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<document id="5B74B744F8CEC210FC054496137B5621" ID-CLB-Dataset="304901" ID-DOI="10.1649/0010-065X(2001)055[0515:EAANGA]2.0.CO;2" ID-GBIF-Dataset="0647875f-6005-4b9f-9388-2318633c573f" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102874" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1630156293176" checkinUser="tatiana" docAuthor="Brown, Kirby W. &amp; Triplehorn, Charles A." docDate="2001" docId="03A95D4BFFEE5023F372FDDC6A769C88" docLanguage="en" docName="TheColeopBull.55.4.515-521.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (4)" docSource="http://dx.doi.org/10.1649/0010-065x(2001)055[0515:eaanga]2.0.co;2" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Epitragosoma arenaria Brown and Triplehorn, new species" docType="treatment" docVersion="2" lastPageNumber="521" masterDocId="FF902533FFEA5025F22BFF9369029D4C" masterDocTitle="EPITRAGOSOMA ARENARIA, A New Genus and Species from Texas (Coleoptera: Tenebrionidae)" masterLastPageNumber="521" masterPageNumber="515" pageNumber="519" updateTime="1730314773783" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C6BF2094A13B876CF58E09AB6A881BFD">EPITRAGOSOMA ARENARIA, A New Genus and Species from Texas (Coleoptera: Tenebrionidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="A6F03B0B62F60ED6F744773C4417891F">Brown, Kirby W.</mods:namePart>
<mods:affiliation id="A6CE52FF4F803553F11F984DB249F8AB">P. O. Box 1842 Paradise, CA 95967, U. S. A.</mods:affiliation>
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<mods:namePart id="81D17359B0B78B2E0CA8AEEF3AB0F132">Triplehorn, Charles A.</mods:namePart>
<mods:affiliation id="1A9A403D7DC2D7E8B04BAA316B329217">Museum of Biological Diversity The Ohio State University 1315 Kinnear Road Columbus, OH 43212 ­ 1192, U. S. A.</mods:affiliation>
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<mods:title id="B8388F726F927DD94CF71FD7392EFC0A">The Coleopterists Bulletin</mods:title>
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<mods:part id="A2612C193A35076880E6AE81CB6851F4">
<mods:date id="4332C572C357CD037F1ACC417090CFAD">2001</mods:date>
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<mods:number id="C3F61EF04E1BD53E9352DEC7E45901F0">2001-12-31</mods:number>
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<treatment id="03A95D4BFFEE5023F372FDDC6A769C88" ID-DOI="http://doi.org/10.5281/zenodo.14014559" ID-Zenodo-Dep="14014559" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88" httpUri="http://treatment.plazi.org/id/03A95D4BFFEE5023F372FDDC6A769C88" lastPageId="6" lastPageNumber="521" pageId="4" pageNumber="519">
<subSubSection id="C31ABFD6FFEE5021F372FDDC6ADB9F2F" box="[345,985,590,612]" pageId="4" pageNumber="519" type="nomenclature">
<paragraph id="8BBFEC5DFFEE5021F372FDDC6ADB9F2F" blockId="4.[345,985,590,638]" box="[345,985,590,612]" pageId="4" pageNumber="519">
<heading id="D0F75B31FFEE5021F372FDDC6ADB9F2F" box="[345,985,590,612]" pageId="4" pageNumber="519">
<taxonomicName id="4C0097DEFFEE5021F372FDDC6A489F28" authority="Brown and Triplehorn" authorityName="Brown and Triplehorn" box="[345,842,591,612]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="species" species="arenaria" status="sp. nov.">
<emphasis id="B974304FFFEE5021F372FDDC6B539F28" box="[345,593,591,612]" italics="true" pageId="4" pageNumber="519">Epitragosoma arenaria</emphasis>
Brown and Triplehorn
</taxonomicName>
,
<emphasis id="B974304FFFEE5021F17CFDDD6ADB9F2F" bold="true" box="[855,985,590,611]" pageId="4" pageNumber="519">
<taxonomicNameLabel id="A2478D34FFEE5021F17CFDDD6ADB9F2F" box="[855,985,590,611]" pageId="4" pageNumber="519" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFEE5021F04CFDFA6BEF9874" pageId="4" pageNumber="519" type="description">
<paragraph id="8BBFEC5DFFEE5021F04CFDFA6BC99F32" blockId="4.[345,985,590,638]" box="[615,715,617,638]" pageId="4" pageNumber="519">
<figureCitation id="133BF0D8FFEE5021F04CFDFA6BC99F32" box="[615,715,617,638]" captionStart-0="Fig" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="1.[269,303,1483,1502]" captionStartId-1="2.[272,315,931,950]" captionStartId-2="4.[272,315,1608,1627]" captionTargetBox-0="[242,1078,269,1455]" captionTargetBox-1="[252,1080,268,903]" captionTargetId-0="figure-11@1.[239,1084,265,1462]" captionTargetId-1="figure-12@2.[248,1083,265,910]" captionTargetId-2="figure-10@3.[246,1078,265,1644]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Epitragosoma arenaria dorsal habitus, female." captionText-1="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." captionText-2="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi-0="http://doi.org/10.5281/zenodo.10102876" figureDoi-1="http://doi.org/10.5281/zenodo.10102878" figureDoi-2="http://doi.org/10.5281/zenodo.14014557" httpUri-0="https://zenodo.org/record/10102876/files/figure.png" httpUri-1="https://zenodo.org/record/10102878/files/figure.png" httpUri-2="https://zenodo.org/record/14014557/files/figure.png" pageId="4" pageNumber="519">Figs. 18</figureCitation>
