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Milléo
<emphasis id="B976EAB3FFB6FFD4FE6E86B22164FC3C" box="[504,555,980,1002]" italics="true" pageId="1" pageNumber="253">et al.</emphasis>
, 2004
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274.
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<emphasis id="B976EAB3FFB6FFD4FF0186952246FBDE" box="[151,265,1010,1032]" italics="true" pageId="1" pageNumber="253">Menoscelis</emphasis>
(
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)
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:
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, 233234.
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<emphasis id="B976EAB3FFB6FFD4FF01817623BCFBF1" box="[151,243,1041,1063]" italics="true" pageId="1" pageNumber="253">Thalassa</emphasis>
(
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)
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:
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: 63
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.
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<emphasis id="B976EAB3FFB6FFD4FF018133227DFBBB" bold="true" box="[151,306,1108,1133]" pageId="1" pageNumber="253">
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species.
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<taxonomicName id="4C024D22FFB6FFD4FEAF81332188FBBB" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" box="[313,711,1108,1133]" class="Insecta" family="Coccinellidae" genus="Menoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="species" species="saginata">
<emphasis id="B976EAB3FFB6FFD4FEAF81332150FBBA" box="[313,543,1108,1133]" italics="true" pageId="1" pageNumber="253">Menoscelis saginata</emphasis>
<bibRefCitation id="EF934B50FFB6FFD4FDB381332188FBBB" author="Mulsant" box="[549,711,1108,1133]" pageId="1" pageNumber="253" refString="Mulsant, E. (1850) Species des Coleopteres trimeres securipalpes. Annales des Sciences Physiques et Naturelles d' Agriculture et d' Industrie, 2, 1 - 1104. [Lyon] http: // dx. doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
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, by subsequent designation of
<bibRefCitation id="EF934B50FFB6FFD4FBB48133278CFBBA" author="Crotch" box="[1058,1219,1108,1133]" pageId="1" pageNumber="253" refString="Crotch, G. R. (1874) A Revision of the Coleopterous Family Coccinellidae. E. W. Janson, London, 311 pp." type="book" year="1874">Crotch (1874)</bibRefCitation>
.
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<emphasis id="B976EAB3FFB6FFD4FF5181102275FB46" bold="true" box="[199,314,1143,1168]" pageId="1" pageNumber="253">Diagnosis</emphasis>
.
<taxonomicName id="4C024D22FFB6FFD4FED18110228AFB46" box="[327,453,1143,1168]" class="Insecta" family="Coccinellidae" genus="Menoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAB3FFB6FFD4FED18110228AFB46" box="[327,453,1143,1168]" italics="true" pageId="1" pageNumber="253">Menoscelis</emphasis>
</taxonomicName>
is a Neotropical genus that is structurally similar to
<taxonomicName id="4C024D22FFB6FFD4FB9A811E2651FB46" authority="Weise, 1926" authorityName="Weise" authorityYear="1926" box="[1036,1310,1143,1168]" class="Insecta" family="Coccinellidae" genus="Diazonema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAB3FFB6FFD4FB9A811E27C3FB46" box="[1036,1164,1145,1168]" italics="true" pageId="1" pageNumber="253">Diazonema</emphasis>
Weise, 1926
</taxonomicName>
, but differs from all other genera of the tribe in the highly modified protibia, angulate near its base (
<figureCitation id="13392A24FFB6FFD4FB2281FB27B2FB63" box="[1204,1277,1180,1205]" captionStart="FIGURE 5" captionStartId="10.[151,250,1174,1196]" captionTargetBox="[151,1435,193,1151]" captionTargetId="figure@10.[151,1436,193,1152]" captionTargetPageId="10" captionText="FIGURE 5. Menoscelis saginata Mulsant, 1850. A. head frontal view; B. antenna; C. prosternum; D E. anterior leg in dorsal (D) and ventral (E) views. F. metasternun; G. abdomen; H. eplipleuron." httpUri="https://zenodo.org/record/262126/files/figure.png" pageId="1" pageNumber="253">Figs 5</figureCitation>
DE) and the transverse female genital plate with spoonlike setae near the apex.
