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<document id="5215A20D93F635892C8070878867B455" ID-CLB-Dataset="53777" ID-DOI="10.11646/zootaxa.5222.2.4" ID-GBIF-Dataset="c60dc4ba-6d53-4a47-9460-80703bcfedce" ID-ISSN="1175-5326" ID-Zenodo-Dep="7456681" ID-ZooBank="55F69A43-2837-4E5F-936D-5ADD23BA5790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1671438652947" checkinUser="plazi" docAuthor="Yasunaga, Tomohide &amp; Duwal, Ram Keshari" docDate="2022" docId="03AB87FF6B10175E819BFDE4FA5DF9BA" docLanguage="en" docName="zootaxa.5222.2.4.pdf" docOrigin="Zootaxa 5222 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sejanus Distant 1910" docType="treatment" docVersion="5" lastPageNumber="158" masterDocId="FF92FF876B11175D810CFF8EFFC1FFB9" masterDocTitle="Two new unique hibiscus-inhabiting species of the plant bug genus Sejanus Distant from Japan and Taiwan (Hemiptera: Heteroptera: Miridae: Phylinae)" masterLastPageNumber="167" masterPageNumber="155" pageNumber="156" updateTime="1699413651002" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="060068C61658AA8AFEA28741A91BFEAE">Two new unique hibiscus-inhabiting species of the plant bug genus Sejanus Distant from Japan and Taiwan (Hemiptera: Heteroptera: Miridae: Phylinae)</mods:title>
</mods:titleInfo>
<mods:name id="BF79A6160A05A22D919CB36B8C3819A3" type="personal">
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<mods:roleTerm id="0D350E56DC3AE717F96A1F8737C61DEA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC1BED9CC580066D1247973A61A08E6F">Yasunaga, Tomohide</mods:namePart>
<mods:affiliation id="EF610879F92B8F4E1DC3CA4E0197EBE6">Research Associate, Division of Invertebrate Zoology, American Museum of Natural History, Central Park at 79 th Street, New York, NY 10024, c / o Nameshi 2 - 33 - 2, Nagasaki 852 - 8061, Japan.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="3B2BC5F15D3387346B9E2741B6949FFA">Duwal, Ram Keshari</mods:namePart>
<mods:nameIdentifier id="86CFFD9D655E573A3F14234CCF2507F4" type="ORCID">0000-0002-2068-461X</mods:nameIdentifier>
<mods:affiliation id="D1CCA2E967F4816B1A70B9F24DCE3621">Visiting Researcher, Agriculture &amp; Agri-Food Canada Environmental Health, K. W. Neatby: Bldg # 20, 960 Carling Avenue, Central Experimental Farm, Ottawa Ontario, Canada K 1 A 0 C 6. ramkeshariduwal @ gmail. com; https: // orcid. org / 0000 - 0002 - 2068 - 461 X</mods:affiliation>
<mods:nameIdentifier id="92857E8114646B411028A0F8D9CB5FF9" type="email">ramkeshariduwal@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="3CFA03C5BF89AE62C82F590DA5A23008">text</mods:typeOfResource>
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<mods:title id="C4AD2D76C572303B9143B54A74F4D537">Zootaxa</mods:title>
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<mods:part id="416DCB6173C6B9C2BEB7A67029440A50">
<mods:date id="913BAA0424697F249B184F4121A422AA">2022</mods:date>
<mods:detail id="ED5D69562508D2C94661E61E6E4EA855" type="pubDate">
<mods:number id="E2403B2DFA2DCC16938F810B31B764E5">2022-12-19</mods:number>
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<mods:detail id="C26C9AF2854141CF3D94CE54F0C52CEF" type="volume">
<mods:number id="69C98BF0C44AA33EA2436992BFA62A0C">5222</mods:number>
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<mods:classification id="466ABB042A00F4C0F5159506D75E0A6C">journal article</mods:classification>
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<mods:identifier id="BB9F5D072ADB25A6F0105CA9EE6E0A8A" type="DOI">10.11646/zootaxa.5222.2.4</mods:identifier>
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<treatment id="03AB87FF6B10175E819BFDE4FA5DF9BA" ID-DOI="http://doi.org/10.5281/zenodo.7456675" ID-GBIF-Taxon="206462451" ID-Zenodo-Dep="7456675" LSID="urn:lsid:plazi:treatment:03AB87FF6B10175E819BFDE4FA5DF9BA" httpUri="http://treatment.plazi.org/id/03AB87FF6B10175E819BFDE4FA5DF9BA" lastPageId="3" lastPageNumber="158" pageId="1" pageNumber="156">
<subSubSection id="C31865626B10175C819BFDE4FC9DFD3D" box="[151,860,618,645]" pageId="1" pageNumber="156" type="multiple">
<paragraph id="8BBD36E96B10175C819BFDE4FC9DFD3D" blockId="1.[151,860,618,645]" box="[151,860,618,645]" pageId="1" pageNumber="156">
<heading id="D0F581856B10175C819BFDE4FC9DFD3D" bold="true" box="[151,860,618,645]" fontSize="11" level="1" pageId="1" pageNumber="156" reason="1">
<emphasis id="B976EAFB6B10175C819BFDE4FC9DFD3D" bold="true" box="[151,860,618,645]" pageId="1" pageNumber="156">
Checklist of
<taxonomicName id="4C024D6A6B10175C8038FDE4FE32FD3D" box="[308,499,618,644]" pageId="1" pageNumber="156">Sejanus species</taxonomicName>
known in
<collectingCountry id="F31576796B10175C837AFDE5FD02FD3C" box="[630,707,619,645]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
and
<collectingCountry id="F31576796B10175C820CFDE4FC9DFD3D" box="[768,860,618,644]" name="Taiwan" pageId="1" pageNumber="156">Taiwan</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31865626B10175C819BFD3DFD29FA78" pageId="1" pageNumber="156" type="synonymic_list">
<paragraph id="8BBD36E96B10175C819BFD3DFDFCFD74" blockId="1.[151,1437,691,1473]" box="[151,573,691,717]" pageId="1" pageNumber="156">
Tribe
<taxonomicName id="4C024D6A6B10175C81D6FD3DFDFCFD74" authority="Schuh, 1974" authorityName="Schuh" authorityYear="1974" box="[218,573,691,717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="tribe" tribe="Leucophoropterini">Leucophoropterini Schuh, 1974</taxonomicName>
</paragraph>
<paragraph id="8BBD36E96B10175C819BFD59FDA4FD48" blockId="1.[151,1437,691,1473]" box="[151,613,727,753]" pageId="1" pageNumber="156">
Subtribe
<taxonomicName id="4C024D6A6B10175C81F1FD59FDA4FD48" authority="Schuh, 1974" authorityName="Schuh" authorityYear="1974" box="[253,613,727,753]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="subTribe" subTribe="Leucophoropterina">Leucophoropterina Schuh, 1974</taxonomicName>
</paragraph>
<paragraph id="8BBD36E96B10175C819BFD75FE1AFCAC" blockId="1.[151,1437,691,1473]" box="[151,475,763,789]" pageId="1" pageNumber="156">
Genus
<taxonomicName id="4C024D6A6B10175C81EAFD75FE1AFCAC" authority="Distant, 1910" authorityName="Distant" authorityYear="1910" box="[230,475,763,789]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAFB6B10175C81EAFD75FEFCFCAC" box="[230,317,763,789]" italics="true" pageId="1" pageNumber="156">Sejanus</emphasis>
Distant, 1910
</taxonomicName>
</paragraph>
<paragraph id="8BBD36E96B10175C819BFC91FBB3FC80" blockId="1.[151,1437,691,1473]" box="[151,1138,799,825]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFC91FE76FC80" authority="Yasunaga, 2001" authorityName="Yasunaga" authorityYear="2001" box="[151,439,799,825]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="amami">
<emphasis id="B976EAFB6B10175C819BFC91FF3EFC81" box="[151,255,799,825]" italics="true" pageId="1" pageNumber="156">S. amami</emphasis>
Yasunaga, 2001
</taxonomicName>
—
<emphasis id="B976EAFB6B10175C80D8FC91FD9CFC80" box="[468,605,799,825]" italics="true" pageId="1" pageNumber="156">Distribution</emphasis>
:
<collectingCountry id="F31576796B10175C8360FC91FD6DFC80" box="[620,684,799,825]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(Amami-Oshima Is.)—
<emphasis id="B976EAFB6B10175C82B4FC91FC2CFC81" box="[952,1005,799,824]" italics="true" pageId="1" pageNumber="156">Host</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFCCDFA56FCE4" blockId="1.[151,1437,691,1473]" box="[151,1431,835,861]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFCCDFD8BFCE4" authority="Yasunaga &amp; Duwal, 2020" authorityName="Yasunaga &amp; Duwal" authorityYear="2020" box="[151,586,835,861]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="azumanus">
<emphasis id="B976EAFB6B10175C819BFCCDFEE2FCE5" box="[151,291,835,861]" italics="true" pageId="1" pageNumber="156">S. azumanus</emphasis>
Yasunaga &amp; Duwal, 2020
</taxonomicName>
—
