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<document id="627858A7ACFA42970CCBC554B225AC1E" ID-CLB-Dataset="279456" ID-DOI="10.11646/zootaxa.5374.4.1" ID-GBIF-Dataset="6f75d877-d9e0-49de-8ce6-b739b532136a" ID-ISSN="1175-5326" ID-Zenodo-Dep="10158722" ID-ZooBank="EEFF13BD-664A-4FD8-9FED-4E4B21A475E4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1700469135046" checkinUser="plazi" docAuthor="Ahrens, Dirk, Lukic, Daniel, Pham, Phu, Li, Wei &amp; Liu, Wangang" docDate="2023" docId="03B73169BE7F2740D9E40BAEFB8DFE00" docLanguage="en" docName="zootaxa.5374.4.1.pdf" docOrigin="Zootaxa 5374 (4)" docSource="https://mapress.com/zt/article/download/zootaxa.5374.4.1/52301" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetraserica fabriziae Liu, Li &amp; Ahrens 2023, new species" docType="treatment" docVersion="7" lastPageNumber="464" masterDocId="FF8E4911BE73274DD9730A19FFFAFFD1" masterDocTitle="Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)" masterLastPageNumber="486" masterPageNumber="451" pageNumber="463" updateTime="1732584714017" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8907E4CDBE4ACF02071B58FBF76833EF">Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)</mods:title>
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<mods:affiliation id="D5870922BBE0B69FA047E824DB584D86">Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xian 710061, P. R. China. &amp; liweiv @ stumail. nwu. edu. cn; https: // orcid. org / 0009 - 0001 - 0195 - 7889</mods:affiliation>
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<taxonomicName id="4C1EFBFCBE7F2741D9E40BAEFD94FE01" ID-CoL="BW5GK" authority="Liu, Li &amp; Ahrens" authorityName="Liu, Li &amp; Ahrens" authorityYear="2023" box="[151,622,438,465]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="fabriziae" status="sp. nov.">
<emphasis id="B96A5C6DBE7F2741D9E40BAEFE75FE00" bold="true" box="[151,399,438,465]" italics="true" pageId="12" pageNumber="463">Tetraserica fabriziae</emphasis>
Liu, Li
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Ahrens
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<emphasis id="B96A5C6DBE7F2741D9E4083AFE30FDED" bold="true" box="[151,458,546,573]" pageId="12" pageNumber="463">Type material examined.</emphasis>
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1 ♂,
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Description of the
<typeStatus id="54A53EDDBE7F2741D8DD08AAFDEDFD1C" box="[430,535,691,717]" pageId="12" pageNumber="463" type="holotype">holotype</typeStatus>
.
</emphasis>
Length:
<quantity id="4CE62D9ABE7F2741DBFA08ABFD1EFD1C" box="[649,740,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="463" unit="mm" value="8.5">8.5 mm</quantity>
, length of elytra:
<quantity id="4CE62D9ABE7F2741DACA08AAFBEEFD1C" box="[953,1044,691,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="12" pageNumber="463" unit="mm" value="6.2">6.2 mm</quantity>
, width:
<quantity id="4CE62D9ABE7F2741DD0508ABFB29FD1C" box="[1142,1235,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="463" unit="mm" value="4.5">4.5 mm</quantity>
. Body oval, dark reddish brown, labroclypeus and dorsal surface shiny, glabrous.
</paragraph>
<paragraph id="8BA1807FBE7F2741D9B408E2FA91FBE4" blockId="12.[151,1437,546,2013]" pageId="12" pageNumber="463">
Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex, anterior angles strongly convex, anterior margin shallowly emarginate medially, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed; surface moderately convex medially, coarsely and moderate densely punctate, without setae; frontoclypeal suture feebly impressed and moderately angled medially; smooth area in front of eye approximately four times as wide as long; ocular canthus moderate and triangular (1/2.5 of ocular diameter), weakly punctate, with one terminal seta. Frons weakly shiny and punctate, bearing only a few erect setae beside eyes. Eyes moderately large, ratio of diameter/ interocular width: 0.69. Antenna yellowish brown, with ten antennomeres, club in male with four antennomeres, slightly longer than remaining antennomeres combined, first joint of club approximately as long as club.
<collectingRegion id="49DA4E9DBE7F2741DBF00E02FD18FBE4" box="[643,738,1051,1077]" country="Myanmar" name="Chin" pageId="12" pageNumber="463">Mentum</collectingRegion>
anteriorly weakly elevated and weakly convex in middle.