<taxonomicName id="4C024D22FFB6FFD4FC1C81A62745FB0E" box="[906,1034,1217,1240]" class="Insecta" family="Coccinellidae" genus="Diazonema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAB3FFB6FFD4FC1C81A62745FB0E" box="[906,1034,1217,1240]" italics="true" pageId="1" pageNumber="253">Diazonema</emphasis>
</taxonomicName>
presents a combination of short and compressed protibia, and a transverse female genital plate lacking spoonlike setae near the apex.
</paragraph>
<paragraph id="8BBD36A1FFB6FFD4FF51806F2034F906" blockId="1.[151,1437,1108,1816]" pageId="1" pageNumber="253">
In addition to the generic characters described by
<bibRefCitation id="EF934B50FFB6FFD4FD6E80602764FAF6" author="Gordon" box="[760,1067,1287,1312]" pageId="1" pageNumber="253" refString="Gordon, R. D. &amp; Canepari, C. (2008) South American Coccinellidae (Coleoptera), Part XI: A systematic revision of Hyperaspidini (Hyperaspidinae). Annali del Museo Civico di Storia Naturale &quot; G. Doria &quot;, 99, 245 - 512." type="journal article" year="2008">Gordon &amp; Canepari (2008)</bibRefCitation>
, species of this genus have other useful characters that aid in their recognition: body usually large (
<quantity id="4CFA9B44FFB6FFD4FCE7804B20B4FA92" box="[881,1019,1324,1349]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.5" metricValueMin="4.5" pageId="1" pageNumber="253" unit="mm" value="6.0" valueMax="7.5" valueMin="4.5">4.57.5 mm</quantity>
) and rounded, moderately to strongly convex. Dorsal surface glabrous or pubescent, maculate or not. Prosternum process about half the coxal diamenter, surface with complete or incomplete carinae joined anteriorly, forming a triangle (
<figureCitation id="13392A24FFB6FFD4FBA580132735FA5B" box="[1075,1146,1396,1421]" captionStart="FIGURE 5" captionStartId="10.[151,250,1174,1196]" captionTargetBox="[151,1435,193,1151]" captionTargetId="figure@10.[151,1436,193,1152]" captionTargetPageId="10" captionText="FIGURE 5. Menoscelis saginata Mulsant, 1850. A. head frontal view; B. antenna; C. prosternum; D E. anterior leg in dorsal (D) and ventral (E) views. F. metasternun; G. abdomen; H. eplipleuron." httpUri="https://zenodo.org/record/262126/files/figure.png" pageId="1" pageNumber="253">Figs 5</figureCitation>
C, 6E). Prosternal carinae often joined, sometimes connected to the anterior margin of the prosternum by a single carina (
<figureCitation id="13392A24FFB6FFD4FB5480F02648FA66" box="[1218,1287,1431,1456]" captionStart="FIGURE 8" captionStartId="13.[151,250,1742,1764]" captionTargetBox="[151,1436,195,1718]" captionTargetId="figure@13.[151,1436,195,1718]" captionTargetPageId="13" captionText="FIGURE 8. Prosternal process of Menoscelis. A B. Menoscelis insignis Mulsant, 1850 female (A) and male (B). C D. M. saginata Mulsant, 1850 female (C) and male (D); E F. M. stephani Corrêa et al., 2011 female (E) and male (F); G. M. brunella Corrêa et al., 2011 female." httpUri="https://zenodo.org/record/262129/files/figure.png" pageId="1" pageNumber="253">Figs 8</figureCitation>
AG, 9AE). Epipleura large, external margin explanate (
<figureCitation id="13392A24FFB6FFD4FD1E80DB219FFA03" box="[648,720,1468,1493]" captionStart="FIGURE 5" captionStartId="10.[151,250,1174,1196]" captionTargetBox="[151,1435,193,1151]" captionTargetId="figure@10.[151,1436,193,1152]" captionTargetPageId="10" captionText="FIGURE 5. Menoscelis saginata Mulsant, 1850. A. head frontal view; B. antenna; C. prosternum; D E. anterior leg in dorsal (D) and ventral (E) views. F. metasternun; G. abdomen; H. eplipleuron." httpUri="https://zenodo.org/record/262126/files/figure.png" pageId="1" pageNumber="253">Figs 5</figureCitation>
