634,842,635,658]" gridcol="3" gridrow="7" pageId="14" pageNumber="15">11.8</td>
<td id="76FF5D97496F78D1FF3B8D11FBF6F0DC" box="[874,1039,635,658]" gridcol="4" gridrow="7" pageId="14" pageNumber="15">11.6</td>
<td id="76FF5D97496F78D1F8618D11FB25F0DC" box="[1072,1244,635,658]" gridcol="5" gridrow="7" pageId="14" pageNumber="15">12.0</td>
<td id="76FF5D97496F78D1F8AD8D11FA75F0DC" box="[1276,1420,635,658]" gridcol="6" gridrow="7" pageId="14" pageNumber="15">0.6</td>
</tr>
<tr id="352E34EB496F78D1FCF48DC1FA75F08C" box="[165,1420,683,706]" gridrow="8" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48DC1FF20F08C" box="[165,217,683,706]" gridcol="0" gridrow="8" pageId="14" pageNumber="15">CL</th>
<td id="76FF5D97496F78D1FCAA8DC1FE9AF08C" box="[251,355,683,706]" gridcol="1" gridrow="8" pageId="14" pageNumber="15">23.6</td>
<td id="76FF5D97496F78D1FDD78DC1FDA1F08C" box="[390,600,683,706]" gridcol="2" gridrow="8" pageId="14" pageNumber="15">23.5</td>
<td id="76FF5D97496F78D1FE2B8DC1FCB3F08C" box="[634,842,683,706]" gridcol="3" gridrow="8" pageId="14" pageNumber="15">24.8</td>
<td id="76FF5D97496F78D1FF3B8DC1FBF6F08C" box="[874,1039,683,706]" gridcol="4" gridrow="8" pageId="14" pageNumber="15">23.8</td>
<td id="76FF5D97496F78D1F8618DC1FB25F08C" box="[1072,1244,683,706]" gridcol="5" gridrow="8" pageId="14" pageNumber="15">23.8</td>
<td id="76FF5D97496F78D1F8AD8DC1FA75F08C" box="[1276,1420,683,706]" gridcol="6" gridrow="8" pageId="14" pageNumber="15">1.1</td>
</tr>
<tr id="352E34EB496F78D1FCF48DB1FA75F0BC" box="[165,1420,731,754]" gridrow="9" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48DB1FF20F0BC" box="[165,217,731,754]" gridcol="0" gridrow="9" pageId="14" pageNumber="15">BD</th>
<td id="76FF5D97496F78D1FCAA8DB1FE9AF0BC" box="[251,355,731,754]" gridcol="1" gridrow="9" pageId="14" pageNumber="15">20.3</td>
<td id="76FF5D97496F78D1FDD78DB1FDA1F0BC" box="[390,600,731,754]" gridcol="2" gridrow="9" pageId="14" pageNumber="15">18.3</td>
<td id="76FF5D97496F78D1FE2B8DB1FCB3F0BC" box="[634,842,731,754]" gridcol="3" gridrow="9" pageId="14" pageNumber="15">16.9</td>
<td id="76FF5D97496F78D1FF3B8DB1FBF6F0BC" box="[874,1039,731,754]" gridcol="4" gridrow="9" pageId="14" pageNumber="15">17.3</td>
<td id="76FF5D97496F78D1F8618DB1FB25F0BC" box="[1072,1244,731,754]" gridcol="5" gridrow="9" pageId="14" pageNumber="15">18.1</td>
<td id="76FF5D97496F78D1F8AD8DB1FA75F0BC" box="[1276,1420,731,754]" gridcol="6" gridrow="9" pageId="14" pageNumber="15">1.3</td>
</tr>
<tr id="352E34EB496F78D1FCF48C61FA75F16C" box="[165,1420,779,802]" gridrow="10" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48C61FF20F16C" box="[165,217,779,802]" gridcol="0" gridrow="10" pageId="14" pageNumber="15">BA</th>
<td id="76FF5D97496F78D1FCAA8C61FE9AF16C" box="[251,355,779,802]" gridcol="1" gridrow="10" pageId="14" pageNumber="15">20.1</td>
<td id="76FF5D97496F78D1FDD78C61FDA1F16C" box="[390,600,779,802]" gridcol="2" gridrow="10" pageId="14" pageNumber="15">19.0</td>
<td id="76FF5D97496F78D1FE2B8C61FCB3F16C" box="[634,842,779,802]" gridcol="3" gridrow="10" pageId="14" pageNumber="15">17.2</td>
<td id="76FF5D97496F78D1FF3B8C61FBF6F16C" box="[874,1039,779,802]" gridcol="4" gridrow="10" pageId="14" pageNumber="15">17.7</td>
<td id="76FF5D97496F78D1F8618C61FB25F16C" box="[1072,1244,779,802]" gridcol="5" gridrow="10" pageId="14" pageNumber="15">18.7</td>