</paragraph>
<paragraph id="8BBFEC5DFFEE5021F33BFD0E6BEF9874" blockId="4.[249,1083,669,1336]" pageId="4" pageNumber="519">
<emphasis id="B974304FFFEE5021F33BFD0E688B9FFC" bold="true" box="[272,393,669,688]" pageId="4" pageNumber="519">Description.</emphasis>
Female (
<figureCitation id="133BF0D8FFEE5021F3C0FD0E6B259FFC" box="[491,551,669,688]" captionStart="Fig" captionStartId="1.[269,303,1483,1502]" captionTargetBox="[242,1078,269,1455]" captionTargetId="figure-11@1.[239,1084,265,1462]" captionTargetPageId="1" captionText="Fig. 1. Epitragosoma arenaria dorsal habitus, female." figureDoi="http://doi.org/10.5281/zenodo.10102876" httpUri="https://zenodo.org/record/10102876/files/figure.png" pageId="4" pageNumber="519">Fig. 1</figureCitation>
). Moderately shining reddish­brown with an aeneous tinge, clothed with short, fine but conspicuous, appressed, grayish­silver pubescence.
<emphasis id="B974304FFFEE5021F2D2FD5E68319FAC" box="[249,307,717,736]" italics="true" pageId="4" pageNumber="519">Head:</emphasis>
(
<figureCitation id="133BF0D8FFEE5021F369FD5E687F9FAC" box="[322,381,717,736]" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="4" pageNumber="519">Fig. 5</figureCitation>
) slightly broader than long; temporal area scarcely swollen behind eyes, much less than one half the length of longitudinal diameter of eye; head surface coarsely and densely punctured, each puncture with a conspicuous appressed seta.
<emphasis id="B974304FFFEE5021F1FEFD6E6D3B9E5C" box="[981,1081,765,784]" italics="true" pageId="4" pageNumber="519">Pronotum:</emphasis>
slightly broader than long, about equally wide at middle and at hind angles; lateral margins distinctly sinuate in basal one third with basal angles notably everted and apical angles not to slightly everted; surface coarsely and densely punctured, each puncture with a prominent, appresed seta, setae of anterior portion of disc directed caudad, lateral setae directed mediad, forming a swirling dendritic effect over much of pronotum with a faint impunctate medial line.
<emphasis id="B974304FFFEE5021F030FC1E6B5E9EEC" box="[539,604,909,928]" italics="true" pageId="4" pageNumber="519">Elytra:</emphasis>
surface finely and shallowly but densely punctured with a fine pattern of microreticulations between punctures; each puncture with a conspicuous, appressed seta directed caudad (setae somewhat shorter and finer than those of head and pronotum); faint dark subcutaneous longitudinal lines are visible but pattern of punctures is random.
<emphasis id="B974304FFFEE5021F3C9FC7E6B7E994C" box="[482,636,1005,1024]" italics="true" pageId="4" pageNumber="519">Ventral surface:</emphasis>
of pronotum coarsely and rugosely punctured; punctures progressively less dense on prosternum and prosternal process, each puncture with an appressed seta similar to pronotum; metasternum with large, shallow, contiguous punctures; abdominal sterna finely and rugosely punctured, clothed with appressed setae that are similar in size but less conspicuous than those of elytra.
<emphasis id="B974304FFFEE5021F17AFBDE6A86992C" box="[849,900,1101,1120]" italics="true" pageId="4" pageNumber="519">Legs:</emphasis>
densely punctured, each puncture bearing an appressed grayish­silver seta; femora smooth; tibae rugose, bearing numerous ventral and latero­ventral stout orange spines especially towards apices; tarsi with dorsal and lateral surfaces similar to tibiae with coarse, stout setae confined to apices and ventro­apical surfaces of pre­apical segments except for metatarsus (
<figureCitation id="133BF0D8FFEE5021F63FFB3E68079994" captionStart="Figs" captionStartId="4.[272,315,1608,1627]" captionTargetId="figure-10@3.[246,1078,265,1644]" captionTargetPageId="3" captionText="Figs. 48. Epitragosoma arenaria. 4) prosternal process, ventral view; 5) head, dorsal; 6) antenna; 7) metatarsus; 8) metathoracic wing." figureDoi="http://doi.org/10.5281/zenodo.14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="4" pageNumber="519">Fig. 7</figureCitation>
) with a more general ventral distribution of stout setae; all apical tarsal segments with two regular vetral rows of stout setae.
