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<document id="34435ED97C479792144F75E287FB97FC" ID-CLB-Dataset="4268" ID-DOI="10.18590/euscorpius.2018.vol2018.iss265.1" ID-GBIF-Dataset="1fb56719-70a0-461c-a6e3-642f454d1459" ID-ISSN="1536-9307" ID-Zenodo-Dep="5510059" ID-ZooBank="EC419E00-B4BB-4178-8A95-C637514C8653" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617134773086" checkinUser="carolina" docAuthor="Kovařík, František, Yağmur, Ersen Aydin &amp; Moradi, Mohammed" docDate="2018" docId="03AB87C64E1247079543CE5FDFFA3A51" docLanguage="en" docName="Euscorpius.265.1-14.pdf" docOrigin="Euscorpius 265" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Hottentotta sistanensis Kovařík &amp; Yağmur &amp; Moradi 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="FF92FFBE4E18470C9400C95DDA5A3F60" masterDocTitle="Two new Hottentotta species from Iran, with a review of Hottentotta saulcyi (Scorpiones: Buthidae)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="9" updateTime="1698935412841" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="97C9DDEDF2D9B3B25753EE772346FE96">Two new Hottentotta species from Iran, with a review of Hottentotta saulcyi (Scorpiones: Buthidae)</mods:title>
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<taxonomicName id="4C024D534E1247069543CE5FD80F387C" ID-CoL="3MQYD" authority="Kovařík &amp; Yağmur &amp; Moradi, 2018" authorityName="Kovařík &amp; Yağmur &amp; Moradi" authorityYear="2018" box="[323,597,1794,1820]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sistanensis" status="sp. nov.">Hottentotta sistanensis</taxonomicName>
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<figureCitation id="13392A554E124706955BCE7EDB86385A" box="[347,476,1827,1850]" captionStart="Figures 2145" captionStartId="6.[192,278,1686,1709]" captionTargetBox="[197,1433,201,1654]" captionTargetId="figure-21@6.[192,1439,199,1656]" captionTargetPageId="6" captionText="Figures 2145: Pedipalp segments. Figures 2132: Hottentotta navidpouri sp. n. Figures 2122. Paratype female, pedipalp chela, dorsal (21) and external (22) views. Figures 2332. Holotype male. Pedipalp chela, dorsal (23), external (24), and ventral (25) views. Pedipalp patella, dorsal (26), external (27), and ventral (28) views. Pedipalp femur and trochanter, internal (29), dorsal (30), and ventral (31) views. Pedipalp movable fingers (32). Figures 3345: Hottentotta sistanensis sp. n. Figures 3334. Paratype female, pedipalp chela, dorsal (33) and external (34) views. Figures 3545. Holotype male. Pedipalp chela, dorsal (35), external (36), and ventral (37) views. Pedipalp patella, dorsal (38), external (39), and ventral (40) views. Pedipalp femur and trochanter, internal (41), dorsal (42), and ventral (43) views. Pedipalp movable (44) and fixed (45) fingers. The trichobothrial pattern is indicated in Figures 2427, 2930, 3639, and 4142." figureDoi="http://doi.org/10.5281/zenodo.5510073" httpUri="https://zenodo.org/record/5510073/files/figure.png" pageId="10" pageNumber="9">Figs. 3345</figureCitation>
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<figureCitation id="13392A554E12470695E6CE7ED873385A" box="[486,553,1827,1850]" captionStart-0="Figures 4656" captionStart-1="Figures 5760" captionStart-2="Figures 6168" captionStart-3="Figures 6976" captionStartId-0="7.[192,278,1696,1719]" captionStartId-1="9.[192,278,1742,1765]" captionStartId-2="10.[192,278,1592,1615]" captionStartId-3="12.[192,278,1682,1705]" captionTargetBox-0="[211,1402,178,1673]" captionTargetBox-1="[193,1427,217,1694]" captionTargetBox-2="[191,1440,191,1562]" captionTargetBox-3="[197,1429,171,1632]" captionTargetId-0="figure-6@7.[210,1417,167,1680]" captionTargetId-1="figure-6@9.[192,1439,209,1704]" captionTargetId-2="figure-21@10.[192,1439,192,1562]" captionTargetId-3="figure-21@12.