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<document id="24A37552DF2CADA4C672D73E759FAAC8" ID-DOI="10.1080/00222933.2024.2373490" ID-ISSN="1464-5262" ID-Zenodo-Dep="13758245" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1726223587760" checkinUser="plazi" docAuthor="Haitlinger, Ryszard" docDate="2024" docId="03AB87D96B60FF942684CCF8C11394FA" docLanguage="en" docName="JNATHIST.58.29-32.1178-1259.pdf" docOrigin="Journal of Natural History 58 (29 - 32)" docSource="http://dx.doi.org/10.1080/00222933.2024.2373490" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Heterocheylidae Tragardh 1940" docType="treatment" docVersion="4" lastPageNumber="1243" masterDocId="FF92FFA16B21FFD6272DC835C2699246" masterDocTitle="Arthropods associated with world passalid beetles with particular reference to the Diarthrophallidae (Acari: Mesostigmata)" masterLastPageNumber="1259" masterPageNumber="1178" pageNumber="1243" updateTime="1726586162754" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4B03741DA8475C82A3A7E631A16E6FBE">Arthropods associated with world passalid beetles with particular reference to the Diarthrophallidae (Acari: Mesostigmata)</mods:title>
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<mods:date id="6706159F28B07B30DFB8D47128FBF33D">2024</mods:date>
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<mods:identifier id="EF5C8FA8C497012AE8EF71407A2769A9" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03AB87D96B60FF942684CCF8C11394FA" LSID="urn:lsid:plazi:treatment:03AB87D96B60FF942684CCF8C11394FA" httpUri="http://treatment.plazi.org/id/03AB87D96B60FF942684CCF8C11394FA" lastPageId="66" lastPageNumber="1243" pageId="65" pageNumber="1243">
<subSubSection id="C31865446B60FF972684CCF8C11396AE" box="[425,890,1229,1256]" pageId="65" pageNumber="1243" type="nomenclature">
<paragraph id="8BBD36CF6B60FF972684CCF8C11396AE" blockId="65.[425,890,1229,1256]" box="[425,890,1229,1256]" pageId="65" pageNumber="1243">
<heading id="D0F581A36B60FF972684CCF8C11396AE" bold="true" box="[425,890,1229,1256]" centered="true" fontSize="11" level="2" pageId="65" pageNumber="1243" reason="3">
<taxonomicName id="4C024D4C6B60FF972684CCF8C11396AE" ID-CoL="6255N" authority="Tragardh, 1940" authorityName="Tragardh" authorityYear="1940" box="[425,890,1229,1256]" class="Arachnida" family="Heterocheylidae" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">
<emphasis id="B976EADD6B60FF972684CCF8C11396AE" bold="true" box="[425,890,1229,1256]" pageId="65" pageNumber="1243">HETEROCHEYLIDAE Trägårdh, 1940</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C31865446B60FF9427EDCD37C11394FA" lastPageId="66" lastPageNumber="1244" pageId="65" pageNumber="1243" type="discussion">
<paragraph id="8BBD36CF6B60FF9427EDCD37C11394FA" blockId="65.[160,1156,1282,1723]" lastBlockId="66.[160,1156,1213,1724]" lastPageId="66" lastPageNumber="1244" pageId="65" pageNumber="1243">
A small family with one genus and 26 species. All species are associated with the
<taxonomicName id="4C024D4C6B60FF97278DCD10C37A9778" baseAuthorityName="Ermilov and Frolov" baseAuthorityYear="2021" box="[160,275,1317,1342]" class="Insecta" family="Passalidae" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
(
<bibRefCitation id="EF934B3E6B60FF97260DCD10C3899778" author="Lombardini G" box="[288,480,1317,1342]" pageId="65" pageNumber="1243" pagination="44 - 46" refId="ref37424" refString="Lombardini G. 1926. Duo nova genera acarorum. Bollet Soc Entomol Italiana. 58: 44 - 46." type="journal article" year="1926">Lombardini 1926</bibRefCitation>