<td id="76FF5D97496F78D1F8AD8C61FA75F16C" box="[1276,1420,779,802]" gridcol="6" gridrow="10" pageId="14" pageNumber="15">1.3</td>
</tr>
<tr id="352E34EB496F78D1FCF48C51FA75F11C" box="[165,1420,827,850]" gridrow="11" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48C51FF20F11C" box="[165,217,827,850]" gridcol="0" gridrow="11" pageId="14" pageNumber="15">E</th>
<td id="76FF5D97496F78D1FCAA8C51FE9AF11C" box="[251,355,827,850]" gridcol="1" gridrow="11" pageId="14" pageNumber="15">8.0</td>
<td id="76FF5D97496F78D1FDD78C51FDA1F11C" box="[390,600,827,850]" gridcol="2" gridrow="11" pageId="14" pageNumber="15">9.2</td>
<td id="76FF5D97496F78D1FE2B8C51FCB3F11C" box="[634,842,827,850]" gridcol="3" gridrow="11" pageId="14" pageNumber="15">9.7</td>
<td id="76FF5D97496F78D1FF3B8C51FBF6F11C" box="[874,1039,827,850]" gridcol="4" gridrow="11" pageId="14" pageNumber="15">9.8</td>
<td id="76FF5D97496F78D1F8618C51FB25F11C" box="[1072,1244,827,850]" gridcol="5" gridrow="11" pageId="14" pageNumber="15">9.3</td>
<td id="76FF5D97496F78D1F8AD8C51FA75F11C" box="[1276,1420,827,850]" gridcol="6" gridrow="11" pageId="14" pageNumber="15">0.5</td>
</tr>
<tr id="352E34EB496F78D1FCF48C01FA75F1CC" box="[165,1420,875,898]" gridrow="12" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48C01FF20F1CC" box="[165,217,875,898]" gridcol="0" gridrow="12" pageId="14" pageNumber="15">I</th>
<td id="76FF5D97496F78D1FCAA8C01FE9AF1CC" box="[251,355,875,898]" gridcol="1" gridrow="12" pageId="14" pageNumber="15">13.1</td>
<td id="76FF5D97496F78D1FDD78C01FDA1F1CC" box="[390,600,875,898]" gridcol="2" gridrow="12" pageId="14" pageNumber="15">11.9</td>
<td id="76FF5D97496F78D1FE2B8C01FCB3F1CC" box="[634,842,875,898]" gridcol="3" gridrow="12" pageId="14" pageNumber="15">12.1</td>
<td id="76FF5D97496F78D1FF3B8C01FBF6F1CC" box="[874,1039,875,898]" gridcol="4" gridrow="12" pageId="14" pageNumber="15">11.6</td>
<td id="76FF5D97496F78D1F8618C01FB25F1CC" box="[1072,1244,875,898]" gridcol="5" gridrow="12" pageId="14" pageNumber="15">12.0</td>
<td id="76FF5D97496F78D1F8AD8C01FA75F1CC" box="[1276,1420,875,898]" gridcol="6" gridrow="12" pageId="14" pageNumber="15">0.8</td>
</tr>
<tr id="352E34EB496F78D1FCF48CF1FA75F1FC" box="[165,1420,923,946]" gridrow="13" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48CF1FF20F1FC" box="[165,217,923,946]" gridcol="0" gridrow="13" pageId="14" pageNumber="15">D</th>
<td id="76FF5D97496F78D1FCAA8CF1FE9AF1FC" box="[251,355,923,946]" gridcol="1" gridrow="13" pageId="14" pageNumber="15">12</td>
<td id="76FF5D97496F78D1FDD78CF1FDA1F1FC" box="[390,600,923,946]" gridcol="2" gridrow="13" pageId="14" pageNumber="15">11.8</td>
<td id="76FF5D97496F78D1FE2B8CF1FCB3F1FC" box="[634,842,923,946]" gridcol="3" gridrow="13" pageId="14" pageNumber="15">11.7</td>
<td id="76FF5D97496F78D1FF3B8CF1FBF6F1FC" box="[874,1039,923,946]" gridcol="4" gridrow="13" pageId="14" pageNumber="15">11.5</td>
<td id="76FF5D97496F78D1F8618CF1FB25F1FC" box="[1072,1244,923,946]" gridcol="5" gridrow="13" pageId="14" pageNumber="15">11.8</td>
<td id="76FF5D97496F78D1F8AD8CF1FA75F1FC" box="[1276,1420,923,946]" gridcol="6" gridrow="13" pageId="14" pageNumber="15">0.7</td>
</tr>