<collectingCountry id="F31576796B10175C836AFCCDFD67FCE4" box="[614,678,835,861]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(northern Honshu)—
<taxonomicName id="4C024D6A6B10175C8295FCCDFA52FCE4" authority="Thunb." authorityName="Thunb." box="[921,1427,835,861]" class="Magnoliopsida" family="Elaeagnaceae" genus="Elaeagnus" kingdom="Plantae" order="Rosales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="umbellata">
<emphasis id="B976EAFB6B10175C8295FCCDFB47FCE4" box="[921,1158,835,861]" italics="true" pageId="1" pageNumber="156">Elaeagnus umbellata</emphasis>
Thunb. (Elaeagnaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFCE9FBA4FC38" blockId="1.[151,1437,691,1473]" box="[151,1125,871,897]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFCE9FE1CFC38" authority="Yasunaga, 2001" authorityName="Yasunaga" authorityYear="2001" box="[151,477,871,897]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="breviniger">
<emphasis id="B976EAFB6B10175C819BFCE9FEE6FC38" box="[151,295,871,897]" italics="true" pageId="1" pageNumber="156">S. breviniger</emphasis>
Yasunaga, 2001
</taxonomicName>
—
<collectingCountry id="F31576796B10175C830DFCE9FD80FC38" box="[513,577,871,897]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(
<collectingRegion id="49C6F80B6B10175C835CFCE9FD76FC38" box="[592,695,871,897]" country="Japan" name="Okinawa" pageId="1" pageNumber="156">Okinawa</collectingRegion>
Is.).—
<taxonomicName id="4C024D6A6B10175C8209FCE9FBA3FC38" box="[773,1122,871,897]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B976EAFB6B10175C8209FCE9FCA8FC38" box="[773,873,871,897]" italics="true" pageId="1" pageNumber="156">Mallotus</emphasis>
spp. (Euphorbiaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFC05FAF0FC1C" blockId="1.[151,1437,691,1473]" box="[151,1329,907,933]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFC05FE34FC1C" authority="Yasunaga &amp; Duwal" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[151,501,907,933]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="epicurus" status="sp. nov.">
<emphasis id="B976EAFB6B10175C819BFC05FED2FC1D" box="[151,275,907,933]" italics="true" pageId="1" pageNumber="156">S. epicurus</emphasis>
Yasunaga &amp; Duwal
</taxonomicName>
<emphasis id="B976EAFB6B10175C80F0FC05FD96FC1C" bold="true" box="[508,599,907,933]" pageId="1" pageNumber="156">
<taxonomicNameLabel id="A24557806B10175C80F0FC05FDF8FC1C" box="[508,569,907,933]" pageId="1" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
—
</emphasis>
<collectingCountry id="F31576796B10175C835BFC05FD68FC1C" box="[599,681,907,933]" name="Taiwan" pageId="1" pageNumber="156">Taiwan</collectingCountry>
(
<collectingRegion id="49C6F80B6B10175C83BBFC05FCE0FC1C" box="[695,801,907,933]" country="Taiwan" name="Pingtung" pageId="1" pageNumber="156">Pingtung</collectingRegion>
).—
<taxonomicName id="4C024D6A6B10175C8246FC05FAECFC1C" authority="S. Y. Hu" authorityName="S. Y. Hu" box="[842,1325,907,933]" class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="taiwanensis">
<emphasis id="B976EAFB6B10175C8246FC05FBF6FC1D" box="[842,1079,907,933]" italics="true" pageId="1" pageNumber="156">Hibiscus taiwanensis</emphasis>
S.Y. Hu (Malvaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFC21FEB4FC54" blockId="1.[151,1437,691,1473]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFC21FE22FC70" authority="Yasunaga, 2001" authorityName="Yasunaga" authorityYear="2001" box="[151,483,943,969]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="juglandis">
<emphasis id="B976EAFB6B10175C819BFC21FEE0FC70" box="[151,289,943,969]" italics="true" pageId="1" pageNumber="156">S. juglandis</emphasis>
Yasunaga, 2001
</taxonomicName>
—
<collectingCountry id="F31576796B10175C8300FC21FD8DFC70" box="[524,588,943,969]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(
<collectingRegion id="49C6F80B6B10175C836CFC21FD12FC70" box="[608,723,943,969]" country="Japan" name="Hokkaido" pageId="1" pageNumber="156">Hokkaido</collectingRegion>
, Honshu, Shikoku, Kyushu).—
<taxonomicName id="4C024D6A6B10175C854EFC21FEB3FC54" authority="Maxim." authorityName="Maxim." class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="mandshurica">
<emphasis id="B976EAFB6B10175C854EFC21FAF9FC70" box="[1090,1336,943,969]" italics="true" pageId="1" pageNumber="156">Juglans mandshurica</emphasis>
Maxim. (Juglandaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFC79FCD6FB8C" blockId="1.[151,1437,691,1473]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFC79FD62FBA8" authority="Yasunaga, Ishikawa &amp; Ito, 2013" authorityName="Yasunaga, Ishikawa &amp; Ito" authorityYear="2013" box="[151,675,1015,1041]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="komabanus">
<emphasis id="B976EAFB6B10175C819BFC79FEF5FBA8" box="[151,308,1015,1041]" italics="true" pageId="1" pageNumber="156">S. komabanus</emphasis>
<bibRefCitation id="EF934B186B10175C8036FC79FD62FBA8" author="Yasunaga, T. &amp; Ishikawa, T. &amp; Ito, M." box="[314,675,1015,1041]" pageId="1" pageNumber="156" pagination="151 - 160" refId="ref7879" refString="Yasunaga, T., Ishikawa, T. &amp; Ito, M. (2013) Two new species of the plant bug genus Sejanus Distant from Japan (Heteroptera: Miridae: Phylinae: Leucophoropterini), inhabiting urbanized environments or gardens. Tijdschrift voor Entomologie, 156, 151 - 160. https: // doi. org / 10.1163 / 22119434 - 00002025" type="journal article" year="2013">Yasunaga, Ishikawa &amp; Ito, 2013</bibRefCitation>
</taxonomicName>
—
<collectingCountry id="F31576796B10175C83B3FC79FD3EFBA8" box="[703,767,1015,1041]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(Honshu),
<collectingCountry id="F31576796B10175C8273FC79FC02FBA8" box="[895,963,1015,1041]" name="South Korea" pageId="1" pageNumber="156">Korea</collectingCountry>
(Chungcheongmam-do,
<collectingRegion id="49C6F80B6B10175C85D0FC79FAB2FBA8" box="[1244,1395,1015,1041]" country="South Korea" name="Gyeonggi" pageId="1" pageNumber="156">Gyeonggi-do</collectingRegion>
).—
<taxonomicName id="4C024D6A6B10175C81CAFB95FCD2FB8C" authority="(Lf.) Mull. Arg." authorityName="Mull. Arg." baseAuthorityName="Lf." box="[198,787,1051,1077]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="japonicus">
<emphasis id="B976EAFB6B10175C81CAFB95FE5CFB8D" box="[198,413,1051,1077]" italics="true" pageId="1" pageNumber="156">Mallotus japonicus</emphasis>
(Lf.) Müll.Arg. (Euphorbiaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFBB1FEFDFBC4" blockId="1.[151,1437,691,1473]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFBB1FE1DFBE0" authority="Schuh, 1984" authorityName="Schuh" authorityYear="1984" box="[151,476,1087,1113]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="neofunereus">
<emphasis id="B976EAFB6B10175C819BFBB1FE81FBE0" box="[151,320,1087,1113]" italics="true" pageId="1" pageNumber="156">S. neofunereus</emphasis>
<bibRefCitation id="EF934B186B10175C8041FBB1FE1DFBE0" author="Schuh, R. T." box="[333,476,1087,1113]" pageId="1" pageNumber="156" pagination="1 - 476" refId="ref7217" refString="Schuh, R. T. (1984) Revision of the Phylinae (Hemiptera, Miridae) of the Indo-Pacific. Bulletin of the American Museum of Natural History, 177, 1 - 476." type="journal article" year="1984">Schuh, 1984</bibRefCitation>
</taxonomicName>
—
<collectingCountry id="F31576796B10175C80F5FBB1FDFBFBE0" box="[505,570,1087,1113]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(Ryukyus:
<collectingRegion id="49C6F80B6B10175C83CAFBB1FCB0FBE0" box="[710,881,1087,1113]" country="Japan" name="Okinawa" pageId="1" pageNumber="156">Okinawa Pref.</collectingRegion>
),
<collectingCountry id="F31576796B10175C8285FBB1FBD4FBE0" box="[905,1045,1087,1113]" name="Hong Kong" pageId="1" pageNumber="156">Hong Kong</collectingCountry>
,
<collectingCountry id="F31576796B10175C8529FBB1FB67FBE0" box="[1061,1190,1087,1113]" name="Philippines" pageId="1" pageNumber="156">Philippines</collectingCountry>
,