;
<bibRefCitation id="EF934B3E6B60FF9726C1CD10C0F29778" author="Tragardh I" box="[492,667,1317,1342]" pageId="65" pageNumber="1243" pagination="104 - 110" refId="ref39163" refString="Tragardh I. 1950 a. Description of a new species of Heterocheylus Lombardini from Africa, with notes on the classification of the Pseudocheyletidae. Entomologisk Tidskrift. 71: 104 - 110." type="journal article" year="1950">Trägårdh 1950a</bibRefCitation>
;
<bibRefCitation id="EF934B3E6B60FF972585CD10C10B9778" author="Womersley H" box="[680,866,1317,1342]" pageId="65" pageNumber="1243" pagination="21 - 24" refId="ref40049" refString="Womersley H. 1960. Some Acarina from Australia and New Guinea paraphagic upon millipedes and cockroaches and beetles of the family Passalidae. Trans Roy Soc South Australia. 83: 21 - 24." type="journal article" year="1960">Womersley 1960</bibRefCitation>
;
<bibRefCitation id="EF934B3E6B60FF972443CD10C2BE9727" author="Schuster RO &amp; Lavoipierre MJ" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre 1970</bibRefCitation>
;
<bibRefCitation id="EF934B3E6B60FF9727C9CD7DC31A9727" author="Reinert JA" box="[228,371,1352,1377]" pageId="65" pageNumber="1243" pagination="273 - 276" refId="ref38080" refString="Reinert JA. 1973. Parasites associated with Popilius disjunctus in South Carolina (Coleoptera: Passalidae). Florida Entomologist. 56: 273 - 276. doi: 10.2307 / 3493803." type="journal article" year="1973">Reinert 1973</bibRefCitation>
;
<bibRefCitation id="EF934B3E6B60FF9726ADCD7DC0CD9727" author="Delfinado MD &amp; Baker EW" box="[384,676,1352,1377]" pageId="65" pageNumber="1243" pagination="49 - 59" refId="ref35683" refString="Delfinado MD, Baker EW. 1975. Mites (Acarina) associated with Popilius disjunctus (Illiger) (Coleoptera: Passalidae) in Eastern United States. J New York Entomol Soc. 83: 49 - 59." type="journal article" year="1975">Delfinado and Baker 1975</bibRefCitation>
; Barrios-Torres and Villegas-Gusmán 2015;
<bibRefCitation id="EF934B3E6B60FF97278DCD5FC33597C5" author="Katlav A &amp; Riegler M &amp; Seeman OD" box="[160,348,1386,1411]" pageId="65" pageNumber="1243" pagination="401 - 421" refId="ref37175" refString="Katlav A, Riegler M, Seeman OD. 2020. Tiny hitchhikers and parasites: a review of Australian heterostigmatic mites (Acari: Prostigmata) insects, with description of three new species. Austral Entomol. 59: 401 - 421. doi: 10.1111 / aen. 12484." type="journal article" year="2020">
Katlav
<emphasis id="B976EADD6B60FF9727C7CD5FC37097C5" box="[234,281,1386,1411]" italics="true" pageId="65" pageNumber="1243">et al</emphasis>
. 2020
</bibRefCitation>
). Twenty-one species of
<taxonomicName id="4C024D4C6B60FF972542CD5FC14F97C5" authorityName="Tragardh" authorityYear="1940" box="[623,806,1386,1411]" class="Arachnida" family="Heterocheylidae" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">Heterocheylidae</taxonomicName>
were collected from 56 species of
<taxonomicName id="4C024D4C6B60FF9727EFCDB8C35F97E0" baseAuthorityName="Ermilov and Frolov" baseAuthorityYear="2021" box="[194,310,1421,1446]" class="Insecta" family="Passalidae" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
belonging to 24 genera. Five species were collected from unidentified
<taxonomicName id="4C024D4C6B60FF97278DCD85C373978F" baseAuthorityName="Ermilov and Frolov" baseAuthorityYear="2021" box="[160,282,1456,1481]" class="Insecta" family="Passalidae" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
.
