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<document id="A0DB93F32771B9C487E99290807CCD5F" ID-DOI="10.11646/zootaxa.3077.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5243536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629750005925" checkinUser="felipe" docAuthor="Peck, Stewart B. &amp; Cook, Joyce" docDate="2011" docId="03B375533D65A1609DC0FCFEFC430E16" docLanguage="en" docName="zt03077p118.pdf" docOrigin="Zootaxa 3077 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3077.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Pinodytes borealis Peck &amp; Cook 2011, new species" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="FF8A0D2B3D74A1729D57FF87FFB30F0A" masterDocTitle="Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077" masterLastPageNumber="118" masterPageNumber="1" pageNumber="18" updateTime="1699105988326" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2C163E19F0580E834D5F20792898A6E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2BD11ACD9F878A531F93F502E9E7E7A7">
<mods:title id="070A6980DD5798F02751B11380A528C5">Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077</mods:title>
</mods:titleInfo>
<mods:name id="EB2E52479B2BD0C3E46F19D0EC731C95" type="personal">
<mods:role id="4234C8436684154ECEF1E961616F655E">
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</mods:role>
<mods:namePart id="6B7F4126A01CBDC8FE0C3D273CA8C65D">Peck, Stewart B.</mods:namePart>
</mods:name>
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<mods:role id="9705FDEF013BB20428F90EDDA45F52BD">
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<mods:namePart id="467204658A68A3F3CE7DE077DB87CA50">Cook, Joyce</mods:namePart>
</mods:name>
<mods:typeOfResource id="4BE7AB4DDD9F7032BAFD08E0369C04DE">text</mods:typeOfResource>
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<mods:title id="C71E79DC74401E0F195E0BBD83431A96">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9B0803F1D676FF105E0725B51C359B18">
<mods:date id="744FCA4E21A076D496A07E8DC62EA86A">2011</mods:date>
<mods:detail id="BF1EA9197CB5F3691A3CC8153E4ED0D7" type="pubDate">
<mods:number id="130FB3C6CC5FBD01C7AD8B642DD8A96D">2011-10-28</mods:number>
</mods:detail>
<mods:detail id="F594E7A981A0267FB8BC2E67EA31F508" type="volume">
<mods:number id="08EF595A6E5A6CCC2BF1C300F584FE10">3077</mods:number>
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<mods:number id="F266E783268E48481190152D8337E0C9">1</mods:number>
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<mods:url id="B8D3F4C13B37E177DA3475365DD18457">https://biotaxa.org/Zootaxa/article/view/zootaxa.3077.1.1</mods:url>
</mods:location>
<mods:classification id="957777FD970822D62FF549EF40A4ACFE">journal article</mods:classification>
<mods:identifier id="7B4B0E0A59D6E8FE1D1984E3D83D5381" type="DOI">10.11646/zootaxa.3077.1.1</mods:identifier>
<mods:identifier id="BD8DADA232AF9BBEF90DCD8E72BCB31A" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="329D3A09A8BC9C8D2B257EF0D7175A22" type="Zenodo-Dep">5243536</mods:identifier>
</mods:mods>
<treatment id="03B375533D65A1609DC0FCFEFC430E16" ID-DOI="http://doi.org/10.5281/zenodo.5243795" ID-Zenodo-Dep="5243795" LSID="urn:lsid:plazi:treatment:03B375533D65A1609DC0FCFEFC430E16" httpUri="http://treatment.plazi.org/id/03B375533D65A1609DC0FCFEFC430E16" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<subSubSection id="C30097CE3D65A1639DC0FCFEFD0C0C99" box="[151,703,889,915]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BA5C4453D65A1639DC0FCFEFD0C0C99" blockId="17.[151,703,889,948]" box="[151,703,889,915]" pageId="17" pageNumber="18">
<heading id="D0ED73293D65A1639DC0FCFEFD0C0C99" bold="true" box="[151,703,889,915]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="B96E18573D65A1639DC0FCFEFD0C0C99" bold="true" box="[151,703,889,915]" pageId="17" pageNumber="18">