<tr id="352E34EB496F78D1FCF48CA1FA75F1AC" box="[165,1420,971,994]" gridrow="14" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48CA1FF20F1AC" box="[165,217,971,994]" gridcol="0" gridrow="14" pageId="14" pageNumber="15">A</th>
<td id="76FF5D97496F78D1FCAA8CA1FE9AF1AC" box="[251,355,971,994]" gridcol="1" gridrow="14" pageId="14" pageNumber="15">14</td>
<td id="76FF5D97496F78D1FDD78CA1FDA1F1AC" box="[390,600,971,994]" gridcol="2" gridrow="14" pageId="14" pageNumber="15">13.9</td>
<td id="76FF5D97496F78D1FE2B8CA1FCB3F1AC" box="[634,842,971,994]" gridcol="3" gridrow="14" pageId="14" pageNumber="15">13.7</td>
<td id="76FF5D97496F78D1FF3B8CA1FBF6F1AC" box="[874,1039,971,994]" gridcol="4" gridrow="14" pageId="14" pageNumber="15">14.0</td>
<td id="76FF5D97496F78D1F8618CA1FB25F1AC" box="[1072,1244,971,994]" gridcol="5" gridrow="14" pageId="14" pageNumber="15">13.9</td>
<td id="76FF5D97496F78D1F8AD8CA1FA75F1AC" box="[1276,1420,971,994]" gridcol="6" gridrow="14" pageId="14" pageNumber="15">0.5</td>
</tr>
<tr id="352E34EB496F78D1FCF48C91FA75F65C" box="[165,1420,1019,1042]" gridrow="15" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48C91FF20F65C" box="[165,217,1019,1042]" gridcol="0" gridrow="15" pageId="14" pageNumber="15">D/A</th>
<td id="76FF5D97496F78D1FCAA8C91FE9AF65C" box="[251,355,1019,1042]" gridcol="1" gridrow="15" pageId="14" pageNumber="15">-1</td>
<td id="76FF5D97496F78D1FDD78C91FDA1F65C" box="[390,600,1019,1042]" gridcol="2" gridrow="15" pageId="14" pageNumber="15">-0.5</td>
<td id="76FF5D97496F78D1FE2B8C91FCB3F65C" box="[634,842,1019,1042]" gridcol="3" gridrow="15" pageId="14" pageNumber="15">-0.3</td>
<td id="76FF5D97496F78D1FF3B8C91FBF6F65C" box="[874,1039,1019,1042]" gridcol="4" gridrow="15" pageId="14" pageNumber="15">-1.0</td>
<td id="76FF5D97496F78D1F8618C91FB25F65C" box="[1072,1244,1019,1042]" gridcol="5" gridrow="15" pageId="14" pageNumber="15">-0.5</td>
<td id="76FF5D97496F78D1F8AD8C91FA75F65C" box="[1276,1420,1019,1042]" gridcol="6" gridrow="15" pageId="14" pageNumber="15">0.8</td>
</tr>
<tr id="352E34EB496F78D1FCF48B41FA75F60C" box="[165,1420,1067,1090]" gridrow="16" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48B41FF20F60C" box="[165,217,1067,1090]" gridcol="0" gridrow="16" pageId="14" pageNumber="15">LLS</th>
<td id="76FF5D97496F78D1FCAA8B41FE9AF60C" box="[251,355,1067,1090]" gridcol="1" gridrow="16" pageId="14" pageNumber="15">25 +2</td>
<td id="76FF5D97496F78D1FDD78B41FDA1F60C" box="[390,600,1067,1090]" gridcol="2" gridrow="16" pageId="14" pageNumber="15">24.5 + 2.2</td>
<td id="76FF5D97496F78D1FE2B8B41FCB3F60C" box="[634,842,1067,1090]" gridcol="3" gridrow="16" pageId="14" pageNumber="15">24.7 + 2.3</td>
<td id="76FF5D97496F78D1FF3B8B41FBF6F60C" box="[874,1039,1067,1090]" gridcol="4" gridrow="16" pageId="14" pageNumber="15">24.5 + 2.0</td>
<td id="76FF5D97496F78D1F8618B41FB25F60C" box="[1072,1244,1067,1090]" gridcol="5" gridrow="16" pageId="14" pageNumber="15">24.6 + 2.2</td>
<td id="76FF5D97496F78D1F8AD8B41FA75F60C" box="[1276,1420,1067,1090]" gridcol="6" gridrow="16" pageId="14" pageNumber="15">1.0 + 0.4</td>
</tr>
<tr id="352E34EB496F78D1FCF48B31FA75F63C" box="[165,1420,1115,1138]" gridrow="17" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48B31FF20F63C" box="[165,217,1115,1138]" gridcol="0" gridrow="17" pageId="14" pageNumber="15">TS</th>