[192,1439,168,1653]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="12" captionText-0="Figures 4656: Hottentotta saulcyi (4651), H. navidpouri sp. n. (5254), and H. sistanensis sp. n. (5556), pedipalp chela dorsal. Figure 4647. H. saulcyi, male (46) and female (47) from Turkey, Şirnak Province, Kagrik Village, 37°25'39&quot;N 42°14'13.4&quot;E. Figure 48. H. saulcyi, female from Iraq, Dohuk Province, 17 km E of Zakho City, 37°11'12&quot;N 42°54'55&quot;E. Figure 49. H. saulcyi, female from Iran, Ilam Province, 30 km NW Ilam, 33°43'N 46°41'E. Figure 50. H. saulcyi, female from Iran, Lorestan Province, Aligudarz, Shoolabad Village, 33°11'44&quot;N 49°11'31&quot;E. Figure 51. H. saulcyi, female from Iran, Bachtaran Province, Bisotul, 34°23'31&quot;N 47°26'05&quot;E. Figure 52. H. navidpouri sp. n. male paratype from Iran, Hormozgan Province, Bandar Abbas to Lengeh Port road, 26°40'N 55°04'E. Figure 5354. H. navidpouri sp. n. male holotype (53) and female paratype (54) from Iran, Hormozgan Province, Lengeh, 27°08'N 55°49'E. Figure 5556. H. sistanensis sp. n. male holotype (55) and female paratype from type locality (56)." captionText-1="Figures 5760: Hottentotta sistanensis sp. n. Figures 5758. Holotype male, dorsal (57) and ventral (58) views. Figures 59 60. Paratype female, dorsal (59) and ventral (60) views. Scale bar = 10 mm." captionText-2="Figures 6168: Hottentotta sistanensis sp. n. Figures 61, 6468. Holotype male, chelicerae, carapace and tergites IIV (61), sternopectinal region and sternites IIIV and VII (64), and distal segments of legs IIV (6568), retrolateral view. Figures 62 63. Paratype female, chelicerae, carapace and tergites IIV (62), sternopectinal region and sternite III (63)." captionText-3="Figures 6976: Hottentotta sistanensis sp. n. Figures 69, 7173. Holotype male, telson lateral (69), metasoma and telson, lateral (71), dorsal (72), and ventral (73) views. Figures 70, 7476. Paratype female, telson lateral (70), metasoma and telson, lateral (74), dorsal (75), and ventral (76) views. Scale bar = 10 mm (7176)." figureDoi-0="http://doi.org/10.5281/zenodo.5510077" figureDoi-1="http://doi.org/10.5281/zenodo.5510079" figureDoi-2="http://doi.org/10.5281/zenodo.5510083" figureDoi-3="http://doi.org/10.5281/zenodo.5510085" httpUri-0="https://zenodo.org/record/5510077/files/figure.png" httpUri-1="https://zenodo.org/record/5510079/files/figure.png" httpUri-2="https://zenodo.org/record/5510083/files/figure.png" httpUri-3="https://zenodo.org/record/5510085/files/figure.png" pageId="10" pageNumber="9">5576</figureCitation>
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<emphasis id="B976EAC24E12470694C3CE1FD8933818" bold="true" pageId="10" pageNumber="9">http://zoobank.org/urn:lsid:zoobank.org:act:76C5B9 ED-107C-495F-B6CB-342A2EA02840</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C318655B4E13470794F0C8EBD85D3E8C" pageId="11" pageNumber="10" type="etymology">
<paragraph id="8BBD36D04E13470794F0C8EBD85D3E8C" blockId="11.[192,801,438,492]" pageId="11" pageNumber="10">ETYMOLOGY. Named after the Sistan region of the Iranian province of occurence.</paragraph>
</subSubSection>
<subSubSection id="C318655B4E13470794F0CB51D87A3B84" pageId="11" pageNumber="10" type="diagnosis">
<paragraph id="8BBD36D04E13470794F0CB51D87A3B84" blockId="11.[192,803,524,1252]" pageId="11" pageNumber="10">
DIAGNOSIS. Total length of adults
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on fixed finger of pedipalp situated between trichobothria
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and
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or
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and
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. Male with slightly narrower metasomal segments, width of pedipalp chela same in both sexes. Pectinal teeth number
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males,
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females. Nearly entire body hirsute, pedipalps, dorsal surface of mesosoma, legs, lateral and ventral surfaces of metasomal segments, and vesicle hirsute rather densely. Hairs on patella of pedipalps are long. Chelicerae brown to black, reticulate. Color yellow to yellowish green or brown, except black central part of carapace and tergites IVI, telson, posterior part of fourth and whole fifth metasomal segment. Femur of pedipalp with 4 carinae. Patella with 8 carinae. Chela lacks carinae. Movable fingers of pedipalps with 1516 rows of granules and 5 or 6 terminal granules. Seventh sternite with 4 well marked carinae. First metasomal segment with 10 carinae; second with 8 or 10 carinae; third and fourth with 8 carinae; fifth with 5 carinae, 3 ventral (one median, two lateral) and two dorsal. All metasomal segments generally smooth, without granules between carinae. All metasomal segments of both sexes longer than wide. Metasomal segment II length/width ratio more than 1.6.