<taxonomicName id="4C1ABFC63D65A1639DC0FCFEFD910C99" ID-CoL="4HZC7" authority="Peck &amp; Cook" authorityName="Peck &amp; Cook" authorityYear="2011" box="[151,546,889,915]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
<emphasis id="B96E18573D65A1639DC0FCFEFEC00C99" bold="true" box="[151,371,889,915]" italics="true" pageId="17" pageNumber="18">Pinodytes borealis</emphasis>
Peck &amp; Cook
</taxonomicName>
,
<taxonomicNameLabel id="A25DA52C3D65A1639F78FCFEFD0C0C99" box="[559,703,889,915]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D65A1639DC0FC1BFEF90CBE" box="[151,330,924,948]" pageId="17" pageNumber="18" type="description">
<paragraph id="8BA5C4453D65A1639DC0FC1BFEF90CBE" blockId="17.[151,703,889,948]" box="[151,330,924,948]" pageId="17" pageNumber="18">
(
<figureCitation id="1321D8C03D65A1639DC8FC1BFF5D0CBE" box="[159,238,924,948]" captionStart="FIGURES 2–10" captionStartId="53.[151,264,1927,1950]" captionTargetBox="[191,1402,155,1906]" captionTargetId="figure-93@53.[185,1402,153,1906]" captionTargetPageId="53" captionText="FIGURES 2–10. Pinodytes habitus photos. 2. P.newtoni, total length 1.38 mm. 3. P. borealis, total length 2.10 mm. 4. P. punctatus, total length 1.36 mm. 5. P.colorado, total length 1.66 mm. 6. P.pusio, total length 1.54 mm. 7. P. ovatus, total length 1.34 mm. 8. P. chandleri, total length 1.54 mm. 9. P. constrictus, total length 1.48 mm. 10. P.contortus, total length 1.16 mm." figureDoi="http://doi.org/10.5281/zenodo.5243541" httpUri="https://zenodo.org/record/5243541/files/figure.png" pageId="17" pageNumber="18">Figs. 3</figureCitation>
,
<figureCitation id="1321D8C03D65A1639DADFC1BFEF10CBE" box="[250,322,924,948]" captionStart-0="FIGURE 52" captionStart-1="FIGURES 53–61" captionStartId-0="59.[151,250,1193,1216]" captionStartId-1="60.[151,263,1796,1819]" captionTargetBox-0="[166,1420,193,1173]" captionTargetBox-1="[195,1398,201,1765]" captionTargetId-0="figure-83@59.[166,1420,193,1173]" captionTargetId-1="figure-84@60.[183,1404,193,1775]" captionTargetPageId-0="59" captionTargetPageId-1="60" captionText-0="FIGURE 52. Distributions of basal and monobasic species groups of Pinodytes in western United States. P. newtoni, closed circles in Missouri, Arkansas, and Texas. P. borealis, closed circles in inset map of Alaska. P. punctatus, open squares in eastern Washington and northern Idaho. P.colorado, open circles in western Colorado. Note: in this and all following maps, small open circles represent major cities." captionText-1="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi-0="http://doi.org/10.5281/zenodo.5243561" figureDoi-1="http://doi.org/10.5281/zenodo.5243563" httpUri-0="https://zenodo.org/record/5243561/files/figure.png" httpUri-1="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">52–61</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D65A1639DC0FC63FC8C0BBA" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BA5C4453D65A1639DC0FC63FCFB0B86" blockId="17.[151,1436,996,1992]" pageId="17" pageNumber="18">
Type material.
<materialsCitation id="3B72CE183D65A1639C19FC63FE010B2A" collectionCode="SBPC" collectorName="S. &amp; J. Peck" country="United States of America" location="Alaska" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Alaska" typeStatus="holotype">
<emphasis id="B96E18573D65A1639C19FC63FE080CF6" bold="true" box="[334,443,996,1020]" pageId="17" pageNumber="18">
<typeStatus id="54A17AE73D65A1639C19FC63FE080CF6" box="[334,443,996,1020]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
: male (
<collectionCode id="ED0B5C803D65A1639F46FC63FDE80CF6" box="[529,603,996,1020]" pageId="17" pageNumber="18">SBPC</collectionCode>
).