<td id="76FF5D97496F78D1FCAA8B31FE9AF63C" box="[251,355,1115,1138]" gridcol="1" gridrow="17" pageId="14" pageNumber="15">9</td>
<td id="76FF5D97496F78D1FDD78B31FDA1F63C" box="[390,600,1115,1138]" gridcol="2" gridrow="17" pageId="14" pageNumber="15">8.7</td>
<td id="76FF5D97496F78D1FE2B8B31FCB3F63C" box="[634,842,1115,1138]" gridcol="3" gridrow="17" pageId="14" pageNumber="15">8.3</td>
<td id="76FF5D97496F78D1FF3B8B31FBF6F63C" box="[874,1039,1115,1138]" gridcol="4" gridrow="17" pageId="14" pageNumber="15">8.5</td>
<td id="76FF5D97496F78D1F8618B31FB25F63C" box="[1072,1244,1115,1138]" gridcol="5" gridrow="17" pageId="14" pageNumber="15">8.6</td>
<td id="76FF5D97496F78D1F8AD8B31FA75F63C" box="[1276,1420,1115,1138]" gridcol="6" gridrow="17" pageId="14" pageNumber="15">0.5</td>
</tr>
<tr id="352E34EB496F78D1FCF48BE1FA75F6EC" box="[165,1420,1163,1186]" gridrow="18" pageId="14" pageNumber="15">
<th id="76FF5D97496F78D1FCF48BE1FF20F6EC" box="[165,217,1163,1186]" gridcol="0" gridrow="18" pageId="14" pageNumber="15">CS</th>
<td id="76FF5D97496F78D1FCAA8BE1FE9AF6EC" box="[251,355,1163,1186]" gridcol="1" gridrow="18" pageId="14" pageNumber="15">12</td>
<td id="76FF5D97496F78D1FDD78BE1FDA1F6EC" box="[390,600,1163,1186]" gridcol="2" gridrow="18" pageId="14" pageNumber="15">12.2</td>
<td id="76FF5D97496F78D1FE2B8BE1FCB3F6EC" box="[634,842,1163,1186]" gridcol="3" gridrow="18" pageId="14" pageNumber="15">12.0</td>
<td id="76FF5D97496F78D1FF3B8BE1FBF6F6EC" box="[874,1039,1163,1186]" gridcol="4" gridrow="18" pageId="14" pageNumber="15">12.5</td>
<td id="76FF5D97496F78D1F8618BE1FB25F6EC" box="[1072,1244,1163,1186]" gridcol="5" gridrow="18" pageId="14" pageNumber="15">12.2</td>
<td id="76FF5D97496F78D1F8AD8BE1FA75F6EC" box="[1276,1420,1163,1186]" gridcol="6" gridrow="18" pageId="14" pageNumber="15">0.5</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA136A9496F8720FC978A62FA97F474" blockId="14.[151,1437,1288,2034]" pageId="14" pageNumber="15">
<emphasis id="B96AEABB496F8720FC978A62FEACF76C" bold="true" box="[198,341,1288,1314]" pageId="14" pageNumber="15">Coloration.</emphasis>
Live specimens. Males (
<figureCitation id="13252A2C496F8720FEDC8A62FCEAF76C" box="[653,787,1288,1314]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="12.[151,255,1817,1841]" captionStartId-1="13.[151,255,991,1015]" captionTargetBox-0="[248,1338,1189,1790]" captionTargetBox-1="[246,1341,277,964]" captionTargetId-0="figure@12.[248,1339,1187,1791]" captionTargetId-1="figure@13.[246,1342,277,964]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURE 9. Chromaphyosemion omega, males from the type locality, not preserved. Photographed directly after collection." captionText-1="FIGURE 10. Chromaphyosemion omega, male from the strain C 89 / 35, not preserved. Photo by W. Eigelshofen" httpUri-0="https://zenodo.org/record/178563/files/figure.png" httpUri-1="https://zenodo.org/record/178564/files/figure.png" pageId="14" pageNumber="15">Figs. 9–10</figureCitation>