<taxonomicName id="4C024D4C6B60FF972606CD85C1A9978F" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[299,960,1456,1481]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B976EADD6B60FF972606CD85C055978F" box="[299,572,1456,1481]" italics="true" pageId="65" pageNumber="1243">Heterocheylus proximus</emphasis>
<bibRefCitation id="EF934B3E6B60FF972564CD85C1A9978F" author="Schuster RO &amp; Lavoipierre MJ" box="[585,960,1456,1481]" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D4C6B60FF972325CD85C07097AD" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B976EADD6B60FF972325CD85C6ED978F" box="[1032,1156,1456,1481]" italics="true" pageId="65" pageNumber="1243">H. rugosus</emphasis>
<bibRefCitation id="EF934B3E6B60FF97278DCDE7C07097AD" author="Schuster RO &amp; Lavoipierre MJ" box="[160,537,1490,1515]" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
were collected from 10 species of passalids, and
<taxonomicName id="4C024D4C6B60FF97278DCDC0C0C69448" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[160,687,1525,1550]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="species" species="cavagnaroi">
<emphasis id="B976EADD6B60FF97278DCDC0C3559448" box="[160,316,1525,1550]" italics="true" pageId="65" pageNumber="1243">H. cavagnaroi</emphasis>
<bibRefCitation id="EF934B3E6B60FF972665CDC0C0C69448" author="Schuster RO &amp; Lavoipierre MJ" box="[328,687,1525,1550]" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D4C6B60FF9725DDCDC0C2BE9477" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="species" species="keiferi">
<emphasis id="B976EADD6B60FF9725DDCDC0C13A9448" box="[752,851,1525,1550]" italics="true" pageId="65" pageNumber="1243">H. keiferi</emphasis>
<bibRefCitation id="EF934B3E6B60FF972472CDC0C2BE9477" author="Schuster RO &amp; Lavoipierre MJ" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
were collected from four species of passalids (
<tableCitation id="C68003746B60FF9725D3CE2DC1099477" box="[766,864,1560,1585]" captionStart="Table 2" captionStartId="8.[160,216,142,165]" captionTargetPageId="8" captionText="Table 2. List of passalid species and their arthropods." httpUri="http://table.plazi.org/id/DF7D66476B29FFDE278DC8BBC0DA92E3" pageId="65" pageNumber="1243" tableUuid="DF7D66476B29FFDE278DC8BBC0DA92E3">Tables 2</tableCitation>
,
<tableCitation id="C68003746B60FF972443CE2DC1E09477" box="[878,905,1560,1585]" captionStart="Table 15" captionStartId="66.[160,216,142,165]" captionTargetPageId="66" captionText="Table 15. List of Cunaxidae, Heterocheylidae and Tarsocheylidae found on Passalidae." pageId="65" pageNumber="1243">15</tableCitation>
). Fourteen species of heterocheylids were collected from only one species of host each.