</paragraph>
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DESCRIPTION. The total length is
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, males usually smaller than females. Chelal trichobothrium
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usually located between
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and
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, or may be on level with trichobothrium
<emphasis id="B976EAC24E13470795ADCC3ADB913A1E" box="[429,459,1383,1406]" italics="true" pageId="11" pageNumber="10">est</emphasis>
, rarely between
<emphasis id="B976EAC24E1347079695CC3AD8E93A1E" box="[661,691,1383,1406]" italics="true" pageId="11" pageNumber="10">est</emphasis>
and
<emphasis id="B976EAC24E13470796F8CC3AD9463A1E" box="[760,796,1383,1406]" italics="true" pageId="11" pageNumber="10">esb</emphasis>
. Male has the fingers proximally almost straight and female absolutely steaight, there is not sexual dimorphism in shape of chela of pedipalp. Chelicerae are brown to black, with reticulation, the tips of teeth on cheliceral fingers are black. For the position and distribution of trichobothria see
<figureCitation id="13392A554E1347079579CF42D82F3956" box="[377,629,1567,1590]" captionStart="Figures 2145" captionStartId="6.[192,278,1686,1709]" captionTargetBox="[197,1433,201,1654]" captionTargetId="figure-21@6.[192,1439,199,1656]" captionTargetPageId="6" captionText="Figures 2145: Pedipalp segments. Figures 2132: Hottentotta navidpouri sp. n. Figures 2122. Paratype female, pedipalp chela, dorsal (21) and external (22) views. Figures 2332. Holotype male. Pedipalp chela, dorsal (23), external (24), and ventral (25) views. Pedipalp patella, dorsal (26), external (27), and ventral (28) views. Pedipalp femur and trochanter, internal (29), dorsal (30), and ventral (31) views. Pedipalp movable fingers (32). Figures 3345: Hottentotta sistanensis sp. n. Figures 3334. Paratype female, pedipalp chela, dorsal (33) and external (34) views. Figures 3545. Holotype male. Pedipalp chela, dorsal (35), external (36), and ventral (37) views. Pedipalp patella, dorsal (38), external (39), and ventral (40) views. Pedipalp femur and trochanter, internal (41), dorsal (42), and ventral (43) views. Pedipalp movable (44) and fixed (45) fingers. The trichobothrial pattern is indicated in Figures 2427, 2930, 3639, and 4142." figureDoi="http://doi.org/10.5281/zenodo.5510073" httpUri="https://zenodo.org/record/5510073/files/figure.png" pageId="11" pageNumber="10">Figs. 3639 and 4142.</figureCitation>
</paragraph>
<paragraph id="8BBD36D04E13470794F0CF60D95C386C" blockId="11.[192,802,1291,1896]" pageId="11" pageNumber="10">
COLORATION (
<figureCitation id="13392A554E1347079597CF60D87E3934" box="[407,548,1597,1620]" captionStart-0="Figures 5760" captionStart-1="Figures 6168" captionStartId-0="9.[192,278,1742,1765]" captionStartId-1="10.[192,278,1592,1615]" captionTargetBox-0="[193,1427,217,1694]" captionTargetBox-1="[191,1440,191,1562]" captionTargetId-0="figure-6@9.[192,1439,209,1704]" captionTargetId-1="figure-21@10.[192,1439,192,1562]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figures 5760: Hottentotta sistanensis sp. n. Figures 5758. Holotype male, dorsal (57) and ventral (58) views. Figures 59 60. Paratype female, dorsal (59) and ventral (60) views. Scale bar = 10 mm." captionText-1="Figures 6168: Hottentotta sistanensis sp. n. Figures 61, 6468. Holotype male, chelicerae, carapace and tergites IIV (61), sternopectinal region and sternites IIIV and VII (64), and distal segments of legs IIV (6568), retrolateral view. Figures 62 63. Paratype female, chelicerae, carapace and tergites IIV (62), sternopectinal region and sternite III (63)." figureDoi-0="http://doi.org/10.5281/zenodo.5510079" figureDoi-1="http://doi.org/10.5281/zenodo.5510083" httpUri-0="https://zenodo.org/record/5510079/files/figure.png" httpUri-1="https://zenodo.org/record/5510083/files/figure.png" pageId="11" pageNumber="10">Figs. 5764</figureCitation>
,
<figureCitation id="13392A554E134707963ACF60D8D83934" box="[570,642,1597,1620]" captionStart="Figures 6976" captionStartId="12.[192,278,1682,1705]" captionTargetBox="[197,1429,171,1632]" captionTargetId="figure-21@12.[192,1439,168,1653]" captionTargetPageId="12" captionText="Figures 6976: Hottentotta sistanensis sp. n. Figures 69, 7173. Holotype male, telson lateral (69), metasoma and telson, lateral (71), dorsal (72), and ventral (73) views. Figures 70, 7476. Paratype female, telson lateral (70), metasoma and telson, lateral (74), dorsal (75), and ventral (76) views. Scale bar = 10 mm (7176)." figureDoi="http://doi.org/10.5281/zenodo.5510085" httpUri="https://zenodo.org/record/5510085/files/figure.png" pageId="11" pageNumber="10">6976</figureCitation>
). The central parts of carapace and tergites IVI, telson, posterior part of fourth and whole fifth metasomal segment, and pedipalp chela are reddish black to black; the metasoma I III, femur and patella of pedipalps, margins of tergites I VI, tergite VII, sternites, and legs are yellow to yellowish green or brown. The tarsomeres of legs are yellow.