). Back brownish, scales from point of greatest body height on sides dorsally metallic copper to greenish, sides between black lateral stripes pale greyish with a light blue metallic sheen, ventrally greyish to whitish, from lower head extending ventrally on sides to end of caudal more or less greyish to orange, often also on sides on caudal peduncle behind posterior end of caudal fin base. Yellowish or orange metallic blotch on opercle behind eye between lateral stripes. In fully colored males upper half of sides with metallic copper to greenish color, middle of sides light bluish and lower sides with intensive orange color. Three rows of red dots on scales extending from opercle and dissolve into irregularly distributed red dots on sides, lower row on middle of body splitting in two rows posterior to opercle.
</paragraph>
<paragraph id="8BA136A9496F8720FC978922FAB6F5EC" blockId="14.[151,1437,1288,2034]" pageId="14" pageNumber="15">Dorsal fin greenish or bluish-green, upper half more orange with orange fin streamer. Red dots between fin rays, on upper half of fin often fusing into streaks. Anal fin orange, sometimes basal green or bluish-green band with red dots, anal fin center usually without red markings. Submarginal band red, marginal band blue. Posterior end of fin sometimes with small extension, yellow or bluish. Caudal fin center bluish-green to hyaline orange, in lower part bordered by orange line directly extending into lower fin streamer above red submarginal band, marginal band blue. Upper margin of caudal fin with small blue band, interrupted by red markings; below orange, extending from base of caudal fin straight into upper fin streamer, often conspicuously colored yellow to orange. Red dots and streaks between fin rays. Pelvic fins similarly colored as anal fin, orange with submarginal red and marginal blue bands. Pectoral fins hyaline orange, distally darker.</paragraph>
<paragraph id="8BA136A9496F8721FC9788DAFC73F384" blockId="14.[151,1437,1288,2034]" lastBlockId="15.[151,1437,152,1258]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
Females (
<figureCitation id="13252A2C496F8720FD6B88DAFE6BF584" box="[314,402,1968,1994]" captionStart="FIGURE 11" captionStartId="13.[151,255,1779,1803]" captionTargetBox="[246,1341,1068,1755]" captionTargetId="figure@13.[246,1342,1068,1755]" captionTargetPageId="13" captionText="FIGURE 11. Chromaphyosemion omega, female from the type locality, not preserved. Photographed after some months in the aquarium. Photo by W. Eigelshofen" httpUri="https://zenodo.org/record/178565/files/figure.png" pageId="14" pageNumber="15">Fig. 11</figureCitation>
). Back brown, three dorsolateral rows of scales with metallic copper or greenish color from point of greatest body height to end of caudal peduncle. Scales with darker borders, sometimes with red dots; on sides with two parallel black stripes, between stripes pale greyish or beige, some bluish-green metallic scales on posterior sides, on lower anterior sides orange hue, ventrally light grey to whitish. Some specimens with small red dots on scales, more or less regularly in rows. Orange area on opercle between lateral stripes. All fins hyaline, coloration similar to fins of males except anal fin, greenish, at least lower half orange to yellowish. Dark or red dots in dorsal fin, dots and streaks in caudal fin, in some specimens also red dots in anal fin. Anal and caudal fins with blue marginal band, anal fin with dark red submarginal band. Pelvic fins hyaline greenish in upper half, orange in lower half with bluish margin. Pectoral fins hyaline with slightly dark pigmentation and metallic blue margin on lower distal half.