<emphasis id="B96E18573D65A1639F39FC63FC180CF7" bold="true" box="[622,939,996,1021]" pageId="17" pageNumber="18">
<collectingCountry id="F30D84D53D65A1639F39FC63FCFF0CF7" box="[622,844,996,1021]" name="United States of America" pageId="17" pageNumber="18">UNITED STATES</collectingCountry>
.
<collectingRegion id="49DE0AA73D65A1639E0EFC63FC180CF7" box="[857,939,996,1021]" country="United States of America" name="Alaska" pageId="17" pageNumber="18">Alaska</collectingRegion>
</emphasis>
: Chena Ridge, 5miW Fairbanks, 27.VII.84,
<collectorName id="26EFA1933D65A1639DC0FB8FFE980B2A" box="[151,299,1032,1056]" pageId="17" pageNumber="18">S. &amp; J. Peck</collectorName>
, alder litter
</materialsCitation>
.
<materialsCitation id="3B72CE183D65A1639CE8FB8FFAC10B2A" box="[447,1394,1031,1056]" collectionCode="SBPC" collectorName="S. &amp; J. Peck" country="United States of America" location="Alaska" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Alaska" typeStatus="paratype">
<emphasis id="B96E18573D65A1639CE8FB8FFD850B2A" bold="true" box="[447,566,1032,1056]" pageId="17" pageNumber="18">
<typeStatus id="54A17AE73D65A1639CE8FB8FFD850B2A" box="[447,566,1032,1056]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
(145).
<emphasis id="B96E18573D65A1639FD9FB8FFC6A0B2A" bold="true" box="[654,985,1031,1056]" pageId="17" pageNumber="18">
<collectingCountry id="F30D84D53D65A1639FD9FB8FFCDD0B2A" box="[654,878,1031,1056]" name="United States of America" pageId="17" pageNumber="18">UNITED STATES</collectingCountry>
.
<collectingRegion id="49DE0AA73D65A1639E2AFB80FC670B2A" box="[893,980,1031,1056]" country="United States of America" name="Alaska" pageId="17" pageNumber="18">Alaska</collectingRegion>
:
</emphasis>
same data as holotype, 69 (
<collectionCode id="ED0B5C803D65A1639877FB8FFAD90B2A" box="[1312,1386,1032,1056]" pageId="17" pageNumber="18">SBPC</collectionCode>
)
</materialsCitation>
, 17 (
<collectionCode id="ED0B5C803D65A1639DC8FBABFF450B4E" box="[159,246,1068,1092]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="17" pageNumber="18" type="Museum">FMNH</collectionCode>
);
<materialsCitation id="3B72CE183D65A1639C5EFBABFEA50B62" collectionCode="SBPC" collectorName="S. &amp; J. Peck" country="United States of America" location="Chena River Rd." municipality="Chena Hot Spgs." pageId="17" pageNumber="18" specimenCount="1" stateProvince="Alaska" typeStatus="paratype">
<collectingMunicipality id="6BC15E3F3D65A1639C5EFBABFE7A0B4E" box="[265,457,1068,1092]" pageId="17" pageNumber="18">Chena Hot Spgs.</collectingMunicipality>
, mi 49
<location id="8EC5929E3D65A1639F4BFBABFD680B4E" LSID="urn:lsid:plazi:treatment:03B375533D65A1609DC0FCFEFC430E16:8EC5929E3D65A1639F4BFBABFD680B4E" box="[540,731,1068,1092]" country="United States of America" municipality="Chena Hot Spgs." name="Chena River Rd." pageId="17" pageNumber="18" stateProvince="Alaska">Chena River Rd.</location>
, 28.VII.84, 1000’,
<collectorName id="26EFA1933D65A1639EE6FBABFB8D0B4E" box="[945,1086,1068,1092]" pageId="17" pageNumber="18">S. &amp; J. Peck</collectorName>
, birch-alder litter, S face slope, 24 (
<collectionCode id="ED0B5C803D65A1639D94FBD7FEBE0B62" box="[195,269,1104,1128]" pageId="17" pageNumber="18">SBPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72CE183D65A1639C75FBD7FAB80B62" box="[290,1291,1104,1128]" collectionCode="SBPC" collectorName="S. &amp; J. Peck" country="United States of America" location="Circle Hot Springs" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Alaska" typeStatus="paratype">
<location id="8EC5929E3D65A1639C75FBD7FE4A0B62" LSID="urn:lsid:plazi:treatment:03B375533D65A1609DC0FCFEFC430E16:8EC5929E3D65A1639C75FBD7FE4A0B62" box="[290,505,1104,1128]" country="United States of America" name="Circle Hot Springs" pageId="17" pageNumber="18" stateProvince="Alaska">Circle Hot Springs</location>