H, 6H). Protibia flattened, medially emarginate or with parallel sides, tibial groove deep, long, not flanged posteriorly, angulate near base (
<figureCitation id="13392A24FFB6FFD4FC4D80B8276FFA2E" box="[987,1056,1503,1528]" captionStart="FIGURE 5" captionStartId="10.[151,250,1174,1196]" captionTargetBox="[151,1435,193,1151]" captionTargetId="figure@10.[151,1436,193,1152]" captionTargetPageId="10" captionText="FIGURE 5. Menoscelis saginata Mulsant, 1850. A. head frontal view; B. antenna; C. prosternum; D E. anterior leg in dorsal (D) and ventral (E) views. F. metasternun; G. abdomen; H. eplipleuron." httpUri="https://zenodo.org/record/262126/files/figure.png" pageId="1" pageNumber="253">Figs 5</figureCitation>
DE, 6FG); meso- and metatibia narrow, medially emarginate or with parallel sides, not flanged, with shallow tibial groove (
<figureCitation id="13392A24FFB6FFD4FB198363279BF9CB" box="[1167,1236,1540,1565]" captionStart="FIGURE 5" captionStartId="10.[151,250,1174,1196]" captionTargetBox="[151,1435,193,1151]" captionTargetId="figure@10.[151,1436,193,1152]" captionTargetPageId="10" captionText="FIGURE 5. Menoscelis saginata Mulsant, 1850. A. head frontal view; B. antenna; C. prosternum; D E. anterior leg in dorsal (D) and ventral (E) views. F. metasternun; G. abdomen; H. eplipleuron." httpUri="https://zenodo.org/record/262126/files/figure.png" pageId="1" pageNumber="253">Figs 5</figureCitation>
F,
<date id="FFBC1061FFB6FFD4FB6683632644F9CB" box="[1264,1291,1540,1565]" pageId="1" pageNumber="253">6I</date>
). Abdominal postcoxal line incomplete, recurved or slightly recurved (
<figureCitation id="13392A24FFB6FFD4FCC6834020D1F996" box="[848,926,1575,1600]" captionStart="FIGURE 5" captionStartId="10.[151,250,1174,1196]" captionTargetBox="[151,1435,193,1151]" captionTargetId="figure@10.[151,1436,193,1152]" captionTargetPageId="10" captionText="FIGURE 5. Menoscelis saginata Mulsant, 1850. A. head frontal view; B. antenna; C. prosternum; D E. anterior leg in dorsal (D) and ventral (E) views. F. metasternun; G. abdomen; H. eplipleuron." httpUri="https://zenodo.org/record/262126/files/figure.png" pageId="1" pageNumber="253">Figs 5</figureCitation>
G, 6J). Parameres symmetrical; basal lobe asymmetrical. Sipho slender with well-developed, T-shaped siphonal capsule (
<figureCitation id="13392A24FFB6FFD4FBD9832B27D5F9B2" box="[1103,1178,1612,1637]" captionStart="FIGURE 1" captionStartId="6.[151,250,1927,1949]" captionTargetBox="[302,1284,195,1900]" captionTargetId="figure@6.[302,1284,195,1903]" captionTargetPageId="6" captionText="FIGURE 1. Menoscelis angeloi sp. nov. habitus and genitalia of holotype and paratype A B. male habitus in dorsal (A) and lateral (B) views; C E. female habitus in dorsal (C), lateral (D), and frontal (E) views; F. abdomen. G K. male genitalia, G. tegmen, dorsal; H. same in lateral view; I. sipho; J. female genitalia; K. coxites." httpUri="https://zenodo.org/record/262122/files/figure.png" pageId="1" pageNumber="253">Figs 1</figureCitation>
EG, 4DF). Coxites transverse with spoon-like setae near apex, styles well developed, with or without apical setae; infundibulum lacking; sperm duct simple, uniform in diameter (