<emphasis id="B974304FFFEE5021F04EFB4E6A1999BC" box="[613,795,1245,1264]" italics="true" pageId="4" pageNumber="519">Abdominal tergum:</emphasis>
7 (
<figureCitation id="133BF0D8FFEE5021F117FB4E6A7699BC" box="[828,884,1245,1264]" captionStart="Figs" captionStartId="2.[272,315,931,950]" captionTargetBox="[252,1080,268,903]" captionTargetId="figure-12@2.[248,1083,265,910]" captionTargetPageId="2" captionText="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." figureDoi="http://doi.org/10.5281/zenodo.10102878" httpUri="https://zenodo.org/record/10102878/files/figure.png" pageId="4" pageNumber="519">Fig. 2</figureCitation>
) with two prolonged sharp projections; tergum 8 trapezoidal and with an apico­central uplifted spatulate structure.
<emphasis id="B974304FFFEE5021F300FA9E6860986C" box="[299,354,1293,1312]" italics="true" pageId="4" pageNumber="519">Male:</emphasis>
Similar to female but slightly smaller and more slender. Abdominal tergum 7 weakly bifurcate; tergum 8 normal lunate (
<figureCitation id="133BF0D8FFEE5021F08DFAB66BE29874" box="[678,736,1317,1336]" captionStart="Figs" captionStartId="2.[272,315,931,950]" captionTargetBox="[252,1080,268,903]" captionTargetId="figure-12@2.[248,1083,265,910]" captionTargetPageId="2" captionText="Figs. 23. Epitragosoma arenaria. 2) apical abdominal tergites, female; 3) apical abdominal tergites, male." figureDoi="http://doi.org/10.5281/zenodo.10102878" httpUri="https://zenodo.org/record/10102878/files/figure.png" pageId="4" pageNumber="519">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFEE5021F338FADE6AF5982E" box="[275,1015,1357,1378]" pageId="4" pageNumber="519" type="etymology">
<paragraph id="8BBFEC5DFFEE5021F338FADE6AF5982E" blockId="4.[249,1081,1357,1432]" box="[275,1015,1357,1378]" pageId="4" pageNumber="519">
<emphasis id="B974304FFFEE5021F338FADE6891982E" bold="true" box="[275,403,1357,1378]" pageId="4" pageNumber="519">Etymology.</emphasis>
The name of this species refers to its sand dune habitat.
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFEE5020F338FAFB6A879908" lastPageId="5" lastPageNumber="520" pageId="4" pageNumber="519" type="materials_examined">
<materialsCitation id="3B68E600FFEE5021F338FAFB6BF998F0" collectingDate="1972-07-16" collectorName="C. A. &amp; W. E. &amp; B. W. Triplehorn" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="4" pageNumber="519" specimenCount="1" specimenCount-female="1" stateProvince="Texas" typeStatus="holotype">
<paragraph id="8BBFEC5DFFEE5021F338FAFB6A4F98D4" blockId="4.[249,1081,1357,1432]" pageId="4" pageNumber="519">
<emphasis id="B974304FFFEE5021F338FAFB68B39831" bold="true" box="[275,433,1384,1405]" pageId="4" pageNumber="519">Type locality.</emphasis>
<collectingCountry id="F317ACCDFFEE5021F396FAFB6B059831" box="[445,519,1384,1405]" name="United States of America" pageId="4" pageNumber="519">U.S.A.</collectingCountry>
,
<collectingRegion id="49C422BFFFEE5021F033FAFB6B559831" box="[536,599,1384,1405]" country="United States of America" name="Texas" pageId="4" pageNumber="519">Texas</collectingRegion>
,
<collectingCounty id="62DE94D1FFEE5021F04CFAFB6BF99831" box="[615,763,1384,1405]" pageId="4" pageNumber="519">Ward County</collectingCounty>
,
<location id="8EDFBA86FFEE5021F127FAFB682E98D4" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEE5021F127FAFB682E98D4" country="United States of America" county="Ward County" name="Monahans Sand Hills State Park" pageId="4" pageNumber="519" stateProvince="Texas">Monahans Sand Hills State Park</location>
,
<date id="FFBECA9DFFEE5021F313FA1068C598D4" box="[312,455,1411,1432]" pageId="4" pageNumber="519" value="1972-07-16">
<collectingDate id="EFFA3375FFEE5021F313FA1068C598D4" box="[312,455,1411,1432]" pageId="4" pageNumber="519" value="1972-07-16">16 July 1972</collectingDate>
</date>
,
<collectorName id="26F5898BFFEE5021F3FFFA106B1298D4" box="[468,528,1411,1432]" pageId="4" pageNumber="519">C. A.</collectorName>
,
<collectorName id="26F5898BFFEE5021F035FA106B5898D4" box="[542,602,1411,1432]" pageId="4" pageNumber="519">W. E.</collectorName>
and
<collectorName id="26F5898BFFEE5021F0BAFA106A4B98D4" box="[657,841,1411,1432]" pageId="4" pageNumber="519">B. W. Triplehorn</collectorName>
.