</paragraph>
<paragraph id="8BA136A9496E8721FC978EB2FB61F024" blockId="15.[151,1437,152,1258]" pageId="15" pageNumber="16">Preserved in ethanol. Males. Back and sides brown, both lateral stripes dark brown, belly beige to whitish. Scales on back and sides with dark brown border forming a reticulated pattern. Border of opercle ventrally of lower stripe dark, area below lower lip whitish to beige. Unpaired and pelvic fins hyaline greyish with dark pigmentation at fin rays, dark margin on unpaired and pelvic fins. Pectoral fins hyaline.</paragraph>
<paragraph id="8BA136A9496E8721FC978D12FB77F0F4" blockId="15.[151,1437,152,1258]" pageId="15" pageNumber="16">Females. Similar to males and juveniles. Body brownish, scales on back with dark border, belly beige to whitish. Fins hyaline greyish, slight dark pigmentation and no dark margin on anal fin.</paragraph>
</subSubSection>
<subSubSection id="C3046522496E8721FC978DADFC0FF114" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BA136A9496E8721FC978DADFC0FF114" blockId="15.[151,1437,152,1258]" pageId="15" pageNumber="16">
<emphasis id="B96AEABB496E8721FC978DADFEAAF0AF" bold="true" box="[198,339,711,737]" pageId="15" pageNumber="16">Etymology.</emphasis>
This species is named with regard to
<taxonomicName id="4C1E4D2A496E8721FF798DA2FB98F0AF" box="[808,1121,712,737]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alpha">
<emphasis id="B96AEABB496E8721FF798DA2FB98F0AF" box="[808,1121,712,737]" italics="true" pageId="15" pageNumber="16">Chromaphyosemion alpha</emphasis>
</taxonomicName>
in the sense of
<taxonomicName id="4C1E4D2A496E8721F9798DA2FA9EF0AC" box="[1320,1383,712,738]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alpha">alpha</taxonomicName>
'the beginning'; from Greek:
<taxonomicName id="4C1E4D2A496E8721FDE98D9AFDFFF144" box="[440,518,752,778]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="omega">omega</taxonomicName>
used in the meaning of 'the end'. This is meant in relation to the relative (phylogenetic) position of both species within
<taxonomicName id="4C1E4D2A496E8721FECE8C72FC73F17F" box="[671,906,792,817]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B96AEABB496E8721FECE8C72FC73F17F" box="[671,906,792,817]" italics="true" pageId="15" pageNumber="16">Chromaphyosemion</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1E4D2A496E8721FF868C72FBB8F17F" box="[983,1089,792,817]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="alpha">
<emphasis id="B96AEABB496E8721FF868C72FBB8F17F" box="[983,1089,792,817]" italics="true" pageId="15" pageNumber="16">C. alpha</emphasis>
</taxonomicName>
as the basal species and
<taxonomicName id="4C1E4D2A496E8721F9D08C72FF1CF117" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496E8721F9D08C72FF1CF117" italics="true" pageId="15" pageNumber="16">C. omega</emphasis>
</taxonomicName>
as a more derived species, which is the result of a recent radiation.
</paragraph>
</subSubSection>
<subSubSection id="C3046522496E8721FC978C0DFB10F6A4" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BA136A9496E8721FC978C0DFC41F604" blockId="15.[151,1437,152,1258]" pageId="15" pageNumber="16">
<emphasis id="B96AEABB496E8721FC978C0DFDF8F1CF" bold="true" box="[198,513,871,897]" pageId="15" pageNumber="16">Distribution and habitat.</emphasis>
<taxonomicName id="4C1E4D2A496E8721FE5B8C02FCB5F1CF" box="[522,844,872,897]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496E8721FE5B8C02FCB5F1CF" box="[522,844,872,897]" italics="true" pageId="15" pageNumber="16">Chromaphyosemion omega</emphasis>
</taxonomicName>
is currently known from several places along the road between Kopongo / Somakek and Bonepoupa (see Map 1). Near Somakek it is found in sympatry with
<emphasis id="B96AEABB496E8721FCC68CD2FEB6F19F" box="[151,335,952,977]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C1E4D2A496E8721FCC68CD2FEB2F19F" box="[151,331,952,977]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="riggenbachi">C. riggenbachi</taxonomicName>
.
