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<document id="92FFC3F00D86C562D0CDAF80B9593A21" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF6CCD80FF4CEA32B54FE809" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Herculina polysagittae" docType="treatment" docVersion="8" lastPageNumber="191" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="188" updateTime="1698435542289" updateUser="ExternalLinkService">
<mods:mods id="CC01E870F84C327C6CEAA742D6EFEF69" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="98C5F63529B3FCC3321FAEC379906ABB">
<mods:title id="1443779BD921803F2ADFAB297053D336">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="AFB615110914BBCDF00857F61474AE65" type="personal">
<mods:role id="2E3323B063BFB50A32AAFA63361EE2F5">
<mods:roleTerm id="9D9F5740AFCDA403C0F79024661C7BF1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B4BBAA76596B7F80F742AEFF96993903">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="82295F7349BA040215FFCDE06F544D6A" type="personal">
<mods:role id="24AA71C68D49B8D446365B571BE96CA6">
<mods:roleTerm id="5904F28D1844C7BF2DE8D8A8CD076910">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="063A8E734D65B549D63C2BD69FCC35E4">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9513EC8FCAE4288D4D897E50208C135E">text</mods:typeOfResource>
<mods:relatedItem id="CC1CAC7C66E8FB0ADF8B493528BC08FF" type="host">
<mods:titleInfo id="1D9889323341BA04F65AA34F69D99727">
<mods:title id="D10F8D5BD6679C53B6F1802C916FB678">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="366E554385FBB0B23455530F949ED0CD">
<mods:date id="1CEF119C755FB1F3A4B773905BC55CAD">2016</mods:date>
<mods:detail id="1C375DA9990F7FA65C16B526B9392982" type="volume">
<mods:number id="DA55029A35CFB31C7040DDF150133805">4211</mods:number>
</mods:detail>
<mods:detail id="5DA3711ED0E929CDD23C0C88D29C2FA2" type="issue">
<mods:number id="A5AF1179BCF9A78C596FA79EC9D41EBB">1</mods:number>
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<mods:extent id="3AA689A4AD16E41EB51625B0EA0E3924" unit="page">
<mods:start id="1EE6C04C6E46710678D6D62CD2BEB52E">1</mods:start>
<mods:end id="92BFB2A62598E5A8818DD8B3F9D0BD19">205</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="667E2A78374CA920BF9D9A636D3BA7D2">journal article</mods:classification>
<mods:identifier id="2B4200D149E182600C56845307A85143" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="6F47E142ADFDD06B103E01A9AF4977AD" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="56A070A2D274EDD76865A6FC385478C0" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
<mods:identifier id="4D85519B51F7CBE228F40FDC8EB5A66F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="45B8C0A4962242B693A8FABDDCDAB0C2" type="Zenodo-Dep">272750</mods:identifier>
<mods:identifier id="ECA3681301F5EDBC58F846A07FC64F12" type="ZooBank">6B86C6BA-6AFE-4AAD-870D-04794C138D47</mods:identifier>
</mods:mods>
<treatment id="03B7878CFF6CCD80FF4CEA32B54FE809" ID-DOI="http://doi.org/10.5281/zenodo.6074501" ID-GBIF-Taxon="127419174" ID-Zenodo-Dep="6074501" LSID="urn:lsid:plazi:treatment:03B7878CFF6CCD80FF4CEA32B54FE809" httpUri="http://treatment.plazi.org/id/03B7878CFF6CCD80FF4CEA32B54FE809" lastPageId="190" lastPageNumber="191" pageId="187" pageNumber="188">
<subSubSection id="C3046511FF6CCD85FF4CEA32B043EBCA" pageId="187" pageNumber="188" type="nomenclature">
<paragraph id="8BA1369AFF6CCD85FF4CEA32B315EB2B" blockId="187.[151,614,583,643]" box="[151,614,583,610]" pageId="187" pageNumber="188">
<heading id="D0E981F6FF6CCD85FF4CEA32B315EB2B" bold="true" box="[151,614,583,610]" fontSize="11" level="1" pageId="187" pageNumber="188" reason="1">
<emphasis id="B96AEA88FF6CCD85FF4CEA32B315EB2B" bold="true" box="[151,614,583,610]" pageId="187" pageNumber="188">
<taxonomicName id="4C1E4D19FF6CCD85FF4CEA32B0DAEB28" ID-CoL="849BF" box="[151,425,583,609]" class="Diplopoda" family="Anthroleucosomatidae" genus="Herculina" kingdom="Animalia" order="Chordeumatida" pageId="187" pageNumber="188" phylum="Arthropoda" rank="species" species="polysagittae">
<emphasis id="B96AEA88FF6CCD85FF4CEA32B0DAEB28" bold="true" box="[151,425,583,609]" italics="true" pageId="187" pageNumber="188">Herculina polysagittae</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF6CCD85FF4CEA1FB043EBCA" blockId="187.[151,614,583,643]" box="[151,304,618,643]" pageId="187" pageNumber="188">