<taxonomicName id="4C024D4C6B60FF97248DCE0FC63E9415" authorityName="Tragardh" authorityYear="1940" box="[928,1111,1594,1619]" class="Arachnida" family="Heterocheylidae" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">Heterocheylidae</taxonomicName>
are characterised by a pronounced host specificity. None of the
<taxonomicName id="4C024D4C6B60FF972414CE68C1999430" authorityName="Tragardh" authorityYear="1940" box="[825,1008,1629,1654]" class="Arachnida" family="Heterocheylidae" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="family">Heterocheylidae</taxonomicName>
species were caught on the same host species, except
<taxonomicName id="4C024D4C6B60FF972557CEB5C61794DF" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[634,1150,1664,1689]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="species" species="bengalensis">
<emphasis id="B976EADD6B60FF972557CEB5C17194DF" box="[634,792,1664,1689]" italics="true" pageId="65" pageNumber="1243">H. bengalensis</emphasis>
<bibRefCitation id="EF934B3E6B60FF97240DCEB5C61794DF" author="Schuster RO &amp; Lavoipierre MJ" box="[800,1150,1664,1689]" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
. Some passalids have more than one species:
<taxonomicName id="4C024D4C6B60FF9725BFCE97C63A94FD" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[658,1107,1698,1723]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="65" pageNumber="1243" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B976EADD6B60FF9725BFCE97C09194FD" box="[658,760,1698,1723]" italics="true" pageId="65" pageNumber="1243">H. parvus</emphasis>
<bibRefCitation id="EF934B3E6B60FF9725D0CE97C63A94FD" author="Schuster RO &amp; Lavoipierre MJ" box="[765,1107,1698,1723]" pageId="65" pageNumber="1243" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D4C6B63FF94278DCC88C0359690" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[160,604,1213,1238]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="thai">
<emphasis id="B976EADD6B63FF94278DCC88C2859690" box="[160,236,1213,1238]" italics="true" pageId="66" pageNumber="1244">H. thai</emphasis>
<bibRefCitation id="EF934B3E6B63FF9427DACC88C0359690" author="Schuster RO &amp; Lavoipierre MJ" box="[247,604,1213,1238]" pageId="66" pageNumber="1244" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
were collected from
<taxonomicName id="4C024D4C6B63FF942477CC88C6709690" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[858,1049,1213,1238]" class="Insecta" family="Passalidae" genus="Leptaulax" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B976EADD6B63FF942477CC88C6709690" box="[858,1049,1213,1238]" italics="true" pageId="66" pageNumber="1244">Leptaulax bicolor</emphasis>
</taxonomicName>
;
<taxonomicName id="4C024D4C6B63FF942307CC88C06596BF" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="nixus">
<emphasis id="B976EADD6B63FF942307CC88C6EA9690" box="[1066,1155,1213,1238]" italics="true" pageId="66" pageNumber="1244">H. nixus</emphasis>
<bibRefCitation id="EF934B3E6B63FF94278DCCD5C06596BF" author="Schuster RO &amp; Lavoipierre MJ" box="[160,524,1248,1273]" pageId="66" pageNumber="1244" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D4C6B63FF942562CCD5C65696BF" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[591,1087,1248,1273]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="rugosus">
<emphasis id="B976EADD6B63FF942562CCD5C0A196BF" box="[591,712,1248,1273]" italics="true" pageId="66" pageNumber="1244">H. rugosus</emphasis>
<bibRefCitation id="EF934B3E6B63FF9425FECCD5C65696BF" author="Schuster RO &amp; Lavoipierre MJ" box="[723,1087,1248,1273]" pageId="66" pageNumber="1244" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
were collected from
<taxonomicName id="4C024D4C6B63FF942678CD36C15C975A" authority="(Boisduval, 1835)" baseAuthorityName="Boisduval" baseAuthorityYear="1835" box="[341,821,1283,1308]" class="Insecta" family="Passalidae" genus="Protomocoelus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B976EADD6B63FF942678CD36C009975A" box="[341,608,1283,1308]" italics="true" pageId="66" pageNumber="1244">Protomocoelus australis</emphasis>