, 6.VIII.84,
<collectorName id="26EFA1933D65A1639F2AFBD7FCBE0B62" box="[637,781,1104,1128]" pageId="17" pageNumber="18">S. &amp; J. Peck</collectorName>
, alder-poplar-willow forest litter, 29 (
<collectionCode id="ED0B5C803D65A16399EEFBD7FAB00B62" box="[1209,1283,1104,1128]" pageId="17" pageNumber="18">SBPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B72CE183D65A163984FFBD6FCF70B86" collectionCode="SBPC" collectorName="S. &amp; J. Peck" country="United States of America" location="Livengood" municipality="Lost Creek" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Alaska" typeStatus="paratype">
<collectingMunicipality id="6BC15E3F3D65A163984FFBD6FA2B0B62" box="[1304,1432,1104,1128]" pageId="17" pageNumber="18">Lost Creek</collectingMunicipality>
, 7miN
<location id="8EC5929E3D65A1639D8AFBF3FEEA0B86" LSID="urn:lsid:plazi:treatment:03B375533D65A1609DC0FCFEFC430E16:8EC5929E3D65A1639D8AFBF3FEEA0B86" box="[221,345,1140,1164]" country="United States of America" municipality="Lost Creek" name="Livengood" pageId="17" pageNumber="18" stateProvince="Alaska">Livengood</location>
, 30.VII.84,
<collectorName id="26EFA1933D65A1639CB7FBF3FDDF0B86" box="[480,620,1140,1164]" pageId="17" pageNumber="18">S. &amp; J. Peck</collectorName>
, carrion, 5 (
<collectionCode id="ED0B5C803D65A1639FA5FBF3FC8F0B86" box="[754,828,1140,1164]" pageId="17" pageNumber="18">SBPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA5C4453D65A1639D90FB10FC8C0BBA" blockId="17.[151,1436,996,1992]" box="[199,831,1175,1200]" pageId="17" pageNumber="18">
<emphasis id="B96E18573D65A1639D90FB10FE030BBA" box="[199,432,1175,1200]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">
<emphasis id="B96E18573D65A1639D90FB10FE1A0BBA" bold="true" box="[199,425,1175,1200]" pageId="17" pageNumber="18">Material examined</emphasis>
.
</emphasis>
We have examined
<specimenCount id="9D1C0FCC3D65A1639FC5FB1FFC880BBA" box="[658,827,1176,1200]" pageId="17" pageNumber="18" type="generic">145 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D65A1639D90FB3BFB0508AE" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BA5C4453D65A1639D90FB3BFB300A16" blockId="17.[151,1436,996,1992]" pageId="17" pageNumber="18">
<emphasis id="B96E18573D65A1639D90FB3BFED30BDF" bold="true" box="[199,352,1212,1237]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Distribution.</emphasis>
Specimens (
<figureCitation id="1321D8C03D65A1639CA4FB3BFDF50BDE" box="[499,582,1212,1236]" captionStart="FIGURE 52" captionStartId="59.[151,250,1193,1216]" captionTargetBox="[166,1420,193,1173]" captionTargetId="figure-83@59.[166,1420,193,1173]" captionTargetPageId="59" captionText="FIGURE 52. Distributions of basal and monobasic species groups of Pinodytes in western United States. P. newtoni, closed circles in Missouri, Arkansas, and Texas. P. borealis, closed circles in inset map of Alaska. P. punctatus, open squares in eastern Washington and northern Idaho. P.colorado, open circles in western Colorado. Note: in this and all following maps, small open circles represent major cities." figureDoi="http://doi.org/10.5281/zenodo.5243561" httpUri="https://zenodo.org/record/5243561/files/figure.png" pageId="17" pageNumber="18">Fig. 52</figureCitation>
) are known from the vicinity of Fairbanks, in formerly unglaciated central
<collectingRegion id="49DE0AA73D65A1639DC0FB67FF5B0BF2" box="[151,232,1248,1272]" country="United States of America" name="Alaska" pageId="17" pageNumber="18">Alaska</collectingRegion>
. We interpret these populations to be relictual and indicating a Tertiary distribution, before Pleistocene glaciations caused the extinction of other populations and species in western North America.