<figureCitation id="13252A1FFF6CCD85FF4CEA1FB043EBCA" box="[151,304,618,643]" captionStart-0="FIGURE 157" captionStart-1="FIGURE 158" captionStart-2="FIGURE 159" captionStartId-0="187.[151,250,1986,2008]" captionStartId-1="188.[151,250,1234,1256]" captionStartId-2="189.[151,250,1671,1693]" captionTargetBox-0="[188,1398,1020,1965]" captionTargetBox-1="[198,1388,183,1213]" captionTargetBox-2="[253,1329,408,1618]" captionTargetId-0="figure@187.[188,1398,1020,1965]" captionTargetId-1="figure@188.[198,1388,183,1213]" captionTargetId-2="figure@189.[188,1398,373,1665]" captionTargetPageId-0="187" captionTargetPageId-1="188" captionTargetPageId-2="189" captionText-0="FIGURE 157. Herculina polysagittae gen. et sp. nov., paratype female habitus. Scale line = 1 mm." captionText-1="FIGURE 158. Herculina polysagittae gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 6. D. Leg 7. E. Leg 10. F. Leg 11. Scale lines = 0.2 mm." captionText-2="FIGURE 159. Herculina polysagittae gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, anterior view. c = coxite; t = telopodite; af = arrow-like filaments; cp = coxal process; cv = coxal vesicle; dl = denticulate lamella = ll = lateral lamella; ml = mesal lamella; sp = sternal plate; ss = sternal sac; lol = longitudinal lamella. Scale line = 0.3 mm." httpUri-0="https://zenodo.org/record/272906/files/figure.png" httpUri-1="https://zenodo.org/record/272907/files/figure.png" httpUri-2="https://zenodo.org/record/272908/files/figure.png" pageId="187" pageNumber="188">Figs 157–159</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF6CCD85FF4CEAC7B0CEEA5A" pageId="187" pageNumber="188" type="diagnosis">
<paragraph id="8BA1369AFF6CCD85FF4CEAC7B0CEEA5A" blockId="187.[151,1436,690,967]" pageId="187" pageNumber="188">
<emphasis id="B96AEA88FF6CCD85FF4CEAC7B062EB82" bold="true" box="[151,273,690,715]" pageId="187" pageNumber="188">Diagnosis.</emphasis>
Differs from the only other known congener,
<taxonomicName id="4C1E4D19FF6CCD85FCCAEAC1B2B6EB82" box="[785,965,690,715]" class="Diplopoda" family="Anthroleucosomatidae" genus="Herculina" kingdom="Animalia" order="Chordeumatida" pageId="187" pageNumber="188" phylum="Arthropoda" rank="species" species="oligosagittae">
<emphasis id="B96AEA88FF6CCD85FCCAEAC1B2B6EB82" box="[785,965,690,715]" italics="true" pageId="187" pageNumber="188">H. oligosagittae</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF6CCD85FC17EAC7B50AEB82" bold="true" box="[972,1145,690,715]" pageId="187" pageNumber="188">gen. et sp. nov.</emphasis>
, by the shape of the coxal processes of the anterior gonopods, and by the presence of more numerous arrow-shaped filaments on the coxites of the posterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF6CCD85FF1CEB6BB078EA12" pageId="187" pageNumber="188" type="etymology">
<paragraph id="8BA1369AFF6CCD85FF1CEB6BB078EA12" blockId="187.[151,1436,690,967]" pageId="187" pageNumber="188">
<emphasis id="B96AEA88FF6CCD85FF1CEB6BB03DEA7E" bold="true" box="[199,334,798,823]" pageId="187" pageNumber="188">Etymology.</emphasis>
To emphasize the presence of numerous arrow-shaped filaments on the coxites of the posterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF6CCD82FF1CEB13B558EC0E" lastPageId="188" lastPageNumber="189" pageId="187" pageNumber="188" type="materials_examined">
<paragraph id="8BA1369AFF6CCD85FF1CEB13B2BAEAEA" blockId="187.[151,1436,690,967]" pageId="187" pageNumber="188">
<materialsCitation id="3B763CC7FF6CCD85FF1CEB13B2BAEAEA" ID-GBIF-Occurrence="1424566973" collectingDate="1983-05-13" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" elevation="1750" location="Material" municipality="Bakuriani" pageId="187" pageNumber="188" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B96AEA88FF6CCD85FF1CEB13B0FFEA36" bold="true" box="[199,396,870,895]" pageId="187" pageNumber="188">
<location id="8EC16041FF6CCD85FF1CEB13B05CEA36" LSID="urn:lsid:plazi:treatment:03B7878CFF6CCD80FF4CEA32B54FE809:8EC16041FF6CCD85FF1CEB13B05CEA36" box="[199,303,870,895]" country="Georgia" municipality="Bakuriani" name="Material" pageId="187" pageNumber="188">Material</location>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FF6CCD85FE07EB12B359EA36" box="[476,554,870,895]" pageId="187" pageNumber="188" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF6CCD85FDEEEB12B3E9EA36" box="[565,666,870,895]" pageId="187" pageNumber="188" type="female">1 female</specimenCount>
).