(Boisduval, 1835)
</taxonomicName>
; and
<taxonomicName id="4C024D4C6B63FF942456CD36C30A9778" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="keiferi">
<emphasis id="B976EADD6B63FF942456CD36C1B6975A" box="[891,991,1283,1308]" italics="true" pageId="66" pageNumber="1244">H. keiferi</emphasis>
<bibRefCitation id="EF934B3E6B63FF9424C6CD36C30A9778" author="Schuster RO &amp; Lavoipierre MJ" pageId="66" pageNumber="1244" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D4C6B63FF94268FCD10C1FA9778" authority="Schuster and Lavoipierre, 1970" authorityName="Schuster and Lavoipierre" authorityYear="1970" box="[418,915,1317,1342]" class="Arachnida" family="Heterocheyletidae" genus="Heterocheylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B976EADD6B63FF94268FCD10C04E9778" box="[418,551,1317,1342]" italics="true" pageId="66" pageNumber="1244">H. proximus</emphasis>
<bibRefCitation id="EF934B3E6B63FF94251DCD10C1FA9778" author="Schuster RO &amp; Lavoipierre MJ" box="[560,915,1317,1342]" pageId="66" pageNumber="1244" pagination="1 - 42" refId="ref38437" refString="Schuster RO, Lavoipierre MJ. 1970. The mite family Heterocheylidae Tragardh. Occasional Papers of the California Academy of Sciences. 85: 1 - 42." type="journal article" year="1970">Schuster and Lavoipierre, 1970</bibRefCitation>
</taxonomicName>
were collected from
<taxonomicName id="4C024D4C6B63FF94278DCD7DC3E89727" authorityName="Eschscholtz" authorityYear="1828" box="[160,385,1352,1377]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="interstitialis">
<emphasis id="B976EADD6B63FF94278DCD7DC3E89727" box="[160,385,1352,1377]" italics="true" pageId="66" pageNumber="1244">Passalus interstitialis</emphasis>
</taxonomicName>
(
<tableCitation id="C68003746B63FF9426BFCD7DC0659727" box="[402,524,1352,1377]" captionStart-0="Table 2" captionStart-1="Table 3" captionStart-2="Table 4" captionStart-3="Table 5" captionStart-4="Table 6" captionStartId-0="8.[160,216,142,165]" captionStartId-1="37.[160,216,142,165]" captionStartId-2="38.[160,216,142,165]" captionStartId-3="44.[160,216,1270,1293]" captionStartId-4="46.[160,216,971,994]" captionTargetPageId-0="8" captionTargetPageId-1="37" captionTargetPageId-2="38" captionTargetPageId-3="44" captionTargetPageId-4="46" captionText-0="Table 2. List of passalid species and their arthropods." captionText-1="Table 3. List of arthropods and their hosts from Africa." captionText-2="Table 4. List of arthropods and their hosts from America." captionText-3="Table 5. List of arthropods and their hosts from Asia." captionText-4="Table 6. List of arthropods and their hosts from Oceania." httpUri-0="http://table.plazi.org/id/DF7D66476B29FFDE278DC8BBC0DA92E3" httpUri-1="http://table.plazi.org/id/DF7D66476B04FFF3278DC8BBC0D692E3" httpUri-3="http://table.plazi.org/id/DF7D66476B0DFFFA278DCCC3C0D9974B" httpUri-4="http://table.plazi.org/id/DF7D66476B0FFFF8278DCBFEC0BF91A4" pageId="66" pageNumber="1244" tableUuid-0="DF7D66476B29FFDE278DC8BBC0DA92E3" tableUuid-1="DF7D66476B04FFF3278DC8BBC0D692E3" tableUuid-3="DF7D66476B0DFFFA278DCCC3C0D9974B" tableUuid-4="DF7D66476B0FFFF8278DCBFEC0BF91A4">Tables 2–6</tableCitation>
,
<tableCitation id="C68003746B63FF942536CD7DC05F9727" box="[539,566,1352,1377]" captionStart="Table 15" captionStartId="66.[160,216,142,165]" captionTargetPageId="66" captionText="Table 15. List of Cunaxidae, Heterocheylidae and Tarsocheylidae found on Passalidae." pageId="66" pageNumber="1244">15</tableCitation>
). On 26 species of
<taxonomicName id="4C024D4C6B63FF94243BCD7DC1E09727" baseAuthorityName="Ermilov and Frolov" baseAuthorityYear="2021" box="[790,905,1352,1377]" class="Insecta" family="Passalidae" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
only