<collectingCountry id="F31576796B10175C85BAFBB1FAC9FBE0" box="[1206,1288,1087,1113]" name="Taiwan" pageId="1" pageNumber="156">Taiwan</collectingCountry>
(
<collectingRegion id="49C6F80B6B10175C8410FBB1FAB2FBE0" box="[1308,1395,1087,1113]" country="Taiwan" name="Nantou" pageId="1" pageNumber="156">Nantou</collectingRegion>
).— Unknown.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFB09FBA4FB7C" blockId="1.[151,1437,691,1473]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFB09FE04FB18" authority="(Reuter, 1906)" baseAuthorityName="Reuter" baseAuthorityYear="1906" box="[151,453,1159,1185]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B976EAFB6B10175C819BFB09FED4FB19" box="[151,277,1159,1185]" italics="true" pageId="1" pageNumber="156">S. potanini</emphasis>
(Reuter, 1906)
</taxonomicName>
—
<collectingCountry id="F31576796B10175C80EEFB09FDE2FB18" box="[482,547,1159,1185]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(
<collectingRegion id="49C6F80B6B10175C833BFB09FD6BFB18" box="[567,682,1159,1185]" country="Japan" name="Hokkaido" pageId="1" pageNumber="156">Hokkaido</collectingRegion>
, Honshu, Shikoku, Kyushu), Russian Far East,
<collectingCountry id="F31576796B10175C85D3FB09FAE7FB18" box="[1247,1318,1159,1185]" name="South Korea" pageId="1" pageNumber="156">Korea</collectingCountry>
,
<collectingCountry id="F31576796B10175C843AFB09FABAFB18" box="[1334,1403,1159,1185]" name="China" pageId="1" pageNumber="156">China</collectingCountry>
.— Various deciduous broadleaf trees, particularly on willows,
<taxonomicName id="4C024D6A6B10175C826CFB25FBA0FB7C" box="[864,1121,1195,1221]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B976EAFB6B10175C826CFB25FC59FB7C" box="[864,920,1195,1221]" italics="true" pageId="1" pageNumber="156">Salix</emphasis>
spp. (Salicaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFB41FE98FAB4" blockId="1.[151,1437,691,1473]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFB41FE3CFB50" authority="Yasunaga &amp; Duwal" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[151,509,1231,1257]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="species" species="ryujin" status="sp. nov.">
<emphasis id="B976EAFB6B10175C819BFB41FEC1FB51" box="[151,256,1231,1257]" italics="true" pageId="1" pageNumber="156">S. ryujin</emphasis>
Yasunaga &amp; Duwal
</taxonomicName>
<emphasis id="B976EAFB6B10175C8301FB41FDB3FB50" bold="true" box="[525,626,1231,1257]" pageId="1" pageNumber="156">
<taxonomicNameLabel id="A24557806B10175C8301FB41FD95FB50" box="[525,596,1231,1257]" pageId="1" pageNumber="156" rank="species">n. sp.</taxonomicNameLabel>
—
</emphasis>
<collectingCountry id="F31576796B10175C837EFB41FD73FB50" box="[626,690,1231,1257]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(Kyushu:
<collectingRegion id="49C6F80B6B10175C8237FB41FC2FFB50" box="[827,1006,1231,1257]" country="Japan" name="Nagasaki" pageId="1" pageNumber="156">Nagasaki Pref.</collectingRegion>
).
<emphasis id="B976EAFB6B10175C82F5FB41FBD7FB50" box="[1017,1046,1231,1257]" italics="true" pageId="1" pageNumber="156">—</emphasis>
<taxonomicName id="4C024D6A6B10175C851AFB41FE97FAB4" authority="Sieb. et Zucc." authorityName="Sieb. et Zucc." class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="hamabo">
<emphasis id="B976EAFB6B10175C851AFB41FB1EFB50" box="[1046,1247,1231,1257]" italics="true" pageId="1" pageNumber="156">Hibiscus hamabo</emphasis>
Sieb. et Zucc. (Malvaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36E96B10175C819BFA99FD29FA78" blockId="1.[151,1437,691,1473]" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C819BFA99FD44FA88" authority="Yasunaga &amp; Ishikawa, 2013" authorityName="Yasunaga &amp; Ishikawa" authorityYear="2013" box="[151,645,1303,1329]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="vivaricolus">
<emphasis id="B976EAFB6B10175C819BFA99FEF2FA88" box="[151,307,1303,1329]" italics="true" pageId="1" pageNumber="156">S. vivaricolus</emphasis>
<bibRefCitation id="EF934B186B10175C8033FA99FD44FA88" author="Yasunaga, T. &amp; Ishikawa, T. &amp; Ito, M." box="[319,645,1303,1329]" pageId="1" pageNumber="156" pagination="151 - 160" refId="ref7879" refString="Yasunaga, T., Ishikawa, T. &amp; Ito, M. (2013) Two new species of the plant bug genus Sejanus Distant from Japan (Heteroptera: Miridae: Phylinae: Leucophoropterini), inhabiting urbanized environments or gardens. Tijdschrift voor Entomologie, 156, 151 - 160. https: // doi. org / 10.1163 / 22119434 - 00002025" type="journal article" year="2013">Yasunaga &amp; Ishikawa, 2013</bibRefCitation>
</taxonomicName>
—
<collectingCountry id="F31576796B10175C83AEFA99FD23FA88" box="[674,738,1303,1329]" name="Japan" pageId="1" pageNumber="156">Japan</collectingCountry>
(SW Honshu, Shikoku, Kyushu, and Nakadori, Wakamatsu and Yakushima Islands);
<collectingCountry id="F31576796B10175C80D7FAB5FDDEFAEC" box="[475,543,1339,1365]" name="South Korea" pageId="1" pageNumber="156">Korea</collectingCountry>
(
<collectingRegion id="49C6F80B6B10175C8321FAB5FD04FAEC" box="[557,709,1339,1365]" country="South Korea" name="Gyeonggi" pageId="1" pageNumber="156">Gyeonggi-do</collectingRegion>
,
<collectingRegion id="49C6F80B6B10175C83C3FAB5FD3DFAEC" box="[719,764,1339,1365]" country="South Korea" name="Jeju" pageId="1" pageNumber="156">Jeju</collectingRegion>
Island).—
<taxonomicName id="4C024D6A6B10175C827FFAB5FA58FAEC" authority="K. Koch" authorityName="K. Koch" box="[883,1433,1339,1365]" class="Magnoliopsida" family="Elaeocarpaceae" genus="Elaeocarpus" kingdom="Plantae" order="Oxalidales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="zollingeri">
<emphasis id="B976EAFB6B10175C827FFAB5FBB0FAEC" box="[883,1137,1339,1365]" italics="true" pageId="1" pageNumber="156">Elaeocarpus zollingeri</emphasis>
K.Koch (Elaeocarpaceae)
</taxonomicName>
,
<taxonomicName id="4C024D6A6B10175C81CAFAD1FD57FAC0" authority="Thunb." authorityName="Thunb." box="[198,662,1375,1401]" class="Magnoliopsida" family="Oleaceae" genus="Ligustrum" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="japonicum">
<emphasis id="B976EAFB6B10175C81CAFAD1FE79FAC1" box="[198,440,1375,1400]" italics="true" pageId="1" pageNumber="156">Ligustrum japonicum</emphasis>
Thunb. (Oleaceae)
</taxonomicName>
,
<emphasis id="B976EAFB6B10175C83AFFAD1FB8FFAC0" box="[675,1102,1375,1401]" italics="true" pageId="1" pageNumber="156">
<taxonomicName id="4C024D6A6B10175C83AFFAD1FCBEFAC1" box="[675,895,1375,1401]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="japonicus">Mallotus japonicus</taxonomicName>
,
<taxonomicName id="4C024D6A6B10175C8280FAD1FB8FFAC0" box="[908,1102,1375,1401]" class="Magnoliopsida" family="Euphorbiaceae" genus="Triadica" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="sebifera">Triadica sebifera</taxonomicName>
</emphasis>
(L.) Small, (
<taxonomicName id="4C024D6A6B10175C85E8FAD1FA50FAC0" box="[1252,1425,1375,1401]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
),
<taxonomicName id="4C024D6A6B10175C81CAFA0DFD52FA24" authority="Sieb. &amp; Zucc." authorityName="Sieb. &amp; Zucc." box="[198,659,1411,1437]" class="Magnoliopsida" family="Myricaceae" genus="Myrica" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="rubra">
<emphasis id="B976EAFB6B10175C81CAFA0DFE9AFA24" box="[198,347,1411,1437]" italics="true" pageId="1" pageNumber="156">Myrica rubra</emphasis>
Sieb. &amp; Zucc. (Myricaceae)
</taxonomicName>
,
<taxonomicName id="4C024D6A6B10175C8397FA0DFB19FA24" authority="Sieb. &amp; Zucc." authorityName="Sieb. &amp; Zucc." box="[667,1240,1411,1437]" class="Magnoliopsida" family="Rutaceae" genus="Zanthoxylum" kingdom="Plantae" order="Sapindales" pageId="1" pageNumber="156" phylum="Tracheophyta" rank="species" species="ailanthoides">
<emphasis id="B976EAFB6B10175C8397FA0DFC7AFA24" box="[667,955,1411,1437]" italics="true" pageId="1" pageNumber="156">Zanthoxylum ailanthoides</emphasis>
Sieb. &amp; Zucc. (Rutaceae)
</taxonomicName>
; this polyphagous species is considered predominantly predaceous.