<emphasis id="B96AEA88FF6CCD85FD77EB13B253EA36" bold="true" box="[684,800,870,895]" pageId="187" pageNumber="188">
<typeStatus id="54A58838FF6CCD85FD77EB13B268EA36" box="[684,795,870,895]" pageId="187" pageNumber="188" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFF6CCD85FCFDEB12B2DAEA36" box="[806,937,871,895]" name="Georgia" pageId="187" pageNumber="188">GEORGIA</collectingCountry>
: male, S of
<collectingMunicipality id="6BC5ACE0FF6CCD85FBF1EB13B5E8EA36" box="[1066,1179,870,895]" pageId="187" pageNumber="188">Bakuriani</collectingMunicipality>
,
<taxonomicName id="4C1E4D19FF6CCD85FB7DEB12B595EA37" box="[1190,1254,871,894]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="187" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF6CCD85FB7DEB12B595EA37" box="[1190,1254,871,894]" italics="true" pageId="187" pageNumber="188">Pinus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D19FF6CCD85FAD2EB12B43CEA37" box="[1289,1359,871,894]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="187" pageNumber="188" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF6CCD85FAD2EB12B43CEA37" box="[1289,1359,871,894]" italics="true" pageId="187" pageNumber="188">Fagus</emphasis>
</taxonomicName>
forest,
<elevation id="0033D1A9FF6CCD85FF4CEBFFB19FEAEB" box="[151,236,906,931]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="187" pageNumber="188" unit="m" value="1750.0">
<quantity id="4CE69B7FFF6CCD85FF4CEBFFB19FEAEB" box="[151,236,906,931]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="187" pageNumber="188" unit="m" value="1750.0">1750 m</quantity>
</elevation>
asl, litter, logs,
<collectingDate id="EFE4E9B2FF6CCD85FE45EBFEB345EAEA" box="[414,566,907,931]" pageId="187" pageNumber="188" value="1983-05-13">
<date id="FFA0105AFF6CCD85FE45EBFEB345EAEA" box="[414,566,907,931]" pageId="187" pageNumber="188" value="1983-05-13">13 May 1983</date>
</collectingDate>
,
<collectorName id="26EB534CFF6CCD85FD9AEBFEB3A7EAEA" box="[577,724,906,931]" pageId="187" pageNumber="188">S. Golovatch</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF6CCD85FCC9EBFEB218EAEA" box="[786,875,907,931]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="187" pageNumber="188">ZMUM</collectionCode>
ρ3437).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF6CCD85FF1CEBDBB5C6EA8E" blockId="187.[151,1436,690,967]" box="[199,1205,942,967]" pageId="187" pageNumber="188">
<emphasis id="B96AEA88FF6CCD85FF1CEBDBB047EA8E" bold="true" box="[199,308,942,967]" pageId="187" pageNumber="188">Paratype</emphasis>
(total: 1 female).
<materialsCitation id="3B763CC7FF6CCD85FE26EBDAB5C6EA8E" ID-GBIF-Occurrence="1424566975" box="[509,1205,942,967]" collectingDate="1983-05-13" collectionCode="ZMUM" collectorName="S. Golovatch" country="Georgia" elevation="1750" location="Material" municipality="Bakuriani" pageId="187" pageNumber="188" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F309760AFF6CCD85FE26EBDAB3F3EA8E" box="[509,640,943,967]" name="Georgia" pageId="187" pageNumber="188">GEORGIA</collectingCountry>
:
<specimenCount id="9D18FD13FF6CCD85FD51EBDAB39DEA8E" box="[650,750,942,967]" pageId="187" pageNumber="188" type="female">1 female</specimenCount>
, same data as holotype (
<collectionCode id="ED0FAE5FFF6CCD85FC25EBDAB524EA8E" box="[1022,1111,943,967]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="187" pageNumber="188">ZMUM</collectionCode>
ρ3438).