</paragraph>
<paragraph id="8BBD36D04E13470794F0CE49DE063D2E" blockId="11.[192,802,1291,1896]" lastBlockId="11.[830,1441,199,1265]" pageId="11" pageNumber="10">
MESOSOMA AND CARAPACE (
<figureCitation id="13392A554E134707964ACE49D880384B" box="[586,730,1812,1835]" captionStart="Figures 6168" captionStartId="10.[192,278,1592,1615]" captionTargetBox="[191,1440,191,1562]" captionTargetId="figure-21@10.[192,1439,192,1562]" captionTargetPageId="10" captionText="Figures 6168: Hottentotta sistanensis sp. n. Figures 61, 6468. Holotype male, chelicerae, carapace and tergites IIV (61), sternopectinal region and sternites IIIV and VII (64), and distal segments of legs IIV (6568), retrolateral view. Figures 62 63. Paratype female, chelicerae, carapace and tergites IIV (62), sternopectinal region and sternite III (63)." figureDoi="http://doi.org/10.5281/zenodo.5510083" httpUri="https://zenodo.org/record/5510083/files/figure.png" pageId="11" pageNumber="10">Figs. 6164</figureCitation>
). The carapace is carinate and unevenly covered by granules of varying size; much of the granulation is fine, but some granules are larger and distinctly rounded. Tergites IVI bear three carinae and are granulated, with some intercarinal granules small and others larger and rounded. Tergite VII is pentacarinate. The pectinal tooth count is 2630 (28.928 /18/) in males and 2328 (24.55 /24/) in females. The pectinal marginal tips extend to third of the fourth sternite in female and half of the fourth sternite in male. The pectines have three marginal lamellae and seven to nine middle lamellae. The lamellae bear numerous dark long setae, each fulcrum with two to three setae. All sternites are smooth and sparsely hirsute. The seventh sternite bears four granulate carinae. The other sternites bear two furrows.
</paragraph>
<paragraph id="8BBD36D04E134707976ECB0BDF5B3C45" blockId="11.[830,1441,199,1265]" pageId="11" pageNumber="10">
PEDIPALPS (
<figureCitation id="13392A554E13470797F1CB0BDE2C3D0D" box="[1009,1142,598,621]" captionStart="Figures 2145" captionStartId="6.[192,278,1686,1709]" captionTargetBox="[197,1433,201,1654]" captionTargetId="figure-21@6.[192,1439,199,1656]" captionTargetPageId="6" captionText="Figures 2145: Pedipalp segments. Figures 2132: Hottentotta navidpouri sp. n. Figures 2122. Paratype female, pedipalp chela, dorsal (21) and external (22) views. Figures 2332. Holotype male. Pedipalp chela, dorsal (23), external (24), and ventral (25) views. Pedipalp patella, dorsal (26), external (27), and ventral (28) views. Pedipalp femur and trochanter, internal (29), dorsal (30), and ventral (31) views. Pedipalp movable fingers (32). Figures 3345: Hottentotta sistanensis sp. n. Figures 3334. Paratype female, pedipalp chela, dorsal (33) and external (34) views. Figures 3545. Holotype male. Pedipalp chela, dorsal (35), external (36), and ventral (37) views. Pedipalp patella, dorsal (38), external (39), and ventral (40) views. Pedipalp femur and trochanter, internal (41), dorsal (42), and ventral (43) views. Pedipalp movable (44) and fixed (45) fingers. The trichobothrial pattern is indicated in Figures 2427, 2930, 3639, and 4142." figureDoi="http://doi.org/10.5281/zenodo.5510073" httpUri="https://zenodo.org/record/5510073/files/figure.png" pageId="11" pageNumber="10">Figs. 3345</figureCitation>
). The pedipalps are hirsute and almost smooth. The femur bears four granulated carinae. The patella bears eight carinae, ventral and external are smooth, without granules and obsolete; dorsal and internal are granulate. The chela is without carinae. The movable fingers of pedipalps bear 1516 rows of granules and five or six terminal granules.