<figureCitation id="13392A24FFB6FFD4FD5283F3204DF97B" box="[708,770,1684,1709]" captionStart="FIGURE 1" captionStartId="6.[151,250,1927,1949]" captionTargetBox="[302,1284,195,1900]" captionTargetId="figure@6.[302,1284,195,1903]" captionTargetPageId="6" captionText="FIGURE 1. Menoscelis angeloi sp. nov. habitus and genitalia of holotype and paratype A B. male habitus in dorsal (A) and lateral (B) views; C E. female habitus in dorsal (C), lateral (D), and frontal (E) views; F. abdomen. G K. male genitalia, G. tegmen, dorsal; H. same in lateral view; I. sipho; J. female genitalia; K. coxites." httpUri="https://zenodo.org/record/262122/files/figure.png" pageId="1" pageNumber="253">Figs1</figureCitation>
H, 2E, 3F). Spermatheca with rounded cornu and narrow at base; distal capsule adjacent to cornu or sperm duct (
<figureCitation id="13392A24FFB6FFD4FD7183D02061F906" box="[743,814,1719,1744]" captionStart="FIGURE 2" captionStartId="7.[151,250,1926,1948]" captionTargetBox="[154,1434,207,1903]" captionTargetId="figure@7.[151,1436,193,1904]" captionTargetPageId="7" captionText="FIGURE 2. Menoscelis cordata sp. nov. habitus and details of morphology of female holotype. A B. habitus in dorsal (A) and frontal (B) views; C. abdomen; D. post coxal line; E G. female genitalia, F. spermatheca, G. coxites." httpUri="https://zenodo.org/record/262123/files/figure.png" pageId="1" pageNumber="253">Figs 2</figureCitation>
F, 3G).
</paragraph>
</subSubSection>
<subSubSection id="C318652AFFB6FFD4FF5183BB2664F8CE" pageId="1" pageNumber="253" type="discussion">
<paragraph id="8BBD36A1FFB6FFD4FF5183BB2664F8CE" blockId="1.[151,1437,1108,1816]" pageId="1" pageNumber="253">
<emphasis id="B976EAB3FFB6FFD4FF5183BB2274F923" bold="true" box="[199,315,1756,1781]" pageId="1" pageNumber="253">Remarks.</emphasis>
Two species,
<taxonomicName id="4C024D22FFB6FFD4FE4883BA2116F923" box="[478,601,1756,1781]" class="Insecta" family="Coccinellidae" genus="Menoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="species" species="angeloi" status="sp. nov.">
<emphasis id="B976EAB3FFB6FFD4FE4883BA2116F923" box="[478,601,1756,1781]" italics="true" pageId="1" pageNumber="253">M. angeloi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557C8FFB6FFD4FDF483BB21F2F923" box="[610,701,1756,1781]" pageId="1" pageNumber="253" rank="species">
<emphasis id="B976EAB3FFB6FFD4FDF483BB21F2F923" bold="true" box="[610,701,1756,1781]" pageId="1" pageNumber="253">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C024D22FFB6FFD4FD6183BA203BF923" box="[759,884,1756,1781]" class="Insecta" family="Coccinellidae" genus="Menoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="species" species="cordata" status="sp. nov.">
<emphasis id="B976EAB3FFB6FFD4FD6183BA203BF923" box="[759,884,1756,1781]" italics="true" pageId="1" pageNumber="253">M. cordata</emphasis>
</taxonomicName>
<emphasis id="B976EAB3FFB6FFD4FCEA83BB2097F923" bold="true" box="[892,984,1756,1781]" pageId="1" pageNumber="253">
<taxonomicNameLabel id="A24557C8FFB6FFD4FCEA83BB2097F923" box="[892,984,1756,1781]" pageId="1" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, lack the protibial emargination but are considered members of
<taxonomicName id="4C024D22FFB6FFD4FE358398216EF8CE" box="[419,545,1791,1816]" class="Insecta" family="Coccinellidae" genus="Menoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAB3FFB6FFD4FE358398216EF8CE" box="[419,545,1791,1816]" italics="true" pageId="1" pageNumber="253">Menoscelis</emphasis>
</taxonomicName>
due to the described characters and male and female genital patterns.
</paragraph>
</subSubSection>
</treatment>
</document>