</paragraph>
</subSubSection>
<subSubSection id="C31865626B10175C819BF99DFEDBF994" box="[151,282,1555,1581]" pageId="1" pageNumber="156" type="multiple">
<paragraph id="8BBD36E96B10175C819BF99DFEDBF994" blockId="1.[151,282,1555,1581]" box="[151,282,1555,1581]" pageId="1" pageNumber="156">
<heading id="D0F581856B10175C819BF99DFEDBF994" bold="true" box="[151,282,1555,1581]" fontSize="11" level="1" pageId="1" pageNumber="156" reason="1">
<emphasis id="B976EAFB6B10175C819BF99DFEDBF994" bold="true" box="[151,282,1555,1581]" pageId="1" pageNumber="156">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31865626B10175C819BF9D4FE70F9CD" box="[151,433,1626,1653]" pageId="1" pageNumber="156" type="nomenclature">
<paragraph id="8BBD36E96B10175C819BF9D4FE70F9CD" blockId="1.[151,433,1626,1653]" box="[151,433,1626,1653]" pageId="1" pageNumber="156">
<heading id="D0F581856B10175C819BF9D4FE70F9CD" bold="true" box="[151,433,1626,1653]" fontSize="11" level="1" pageId="1" pageNumber="156" reason="1">
<emphasis id="B976EAFB6B10175C819BF9D4FE70F9CD" bold="true" box="[151,433,1626,1653]" pageId="1" pageNumber="156">
Genus
<taxonomicName id="4C024D6A6B10175C81E2F9D5FE70F9CD" ID-CoL="7F6D" ID-ENA="567818" authority="Distant" authorityName="Distant" authorityYear="1910" box="[238,433,1626,1653]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAFB6B10175C81E2F9D5FE8FF9CC" bold="true" box="[238,334,1627,1653]" italics="true" pageId="1" pageNumber="156">Sejanus</emphasis>
Distant
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31865626B10175C819BF92CFD2AF82C" pageId="1" pageNumber="156" type="diagnosis">
<paragraph id="8BBD36E96B10175C819BF92CFD2AF82C" blockId="1.[151,1437,1698,2013]" pageId="1" pageNumber="156">
<emphasis id="B976EAFB6B10175C819BF92CFED5F905" bold="true" box="[151,276,1698,1724]" pageId="1" pageNumber="156">Diagnosis:</emphasis>
This genus was well defined by
<bibRefCitation id="EF934B186B10175C8381F92DFCE7F904" author="Schuh, R. T." box="[653,806,1699,1725]" pageId="1" pageNumber="156" pagination="1 - 476" refId="ref7217" refString="Schuh, R. T. (1984) Revision of the Phylinae (Hemiptera, Miridae) of the Indo-Pacific. Bulletin of the American Museum of Natural History, 177, 1 - 476." type="journal article" year="1984">Schuh (1984)</bibRefCitation>
and Menard &amp; Schuh (2014); distinguished from other genera of
<taxonomicName id="4C024D6A6B10175C8005F949FE18F958" authorityName="Schuh" authorityYear="1974" box="[265,473,1735,1761]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="tribe" tribe="Leucophoropterini">Leucophoropterini</taxonomicName>
principally by the following characters: Body generally fuscous, small-sized (mostly
<quantity id="4CFA9B0C6B10175C819BF965FF37F8BC" box="[151,246,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="156" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
in total length); dorsum well-polished, shining, with uniformly distributed, simple setae; base of cuneus often with ivory (or occasionally orange-red) fascia (cf.
<figureCitation id="13392A6C6B10175C822DF881FC5EF890" box="[801,927,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1907,1931]" captionTargetBox="[151,1436,306,1880]" captionTargetId="figure-87@2.[151,1436,306,1880]" captionTargetPageId="2" captionText="FIGURE 1. Habitat (A–C) and habitus images of living individuals of Sejanus epicurus n. sp. (C–J) and Termatophylum orientale Poppius, female (K, sharing same niche). A–C. Hibiscus taiwanensis, breeding host of Sejanus epicurus n. sp. D. A male adult preyed upon by a spider. E–F. Fourth instar nymph. G. Final (5th) instar nymph. H. Adult female. I–J. Adult males." figureDoi="http://doi.org/10.5281/zenodo.7456683" httpUri="https://zenodo.org/record/7456683/files/figure.png" pageId="1" pageNumber="156">Figs. 1H–J</figureCitation>
,
<figureCitation id="13392A6C6B10175C82A1F881FC0DF890" box="[941,972,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1874,1898]" captionTargetBox="[151,1436,308,1837]" captionTargetId="figure-77@4.[151,1436,308,1850]" captionTargetPageId="4" captionText="FIGURE 2. Habitat (A–C), and habitus images of Sejanus ryujin n. sp. (D–H) and S. epicurus n. sp. (I–J), and Orius strigicollis (Poppius) (5th instar nymph) and an undetermined thrips sharing same niche (K); G–I showing ventral view of fresh specimens. A–C. Hibiscus hamabo, breeding host of S, ryujin n. sp. at the type locality (Ryujin Island, Omura City). D–F, H. Adult female. F–G. Adult males. I. S. epicurus n. sp., male. J. Same, female." figureDoi="http://doi.org/10.5281/zenodo.7456685" httpUri="https://zenodo.org/record/7456685/files/figure.png" pageId="1" pageNumber="156">2F</figureCitation>
) that is interrupted at middle or divided into two spots in some species (
<figureCitation id="13392A6C6B10175C8305F8BDFDB5F8F4" box="[521,628,1843,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1620,1644]" captionTargetBox="[204,1383,273,1597]" captionTargetId="figure-67@5.[204,1383,273,1597]" captionTargetPageId="5" captionText="FIGURE 3. Habittus images of live individuals for Sejanus spp. A. S. breviniger, adult males. B. Same, final instar nymph. C. S. komabanus, male. D. Same, female. E. S. vivaricolus, adults. F. Same, female reared with brine-shrimp eggs. G. Same, third instar nymph. H. Same, final instar nymph. I. S. neofunereus, male (Iriomote Is., Ryukyus, Japan). J. Same (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.7456687" httpUri="https://zenodo.org/record/7456687/files/figure.png" pageId="1" pageNumber="156">Fig. 3I–J</figureCitation>
); C- or J-shaped vesica (endosoma) with secondary gonopore opening at apex; small, thin-rimmed female sclerotized ring; and relatively simple interramal sclerite with comb-shaped microstructures and narrow posterior margin sclerite.
</paragraph>
</subSubSection>
<subSubSection id="C31865626B10175E81CBF810FB21FE14" lastPageId="3" lastPageNumber="158" pageId="1" pageNumber="156" type="discussion">
<paragraph id="8BBD36E96B10175F81CBF810FD9BFEA4" blockId="1.[151,1437,1698,2013]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="157" pageId="1" pageNumber="156">
<emphasis id="B976EAFB6B10175C81CBF810FE8DF801" bold="true" box="[199,332,1950,1976]" pageId="1" pageNumber="156">Discussion.</emphasis>
<taxonomicName id="4C024D6A6B10175C805EF811FDC1F800" authority="Distant" authorityName="Distant" authorityYear="1910" box="[338,512,1951,1977]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAFB6B10175C805EF811FE68F800" box="[338,425,1951,1977]" italics="true" pageId="1" pageNumber="156">Sejanus</emphasis>
Distant
</taxonomicName>
is a typical Indo-Pacific group and more than a few undescribed species are present mainly in the Oriental Region (Yasunaga &amp; Duwal, unpublished data). For temperate and cold temperate climatic zones of
<collectingCountry id="F31576796B13175F81F0FF19FEFDFF08" box="[252,316,151,177]" name="Japan" pageId="2" pageNumber="157">Japan</collectingCountry>
and adjacent regions, the fauna was considered to have been almost perfectly clarified by a series of recent works (e.g.,
<bibRefCitation id="EF934B186B13175F8065FF35FDE8FF6C" author="Yasunaga, T." box="[361,553,187,213]" pageId="2" pageNumber="157" pagination="121 - 126" refId="ref7482" refString="Yasunaga, T. (2001 a) A review of the phyline plant bug genus Sejanus Distant (Heteroptera: Miridae) in Japan, with description of three new species. Entomological Science, 4, 121 - 126." type="journal article" year="2001">Yasunaga, 2001a</bibRefCitation>
;
<bibRefCitation id="EF934B186B13175F833DFF35FCE0FF6C" author="Yasunaga, T. &amp; Ishikawa, T. &amp; Ito, M." box="[561,801,187,213]" pageId="2" pageNumber="157" pagination="151 - 160" refId="ref7879" refString="Yasunaga, T., Ishikawa, T. &amp; Ito, M. (2013) Two new species of the plant bug genus Sejanus Distant from Japan (Heteroptera: Miridae: Phylinae: Leucophoropterini), inhabiting urbanized environments or gardens. Tijdschrift voor Entomologie, 156, 151 - 160. https: // doi. org / 10.1163 / 22119434 - 00002025" type="journal article" year="2013">
Yasunaga
<emphasis id="B976EAFB6B13175F83AFFF35FD12FF6C" box="[675,723,187,213]" italics="true" pageId="2" pageNumber="157">et al</emphasis>
., 2013
</bibRefCitation>
;
<bibRefCitation id="EF934B186B13175F8227FF35FC13FF6C" author="Oh, M. &amp; Duwal, R. K. &amp; Lee, S. H." box="[811,978,187,213]" pageId="2" pageNumber="157" pagination="196 - 203" refId="ref7150" refString="Oh, M., Duwal, R. K. &amp; Lee, S. H. (2020) Taxonomic review of genus Sejanus Distant (Heteroptera: Miridae: Phylinae) from Korea, with a new species. Journal of Asia-Pacific Entomology, 23, 196 - 203. https: // doi. org / 10.1016 / j. aspen. 2019.12.008" type="journal article" year="2020">
Oh
<emphasis id="B976EAFB6B13175F825FFF35FC42FF6C" box="[851,899,187,213]" italics="true" pageId="2" pageNumber="157">et al</emphasis>
., 2020
</bibRefCitation>
; Yasunaga &amp; Duwal, 2021). Nonetheless, recent fieldworks have yielded two unique congeners that inhabit cryptic underside of hibiscus bracts and flower buds in southwestern
<collectingCountry id="F31576796B13175F8087FE8DFE0AFEA4" box="[395,459,259,285]" name="Japan" pageId="2" pageNumber="157">Japan</collectingCountry>
and
<collectingCountry id="F31576796B13175F830CFE8DFD97FEA4" box="[512,598,259,285]" name="Taiwan" pageId="2" pageNumber="157">Taiwan</collectingCountry>
.