</emphasis>
The latter species also occurs in rivers within the distribution range of
<taxonomicName id="4C1E4D2A496E8721F8CD8CD2FAF5F19F" box="[1180,1292,952,977]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496E8721F8CD8CD2FAF5F19F" box="[1180,1292,952,977]" italics="true" pageId="15" pageNumber="16">C. omega</emphasis>
</taxonomicName>
but extends further north, south and east of the latter. To the west and northwest,
<taxonomicName id="4C1E4D2A496E8721FFB98C8AFBA5F1B7" box="[1000,1116,992,1017]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496E8721FFB98C8AFBA5F1B7" box="[1000,1116,992,1017]" italics="true" pageId="15" pageNumber="16">C. omega</emphasis>
</taxonomicName>
is replaced by the closely related
<taxonomicName id="4C1E4D2A496E8721FCA28B62FE4BF66F" box="[243,434,1032,1057]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="koungueense">
<emphasis id="B96AEABB496E8721FCA28B62FE4BF66F" box="[243,434,1032,1057]" italics="true" pageId="15" pageNumber="16">C. koungueense</emphasis>
</taxonomicName>
. Near Bonepoupa both species are found in close vicinity between the localities Mangoule and KV 03/17 (
<taxonomicName id="4C1E4D2A496E8721FD9D8B5AFD7CF607" box="[460,645,1072,1097]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="koungueense">
<emphasis id="B96AEABB496E8721FD9D8B5AFD7CF607" box="[460,645,1072,1097]" italics="true" pageId="15" pageNumber="16">C. koungueense</emphasis>
</taxonomicName>
) and C 01/62 (
<taxonomicName id="4C1E4D2A496E8721FF668B5AFC5EF607" box="[823,935,1072,1097]" family="Nothobranchiidae" genus="Chromaphyosemion" higherTaxonomySource="CoL" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="omega">
<emphasis id="B96AEABB496E8721FF668B5AFC5EF607" box="[823,935,1072,1097]" italics="true" pageId="15" pageNumber="16">C. omega</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA136A9496E8721FCC68B32FB10F6A4" blockId="15.[151,1437,152,1258]" pageId="15" pageNumber="16">
The
<typeStatus id="54A5880B496E8721FC9F8B32FEF8F63C" box="[206,257,1112,1138]" pageId="15" pageNumber="16">type</typeStatus>
locality is a small river in the forest with a maximal depth of approximately
<quantity id="4CE69B4C496E8721F8F58B32FB10F63C" box="[1188,1257,1112,1138]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="m" value="0.2">0.2 m</quantity>
and up to
<quantity id="4CE69B4C496E8721F93C8B32FA65F63C" box="[1389,1436,1112,1138]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="16" unit="m" value="3.0">3 m</quantity>
wide. The bottom consists of sand and gravel and lacking aquatic plants. The water temperature was 26.6 °C, air temperature 30°C,
<collectingCountry id="F3097639496E8721FDCC8BC2FE38F68C" box="[413,449,1192,1218]" name="Philippines" pageId="15" pageNumber="16">pH</collectingCountry>
6 and conductivity ~ 10µS/cm. Syntopic fishes were
<taxonomicName id="4C1E4D2A496E8721F8668BC2FAB7F68F" box="[1079,1358,1192,1217]" class="Actinopterygii" family="Nothobranchiidae" genus="Epiplatys" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="infrafasciatus">
<emphasis id="B96AEABB496E8721F8668BC2FAB7F68F" box="[1079,1358,1192,1217]" italics="true" pageId="15" pageNumber="16">Epiplatys infrafasciatus</emphasis>
</taxonomicName>
, '
<emphasis id="B96AEABB496E8721F9328BC2FEB9F6A7" italics="true" pageId="15" pageNumber="16">Aphyosemion' ahli</emphasis>
,'
<emphasis id="B96AEABB496E8721FD058BBAFDEAF6A7" box="[340,531,1232,1257]" italics="true" pageId="15" pageNumber="16">A.' franzwerneri</emphasis>
,
<taxonomicName id="4C1E4D2A496E8721FE708BBAFD5EF6A7" box="[545,679,1232,1257]" class="Actinopterygii" family="Poeciliidae" genus="Procatopus" kingdom="Animalia" order="Cyprinodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B96AEABB496E8721FE708BBAFD5EF6A7" box="[545,679,1232,1257]" italics="true" pageId="15" pageNumber="16">Procatopus</emphasis>
</taxonomicName>
sp. and an undetermined juvenile clariid species.
</paragraph>
</subSubSection>
</treatment>
</document>
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