</materialsCitation>
</paragraph>
<caption id="DF616612FF6CCD85FF4CEFB7B5FBEE91" box="[151,1160,1986,2008]" httpUri="https://zenodo.org/record/272906/files/figure.png" pageId="187" pageNumber="188" targetBox="[188,1398,1020,1965]" targetPageId="187">
<paragraph id="8BA1369AFF6CCD85FF4CEFB7B5FBEE91" blockId="187.[151,1160,1986,2008]" box="[151,1160,1986,2008]" pageId="187" pageNumber="188">
<emphasis id="B96AEA88FF6CCD85FF4CEFB7B05FEE9E" bold="true" box="[151,300,1986,2008]" pageId="187" pageNumber="188">FIGURE 157.</emphasis>
<taxonomicName id="4C1E4D19FF6CCD85FEE9EFB7B369EE91" box="[306,538,1986,2008]" class="Diplopoda" family="Anthroleucosomatidae" genus="Herculina" kingdom="Animalia" order="Chordeumatida" pageId="187" pageNumber="188" phylum="Arthropoda" rank="species" species="polysagittae">
<emphasis id="B96AEA88FF6CCD85FEE9EFB7B369EE91" box="[306,538,1986,2008]" italics="true" pageId="187" pageNumber="188">Herculina polysagittae</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF6CCD85FDFBEFB7B3CEEE9E" bold="true" box="[544,701,1986,2007]" pageId="187" pageNumber="188">gen. et sp. nov.</emphasis>
, paratype female habitus. Scale line = 1 mm.
</paragraph>
</caption>
<caption id="DF616612FF6BCD82FF4CECA7B0B5EC4E" httpUri="https://zenodo.org/record/272907/files/figure.png" pageId="188" pageNumber="189" targetBox="[198,1388,183,1213]" targetPageId="188">
<paragraph id="8BA1369AFF6BCD82FF4CECA7B0B5EC4E" blockId="188.[151,1434,1234,1287]" pageId="188" pageNumber="189">
<emphasis id="B96AEA88FF6BCD82FF4CECA7B05EEDAE" bold="true" box="[151,301,1234,1256]" pageId="188" pageNumber="189">FIGURE 158.</emphasis>
<taxonomicName id="4C1E4D19FF6BCD82FEEFECA7B36EEDA1" box="[308,541,1234,1256]" class="Diplopoda" family="Anthroleucosomatidae" genus="Herculina" kingdom="Animalia" order="Chordeumatida" pageId="188" pageNumber="189" phylum="Arthropoda" rank="species" species="polysagittae">
<emphasis id="B96AEA88FF6BCD82FEEFECA7B36EEDA1" box="[308,541,1234,1256]" italics="true" pageId="188" pageNumber="189">Herculina polysagittae</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF6BCD82FDFFECA7B3B7EDAE" bold="true" box="[548,708,1234,1255]" pageId="188" pageNumber="189">gen. et sp. nov.</emphasis>
, holotype male.
<emphasis id="B96AEA88FF6BCD82FCAAECA7B2F9EDAE" bold="true" box="[881,906,1234,1255]" pageId="188" pageNumber="189">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FF6BCD82FC00ECA7B281EDAE" bold="true" box="[987,1010,1234,1255]" pageId="188" pageNumber="189">B.</emphasis>
Leg 4.
<emphasis id="B96AEA88FF6BCD82FB99ECA7B529EDA1" bold="true" box="[1090,1114,1234,1256]" pageId="188" pageNumber="189">C.</emphasis>
Leg 6.
<emphasis id="B96AEA88FF6BCD82FB70ECA7B5B7EDAE" bold="true" box="[1195,1220,1234,1255]" pageId="188" pageNumber="189">D.</emphasis>
Leg 7.
<emphasis id="B96AEA88FF6BCD82FACFECA7B45FEDAE" bold="true" box="[1300,1324,1234,1255]" pageId="188" pageNumber="189">E.</emphasis>
Leg 10.
<emphasis id="B96AEA88FF6BCD82FA53ECA7B4E9EDAE" bold="true" box="[1416,1434,1234,1255]" pageId="188" pageNumber="189">F.</emphasis>
Leg 11. Scale lines = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF6BCD82FF1CED5BB558EC0E" blockId="188.[151,1437,1326,2035]" box="[199,1067,1326,1351]" pageId="188" pageNumber="189">
<materialsCitation id="3B763CC7FF6BCD82FF1CED5BB558EC0E" ID-GBIF-Occurrence="1424567003" box="[199,1067,1326,1351]" country="Georgia" elevation="1750" location="Bakuriani" pageId="188" pageNumber="189" specimenCount="1" typeStatus="holotype">
<emphasis id="B96AEA88FF6BCD82FF1CED5BB016EC0E" bold="true" box="[199,357,1326,1351]" pageId="188" pageNumber="189">
<typeStatus id="54A58838FF6BCD82FF1CED5BB071EC0E" box="[199,258,1326,1351]" pageId="188" pageNumber="189">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFF6BCD82FEB7ED5AB09CEC0E" box="[364,495,1327,1351]" name="Georgia" pageId="188" pageNumber="189">GEORGIA</collectingCountry>
: S of
<location id="8EC16041FF6BCD82FDF6ED5BB3EDEC0E" LSID="urn:lsid:plazi:treatment:03B7878CFF6CCD80FF4CEA32B54FE809:8EC16041FF6BCD82FDF6ED5BB3EDEC0E" box="[557,670,1326,1351]" country="Georgia" name="Bakuriani" pageId="188" pageNumber="189">Bakuriani</location>
,
<taxonomicName id="4C1E4D19FF6BCD82FD71ED5AB399EC0F" box="[682,746,1327,1350]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="188" pageNumber="189" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF6BCD82FD71ED5AB399EC0F" box="[682,746,1327,1350]" italics="true" pageId="188" pageNumber="189">Pinus</emphasis>
</taxonomicName>
&amp;