</paragraph>
<paragraph id="8BBFEC5DFFEE5021F33BFA3A6BF998F0" blockId="4.[272,763,1449,1468]" box="[272,763,1449,1468]" pageId="4" pageNumber="519">
<typeStatus id="54BB52FFFFEE5021F33BFA3A686D98F7" box="[272,367,1449,1468]" pageId="4" pageNumber="519" type="holotype">HOLOTYPE</typeStatus>
.
<specimenCount id="9D0627D4FFEE5021F357FA3A68C198F0" box="[380,451,1449,1468]" pageId="4" pageNumber="519" type="female">Female</specimenCount>
. Length
<quantity id="4CF841B8FFEE5021F03CFA3A6B6098F0" box="[535,610,1449,1468]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="4" pageNumber="519" unit="mm" value="9.6">9.6 mm</quantity>
, width
<quantity id="4CF841B8FFEE5021F080FA3A6BF598F0" box="[683,759,1449,1468]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="519" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
</materialsCitation>
<paragraph id="8BBFEC5DFFEE5021F338FA426D1E98AA" blockId="4.[275,1052,1489,1510]" box="[275,1052,1489,1510]" pageId="4" pageNumber="519">
<materialsCitation id="3B68E600FFEE5021F338FA426D1E98AA" box="[275,1052,1489,1510]" collectingDate="1972-07-16" collectorName="C. A. &amp; W. E. &amp; B. W. Triplehorn" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="4" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Texas" typeStatus="allotype">
<emphasis id="B974304FFFEE5021F338FA42687B98AA" bold="true" box="[275,377,1489,1510]" pageId="4" pageNumber="519">
<typeStatus id="54BB52FFFFEE5021F338FA42687198AA" box="[275,371,1489,1510]" pageId="4" pageNumber="519" type="allotype">Allotype</typeStatus>
.
</emphasis>
<specimenCount id="9D0627D4FFEE5021F3AAFA4268BB98AA" box="[385,441,1489,1510]" pageId="4" pageNumber="519" type="male">Male</specimenCount>
, same data as Holotype. Length
<quantity id="4CF841B8FFEE5021F137FA426A7298AA" box="[796,880,1489,1510]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="4" pageNumber="519" unit="mm" value="9.3">9.3 mm</quantity>
, width
<quantity id="4CF841B8FFEE5021F1E9FA426D1A98AA" box="[962,1048,1489,1510]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="519" unit="mm" value="4.2">4.2 mm</quantity>
.
</materialsCitation>
</paragraph>
<caption id="DF7FBCD5FFEE5021F33BF9DB6BED9B38" ID-DOI="http://doi.org/10.5281/zenodo.14014557" ID-Zenodo-Dep="14014557" httpUri="https://zenodo.org/record/14014557/files/figure.png" pageId="4" pageNumber="519" startId="4.[272,315,1608,1627]" targetBox="[246,1078,265,1644]" targetPageId="3">
<paragraph id="8BBFEC5DFFEE5021F33BF9DB6BED9B38" blockId="4.[249,1082,1608,1652]" pageId="4" pageNumber="519">
<emphasis id="B974304FFFEE5021F33BF9DB686D9B17" bold="true" box="[272,367,1608,1627]" pageId="4" pageNumber="519">Figs. 48.</emphasis>
<taxonomicName id="4C0097DEFFEE5021F3ADF9DA6B609B10" box="[390,610,1609,1628]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="519" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B974304FFFEE5021F3ADF9DA6B609B10" box="[390,610,1609,1628]" italics="true" pageId="4" pageNumber="519">Epitragosoma arenaria</emphasis>
</taxonomicName>
.
<emphasis id="B974304FFFEE5021F044F9DB6B819B17" bold="true" box="[623,643,1608,1627]" pageId="4" pageNumber="519">4)</emphasis>
prosternal process, ventral view;
<emphasis id="B974304FFFEE5021F1EAF9DB6AD79B17" bold="true" box="[961,981,1608,1627]" pageId="4" pageNumber="519">5)</emphasis>
head, dorsal;
<emphasis id="B974304FFFEE5021F30AF9F368369B3F" bold="true" box="[289,308,1632,1651]" pageId="4" pageNumber="519">6)</emphasis>
antenna;
<emphasis id="B974304FFFEE5021F3B8F9F368A49B3F" bold="true" box="[403,422,1632,1651]" pageId="4" pageNumber="519">7)</emphasis>
metatarsus;
<emphasis id="B974304FFFEE5021F034F9F36B309B3F" bold="true" box="[543,562,1632,1651]" pageId="4" pageNumber="519">8)</emphasis>
metathoracic wing.