</paragraph>
<paragraph id="8BA5C4453D65A1639D90FAA0FB0508AE" blockId="17.[151,1436,996,1992]" pageId="17" pageNumber="18">
<emphasis id="B96E18573D65A1639D90FAA0FE6A0A4A" box="[199,473,1319,1344]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">
<emphasis id="B96E18573D65A1639D90FAA0FE600A4A" bold="true" box="[199,467,1319,1344]" pageId="17" pageNumber="18">Diagnostic description</emphasis>
.
</emphasis>
Total length
<quantity id="4CE269A03D65A1639F38FAAFFCA50A4A" box="[623,790,1320,1344]" metricMagnitude="-3" metricUnit="m" metricValue="2.06" metricValueMax="2.1" metricValueMin="2.02" pageId="17" pageNumber="18" unit="mm" value="2.06" valueMax="2.1" valueMin="2.02">2.02–2.10 mm</quantity>
; greatest width
<quantity id="4CE269A03D65A1639E9FFAAFFBDD0A4A" box="[968,1134,1320,1344]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.6" metricValueMin="9.2" pageId="17" pageNumber="18" unit="mm" value="0.94" valueMax="0.96" valueMin="0.92">0.92–0.96 mm</quantity>
. Reddish brown; elongateoval in shape (
<figureCitation id="1321D8C03D65A1639C6BFACBFE320A6E" box="[316,385,1356,1380]" captionStart="FIGURES 2–10" captionStartId="53.[151,264,1927,1950]" captionTargetBox="[191,1402,155,1906]" captionTargetId="figure-93@53.[185,1402,153,1906]" captionTargetPageId="53" captionText="FIGURES 2–10. Pinodytes habitus photos. 2. P.newtoni, total length 1.38 mm. 3. P. borealis, total length 2.10 mm. 4. P. punctatus, total length 1.36 mm. 5. P.colorado, total length 1.66 mm. 6. P.pusio, total length 1.54 mm. 7. P. ovatus, total length 1.34 mm. 8. P. chandleri, total length 1.54 mm. 9. P. constrictus, total length 1.48 mm. 10. P.contortus, total length 1.16 mm." figureDoi="http://doi.org/10.5281/zenodo.5243541" httpUri="https://zenodo.org/record/5243541/files/figure.png" pageId="17" pageNumber="18">Fig. 3</figureCitation>
).
<emphasis id="B96E18573D65A1639CCFFACBFE6B0A6E" bold="true" box="[408,472,1356,1380]" pageId="17" pageNumber="18">Head</emphasis>
. Finely, sparsely punctate, shining, with faint reticulate microsculpture on vertex and sides. Eyes absent. Antennae (
<figureCitation id="1321D8C03D65A1639CBCFAF7FD8F0A82" box="[491,572,1392,1416]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 53</figureCitation>
) with antennomeres 2 and 3 subequal in length; antennomere 5 longer than 4 and 6; antennomere 7 clearly larger than 8; antennomeres 9 and 10 lack visible sensory vesicles.
<emphasis id="B96E18573D65A163999EFA13FAF10AA6" bold="true" box="[1225,1346,1428,1452]" pageId="17" pageNumber="18">Pronotum</emphasis>
. Finely, sparsely punctate, shining, with faint reticulate microsculpture. Slightly wider at middle than at base, narrowing from middle to apex; apical margin weakly emarginate, basal margin straight; apical angles rounded, basal angles nearly right-angled.