<taxonomicName id="4C1E4D19FF6BCD82FCD6ED5AB220EC0F" box="[781,851,1327,1350]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="188" pageNumber="189" phylum="Tracheophyta" rank="genus">
<emphasis id="B96AEA88FF6BCD82FCD6ED5AB220EC0F" box="[781,851,1327,1350]" italics="true" pageId="188" pageNumber="189">Fagus</emphasis>
</taxonomicName>
forest,
<elevation id="0033D1A9FF6BCD82FC73ED5BB28EEC0F" box="[936,1021,1326,1351]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="188" pageNumber="189" unit="m" value="1750.0">
<quantity id="4CE69B7FFF6BCD82FC73ED5BB28EEC0F" box="[936,1021,1326,1351]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="188" pageNumber="189" unit="m" value="1750.0">1750 m</quantity>
</elevation>
asl.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF6BCD80FF1CED27B029E854" lastPageId="190" lastPageNumber="191" pageId="188" pageNumber="189" type="description">
<paragraph id="8BA1369AFF6BCD82FF1CED27B2D1EC22" blockId="188.[151,1437,1326,2035]" box="[199,930,1362,1387]" pageId="188" pageNumber="189">
<emphasis id="B96AEA88FF6BCD82FF1CED27B02AEC22" bold="true" box="[199,345,1362,1387]" pageId="188" pageNumber="189">Description.</emphasis>
Body with 31 segments (including telson) in adults.
</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CED02B2A5ECFB" blockId="188.[151,1437,1326,2035]" pageId="188" pageNumber="189">
MEASUREMENTS.
<typeStatus id="54A58838FF6BCD82FE48ED03B088ECC6" box="[403,507,1398,1423]" pageId="188" pageNumber="189" type="holotype">Holotype</typeStatus>
male
<quantity id="4CE69B7FFF6BCD82FD9FED02B3F4ECC7" box="[580,647,1399,1423]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="188" pageNumber="189" unit="mm" value="9.0">9 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF6BCD82FB09ED02B45EECC7" box="[1234,1325,1399,1423]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="188" pageNumber="189" unit="mm" value="0.7">0.7 mm</quantity>
.
<typeStatus id="54A58838FF6BCD82FAE1ED02B4EFECC6" box="[1338,1436,1399,1423]" pageId="188" pageNumber="189" type="paratype">Paratype</typeStatus>
female
<quantity id="4CE69B7FFF6BCD82FF31EDEFB033ECFB" box="[234,320,1434,1459]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="188" pageNumber="189" unit="mm" value="9.5">9.5 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF6BCD82FCA3EDEEB2A1ECFB" box="[888,978,1435,1459]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="188" pageNumber="189" unit="mm" value="0.7">0.7 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CEDCAB219EC9E" blockId="188.[151,1437,1326,2035]" box="[199,874,1470,1495]" pageId="188" pageNumber="189">
COLORATION (
<figureCitation id="13252A1FFF6BCD82FEB7EDCBB0BEEC9E" box="[364,461,1470,1495]" captionStart="FIGURE 157" captionStartId="187.[151,250,1986,2008]" captionTargetBox="[188,1398,1020,1965]" captionTargetId="figure@187.[188,1398,1020,1965]" captionTargetPageId="187" captionText="FIGURE 157. Herculina polysagittae gen. et sp. nov., paratype female habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272906/files/figure.png" pageId="188" pageNumber="189">Fig. 157</figureCitation>
). Pale yellowish, with a darker head.
</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CED96B5EFEFE6" blockId="188.[151,1437,1326,2035]" pageId="188" pageNumber="189">
HEAD. Flattened in male. Labrum with three medial teeth and 4+4 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 4+4 setae, on each plate arranged in one row. Stipites with
<emphasis id="B96AEA88FF6BCD82FAE9EE72B43EEF57" box="[1330,1357,1543,1566]" italics="true" pageId="188" pageNumber="189">ca</emphasis>
17+17 setae. Antennae
<quantity id="4CE69B7FFF6BCD82FE95EE5EB0D7EF0B" box="[334,420,1579,1603]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="188" pageNumber="189" unit="mm" value="1.4">1.4 mm</quantity>
long in
<typeStatus id="54A58838FF6BCD82FE24EE5FB316EF0A" box="[511,613,1578,1603]" pageId="188" pageNumber="189" type="holotype">holotype</typeStatus>
. Length of antennomeres (in mm): I (0.08), II (0.16), III (0.36), IV (0.19), V (0.32), VI (0.16), VII (0.11) and VIII (0.02). Length/breadth ratios of antennomeres I–VII: I (1), II (2), III (4), IV (2), V (3), VI (1.6) and VII (1.2). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 9–11, arranged in 4 rows in male;
<quantity id="4CE69B7FFF6BCD82FCABEEE2B2A1EFE6" box="[880,978,1686,1711]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" metricValueMax="3.302" metricValueMin="2.794" pageId="188" pageNumber="189" unit="in" value="12.0" valueMax="13.0" valueMin="11.0">11–13 in</quantity>
4 rows in female.