</paragraph>
</caption>
<paragraph id="8BBFEC5DFFEF5020F33BFE9D68CE9E75" blockId="5.[246,1079,270,1652]" pageId="5" pageNumber="520">
<materialsCitation id="3B68E600FFEF5020F33BFE9D68249C72" collectingDate="1972-07-16" collectorName="C. A. &amp; W. E. &amp; B. W. Triplehorn" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="110" stateProvince="Texas" typeStatus="paratype">
<emphasis id="B974304FFFEF5020F33BFE9D68859C6F" bold="true" box="[272,391,270,291]" pageId="5" pageNumber="520">
<typeStatus id="54BB52FFFFEF5020F33BFE9D68809C6F" box="[272,386,270,291]" pageId="5" pageNumber="520" type="paratype">Paratypes</typeStatus>
.
</emphasis>
All from same locality as Holotype: (
<specimenCount id="9D0627D4FFEF5020F101FE9C6A519C68" box="[810,851,271,292]" pageId="5" pageNumber="520">110</specimenCount>
) same data as Holotype
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F319FEBA6A569C72" box="[306,852,297,318]" collectingDate="1968-07-08" collectorName="C. A. &amp; W. E. &amp; B. W. Triplehorn" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="1" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F317FEBA68459C72" box="[316,327,297,318]" pageId="5" pageNumber="520">1</specimenCount>
) same collectors as Holotype except
<date id="FFBECA9DFFEF5020F0FEFEBA6A569C72" box="[725,852,297,318]" pageId="5" pageNumber="520" value="1968-07-08">
<collectingDate id="EFFA3375FFEF5020F0FEFEBA6A569C72" box="[725,852,297,318]" pageId="5" pageNumber="520" value="1968-07-08">8 July 1968</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F14AFEBA6B549C15" collectingDate="1973-07-28" collectorName="C. A. &amp; W. E. &amp; B. W. Triplehorn" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="23" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F143FEBA6A869C72" box="[872,900,297,318]" pageId="5" pageNumber="520">23</specimenCount>
) same collectors as Holotype except
<date id="FFBECA9DFFEF5020F3E2FED76B549C15" box="[457,598,324,345]" pageId="5" pageNumber="520" value="1973-07-28">
<collectingDate id="EFFA3375FFEF5020F3E2FED76B549C15" box="[457,598,324,345]" pageId="5" pageNumber="520" value="1973-07-28">28 July 1973</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F049FED76ABE9C15" box="[610,956,324,345]" collectingDate="1970-09-12" collectorName="A. S. Menke" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="10" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F040FED76B849C15" box="[619,646,324,345]" pageId="5" pageNumber="520">10</specimenCount>
)
<date id="FFBECA9DFFEF5020F0BDFED76A2B9C15" box="[662,809,324,345]" pageId="5" pageNumber="520" value="1970-09-12">
<collectingDate id="EFFA3375FFEF5020F0BDFED76A2B9C15" box="[662,809,324,345]" pageId="5" pageNumber="520" value="1970-09-12">12 Sept. 1970</collectingDate>
</date>
<collectorName id="26F5898BFFEF5020F11AFED76ABE9C15" box="[817,956,324,345]" pageId="5" pageNumber="520">A. S. Menke</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F1ECFED76B289C38" collectingDate="1963-08-03" collectorName="J. G. &amp; B. L. Rozen" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="9" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F1FBFED76ADC9C15" box="[976,990,324,345]" pageId="5" pageNumber="520">9</specimenCount>
)
<date id="FFBECA9DFFEF5020F1C5FED7682F9C38" pageId="5" pageNumber="520" value="1963-08-03">
<collectingDate id="EFFA3375FFEF5020F1C5FED7682F9C38" pageId="5" pageNumber="520" value="1963-08-03">3 Aug. 1963</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F312FECC686F9C38" box="[313,365,351,372]" pageId="5" pageNumber="520">J. G.</collectorName>
and
<collectorName id="26F5898BFFEF5020F38FFECC6B289C38" box="[420,554,351,372]" pageId="5" pageNumber="520">B. L. Rozen</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F01CFECC6AFC9C38" box="[567,1022,351,372]" collectingDate="1982-08-31" collectingDateMax="1982-09-01" collectingDateMin="1982-08-31" collectorName="Rolf L. Aalbu" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="50" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F015FECC6B589C38" box="[574,602,351,372]" pageId="5" pageNumber="520">50</specimenCount>
)
<date id="FFBECA9DFFEF5020F040FECC6A5B9C38" box="[619,857,351,372]" pageId="5" pageNumber="520" value="1982-08-31" valueMax="1982-09-01" valueMin="1982-08-31">