</paragraph>
<paragraph id="8BA1807FBE7F2741D9B40E26FD1AFB38" blockId="12.[151,1437,546,2013]" pageId="12" pageNumber="463">Pronotum strongly transverse, widest just before base, lateral margins in basal quarter strongly convex, in anterior portion moderately convex and evenly convergent anteriorly; anterior angles distinctly produced, posterior angles strongly convex. Anterior margin slightly convex medially, with a fine, complete marginal line. Surface densely and finely punctate, with minute setae in punctures; lateral margins sparsely setose. Hypomeron not carinate. Scutellum triangular, with fine and dense punctures.</paragraph>
<paragraph id="8BA1807FBE7F2741D9B40EEAFE20FAA8" blockId="12.[151,1437,546,2013]" pageId="12" pageNumber="463">Elytra oblong, widest at posterior third, striae weakly impressed, finely and densely punctate; intervals weakly convex, with fine, dense punctures, glabrous except minute setae in punctures; epipleural edge robust, ending at the strongly rounded external apical angle of elytra, epipleura slightly setose, apical border membraneous, with a fine rim of short microtrichomes.</paragraph>
<paragraph id="8BA1807FBE7F2741D9B40F9AFF14F980" blockId="12.[151,1437,546,2013]" pageId="12" pageNumber="463">Ventral surface dull, reddish brown, thorax with large and dense punctures, sparsely setose, metasternum sparsely covered with fine, short, or minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctate, with a transverse row of coarse punctures each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.5. Pygidium weakly convex and dull, coarsely densely punctate, without smooth midline, punctures with fine setae along apical margin.</paragraph>
<paragraph id="8BA1807FBE7F2741D9B40C42FD7FF868" blockId="12.[151,1437,546,2013]" pageId="12" pageNumber="463">
Legs moderately wide; metafemur moderately shiny, anterior margin acute, lacking an adjacent serrated line; ventral posterior margin straight and smooth (
<figureCitation id="13259CFABE7F2741DBEF0C66FD0AF948" box="[668,752,1663,1689]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. AF: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); GL: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="12" pageNumber="463">Fig. 4L</figureCitation>
), moderately widened in apical half; dorsal posterior margin smooth, with moderately dense, short setae basally. Metatibia moderately short, widest at middle, ratio width/ length: 1/ 3.2, Dorsal margin longitudinally convex, with two groups of spines, basal one at first third, apical one at two thirds of metatibial length, basally with a few short single setae; lateral face smooth, sparsely finely punctate; ventral margin finely serrate, with four robust equidistant setae; medial face smooth; apex interiorly near tarsal articulation moderately concave. Tarsomeres in distal half with fine, dense setae ventrally, dorsally smooth; metatarsomeres ventrally with a strong serrated ridge and glabrous, first metatarsomere shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
</paragraph>
<paragraph id="8BA1807FBE7F2741D9B40DDAFD78F80C" blockId="12.[151,1437,546,2013]" box="[199,642,1987,2013]" pageId="12" pageNumber="463">
Aedeagus.
<figureCitation id="13259CFABE7F2741D8370DDAFE4CF80C" box="[324,438,1987,2013]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. AF: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); GL: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="12" pageNumber="463">Fig. 4GJ</figureCitation>
. Habitus:
<figureCitation id="13259CFABE7F2741DB550DDAFD81F80C" box="[550,635,1987,2013]" captionStart="FIGURE 4" captionStartId="11.[152,255,1781,1805]" captionTargetBox="[162,1430,283,1748]" captionTargetId="figure-54@11.[151,1436,273,1758]" captionTargetPageId="11" captionText="FIGURE 4. AF: Tetraserica yucheni Liu, Li &amp; Ahrens, new species (holotype); GL: T. fabriziae Liu, Li &amp; Ahrens, new species (holotype). A, G: aedeagus, left side lateral view; B, C, H, I: parameres, dorsal view; D, J: aedeagus, right side lateral view; E, K: habitus; F, L: hind leg. Scale: 0.5 mm. Legs and habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.10158730" httpUri="https://zenodo.org/record/10158730/files/figure.png" pageId="12" pageNumber="463">Fig. 4K</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7E2740D9B40A8FFBBAFECC" pageId="13" pageNumber="464" type="diagnosis">
<paragraph id="8BA1807FBE7E2740D9B40A8FFBBAFECC" blockId="13.[151,1436,150,465]" pageId="13" pageNumber="464">
<emphasis id="B96A5C6DBE7E2740D9B40A8FFEB8FF61" bold="true" box="[199,322,150,176]" pageId="13" pageNumber="464">Diagnosis.</emphasis>
<taxonomicName id="4C1EFBFCBE7E2740D8380A8EFCF1FF60" authority="Liu, Li &amp; Ahrens" authorityName="Liu, Li &amp; Ahrens" authorityYear="2023" box="[331,779,151,177]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="fabriziae" status="sp. nov.">