</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CEECEB189EFBF" blockId="188.[151,1437,1326,2035]" pageId="188" pageNumber="189">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CEF76B4F9EE76" blockId="188.[151,1437,1326,2035]" pageId="188" pageNumber="189">
BODY SEGMENTS (
<figureCitation id="13252A1FFF6BCD82FE7EEF77B37CEE52" box="[421,527,1794,1819]" captionStart="FIGURE 157" captionStartId="187.[151,250,1986,2008]" captionTargetBox="[188,1398,1020,1965]" captionTargetId="figure@187.[188,1398,1020,1965]" captionTargetPageId="187" captionText="FIGURE 157. Herculina polysagittae gen. et sp. nov., paratype female habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272906/files/figure.png" pageId="188" pageNumber="189">Fig. 157</figureCitation>
). Lateral keels like lateral swellings. Macrochaetae long and trichoid. CIX (pleurotergite 15) ~ 0.85; MIX (pleurotergite 15) ~ 2; PIX (pleurotergite 15) ~ 0.5; MA (pleurotergite 15) ~ 110˚.
</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CEF3EB39FEECF" blockId="188.[151,1437,1326,2035]" pageId="188" pageNumber="189">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<paragraph id="8BA1369AFF6BCD82FF1CEFE6B2D5EEBA" blockId="188.[151,1437,1326,2035]" pageId="188" pageNumber="189">WALKING LEGS. In both sexes, leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae. Leg-pairs 12–15 each with a small, inner, subtriangular, coxal protrusion in males (female not examined).</paragraph>
<paragraph id="8BA1369AFF6ACD83FF1CE8EDB58FE809" blockId="189.[151,1437,151,320]" pageId="189" pageNumber="190">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFF6ACD83FDC0E8E2B3F3E9F9" box="[539,640,151,176]" captionStart="FIGURE 158" captionStartId="188.[151,250,1234,1256]" captionTargetBox="[198,1388,183,1213]" captionTargetId="figure@188.[198,1388,183,1213]" captionTargetPageId="188" captionText="FIGURE 158. Herculina polysagittae gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 6. D. Leg 7. E. Leg 10. F. Leg 11. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272907/files/figure.png" pageId="189" pageNumber="190">Fig. 158</figureCitation>
). Leg-pairs 3–7 enlarged, especially so leg-pairs 5 and 6. Leg-pairs 3 and 4 each with a basal exterior protrusion on prefemur. Leg-pairs 6 the most robust, bizarre, femora extremly enlarged, tarsi with a basal inner protrusion. Leg-pair 5 similar to leg-pair 6, but with smaller podomeres. Leg-pair 7 with a posterior, hook-shaped, coxal horn. Leg-pair 10 with coxal glands and a short subtriangular coxal protrusion. Leg-pair 11 with coxal glands, and an exterior and an interior, rounded, coxal protrusion.
</paragraph>
<caption id="DF616612FF6ACD83FF4CEEF2B503EFB0" httpUri="https://zenodo.org/record/272908/files/figure.png" pageId="189" pageNumber="190" targetBox="[253,1329,408,1618]" targetPageId="189">
<paragraph id="8BA1369AFF6ACD83FF4CEEF2B503EFB0" blockId="189.[151,1435,1671,1785]" pageId="189" pageNumber="190">
<emphasis id="B96AEA88FF6ACD83FF4CEEF2B043EFD5" bold="true" box="[151,304,1671,1693]" pageId="189" pageNumber="190">FIGURE 159.</emphasis>
<taxonomicName id="4C1E4D19FF6ACD83FEE2EEF2B356EFD4" box="[313,549,1671,1693]" class="Diplopoda" family="Anthroleucosomatidae" genus="Herculina" kingdom="Animalia" order="Chordeumatida" pageId="189" pageNumber="190" phylum="Arthropoda" rank="species" species="polysagittae">
<emphasis id="B96AEA88FF6ACD83FEE2EEF2B356EFD4" box="[313,549,1671,1693]" italics="true" pageId="189" pageNumber="190">Herculina polysagittae</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF6ACD83FDF5EEF2B3A5EFD5" bold="true" box="[558,726,1671,1692]" pageId="189" pageNumber="190">gen. et sp. nov.</emphasis>
, holotype male gonopods.