<collectingDate id="EFFA3375FFEF5020F040FECC6A5B9C38" box="[619,857,351,372]" pageId="5" pageNumber="520" value="1982-08-31" valueMax="1982-09-01" valueMin="1982-08-31">31 Aug.1 Sept. 1982</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F14EFECC6AFC9C38" box="[869,1022,351,372]" pageId="5" pageNumber="520">Rolf L. Aalbu</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F620FECC6B8A9CC2" collectingDate="1986-09-30" collectingDateMax="1982-09-01" collectingDateMin="1982-08-31" collectorName="J. T. &amp; E. A. Doyen" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="32" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F639FECC6D2C9C38" box="[1042,1070,351,372]" pageId="5" pageNumber="520">32</specimenCount>
)
<date id="FFBECA9DFFEF5020F2DDFEEA68909CC2" box="[246,402,377,398]" pageId="5" pageNumber="520" value="1986-09-30">
<collectingDate id="EFFA3375FFEF5020F2DDFEEA68909CC2" box="[246,402,377,398]" pageId="5" pageNumber="520" value="1986-09-30">30 Sept. 1986</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F38AFEEA68D29CC2" box="[417,464,377,398]" pageId="5" pageNumber="520">J. T.</collectorName>
&amp;
<collectorName id="26F5898BFFEF5020F3D2FEEA6B8A9CC2" box="[505,648,377,398]" pageId="5" pageNumber="520">E. A. Doyen</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F0BCFEEA682A9CE5" collectingDate="1987-08-02" collectingDateMax="1982-09-01" collectingDateMin="1982-08-31" collectingMethod="night sweep" collectorName="C. B. Barr" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="1" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F08AFEEA6BAE9CC2" box="[673,684,377,398]" pageId="5" pageNumber="520">1</specimenCount>
)
<date id="FFBECA9DFFEF5020F0EBFEEA6A499CC2" box="[704,843,377,398]" pageId="5" pageNumber="520" value="1987-08-02">
<collectingDate id="EFFA3375FFEF5020F0EBFEEA6A499CC2" box="[704,843,377,398]" pageId="5" pageNumber="520" value="1987-08-02">2 Aug. 1987</collectingDate>
</date>
(
<collectingMethod id="5241944AFFEF5020F176FEEA6AE79CC2" box="[861,997,377,398]" pageId="5" pageNumber="520">night sweep</collectingMethod>
),
<collectorName id="26F5898BFFEF5020F1D0FEEA682A9CE5" pageId="5" pageNumber="520">C. B. Barr</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F31DFE076B849CE5" box="[310,646,404,425]" collectingDate="1977-07-19" collectingDateMax="1982-09-01" collectingDateMin="1982-08-31" collectingMethod="night sweep" collectorName="A. V. Evans" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="1" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F36BFE0768499CE5" box="[320,331,404,425]" pageId="5" pageNumber="520">1</specimenCount>
)
<date id="FFBECA9DFFEF5020F374FE0768F29CE5" box="[351,496,404,425]" pageId="5" pageNumber="520" value="1977-07-19">
<collectingDate id="EFFA3375FFEF5020F374FE0768F29CE5" box="[351,496,404,425]" pageId="5" pageNumber="520" value="1977-07-19">19 July 1977</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F3D4FE076B849CE5" box="[511,646,404,425]" pageId="5" pageNumber="520">A. V. Evans</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F0BFFE076D309CE5" box="[660,1074,404,425]" collectingDate="1959-09-13" collectingDateMax="1959-09-14" collectingDateMin="1959-09-13" collectingMethod="night sweep" collectorName="E. R. Tinkham" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="1" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F0B5FE076BAB9CE5" box="[670,681,404,425]" pageId="5" pageNumber="520">1</specimenCount>
)
<date id="FFBECA9DFFEF5020F096FE076A829CE5" box="[701,896,404,425]" pageId="5" pageNumber="520" value="1959-09-13" valueMax="1959-09-14" valueMin="1959-09-13">
<collectingDate id="EFFA3375FFEF5020F096FE076A829CE5" box="[701,896,404,425]" pageId="5" pageNumber="520" value="1959-09-13" valueMax="1959-09-14" valueMin="1959-09-13">1314 Sept. 1959</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F1A5FE076D309CE5" box="[910,1074,404,425]" pageId="5" pageNumber="520">E. R. Tinkham</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F2DDFE3C6B779C88" box="[246,629,431,452]" collectingDate="1982-08-21" collectingDateMax="1982-08-22" collectingDateMin="1982-08-21" collectingMethod="night sweep" collectorName="R. Turnbow" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="16" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F2D5FE3C681B9C88" box="[254,281,431,452]" pageId="5" pageNumber="520">16</specimenCount>