<emphasis id="B96E18573D65A1639C2DF987FE750912" bold="true" box="[378,454,1536,1560]" pageId="17" pageNumber="18">Elytra</emphasis>
. Strial punctures in rows, weakly impressed medially in basal one-half; interstrial punctures variable in size and density; shining. Joined elytra widest near middle, wider than pronotum, sides moderately rounded, rounded at apex.
<emphasis id="B96E18573D65A1639C87F9CFFDBB096A" bold="true" box="[464,520,1608,1632]" pageId="17" pageNumber="18">Legs</emphasis>
. Protibia (
<figureCitation id="1321D8C03D65A1639FD2F9CFFD69096A" box="[645,730,1608,1632]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 54</figureCitation>
) moderately slender; outer margin with spine one-fifth from apex; apex spinose; inner margin with fine spines on apical two-thirds. Mesotibia (
<figureCitation id="1321D8C03D65A1639979F9EBFBCD098E" box="[1070,1150,1644,1668]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 55</figureCitation>
) moderately slender; apical one-half of inner margin finely spinose, outer margin and apex spinose. Metatibia (
<figureCitation id="1321D8C03D65A1639922F917FB7B09A2" box="[1141,1224,1680,1704]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 56</figureCitation>
) elongate, narrow, straight; fine spines on apical one-half of inner margin; spinose at apex. Metafemur (
<figureCitation id="1321D8C03D65A1639907F933FB1109C6" box="[1104,1186,1716,1740]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 56</figureCitation>
) slender. Male protarsomeres (
<figureCitation id="1321D8C03D65A1639C53F95FFEEB09FA" box="[260,344,1752,1776]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 54</figureCitation>
) and mesotarsomeres (
<figureCitation id="1321D8C03D65A1639F36F95FFD0709FA" box="[609,692,1752,1776]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 55</figureCitation>
) little expanded, bearing elongate setae laterally and thin, broad, colorless phanerae ventrally in two rows.
<emphasis id="B96E18573D65A1639F39F97BFD0F081E" bold="true" box="[622,700,1788,1812]" pageId="17" pageNumber="18">Venter</emphasis>
. Mesoventrite (
<figureCitation id="1321D8C03D65A1639E38F97BFC72081E" box="[879,961,1788,1812]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 61</figureCitation>
) carinate; longitudinal carina expanded at middle, not excavated behind transverse carina.
<emphasis id="B96E18573D65A1639FEFF8A7FCD00832" bold="true" box="[696,867,1823,1848]" pageId="17" pageNumber="18">Male genitalia</emphasis>
. Median lobe of aedeagus (
<figureCitation id="1321D8C03D65A16399C9F8A7FA950832" box="[1182,1318,1824,1848]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Figs. 57, 59</figureCitation>
) elongate, broad, with apex flattened and dorsoventrally sinuate. Everted internal sac (
<figureCitation id="1321D8C03D65A1639EA2F8C3FBFB0856" box="[1013,1096,1860,1884]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 58</figureCitation>
) with a bifurcated sclerotized structure, a single elongate spine, and numerous small spines. Parameres (
<figureCitation id="1321D8C03D65A1639E8EF8EFFBD2088A" box="[985,1121,1896,1920]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Figs. 57, 59</figureCitation>
) straight, not reaching apex of median lobe, each with two apical setae.
<emphasis id="B96E18573D65A1639FD6F80BFCAD08AF" bold="true" box="[641,798,1932,1957]" pageId="17" pageNumber="18">Spermatheca</emphasis>
. Elongate (
<figureCitation id="1321D8C03D65A1639EC9F80BFC4108AE" box="[926,1010,1932,1956]" captionStart="FIGURES 53–61" captionStartId="60.[151,263,1796,1819]" captionTargetBox="[195,1398,201,1765]" captionTargetId="figure-84@60.[183,1404,193,1775]" captionTargetPageId="60" captionText="FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56)." figureDoi="http://doi.org/10.5281/zenodo.5243563" httpUri="https://zenodo.org/record/5243563/files/figure.png" pageId="17" pageNumber="18">Fig. 60</figureCitation>
), tubular, curved.