</paragraph>
<caption id="DF7D66616B13175F819BF8FDFF1BF84E" ID-DOI="http://doi.org/10.5281/zenodo.7456683" ID-Zenodo-Dep="7456683" httpUri="https://zenodo.org/record/7456683/files/figure.png" pageId="2" pageNumber="157" startId="2.[151,250,1907,1931]" targetBox="[151,1436,306,1880]" targetPageId="2">
<paragraph id="8BBD36E96B13175F819BF8FDFF1BF84E" blockId="2.[151,1437,1907,2039]" pageId="2" pageNumber="157">
<emphasis id="B976EAFB6B13175F819BF8FDFED6F832" bold="true" box="[151,279,1907,1931]" pageId="2" pageNumber="157">FIGURE 1.</emphasis>
Habitat (A–C) and habitus images of living individuals of
<taxonomicName id="4C024D6A6B13175F8282F8FDFBFFF832" authority="Yasunaga &amp; Duwal, 2022" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[910,1086,1907,1931]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="epicurus" status="sp. nov.">
<emphasis id="B976EAFB6B13175F8282F8FDFBFFF832" box="[910,1086,1907,1931]" italics="true" pageId="2" pageNumber="157">Sejanus epicurus</emphasis>
</taxonomicName>
<emphasis id="B976EAFB6B13175F8544F8FDFB45F832" bold="true" box="[1096,1156,1907,1931]" pageId="2" pageNumber="157">
<taxonomicNameLabel id="A24557806B13175F8544F8FDFB45F832" box="[1096,1156,1907,1931]" pageId="2" pageNumber="157" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(C–J) and
<taxonomicName id="4C024D6A6B13175F85F2F8FDFE8EF816" authority="Poppius" authorityName="Poppius" authorityYear="1915" class="Insecta" family="Miridae" genus="Termatophylum" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="orientale">
<emphasis id="B976EAFB6B13175F85F2F8FDFF33F816" italics="true" pageId="2" pageNumber="157">Termatophylum orientale</emphasis>
Poppius
</taxonomicName>
, female (K, sharing same niche).
<emphasis id="B976EAFB6B13175F83BCF819FD29F816" bold="true" box="[688,744,1943,1967]" pageId="2" pageNumber="157">A–C.</emphasis>
<taxonomicName id="4C024D6A6B13175F83FCF819FC08F816" box="[752,969,1943,1967]" class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="2" pageNumber="157" phylum="Tracheophyta" rank="species" species="taiwanensis">
<emphasis id="B976EAFB6B13175F83FCF819FC08F816" box="[752,969,1943,1967]" italics="true" pageId="2" pageNumber="157">Hibiscus taiwanensis</emphasis>
</taxonomicName>
, breeding host of
<taxonomicName id="4C024D6A6B13175F8585F819FAF6F816" authority="Yasunaga &amp; Duwal, 2022" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[1161,1335,1943,1967]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="157" phylum="Arthropoda" rank="species" species="epicurus" status="sp. nov.">
<emphasis id="B976EAFB6B13175F8585F819FAF6F816" box="[1161,1335,1943,1967]" italics="true" pageId="2" pageNumber="157">Sejanus epicurus</emphasis>
</taxonomicName>
<emphasis id="B976EAFB6B13175F844CF819FA5DF816" bold="true" box="[1344,1436,1943,1967]" pageId="2" pageNumber="157">
<taxonomicNameLabel id="A24557806B13175F844CF819FABAF816" box="[1344,1403,1943,1967]" pageId="2" pageNumber="157" rank="species">n. sp.</taxonomicNameLabel>
D.
</emphasis>
A male adult preyed upon by a spider.
<emphasis id="B976EAFB6B13175F8323F835FDA1F86A" bold="true" box="[559,608,1979,2003]" pageId="2" pageNumber="157">E–F.</emphasis>
Fourth instar nymph.
<emphasis id="B976EAFB6B13175F8247F835FCA4F86A" bold="true" box="[843,869,1979,2003]" pageId="2" pageNumber="157">G.</emphasis>
Final (5th) instar nymph.
<emphasis id="B976EAFB6B13175F8575F835FB55F86A" bold="true" box="[1145,1172,1979,2003]" pageId="2" pageNumber="157">H.</emphasis>
Adult female.
<emphasis id="B976EAFB6B13175F843CF835FA9BF86A" bold="true" box="[1328,1370,1979,2003]" pageId="2" pageNumber="157">I–J.</emphasis>
Adult males.
</paragraph>
</caption>
<paragraph id="8BBD36E96B12175E81CBFF19FB21FE14" blockId="3.[151,1437,151,429]" pageId="3" pageNumber="158">
Several species of phyline plant bugs, e.g.
<taxonomicName id="4C024D6A6B12175E83DBFF19FB81FF08" authority="Yasunaga" authorityName="Yasunaga" box="[727,1088,151,177]" class="Insecta" family="Miridae" genus="Campylomma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="astica">
<emphasis id="B976EAFB6B12175E83DBFF19FC05FF09" box="[727,964,151,177]" italics="true" pageId="3" pageNumber="158">Campylomma astica</emphasis>
Yasunaga
</taxonomicName>
,
<taxonomicName id="4C024D6A6B12175E855EFF19FAB0FF08" authority="Yasunaga" authorityName="Yasunaga" box="[1106,1393,151,177]" class="Insecta" family="Miridae" genus="Campylomma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="hibiscicola">
<emphasis id="B976EAFB6B12175E855EFF19FB34FF08" box="[1106,1269,151,177]" italics="true" pageId="3" pageNumber="158">C. hibiscicola</emphasis>
Yasunaga
</taxonomicName>
,
<taxonomicName id="4C024D6A6B12175E848FFF19FE55FF6C" authority="Yasunaga" authorityName="Yasunaga" authorityYear="2016" class="Insecta" family="Miridae" genus="Campylomma" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="seunghwani">
<emphasis id="B976EAFB6B12175E848FFF19FEDCFF6C" italics="true" pageId="3" pageNumber="158">C. seunghwani</emphasis>
Yasunaga
</taxonomicName>
and
<taxonomicName id="4C024D6A6B12175E80DFFF35FCA8FF6C" authority="Yasunaga" authorityName="Yasunaga" authorityYear="2010" box="[467,873,187,213]" class="Insecta" family="Miridae" genus="Decomioides" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="verecundus">
<emphasis id="B976EAFB6B12175E80DFFF35FD2EFF6C" box="[467,751,187,213]" italics="true" pageId="3" pageNumber="158">Decomioides verecundus</emphasis>
Yasunaga
</taxonomicName>
, were documented to prefer similarly concealed habitats on
<taxonomicName id="4C024D6A6B12175E8011FF51FE26FF40" box="[285,487,223,249]" class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="species" species="tiliaceus">
<emphasis id="B976EAFB6B12175E8011FF51FE26FF40" box="[285,487,223,249]" italics="true" pageId="3" pageNumber="158">Hibiscus tiliaceus</emphasis>
</taxonomicName>
L. (Yasunaga, 2021) or
<emphasis id="B976EAFB6B12175E820EFF51FCB6FF41" box="[770,887,223,248]" italics="true" pageId="3" pageNumber="158">Macarang</emphasis>
a spp. (
<taxonomicName id="4C024D6A6B12175E82C7FF51FBB4FF40" box="[971,1141,223,249]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
) (
<bibRefCitation id="EF934B186B12175E8581FF51FA86FF40" author="Yasunaga, T." box="[1165,1351,223,249]" pageId="3" pageNumber="158" pagination="50 - 92" refId="ref7594" refString="Yasunaga, T. (2010) Plant bugs of the tribe Phylini in Thailand (Heteroptera: Miridae: Phylinae), with descriptions of six new species from additional areas in tropical and subtropical Asia. Entomologica Americana, 116, 50 - 92. https: // doi. org / 10.1664 / 10 - RA- 006.1" type="journal article" year="2010">Yasunaga, 2010</bibRefCitation>
;
<bibRefCitation id="EF934B186B12175E8458FF51FA50FF40" author="Yasunaga, T." box="[1364,1425,223,249]" pageId="3" pageNumber="158" pagination="459 - 465" refId="ref7654" refString="Yasunaga, T. (2016) A review of the plant bug genus Campylomma Reuter from Indochina (Heteroptera: Miridae: Phylinae: Nasocorini), with description of a new species cryptically inhabiting Macaranga bracts (Euphorbiaceae) in Thailand. Journal of Asia-Pacific Entomology, 19, 459 - 465." type="journal article" year="2016">2016</bibRefCitation>
). Within such niches, thrips (
<taxonomicName id="4C024D6A6B12175E80B2FE8DFD9CFEA4" authorityName="Haliday" authorityYear="1836" box="[446,605,259,285]" class="Insecta" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="order">Thysanoptera</taxonomicName>
), cicadellid leafhoppers(Auchenorrhyncha) and/or aphids (Sternorrhyncha) were observed to co-occur and are assumed to be prey of these phylines (e.g.