</paragraph>
<paragraph id="8BBD36D04E134707976ECA71DEB93CBD" blockId="11.[830,1441,199,1265]" pageId="11" pageNumber="10">
LEGS (
<figureCitation id="13392A554E13470797B7CA71DE643C23" box="[951,1086,812,835]" captionStart="Figures 6168" captionStartId="10.[192,278,1592,1615]" captionTargetBox="[191,1440,191,1562]" captionTargetId="figure-21@10.[192,1439,192,1562]" captionTargetPageId="10" captionText="Figures 6168: Hottentotta sistanensis sp. n. Figures 61, 6468. Holotype male, chelicerae, carapace and tergites IIV (61), sternopectinal region and sternites IIIV and VII (64), and distal segments of legs IIV (6568), retrolateral view. Figures 62 63. Paratype female, chelicerae, carapace and tergites IIV (62), sternopectinal region and sternite III (63)." figureDoi="http://doi.org/10.5281/zenodo.5510083" httpUri="https://zenodo.org/record/5510083/files/figure.png" pageId="11" pageNumber="10">Figs. 6568</figureCitation>
). The tarsomeres bear two rows of short and strong spiniform setae on the ventral surface and numerous macrosetae on the other surfaces. Pedal spur of legs without setae. Femur and patella with carinae indicated. Tibial spurs present and long on third and fourth legs and absent in the other legs.
</paragraph>
<paragraph id="8BBD36D04E134707976ECAB9D9C33B91" blockId="11.[830,1441,199,1265]" pageId="11" pageNumber="10">
METASOMA AND TELSON (
<figureCitation id="13392A554E1347079095CAB9DF443C9B" box="[1173,1310,996,1019]" captionStart="Figures 6976" captionStartId="12.[192,278,1682,1705]" captionTargetBox="[197,1429,171,1632]" captionTargetId="figure-21@12.[192,1439,168,1653]" captionTargetPageId="12" captionText="Figures 6976: Hottentotta sistanensis sp. n. Figures 69, 7173. Holotype male, telson lateral (69), metasoma and telson, lateral (71), dorsal (72), and ventral (73) views. Figures 70, 7476. Paratype female, telson lateral (70), metasoma and telson, lateral (74), dorsal (75), and ventral (76) views. Scale bar = 10 mm (7176)." figureDoi="http://doi.org/10.5281/zenodo.5510085" httpUri="https://zenodo.org/record/5510085/files/figure.png" pageId="11" pageNumber="10">Figs. 6976</figureCitation>
). All metasomal segments are hirsute rather densely. All metasomal segments of both sexes longer than wide. The segment I bears 10 carinae; segment II bears 10 or 8 carinae; segments III and IV bear 8 carinae; and segment V bear five carinae, three ventral and two dorsal. All carinae are with consistent denticles. The telson is only sparsely hirsute, bulbous, bumpy and finely granulated, more in females.
</paragraph>
<paragraph id="8BBD36D04E134707976ECC47DFFA3A51" blockId="11.[878,1440,1306,1329]" box="[878,1440,1306,1329]" pageId="11" pageNumber="10">
AFFINITIES. See affinities under
<taxonomicName id="4C024D534E13470790C7CC47DF073A51" authority="Kovařík &amp; Yağmur &amp; Moradi, 2018" authorityName="Kovařík &amp; Yağmur &amp; Moradi" authorityYear="2018" box="[1223,1373,1306,1329]" class="Arachnida" family="Buthidae" genus="Hottentotta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="navidpouri" status="sp. nov.">
<emphasis id="B976EAC24E13470790C7CC47DF073A51" box="[1223,1373,1306,1329]" italics="true" pageId="11" pageNumber="10">H. navidpouri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557B94E1347079164CC47DFFA3A51" box="[1380,1440,1306,1329]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B976EAC24E1347079164CC47DF273A51" bold="true" box="[1380,1405,1306,1329]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="B976EAC24E134707918ACC47DFC33A51" bold="true" box="[1418,1433,1306,1329]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>