)
<date id="FFBECA9DFFEF5020F301FE3C68E49C88" box="[298,486,431,452]" pageId="5" pageNumber="520" value="1982-08-21" valueMax="1982-08-22" valueMin="1982-08-21">
<collectingDate id="EFFA3375FFEF5020F301FE3C68E49C88" box="[298,486,431,452]" pageId="5" pageNumber="520" value="1982-08-21" valueMax="1982-08-22" valueMin="1982-08-21">2122 Aug. 1982</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F3D8FE3C6B779C88" box="[499,629,431,452]" pageId="5" pageNumber="520">R. Turnbow</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F0AAFE3C6AB49C88" box="[641,950,431,452]" collectingDate="1978-07-06" collectingDateMax="1982-08-22" collectingDateMin="1982-08-21" collectingMethod="night sweep" collectorName="R. Turnbow" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="1" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F0A0FE3C6B949C88" box="[651,662,431,452]" pageId="5" pageNumber="520">1</specimenCount>
)
<date id="FFBECA9DFFEF5020F08CFE3C6A2A9C88" box="[679,808,431,452]" pageId="5" pageNumber="520" value="1978-07-06">
<collectingDate id="EFFA3375FFEF5020F08CFE3C6A2A9C88" box="[679,808,431,452]" pageId="5" pageNumber="520" value="1978-07-06">6 July 1978</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F11FFE3C6AB49C88" box="[820,950,431,452]" pageId="5" pageNumber="520">R. Turnbow</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F1EAFE3C6B869C92" collectingDate="1976-07-23" collectingDateMax="1982-08-22" collectingDateMin="1982-08-21" collectingMethod="sweeping grass" collectorName="G. H. Nelson" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="6" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F1E1FE3C6ADA9C88" box="[970,984,431,452]" pageId="5" pageNumber="520">6</specimenCount>
)
<date id="FFBECA9DFFEF5020F1C3FE3C68289C92" pageId="5" pageNumber="520" value="1976-07-23">
<collectingDate id="EFFA3375FFEF5020F1C3FE3C68289C92" pageId="5" pageNumber="520" value="1976-07-23">23 July 1976</collectingDate>
</date>
(
<collectingMethod id="5241944AFFEF5020F311FE5A68DD9C92" box="[314,479,457,478]" pageId="5" pageNumber="520">sweeping grass</collectingMethod>
),
<collectorName id="26F5898BFFEF5020F3DFFE5A6B869C92" box="[500,644,457,478]" pageId="5" pageNumber="520">G. H. Nelson</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F0BBFE5A688A9CB5" collectingDate="1975-07-15" collectingDateMax="1975-07-16" collectingDateMin="1975-07-15" collectingMethod="sweeping grass" collectorName="G. H. Nelson" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="10" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F0B3FE5A6BB19C92" box="[664,691,457,478]" pageId="5" pageNumber="520">10</specimenCount>
)
<date id="FFBECA9DFFEF5020F0EFFE5A6A779C92" box="[708,885,457,478]" pageId="5" pageNumber="520" value="1975-07-15" valueMax="1975-07-16" valueMin="1975-07-15">
<collectingDate id="EFFA3375FFEF5020F0EFFE5A6A779C92" box="[708,885,457,478]" pageId="5" pageNumber="520" value="1975-07-15" valueMax="1975-07-16" valueMin="1975-07-15">1516 July 1975</collectingDate>
</date>
(
<collectingMethod id="5241944AFFEF5020F1AEFE5A6D289C92" box="[901,1066,457,478]" pageId="5" pageNumber="520">sweeping grass</collectingMethod>
),
<collectorName id="26F5898BFFEF5020F2DDFE77688A9CB5" box="[246,392,484,505]" pageId="5" pageNumber="520">G. H. Nelson</collectorName>
</materialsCitation>
;
<materialsCitation id="3B68E600FFEF5020F3BDFE776BF99CB5" box="[406,763,484,505]" collectingDate="1972-07-21" collectingDateMax="1975-07-16" collectingDateMin="1975-07-15" collectingMethod="sweeping grass" collectorName="Karl Stephan" country="United States of America" county="Ward County" location="Monahans Sand Hills State Park" pageId="5" pageNumber="520" specimenCount="12" stateProvince="Texas">
(
<specimenCount id="9D0627D4FFEF5020F3B5FE7768BB9CB5" box="[414,441,484,505]" pageId="5" pageNumber="520">12</specimenCount>
)
<date id="FFBECA9DFFEF5020F3E0FE776B589CB5" box="[459,602,484,505]" pageId="5" pageNumber="520" value="1972-07-21">
<collectingDate id="EFFA3375FFEF5020F3E0FE776B589CB5" box="[459,602,484,505]" pageId="5" pageNumber="520" value="1972-07-21">21 July 1972</collectingDate>
</date>
,
<collectorName id="26F5898BFFEF5020F043FE776BF59CB5" box="[616,759,484,505]" pageId="5" pageNumber="520">Karl Stephan</collectorName>
.