<bibRefCitation id="EF934B186B12175E8513FEA9FACEFEF8" author="Nakatani, Y. &amp; Yasunaga T. &amp; Yamada, K." box="[1055,1295,295,321]" pageId="3" pageNumber="158" pagination="41 - 49" refId="ref7107" refString="Nakatani, Y., Yasunaga T. &amp; Yamada, K. (2017) Two species of Deraeocoris Kirschbaum new to Japan (Miridae). Rostria, 61, 41 - 49. [in Japanese with English summary]" type="journal article" year="2017">
Nakatani
<emphasis id="B976EAFB6B12175E8582FEA9FB03FEF8" box="[1166,1218,295,321]" italics="true" pageId="3" pageNumber="158">et al</emphasis>
. 2017
</bibRefCitation>
). Except for some Palearctic members (see above checklist), the breeding host plants of many
<taxonomicName id="4C024D6A6B12175E853EFEC5FB48FEDC" authorityName="Distant" authorityYear="1910" box="[1074,1161,331,357]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAFB6B12175E853EFEC5FB48FEDC" box="[1074,1161,331,357]" italics="true" pageId="3" pageNumber="158">Sejanus</emphasis>
</taxonomicName>
congeners are unknown (Menard &amp; Schuh, 2014). In laboratory tests, both adult and immature forms of
<taxonomicName id="4C024D6A6B12175E8534FEE1FB13FE30" authorityName="Yasunaga &amp; Ishikawa" authorityYear="2013" box="[1080,1234,367,393]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="species" species="vivaricolus">
<emphasis id="B976EAFB6B12175E8534FEE1FB13FE30" box="[1080,1234,367,393]" italics="true" pageId="3" pageNumber="158">S. vivaricolus</emphasis>
</taxonomicName>
can be reared by brine-shrimp eggs (cf.
<figureCitation id="13392A6C6B12175E809AFE1DFE2BFE14" box="[406,490,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1620,1644]" captionTargetBox="[204,1383,273,1597]" captionTargetId="figure-67@5.[204,1383,273,1597]" captionTargetPageId="5" captionText="FIGURE 3. Habittus images of live individuals for Sejanus spp. A. S. breviniger, adult males. B. Same, final instar nymph. C. S. komabanus, male. D. Same, female. E. S. vivaricolus, adults. F. Same, female reared with brine-shrimp eggs. G. Same, third instar nymph. H. Same, final instar nymph. I. S. neofunereus, male (Iriomote Is., Ryukyus, Japan). J. Same (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.7456687" httpUri="https://zenodo.org/record/7456687/files/figure.png" pageId="3" pageNumber="158">Fig. 3F</figureCitation>
); we consider that most
<taxonomicName id="4C024D6A6B12175E83F7FE1DFC6BFE14" box="[763,938,403,429]" pageId="3" pageNumber="158">
<emphasis id="B976EAFB6B12175E83F7FE1DFC93FE14" box="[763,850,403,429]" italics="true" pageId="3" pageNumber="158">Sejanus</emphasis>
species
</taxonomicName>
are principally predaceous.
</paragraph>
</subSubSection>
<subSubSection id="C31865626B12175E819BFE71FD20FDA1" box="[151,737,510,537]" pageId="3" pageNumber="158" type="multiple">
<paragraph id="8BBD36E96B12175E819BFE71FD20FDA1" blockId="3.[151,737,510,537]" box="[151,737,510,537]" pageId="3" pageNumber="158">
<heading id="D0F581856B12175E819BFE71FD20FDA1" bold="true" box="[151,737,510,537]" fontSize="11" level="1" pageId="3" pageNumber="158" reason="1">
<emphasis id="B976EAFB6B12175E819BFE71FD20FDA1" bold="true" box="[151,737,510,537]" pageId="3" pageNumber="158">
Key to
<taxonomicName id="4C024D6A6B12175E81FDFE71FE6FFDA1" box="[241,430,510,537]" pageId="3" pageNumber="158">
<emphasis id="B976EAFB6B12175E81FDFE71FE90FDA0" bold="true" box="[241,337,511,537]" italics="true" pageId="3" pageNumber="158">Sejanus</emphasis>
species
</taxonomicName>
from
<collectingCountry id="F31576796B12175E80F7FE71FD89FDA0" box="[507,584,511,537]" name="Japan" pageId="3" pageNumber="158">Japan</collectingCountry>
and
<collectingCountry id="F31576796B12175E8389FE70FD20FDA1" box="[645,737,510,536]" name="Taiwan" pageId="3" pageNumber="158">Taiwan</collectingCountry>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31865626B12175E819BFDCAFA5DF9BA" pageId="3" pageNumber="158" type="key">
<keyStep id="30F62E4C6B12175E819BFDCAFA5DFD08" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFDCAFA5DFDCF" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFDCAFA5DFDCF" pageId="3" pageNumber="158">
1. Hemelytron totally shiny fuscous; base of cuneus with faint, narrow, and indistinct pale (reddish or whitish) fascia or spots if present at all (cf.
<figureCitation id="13392A6C6B12175E8085FDEFFE13FDCF" box="[393,466,609,631]" captionStart="FIGURE 2" captionStartId="4.[151,250,1874,1898]" captionTargetBox="[151,1436,308,1837]" captionTargetId="figure-77@4.[151,1436,308,1850]" captionTargetPageId="4" captionText="FIGURE 2. Habitat (A–C), and habitus images of Sejanus ryujin n. sp. (D–H) and S. epicurus n. sp. (I–J), and Orius strigicollis (Poppius) (5th instar nymph) and an undetermined thrips sharing same niche (K); G–I showing ventral view of fresh specimens. A–C. Hibiscus hamabo, breeding host of S, ryujin n. sp. at the type locality (Ryujin Island, Omura City). D–F, H. Adult female. F–G. Adult males. I. S. epicurus n. sp., male. J. Same, female." figureDoi="http://doi.org/10.5281/zenodo.7456685" httpUri="https://zenodo.org/record/7456685/files/figure.png" pageId="3" pageNumber="158">Fig. 2E</figureCitation>
,
<figureCitation id="13392A6C6B12175E80D0FDEFFDD5FDCF" box="[476,532,609,630]" captionStart="FIGURE 3" captionStartId="5.[151,250,1620,1644]" captionTargetBox="[204,1383,273,1597]" captionTargetId="figure-67@5.[204,1383,273,1597]" captionTargetPageId="5" captionText="FIGURE 3. Habittus images of live individuals for Sejanus spp. A. S. breviniger, adult males. B. Same, final instar nymph. C. S. komabanus, male. D. Same, female. E. S. vivaricolus, adults. F. Same, female reared with brine-shrimp eggs. G. Same, third instar nymph. H. Same, final instar nymph. I. S. neofunereus, male (Iriomote Is., Ryukyus, Japan). J. Same (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.7456687" httpUri="https://zenodo.org/record/7456687/files/figure.png" pageId="3" pageNumber="158">3C–E</figureCitation>
)......................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BBD36E96B12175E819BFDF1FA5DFD08" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFDF1FA5DFD08" pageId="3" pageNumber="158">
- Hemelytron almost entirely fuscous, except for base of cuneus with defined, white or ivory spots (
<figureCitation id="13392A6C6B12175E8597FDF0FB34FD2A" box="[1179,1269,638,660]" captionStart="FIGURE 3" captionStartId="5.[151,250,1620,1644]" captionTargetBox="[204,1383,273,1597]" captionTargetId="figure-67@5.[204,1383,273,1597]" captionTargetPageId="5" captionText="FIGURE 3. Habittus images of live individuals for Sejanus spp. A. S. breviniger, adult males. B. Same, final instar nymph. C. S. komabanus, male. D. Same, female. E. S. vivaricolus, adults. F. Same, female reared with brine-shrimp eggs. G. Same, third instar nymph. H. Same, final instar nymph. I. S. neofunereus, male (Iriomote Is., Ryukyus, Japan). J. Same (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.7456687" httpUri="https://zenodo.org/record/7456687/files/figure.png" pageId="3" pageNumber="158">Fig. 3I–J</figureCitation>
), or a continuous creamy white fascia (
<figureCitation id="13392A6C6B12175E80A3FD12FDD1FD08" box="[431,528,668,690]" captionStart="FIGURE 1" captionStartId="2.[151,250,1907,1931]" captionTargetBox="[151,1436,306,1880]" captionTargetId="figure-87@2.[151,1436,306,1880]" captionTargetPageId="2" captionText="FIGURE 1. Habitat (A–C) and habitus images of living individuals of Sejanus epicurus n. sp. (C–J) and Termatophylum orientale Poppius, female (K, sharing same niche). A–C. Hibiscus taiwanensis, breeding host of Sejanus epicurus n. sp. D. A male adult preyed upon by a spider. E–F. Fourth instar nymph. G. Final (5th) instar nymph. H. Adult female. I–J. Adult males." figureDoi="http://doi.org/10.5281/zenodo.7456683" httpUri="https://zenodo.org/record/7456683/files/figure.png" pageId="3" pageNumber="158">Fig. 1H–J</figureCitation>
,
<figureCitation id="13392A6C6B12175E8316FD12FDF5FD08" box="[538,564,668,689]" captionStart="FIGURE 2" captionStartId="4.[151,250,1874,1898]" captionTargetBox="[151,1436,308,1837]" captionTargetId="figure-77@4.[151,1436,308,1850]" captionTargetPageId="4" captionText="FIGURE 2. Habitat (A–C), and habitus images of Sejanus ryujin n. sp. (D–H) and S. epicurus n. sp. (I–J), and Orius strigicollis (Poppius) (5th instar nymph) and an undetermined thrips sharing same niche (K); G–I showing ventral view of fresh specimens. A–C. Hibiscus hamabo, breeding host of S, ryujin n. sp. at the type locality (Ryujin Island, Omura City). D–F, H. Adult female. F–G. Adult males. I. S. epicurus n. sp., male. J. Same, female." figureDoi="http://doi.org/10.5281/zenodo.7456685" httpUri="https://zenodo.org/record/7456685/files/figure.png" pageId="3" pageNumber="158">2F</figureCitation>