</paragraph>
</subSubSection>
<subSubSection id="C30097CE3D65A1609D90F837FC430E16" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BA5C4453D65A1639D90F837FAB908C2" blockId="17.[151,1436,996,1992]" box="[199,1290,1968,1992]" pageId="17" pageNumber="18">
<emphasis id="B96E18573D65A1639D90F837FEFE08C2" bold="true" box="[199,333,1968,1992]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Etymology.</emphasis>
The name
<taxonomicName id="4C1ABFC63D65A1639C9CF837FDC208C2" authority=", Latin" authorityName="Latin" box="[459,625,1968,1992]" class="Insecta" family="Leiodidae" genus="Pinodytes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B96E18573D65A1639C9CF837FD9508C2" box="[459,550,1968,1992]" italics="true" pageId="17" pageNumber="18">borealis</emphasis>
, Latin
</taxonomicName>
, northern, refers to the northern distribution of this species.
</paragraph>
<paragraph id="8BA5C4453D66A1609D90FF10FC430E16" blockId="18.[151,1436,151,284]" pageId="18" pageNumber="19">
<emphasis id="B96E18573D66A1609D90FF10FEF90FA5" box="[199,330,151,176]" inLineHeading="true" pageId="18" pageNumber="19" reason="1">
<emphasis id="B96E18573D66A1609D90FF10FEF00FBA" bold="true" box="[199,323,151,176]" pageId="18" pageNumber="19">Bionomics</emphasis>
.
</emphasis>
<materialsCitation id="3B72CE183D66A1609C05FF10FC580E16" location="These" municipality="We" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Alaska">
The collections are all from leaf litter of broadleaf forests (aspens) on loess soils. These soils are deep, well drained, and warmer than other soil
<typeStatus id="54A17AE73D66A1609FE0FF3BFD410FDE" box="[695,754,188,212]" pageId="18" pageNumber="19">types</typeStatus>
in central
<collectingRegion id="49DE0AA73D66A1609E27FF3BFC720FDE" box="[880,961,188,212]" country="United States of America" name="Alaska" pageId="18" pageNumber="19">Alaska</collectingRegion>
.
<collectingMunicipality id="6BC15E3F3D66A1609E99FF3BFC410FD9" box="[974,1010,188,211]" pageId="18" pageNumber="19">We</collectingMunicipality>
suggest that these soil properties are important for the northern survival of the species.
<location id="8EC5929E3D66A1609F85FF58FCA60FFD" LSID="urn:lsid:plazi:treatment:03B375533D65A1609DC0FCFEFC430E16:8EC5929E3D66A1609F85FF58FCA60FFD" box="[722,789,223,247]" municipality="We" name="These" pageId="18" pageNumber="19" stateProvince="Alaska">These</location>
soils are the same as those in which is found the eyeless northern relictual soil staphylinid
<taxonomicName id="4C1ABFC63D66A1609F45FE83FC580E16" authority="Smetana (1986)" authorityName="Smetana" authorityYear="1986" box="[530,1003,260,284]" class="Insecta" family="Staphylinidae" genus="Chionotyphlus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="alaskensis">
<emphasis id="B96E18573D66A1609F45FE83FC9C0E16" box="[530,815,260,284]" italics="true" pageId="18" pageNumber="19">Chionotyphlus alaskensis</emphasis>
<bibRefCitation id="EF8BB9B43D66A1609E60FE83FC580E16" author="Smetana, A." box="[823,1003,260,284]" pageId="18" pageNumber="19" pagination="171 - 187" refId="ref57254" refString="Smetana, A. 1986. Chionotyphylus alaskensis n. g., n. sp., a Tertiary relict from unglaciated interior Alaska (Coleoptera, Staphylinidae). Nouvelle Revue d'Entomologie (n. s.), 3, 171 - 187." type="journal article" year="1986">Smetana (1986)</bibRefCitation>
</taxonomicName>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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