<taxonomicName id="4C024D4C6B63FF9424E1CD7DC6EA9727" authorityName="Tragardh" authorityYear="1940" box="[972,1155,1352,1377]" class="Arachnida" family="Heterocheylidae" kingdom="Animalia" order="Trombidiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="family">Heterocheylidae</taxonomicName>
were found:
<taxonomicName id="4C024D4C6B63FF942615CD5EC08F97C2" authority="(Heller, 1910)" baseAuthorityName="Heller" baseAuthorityYear="1910" box="[312,742,1387,1412]" class="Insecta" family="Passalidae" genus="Aulacocyclus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B976EADD6B63FF942615CD5EC05497C2" box="[312,573,1387,1412]" italics="true" pageId="66" pageNumber="1244">Aulacocyclus papuanus</emphasis>
(Heller, 1910)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9425D8CD5EC67097C2" authority="Kaup, 1868" authorityName="Kaup" authorityYear="1868" box="[757,1049,1387,1412]" class="Arachnida" family="Ixodidae" genus="Aponomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ixodida" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="rosenbergi">
<emphasis id="B976EADD6B63FF9425D8CD5EC1E197C2" box="[757,904,1387,1412]" italics="true" pageId="66" pageNumber="1244">A. rosenbergi</emphasis>
Kaup, 1868
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94230ACD5EC3C097E0" authority="(Kuwert, 1901)" baseAuthorityName="Kuwert" baseAuthorityYear="1901" class="Insecta" family="Passalidae" genus="Aceraius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="illegalis">
<emphasis id="B976EADD6B63FF94230ACD5EC29B97E0" italics="true" pageId="66" pageNumber="1244">Aceraius illegalis</emphasis>
(Kuwert, 1901)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF942695CDB8C14797E0" authority="(Kaup, 1868)" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[440,814,1421,1446]" class="Insecta" family="Passalidae" genus="Cetejus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="peltostictus">
<emphasis id="B976EADD6B63FF942695CDB8C0E797E0" box="[440,654,1421,1446]" italics="true" pageId="66" pageNumber="1244">Cetejus peltostictus</emphasis>
(Kaup, 1868)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF942410CDB8C288978F" authority="(Klug, 1832)" baseAuthorityName="Klug" baseAuthorityYear="1832" class="Insecta" family="Passalidae" genus="Ciceronius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="morbillosus">
<emphasis id="B976EADD6B63FF942410CDB8C65197E0" box="[829,1080,1421,1446]" italics="true" pageId="66" pageNumber="1244">Ciceronius morbillosus</emphasis>
(Klug, 1832)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9427C3CD85C056978F" authority="Kuwert, 1908" authorityName="Kuwert" authorityYear="1908" box="[238,575,1456,1481]" class="Insecta" family="Passalidae" genus="Didimus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="haroldi">
<emphasis id="B976EADD6B63FF9427C3CD85C3F4978F" box="[238,413,1456,1481]" italics="true" pageId="66" pageNumber="1244">Didimus haroldi</emphasis>
Kuwert, 1908
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF942561CD85C1FB978F" authority="Kuwert, 1891" authorityName="Kuwert" authorityYear="1891" box="[588,914,1456,1481]" class="Insecta" family="Passalidae" genus="Gonatas" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="major">
<emphasis id="B976EADD6B63FF942561CD85C099978F" box="[588,752,1456,1481]" italics="true" pageId="66" pageNumber="1244">Gonatas major</emphasis>
Kuwert, 1891
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9424B2CD85C28A97AA" authority="(Kuwert, 1891)" baseAuthorityName="Kuwert" baseAuthorityYear="1891" class="Insecta" family="Passalidae" genus="Gonatas" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="B976EADD6B63FF9424B2CD85C674978F" box="[927,1053,1456,1481]" italics="true" pageId="66" pageNumber="1244">G. minimus</emphasis>