)...................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFD37FA5DFD55" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFD37FA5DFD77" blockId="3.[151,1437,580,1540]" box="[151,1436,697,719]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFD37FA5DFD77" box="[151,1436,697,719]" pageId="3" pageNumber="158">2. Apex of metafemur clearly yellow, yellowish brown or yellow-orange........................................... 3</keyLead>
</paragraph>
<paragraph id="8BBD36E96B12175E819BFD59FA5DFD55" blockId="3.[151,1437,580,1540]" box="[151,1436,726,748]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFD59FA5DFD55" box="[151,1436,726,748]" pageId="3" pageNumber="158">- Metafemur entirely darkened, or its extreme apex only slightly pale............................................. 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFD7AFA5DFC9E" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFD7AFA5DFCB0" blockId="3.[151,1437,580,1540]" box="[151,1436,754,778]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFD7AFA5DFCB0" box="[151,1436,754,778]" pageId="3" pageNumber="158">
3. Antennal segment II pale brown with darkened apical 1/3–1/2 (♁)/ 1/4–1/6 (
<specimenCount id="9D04FD606B12175E82CEFD7AFC13FCB0" box="[962,978,756,777]" pageId="3" pageNumber="158" type="female">♀</specimenCount>
).................................... 4
</keyLead>
</paragraph>
<paragraph id="8BBD36E96B12175E819BFC9FFA5DFC9E" blockId="3.[151,1437,580,1540]" box="[151,1436,783,807]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFC9FFA5DFC9E" box="[151,1436,783,807]" pageId="3" pageNumber="158">
- More than apical 2/3 (♁)/ 1/2 (
<specimenCount id="9D04FD606B12175E8300FC9FFDDDFC9F" box="[524,540,785,806]" pageId="3" pageNumber="158" type="female">♀</specimenCount>
) of antennal segment II darkened.............................................. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFCA0FA5DFCD8" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFCA0FA5DFCFD" blockId="3.[151,1437,580,1540]" box="[151,1436,814,836]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFCA0FA5DFCFD" box="[151,1436,814,836]" pageId="3" pageNumber="158">
4. Male......................................................................................
<taxonomicName id="4C024D6A6B12175E8428FCA1FA5DFCFD" authorityName="Yasunaga &amp; Duwal" authorityYear="2020" box="[1316,1436,815,836]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="azumanus">
<emphasis id="B976EAFB6B12175E8428FCA1FA5DFCFD" box="[1316,1436,815,836]" italics="true" pageId="3" pageNumber="158">S. azumanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBD36E96B12175E819BFCC2FA5DFCD8" blockId="3.[151,1437,580,1540]" box="[151,1436,843,866]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFCC2FA5DFCD8" box="[151,1436,843,866]" pageId="3" pageNumber="158">- Female............................................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFCE7FA5DFC6F" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFCE7FA5DFC25" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
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5. Labium as long as basal width of pronotum or metafemur; associated with
<taxonomicName id="4C024D6A6B12175E82BCFCE7FBBDFCC7" ID-CoL="8V7ZS" box="[944,1148,873,894]" class="Magnoliopsida" family="Elaeagnaceae" genus="Elaeagnus" kingdom="Plantae" order="Rosales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="species" species="umbellata">
<emphasis id="B976EAFB6B12175E82BCFCE7FBBDFCC7" box="[944,1148,873,894]" italics="true" pageId="3" pageNumber="158">Elaeagnus umbellata</emphasis>
</taxonomicName>
in northern Honshu.................................................................................................
<taxonomicName id="4C024D6A6B12175E8428FC09FA5DFC25" authorityName="Yasunaga &amp; Duwal" authorityYear="2020" box="[1316,1436,903,924]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="azumanus">
<emphasis id="B976EAFB6B12175E8428FC09FA5DFC25" box="[1316,1436,903,924]" italics="true" pageId="3" pageNumber="158">S. azumanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBD36E96B12175E819BFC2AFA5DFC6F" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
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- Labium obviously longer than basal width of pronotum or metafemur; currently known to inhabit a halophyte,
<taxonomicName id="4C024D6A6B12175E85FDFC2AFA57FC00" ID-CoL="3LJTB" box="[1265,1430,932,953]" class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="species" species="hamabo">
<emphasis id="B976EAFB6B12175E85FDFC2AFA57FC00" box="[1265,1430,932,953]" italics="true" pageId="3" pageNumber="158">Hibiscus hamabo</emphasis>
</taxonomicName>
, in warm temperate climatic zone...........................................................
<taxonomicName id="4C024D6A6B12175E85EBFC4FFAF9FC6E" authority="Yasunaga &amp; Duwal, 2022" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[1255,1336,961,983]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="ryujin" status="sp. nov.">
<emphasis id="B976EAFB6B12175E85EBFC4FFAF9FC6E" box="[1255,1336,961,983]" italics="true" pageId="3" pageNumber="158">S. ryujin</emphasis>
</taxonomicName>
<emphasis id="B976EAFB6B12175E8432FC4FFAB4FC6F" bold="true" box="[1342,1397,961,982]" pageId="3" pageNumber="158">
<taxonomicNameLabel id="A24557806B12175E8432FC4FFAB4FC6F" box="[1342,1397,961,982]" pageId="3" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
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)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFC50FA5DFBF2" pageId="3" pageNumber="158">
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<keyLead id="30F395DC6B12175E819BFC50FA5DFBA8" pageId="3" pageNumber="158">
6. Body larger; total body length more than
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(♁)/
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(
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); length from apex of clypeus to cuneal fracture greater than 2.0 mm.....................................................................................
<taxonomicName id="4C024D6A6B12175E8438FC72FA5DFBA8" ID-CoL="4WB7P" baseAuthorityName="Reuter" baseAuthorityYear="1906" box="[1332,1436,1020,1041]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="potanini">
<emphasis id="B976EAFB6B12175E8438FC72FA5DFBA8" box="[1332,1436,1020,1041]" italics="true" pageId="3" pageNumber="158">S. potanini</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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- Body smaller; total body length up to
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(♁)/
<quantity id="4CFA9B0C6B12175E83DBFB97FCE3FB97" box="[727,802,1049,1071]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="158" unit="mm" value="2.5">2.5 mm</quantity>
(
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); length from apex of clypeus to cuneal fracture less than
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......................................................................................
<taxonomicName id="4C024D6A6B12175E841BFBB9FA5DFBF2" authorityName="Yasunaga, Ishikawa &amp; Ito" authorityYear="2013" box="[1303,1436,1078,1100]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="komabanus">
<emphasis id="B976EAFB6B12175E841BFBB9FA5DFBF2" box="[1303,1436,1078,1100]" italics="true" pageId="3" pageNumber="158">S. komabanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFBDAFA5DFB1A" pageId="3" pageNumber="158">
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<keyLead id="30F395DC6B12175E819BFBDAFA5CFBD0" box="[151,1437,1108,1130]" pageId="3" pageNumber="158">
7. Male antennal segment II shorter than basal width of pronotum; ventral cleft of scent efferent system elongate.
<taxonomicName id="4C024D6A6B12175E8416FBDAFA5CFBD0" authorityName="Yasunaga &amp; Ishikawa" authorityYear="2013" box="[1306,1437,1108,1129]" class="Magnoliopsida" family="Salicaceae" genus="Salix" kingdom="Plantae" order="Malpighiales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="species" species="vivaricolus">
<emphasis id="B976EAFB6B12175E8416FBDAFA5CFBD0" box="[1306,1437,1108,1129]" italics="true" pageId="3" pageNumber="158">S. vivaricolus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBD36E96B12175E819BFBFFFA5DFB1A" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFBFFFA5DFB1A" pageId="3" pageNumber="158">
- Male antennal segment II longer than basal width of pronotum; ventral cleft of scent efferent system rounded and shallow; currently known only from
<collectingRegion id="49C6F80B6B12175E80E7FB00FD83FB1D" box="[491,578,1166,1188]" country="Japan" name="Okinawa" pageId="3" pageNumber="158">Okinawa</collectingRegion>
Island........................................................