<emphasis id="B96AEA88FF6ACD83FC2CEEF2B563EFD5" bold="true" box="[1015,1040,1671,1692]" pageId="189" pageNumber="190">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FF6ACD83FA5FEEF2B4E8EFD5" bold="true" box="[1412,1435,1671,1692]" pageId="189" pageNumber="190">B.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FF6ACD83FE37EED0B377EFF2" bold="true" box="[492,516,1701,1723]" pageId="189" pageNumber="190">C.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FF6ACD83FCA7EED3B2E6EFF2" bold="true" box="[892,917,1702,1723]" pageId="189" pageNumber="190">D.</emphasis>
Posterior gonopods, anterior view.
<emphasis id="B96AEA88FF6ACD83FADCEED2B461EFF5" box="[1287,1298,1703,1724]" italics="true" pageId="189" pageNumber="190">c</emphasis>
= coxite;
<emphasis id="B96AEA88FF6ACD83FAA5EED2B4F6EFF5" box="[1406,1413,1703,1724]" italics="true" pageId="189" pageNumber="190">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FF6ACD83FED1EEB1B06DEF93" box="[266,286,1732,1754]" italics="true" pageId="189" pageNumber="190">af</emphasis>
= arrow-like filaments;
<emphasis id="B96AEA88FF6ACD83FDCFEEB0B35FEF93" box="[532,556,1733,1754]" italics="true" pageId="189" pageNumber="190">cp</emphasis>
= coxal process;
<emphasis id="B96AEA88FF6ACD83FD04EEB0B385EF93" box="[735,758,1733,1754]" italics="true" pageId="189" pageNumber="190">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FF6ACD83FC79EEB1B2C6EF93" box="[930,949,1732,1754]" italics="true" pageId="189" pageNumber="190">dl</emphasis>
= denticulate lamella=
<emphasis id="B96AEA88FF6ACD83FB7DEEB1B5C7EF93" box="[1190,1204,1732,1754]" italics="true" pageId="189" pageNumber="190">ll</emphasis>
= lateral lamella;
<emphasis id="B96AEA88FF6ACD83FAB5EEB1B4F4EF93" box="[1390,1415,1732,1754]" italics="true" pageId="189" pageNumber="190">ml</emphasis>
= mesal lamella;
<emphasis id="B96AEA88FF6ACD83FEEBEE91B034EFB0" box="[304,327,1764,1785]" italics="true" pageId="189" pageNumber="190">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF6ACD83FE31EE91B08DEFB0" box="[490,510,1764,1785]" italics="true" pageId="189" pageNumber="190">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF6ACD83FD4BEE96B3D8EFB0" box="[656,683,1763,1785]" italics="true" pageId="189" pageNumber="190">lol</emphasis>
= longitudinal lamella. Scale line = 0.3 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF6ACD83FF1CEF53B0DFEEBB" blockId="189.[151,1436,1829,2034]" pageId="189" pageNumber="190">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFF6ACD83FE0CEF50B34EEE77" box="[471,573,1829,1854]" captionStart="FIGURE 159" captionStartId="189.[151,250,1671,1693]" captionTargetBox="[253,1329,408,1618]" captionTargetId="figure@189.[188,1398,373,1665]" captionTargetPageId="189" captionText="FIGURE 159. Herculina polysagittae gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, anterior view. c = coxite; t = telopodite; af = arrow-like filaments; cp = coxal process; cv = coxal vesicle; dl = denticulate lamella = ll = lateral lamella; ml = mesal lamella; sp = sternal plate; ss = sternal sac; lol = longitudinal lamella. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272908/files/figure.png" pageId="189" pageNumber="190">Fig. 159</figureCitation>
A–C). Sternal plate (
<emphasis id="B96AEA88FF6ACD83FCEEEF52B23DEE77" box="[821,846,1831,1854]" italics="true" pageId="189" pageNumber="190">sp</emphasis>
) anteriorly with a nippled sternal sac (
<emphasis id="B96AEA88FF6ACD83FAFDEF52B44FEE77" box="[1318,1340,1831,1854]" italics="true" pageId="189" pageNumber="190">ss</emphasis>
). Coxal processes (
<emphasis id="B96AEA88FF6ACD83FEC1EF3EB047EE2B" box="[282,308,1867,1890]" italics="true" pageId="189" pageNumber="190">cp</emphasis>
) fused in basal third; distal two-thirds divided, but connected by a denticulate lamella (
<emphasis id="B96AEA88FF6ACD83FAADEF3CB4FFEE2B" box="[1398,1420,1865,1890]" italics="true" pageId="189" pageNumber="190">dl</emphasis>
). Posteriorly, lateral edges of coxal processes carrying a very wide lamella divided into a mesal (
<emphasis id="B96AEA88FF6ACD83FB65EF18B5A9EECF" box="[1214,1242,1901,1926]" italics="true" pageId="189" pageNumber="190">ml</emphasis>
) and a lateral (
<emphasis id="B96AEA88FF6ACD83FA58EF18B4E0EECF" box="[1411,1427,1901,1926]" italics="true" pageId="189" pageNumber="190">ll</emphasis>
) part, both parts with setae and denticles. One more longitudinal lamella (
<emphasis id="B96AEA88FF6ACD83FC0AEFE4B29DEEE3" box="[977,1006,1937,1962]" italics="true" pageId="189" pageNumber="190">lol</emphasis>
) present on both mesal sides of coxal processes in posterior view. The latter lamella can only be partly seen in anterior view. Coxal vesicles (
<emphasis id="B96AEA88FF6ACD83FAF7EFC2B436EE87" box="[1324,1349,1975,1998]" italics="true" pageId="189" pageNumber="190">cv</emphasis>
) barely visible in posterior view.