(Kuwert, 1891)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9427DECDE6C06197AA" authority="Gravely, 1918" authorityName="Gravely" authorityYear="1918" box="[243,520,1491,1516]" class="Insecta" family="Passalidae" genus="Gonatas" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B976EADD6B63FF9427DECDE6C33197AA" box="[243,344,1491,1516]" italics="true" pageId="66" pageNumber="1244">G. minor</emphasis>
Gravely, 1918
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94253ACDE6C15B97AA" authority="(Kaup, 1868)" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[535,818,1491,1516]" class="Insecta" family="Passalidae" genus="Gonatas" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="pumilio">
<emphasis id="B976EADD6B63FF94253ACDE6C0E797AA" box="[535,654,1491,1516]" italics="true" pageId="66" pageNumber="1244">G. pumilio</emphasis>
(Kaup, 1868)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94246FCDE6C6E897AA" authority="Kuwert, 1891" authorityName="Kuwert" authorityYear="1891" box="[834,1153,1491,1516]" class="Insecta" family="Passalidae" genus="Gonatas" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="schellongi">
<emphasis id="B976EADD6B63FF94246FCDE6C1BC97AA" box="[834,981,1491,1516]" italics="true" pageId="66" pageNumber="1244">G. schellongi</emphasis>
Kuwert, 1891
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94278DCDC0C0509448" authority="Kaup, 1871" authorityName="Kaup" authorityYear="1871" box="[160,569,1525,1550]" class="Insecta" family="Passalidae" genus="Macrolinus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="rotundifrons">
<emphasis id="B976EADD6B63FF94278DCDC0C3C09448" box="[160,425,1525,1550]" italics="true" pageId="66" pageNumber="1244">Macrolinus rotundifrons</emphasis>
Kaup, 1871
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF942565CDC0C1F99448" authority="Kaup, 1891" authorityName="Kaup" authorityYear="1891" box="[584,912,1525,1550]" class="Insecta" family="Passalidae" genus="Macrolinus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="sulciperfectus">
<emphasis id="B976EADD6B63FF942565CDC0C1699448" box="[584,768,1525,1550]" italics="true" pageId="66" pageNumber="1244">M. sulciperfectus</emphasis>
Kaup, 1891
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9424B2CDC0C3019477" authority="(Percheron, 1844)" baseAuthorityName="Percheron" baseAuthorityYear="1844" class="Insecta" family="Passalidae" genus="Ophrygonius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="cantori">
<emphasis id="B976EADD6B63FF9424B2CDC0C6EA9448" box="[927,1155,1525,1550]" italics="true" pageId="66" pageNumber="1244">Ophrygonius cantori</emphasis>
(Percheron, 1844)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94265ECE2DC1799477" authority="Percheron, 1835" authorityName="Percheron" authorityYear="1835" box="[371,784,1560,1585]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="anguliferus">
<emphasis id="B976EADD6B63FF94265ECE2DC0269477" box="[371,591,1560,1585]" italics="true" pageId="66" pageNumber="1244">Passalus anguliferus</emphasis>
Percheron, 1835
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF942436CE2DC6E99477" authority="Percheron, 1841" authorityName="Percheron" authorityYear="1841" box="[795,1152,1560,1585]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="binominatus">
<emphasis id="B976EADD6B63FF942436CE2DC1D69477" box="[795,959,1560,1585]" italics="true" pageId="66" pageNumber="1244">P. binominatus</emphasis>
Percheron, 1841
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94278DCE0EC3999412" authority="Kuwert, 1891" authorityName="Kuwert" authorityYear="1891" box="[160,496,1595,1620]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="clypelomeleus">
<emphasis id="B976EADD6B63FF94278DCE0EC33B9412" box="[160,338,1595,1620]" italics="true" pageId="66" pageNumber="1244">P. clypelomeleus</emphasis>
Kuwert, 1891
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9426D6CE0EC13C9412" authority="(Kuwert, 1891)" baseAuthorityName="Kuwert" baseAuthorityYear="1891" box="[507,853,1595,1620]" class="Insecta" family="Passalidae" genus="Passalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="considerabilis">