<taxonomicName id="4C024D6A6B12175E842DFB01FA5DFB1A" ID-CoL="4WB6Q" authorityName="Yasunaga" authorityYear="2001" box="[1313,1436,1166,1188]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="breviniger">
<emphasis id="B976EAFB6B12175E842DFB01FA5DFB1A" box="[1313,1436,1166,1188]" italics="true" pageId="3" pageNumber="158">S. breviniger</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFB22FA5DFB67" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFB22FA5CFB78" blockId="3.[151,1437,580,1540]" box="[151,1437,1196,1218]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFB22FA5CFB78" box="[151,1437,1196,1218]" pageId="3" pageNumber="158">
8. All femora creamy yellow, not darkened (
<figureCitation id="13392A6C6B12175E837FFB22FD0BFB78" box="[627,714,1196,1218]" captionStart="FIGURE 2" captionStartId="4.[151,250,1874,1898]" captionTargetBox="[151,1436,308,1837]" captionTargetId="figure-77@4.[151,1436,308,1850]" captionTargetPageId="4" captionText="FIGURE 2. Habitat (A–C), and habitus images of Sejanus ryujin n. sp. (D–H) and S. epicurus n. sp. (I–J), and Orius strigicollis (Poppius) (5th instar nymph) and an undetermined thrips sharing same niche (K); G–I showing ventral view of fresh specimens. A–C. Hibiscus hamabo, breeding host of S, ryujin n. sp. at the type locality (Ryujin Island, Omura City). D–F, H. Adult female. F–G. Adult males. I. S. epicurus n. sp., male. J. Same, female." figureDoi="http://doi.org/10.5281/zenodo.7456685" httpUri="https://zenodo.org/record/7456685/files/figure.png" pageId="3" pageNumber="158">Fig. 2I–J</figureCitation>
).............................................
<taxonomicName id="4C024D6A6B12175E85FAFB22FAA0FB78" authority="Yasunaga &amp; Duwal, 2022" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[1270,1377,1196,1217]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="epicurus" status="sp. nov.">
<emphasis id="B976EAFB6B12175E85FAFB22FAA0FB78" box="[1270,1377,1196,1217]" italics="true" pageId="3" pageNumber="158">S. epicurus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A24557806B12175E846AFB22FA5CFB78" box="[1382,1437,1196,1217]" pageId="3" pageNumber="158" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
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<keyLead id="30F395DC6B12175E819BFB47FA5DFB67" box="[151,1436,1225,1247]" pageId="3" pageNumber="158">- All femora, at least their basal parts, dark brown to fuscous.................................................... 9</keyLead>
</paragraph>
</keyStep>
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<keyLead id="30F395DC6B12175E819BFB68FA5DFB42" box="[151,1436,1254,1276]" pageId="3" pageNumber="158">9. Apex of metafemur yellow or creamy white............................................................... 10</keyLead>
</paragraph>
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<keyLead id="30F395DC6B12175E819BFA8AFA5DFAA0" box="[151,1436,1284,1306]" pageId="3" pageNumber="158">- Metafemur almost totally darkened...................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F62E4C6B12175E819BFAAFFA5DFA37" pageId="3" pageNumber="158">
<paragraph id="8BBD36E96B12175E819BFAAFFA5DFAEA" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFAAFFA5DFAEA" pageId="3" pageNumber="158">
10. Body larger; total body length greater than 3.0 mm; base of cuneus usually with two ivory spots; specialist of a Japanese walnut, inhabiting deciduous forest zone.................................................................
<taxonomicName id="4C024D6A6B12175E8427FAB1FA5DFAEA" ID-CoL="4WB7C" authorityName="Yasunaga" authorityYear="2001" box="[1323,1436,1342,1364]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="juglandis">
<emphasis id="B976EAFB6B12175E8427FAB1FA5DFAEA" box="[1323,1436,1342,1364]" italics="true" pageId="3" pageNumber="158">S. juglandis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="30F395DC6B12175E819BFAD2FA5DFA37" pageId="3" pageNumber="158">
- Body tiny; total body length up to
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; base of cuneus with whitish brown fascia (
<figureCitation id="13392A6C6B12175E853BFAD2FB43FAC8" box="[1079,1154,1372,1394]" captionStart="FIGURE 2" captionStartId="4.[151,250,1874,1898]" captionTargetBox="[151,1436,308,1837]" captionTargetId="figure-77@4.[151,1436,308,1850]" captionTargetPageId="4" captionText="FIGURE 2. Habitat (A–C), and habitus images of Sejanus ryujin n. sp. (D–H) and S. epicurus n. sp. (I–J), and Orius strigicollis (Poppius) (5th instar nymph) and an undetermined thrips sharing same niche (K); G–I showing ventral view of fresh specimens. A–C. Hibiscus hamabo, breeding host of S, ryujin n. sp. at the type locality (Ryujin Island, Omura City). D–F, H. Adult female. F–G. Adult males. I. S. epicurus n. sp., male. J. Same, female." figureDoi="http://doi.org/10.5281/zenodo.7456685" httpUri="https://zenodo.org/record/7456685/files/figure.png" pageId="3" pageNumber="158">Fig. 2F</figureCitation>
); inhabiting coastal
<taxonomicName id="4C024D6A6B12175E8444FAD2FEEEFA37" ID-CoL="3LJTB" class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="species" species="hamabo">
<emphasis id="B976EAFB6B12175E8444FAD2FEEEFA37" italics="true" pageId="3" pageNumber="158">Hibiscus hamabo</emphasis>
</taxonomicName>
...............................................................................
<taxonomicName id="4C024D6A6B12175E85EBFAF7FAF9FA36" authority="Yasunaga &amp; Duwal, 2022" authorityName="Yasunaga &amp; Duwal" authorityYear="2022" box="[1255,1336,1401,1423]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="ryujin" status="sp. nov.">
<emphasis id="B976EAFB6B12175E85EBFAF7FAF9FA36" box="[1255,1336,1401,1423]" italics="true" pageId="3" pageNumber="158">S. ryujin</emphasis>
</taxonomicName>
<emphasis id="B976EAFB6B12175E8432FAF7FAB4FA37" bold="true" box="[1342,1397,1401,1422]" pageId="3" pageNumber="158">
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</emphasis>
(♁)
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11. Antennal segment II longer than
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(♁)/
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(
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); male antennal segment II wholly black; anterior ivory mark of cuneus usually continuous........................................................................
<taxonomicName id="4C024D6A6B12175E844FFA3AFA5DFA70" ID-CoL="4WB6M" authorityName="Yasunaga" authorityYear="2001" box="[1347,1436,1460,1481]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="amami">
<emphasis id="B976EAFB6B12175E844FFA3AFA5DFA70" box="[1347,1436,1460,1481]" italics="true" pageId="3" pageNumber="158">S. amami</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BBD36E96B12175E819BFA5FFA5DF9BA" blockId="3.[151,1437,580,1540]" pageId="3" pageNumber="158">
<keyLead id="30F395DC6B12175E819BFA5FFA5DF9BA" pageId="3" pageNumber="158">
- Antennal segment II &lt;
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; base of male antennal segment II pale; anterior ivory mark of cuneus usually divided into two spots or slightly interrupted medially (
<figureCitation id="13392A6C6B12175E835EFA60FD68F9BA" box="[594,681,1518,1540]" captionStart="FIGURE 3" captionStartId="5.[151,250,1620,1644]" captionTargetBox="[204,1383,273,1597]" captionTargetId="figure-67@5.[204,1383,273,1597]" captionTargetPageId="5" captionText="FIGURE 3. Habittus images of live individuals for Sejanus spp. A. S. breviniger, adult males. B. Same, final instar nymph. C. S. komabanus, male. D. Same, female. E. S. vivaricolus, adults. F. Same, female reared with brine-shrimp eggs. G. Same, third instar nymph. H. Same, final instar nymph. I. S. neofunereus, male (Iriomote Is., Ryukyus, Japan). J. Same (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.7456687" httpUri="https://zenodo.org/record/7456687/files/figure.png" pageId="3" pageNumber="158">Fig. 3I–J</figureCitation>
)..................................................
<taxonomicName id="4C024D6A6B12175E841CFA61FA5DF9BA" ID-CoL="4WB7H" authorityName="Schuh" authorityYear="1984" box="[1296,1436,1518,1540]" class="Insecta" family="Miridae" genus="Sejanus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="neofunereus">
<emphasis id="B976EAFB6B12175E841CFA61FA5DF9BA" box="[1296,1436,1518,1540]" italics="true" pageId="3" pageNumber="158">S. neofunereus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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