</paragraph>
<paragraph id="8BA1369AFF69CD80FF1CE8EDB029E854" blockId="190.[151,1436,151,320]" pageId="190" pageNumber="191">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFF69CD80FE02E8E2B34EE9F9" box="[473,573,151,176]" captionStart="FIGURE 159" captionStartId="189.[151,250,1671,1693]" captionTargetBox="[253,1329,408,1618]" captionTargetId="figure@189.[188,1398,373,1665]" captionTargetPageId="189" captionText="FIGURE 159. Herculina polysagittae gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, anterior view. c = coxite; t = telopodite; af = arrow-like filaments; cp = coxal process; cv = coxal vesicle; dl = denticulate lamella = ll = lateral lamella; ml = mesal lamella; sp = sternal plate; ss = sternal sac; lol = longitudinal lamella. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272908/files/figure.png" pageId="190" pageNumber="191">Fig. 159</figureCitation>
D). Coxites (
<emphasis id="B96AEA88FF69CD80FD09E8ECB3ADE9F9" box="[722,734,153,176]" italics="true" pageId="190" pageNumber="191">c</emphasis>
) divided. Laterally, with a small telopodite (
<emphasis id="B96AEA88FF69CD80FB3EE8ECB59EE9F9" box="[1253,1261,153,176]" italics="true" pageId="190" pageNumber="191">t</emphasis>
) with a single, very long, arrow-shaped filament (
<emphasis id="B96AEA88FF69CD80FDE2E8C9B33CE99D" box="[569,591,188,212]" italics="true" pageId="190" pageNumber="191">af</emphasis>
). The most characteristic parts of coxites are numerous arrow-shaped filaments (
<emphasis id="B96AEA88FF69CD80FEC9E895B05BE9B1" box="[274,296,224,248]" italics="true" pageId="190" pageNumber="191">af</emphasis>
). Posteriorly, coxites with lamellar coxal processes (
<emphasis id="B96AEA88FF69CD80FC56E894B2D4E9B1" box="[909,935,225,248]" italics="true" pageId="190" pageNumber="191">cp</emphasis>
) with setae. Coxal vesicles (
<emphasis id="B96AEA88FF69CD80FB22E894B461E9B1" box="[1273,1298,225,248]" italics="true" pageId="190" pageNumber="191">cv</emphasis>
) located on anteromesal side.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF69CD80FF1CE952B54FE809" box="[199,1084,295,320]" pageId="190" pageNumber="191" type="distribution">
<paragraph id="8BA1369AFF69CD80FF1CE952B54FE809" blockId="190.[151,1436,151,320]" box="[199,1084,295,320]" pageId="190" pageNumber="191">
<emphasis id="B96AEA88FF69CD80FF1CE952B013E809" bold="true" box="[199,352,295,320]" pageId="190" pageNumber="191">Distribution.</emphasis>
<collectingRegion id="49DAF878FF69CD80FEBCE952B0B2E809" box="[359,449,295,320]" country="United States of America" name="Georgia" pageId="190" pageNumber="191">
<collectingCountry id="F309760AFF69CD80FEBCE952B0B2E809" box="[359,449,295,320]" name="Georgia" pageId="190" pageNumber="191">Georgia</collectingCountry>
</collectingRegion>
(known only from
<typeStatus id="54A58838FF69CD80FD40E95DB3B8E809" box="[667,715,296,320]" pageId="190" pageNumber="191">type</typeStatus>
locality) (
<figureCitation id="13252A1FFF69CD80FCE4E952B2ECE809" box="[831,927,295,320]" captionStart="FIGURE 170" captionStartId="198.[151,250,1064,1086]" captionTargetBox="[188,1398,193,1043]" captionTargetId="figure@198.[188,1398,193,1043]" captionTargetPageId="198" captionText="FIGURE 170. Distribution of the Caucasian Anthroleucosomatidae: the Dentatosoma, the Herculina, the Ratcheuma and the Vegrandosoma complexes." httpUri="https://zenodo.org/record/272919/files/figure.png" pageId="190" pageNumber="191">Fig. 170</figureCitation>
, red triangle).
</paragraph>
</subSubSection>
</treatment>
</document>
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