<emphasis id="B96A5C6DBE7E2740D8380A8EFDCCFF60" box="[331,566,151,177]" italics="true" pageId="13" pageNumber="464">Tetraserica fabriziae</emphasis>
Liu, Li
<emphasis id="B96A5C6DBE7E2740DBEB0A8EFD54FF60" box="[664,686,151,177]" italics="true" pageId="13" pageNumber="464">&amp;</emphasis>
Ahrens
</taxonomicName>
,
<taxonomicNameLabel id="A259E116BE7E2740DA6A0A8EFC5BFF60" box="[793,929,151,177]" pageId="13" pageNumber="464" rank="species">new species</taxonomicNameLabel>
differs from
<taxonomicName id="4C1EFBFCBE7E2740DD4F0A8EFE6EFF04" authority="Liu, Fabrizi, Bai, Yang &amp; Ahrens, 2014" authorityName="Liu, Fabrizi, Bai, Yang &amp; Ahrens" authorityYear="2014" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis id="B96A5C6DBE7E2740DD4F0A8EFB35FF60" box="[1084,1231,151,177]" italics="true" pageId="13" pageNumber="464">T. longipenis</emphasis>
<bibRefCitation id="EF8FFD8EBE7E2740DDAB0A8EFE6EFF04" author="Liu, W. &amp; Fabrizi, S. &amp; Bai, M. &amp; Yang, X. - K. &amp; Ahrens, D." pageId="13" pageNumber="464" pagination="83 - 121" refId="ref44587" refString="Liu, W., Fabrizi, S., Bai, M., Yang, X. - K. &amp; Ahrens, D. (2014) A taxonomic review of the Tetraserica Ahrens, 2004 species of China (Coleoptera, Scarabaeidae, Sericini). ZooKeys, 448, 83 - 121. https: // doi. org / 10.3897 / zookeys. 443.8429" type="journal article" year="2014">Liu, Fabrizi, Bai, Yang &amp; Ahrens, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1EFBFCBE7E2740D8A20AA2FC9BFF04" authority="Kobayashi, 2017" authorityName="Kobayashi" authorityYear="2017" box="[465,865,187,213]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="angkhangensis">
<emphasis id="B96A5C6DBE7E2740D8A20AA2FD6CFF04" box="[465,662,187,213]" italics="true" pageId="13" pageNumber="464">T. angkhangensis</emphasis>
<bibRefCitation id="EF8FFD8EBE7E2740DBD30AA2FC9BFF04" author="Kobayashi, H." box="[672,865,187,213]" pageId="13" pageNumber="464" pagination="33 - 45" refId="ref44520" refString="Kobayashi, H. (2017) Notes on the genus Tetraserica (Coleoptera, Scarabaeidae, Melolonthinae, Sericini) from Thailand. Kogane, 20, 33 - 45." type="journal article" year="2017">Kobayashi, 2017</bibRefCitation>
</taxonomicName>
by the evenly moderately narrow right paramere which is apically not widened. In this new species, the median phallobasal lamina is straight as in
<taxonomicName id="4C1EFBFCBE7E2740DDA40AC6FA9DFF28" authorityName="Liu, Fabrizi, Bai, Yang &amp; Ahrens" authorityYear="2014" box="[1239,1383,223,249]" class="Insecta" family="Scarabaeidae" genus="Tetraserica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="longipenis">
<emphasis id="B96A5C6DBE7E2740DDA40AC6FA9DFF28" box="[1239,1383,223,249]" italics="true" pageId="13" pageNumber="464">T. longipenis</emphasis>
</taxonomicName>
, and the moderately long dorsal lobe of left paramere is at apex slightly hooked dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7E2740D9B40B3FFF3EFE58" pageId="13" pageNumber="464" type="description">
<paragraph id="8BA1807FBE7E2740D9B40B3FFF3EFE58" blockId="13.[151,1436,150,465]" pageId="13" pageNumber="464">
<emphasis id="B96A5C6DBE7E2740D9B40B3FFEC5FE91" bold="true" box="[199,319,294,320]" pageId="13" pageNumber="464">Variation.</emphasis>
Length:
<quantity id="4CE62D9ABE7E2740D8D00B3FFDD6FE90" box="[419,556,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="13" pageNumber="464" unit="mm" value="8.3" valueMax="8.5" valueMin="8.1">8.18.5 mm</quantity>
, length of elytra:
<quantity id="4CE62D9ABE7E2740DB830B3EFC83FE90" box="[752,889,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.3" metricValueMin="6.2" pageId="13" pageNumber="464" unit="mm" value="6.25" valueMax="6.3" valueMin="6.2">6.26.3 mm</quantity>
, width:
<quantity id="4CE62D9ABE7E2740DAA00B3FFBA6FE90" box="[979,1116,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.8" metricValueMin="4.5" pageId="13" pageNumber="464" unit="mm" value="4.65" valueMax="4.8" valueMin="4.5">4.54.8 mm</quantity>
. Female: Antennal club with three antennomeres, as long as remaining antennomeres combined. Eyes slightly smaller than in male. Pygidium flat.
</paragraph>
</subSubSection>
<subSubSection id="C304D3F4BE7E2740D9B40B8AFB8DFE00" pageId="13" pageNumber="464" type="etymology">
<paragraph id="8BA1807FBE7E2740D9B40B8AFB8DFE00" blockId="13.[151,1436,150,465]" pageId="13" pageNumber="464">
<emphasis id="B96A5C6DBE7E2740D9B40B8AFEB4FE7C" bold="true" box="[199,334,403,429]" pageId="13" pageNumber="464">Etymology.</emphasis>
The new species is dedicated to Silvia Fabrizi, who we keep all in warmest memory and who greatly contributed to the knowledge of Sericini chafers (noun in genitive singular case).
</paragraph>
</subSubSection>
</treatment>
</document>