<emphasis id="B976EADD6B63FF9426D6CE0EC0C19412" box="[507,680,1595,1620]" italics="true" pageId="66" pageNumber="1244">P. considerabilis</emphasis>
(Kuwert, 1891)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94244DCE0EC2B09430" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" class="Arachnida" family="Chernetidae" genus="Phymatochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="mucronatus">
<emphasis id="B976EADD6B63FF94244DCE0EC1949412" box="[864,1021,1595,1620]" italics="true" pageId="66" pageNumber="1244">P. mucronatus</emphasis>
Burmeister, 1847
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9427CBCE68C0739430" authority="Percheron, 1835" authorityName="Percheron" authorityYear="1835" box="[230,538,1629,1654]" class="Arachnida" family="Chernetidae" genus="Phymatochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="plicatus">
<emphasis id="B976EADD6B63FF9427CBCE68C33F9430" box="[230,342,1629,1654]" italics="true" pageId="66" pageNumber="1244">P. plicatus</emphasis>
Percheron, 1835
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94250ACE68C14C9430" authority="Bates, 1886" authorityName="Bates" authorityYear="1886" box="[551,805,1629,1654]" class="Arachnida" family="Brachychthoniidae" genus="Poecilochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis id="B976EADD6B63FF94250ACE68C0F19430" box="[551,664,1629,1654]" italics="true" pageId="66" pageNumber="1244">P. spiniger</emphasis>
Bates, 1886
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94241FCE68C2B394DF" authority="Luederwaldt, 1937" authorityName="Luederwaldt" authorityYear="1931" class="Insecta" family="Passalidae" genus="Paxillus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="pleuralis">
<emphasis id="B976EADD6B63FF94241FCE68C1819430" box="[818,1000,1629,1654]" italics="true" pageId="66" pageNumber="1244">Paxillus pleuralis</emphasis>
Luederwaldt, 1937
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF9427C5CEB5C04D94DF" authority="(Kaup, 1866)" baseAuthorityName="Kaup" baseAuthorityYear="1866" box="[232,548,1664,1689]" class="Insecta" family="Passalidae" genus="Popilius" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="haagi">
<emphasis id="B976EADD6B63FF9427C5CEB5C3EC94DF" box="[232,389,1664,1689]" italics="true" pageId="66" pageNumber="1244">Popilius haagi</emphasis>
(Kaup, 1866)
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF94251ECEB5C1C494DF" authority="Klug, 1832" authorityName="Klug" authorityYear="1832" box="[563,941,1664,1689]" class="Insecta" family="Passalidae" genus="Solenocyclus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="exaratus">
<emphasis id="B976EADD6B63FF94251ECEB5C14C94DF" box="[563,805,1664,1689]" italics="true" pageId="66" pageNumber="1244">Solenocyclus exaratus</emphasis>
Klug, 1832
</taxonomicName>
,
<taxonomicName id="4C024D4C6B63FF942491CEB5C3D394FA" authority="(Percheron, 1841)" baseAuthorityName="Percheron" baseAuthorityYear="1841" class="Insecta" family="Passalidae" genus="Stephanocephalus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="hostilis">
<emphasis id="B976EADD6B63FF942491CEB5C28294FA" italics="true" pageId="66" pageNumber="1244">Stephanocephalus hostilis</emphasis>
(Percheron, 1841)
</taxonomicName>
, and
<taxonomicName id="4C024D4C6B63FF9426DACE96C11F94FA" authority="(Kaup, 1868)" baseAuthorityName="Kaup" baseAuthorityYear="1868" box="[503,886,1699,1724]" class="Insecta" family="Passalidae" genus="Trichostigmus" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="1244" phylum="Arthropoda" rank="species" species="thoreyi">
<emphasis id="B976EADD6B63FF9426DACE96C08994FA" box="[503,736,1699,1724]" italics="true" pageId="66" pageNumber="1244">Trichostigmus thoreyi</emphasis>
(Kaup, 1868)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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