</materialsCitation>
<emphasis id="B974304FFFEF5020F128FE776AA79CB5" box="[771,933,484,505]" italics="true" pageId="5" pageNumber="520">Measurements:</emphasis>
Length:
<quantity id="4CF841B8FFEF5020F623FE7768559F58" metricMagnitude="-3" metricUnit="m" metricValue="9.899999999999999" metricValueMax="10.7" metricValueMin="9.1" pageId="5" pageNumber="520" unit="mm" value="9.899999999999999" valueMax="10.7" valueMin="9.1">9.1 10.7 mm</quantity>
; width: 4.0
<quantity id="4CF841B8FFEF5020F3F5FE6C6B319F58" box="[478,563,511,532]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="520" unit="mm" value="4.9">4.9 mm</quantity>
.
<typeStatus id="54BB52FFFFEF5020F015FE6C6BA09F58" box="[574,674,511,532]" pageId="5" pageNumber="520" type="holotype">Holotype</typeStatus>
,
<typeStatus id="54BB52FFFFEF5020F086FE6C6A0A9F58" box="[685,776,511,532]" pageId="5" pageNumber="520" type="allotype">Allotype</typeStatus>
and
<typeStatus id="54BB52FFFFEF5020F115FE6C6AA69F58" box="[830,932,511,532]" pageId="5" pageNumber="520" type="paratype">Paratypes</typeStatus>
are deposited in the
<collectingCountry id="F317ACCDFFEF5020F316FD8A68C99F62" box="[317,459,537,558]" name="United States of America" pageId="5" pageNumber="520">United States</collectingCountry>
National Museum. Additional
<typeStatus id="54BB52FFFFEF5020F137FD8A6A809F62" box="[796,898,537,558]" pageId="5" pageNumber="520" type="paratype">Paratypes</typeStatus>
are deposited in the
<collectingRegion id="49C422BFFFEF5020F334FDA768929F05" box="[287,400,564,585]" country="United States of America" name="Ohio" pageId="5" pageNumber="520">Ohio State</collectingRegion>
University Museum of Biological Diversity;
<collectingMunicipality id="6BDB7627FFEF5020F153FDA768C39F28" pageId="5" pageNumber="520">Peabody Museum of Natural History</collectingMunicipality>
,
<location id="8EDFBA86FFEF5020F3E4FDDC6B789F28" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F3E4FDDC6B789F28" box="[463,634,591,612]" country="United States of America" county="University Museum of Biological Diversity" municipality="Peabody Museum of Natural History" name="Yale University" pageId="5" pageNumber="520" stateProvince="California">Yale University</location>
; Museum of Comparative Zoology, Harvard University;
<location id="8EDFBA86FFEF5020F380FDFA6A479F32" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F380FDFA6A479F32" box="[427,837,617,638]" country="United States of America" county="University Museum of Biological Diversity" municipality="Peabody Museum of Natural History" name="American Museum of Natural History" pageId="5" pageNumber="520" stateProvince="California">American Museum of Natural History</location>
;
<location id="8EDFBA86FFEF5020F17AFDFA6ABD9F32" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F17AFDFA6ABD9F32" box="[849,959,617,638]" country="United States of America" county="University Museum of Biological Diversity" municipality="Peabody Museum of Natural History" name="University" pageId="5" pageNumber="520" stateProvince="California">University</location>
of
<collectingRegion id="49C422BFFFEF5020F1CEFDFA681A9FD5" country="United States of America" name="California" pageId="5" pageNumber="520">California</collectingRegion>
,
<collectorName id="26F5898BFFEF5020F30CFD1768879FD5" box="[295,389,644,665]" pageId="5" pageNumber="520">Berkeley</collectorName>
and
<collectorName id="26F5898BFFEF5020F3EBFD176B029FD5" box="[448,512,644,665]" pageId="5" pageNumber="520">Davis</collectorName>
;
<collectingRegion id="49C422BFFFEF5020F024FD176B499FD5" box="[527,587,644,665]" country="United States of America" name="Texas" pageId="5" pageNumber="520">Texas</collectingRegion>
<collectorName id="26F5898BFFEF5020F07DFD176A019FD5" box="[598,771,644,665]" pageId="5" pageNumber="520">Tech University</collectorName>
, Lubbock,
<collectingRegion id="49C422BFFFEF5020F154FD176ABD9FD5" box="[895,959,644,665]" country="United States of America" name="Texas" pageId="5" pageNumber="520">Texas</collectingRegion>
;
<collectingRegion id="49C422BFFFEF5020F1E6FD176D1A9FD5" box="[973,1048,644,665]" country="United States of America" name="Florida" pageId="5" pageNumber="520">Florida</collectingRegion>
<collectorName id="26F5898BFFEF5020F608FD1768A69FF8" pageId="5" pageNumber="520">A &amp; M University</collectorName>
,
<location id="8EDFBA86FFEF5020F39AFD0C6B2E9FF8" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F39AFD0C6B2E9FF8" box="[433,556,671,692]" country="United States of America" name="Tallahassee" pageId="5" pageNumber="520" stateProvince="Florida">Tallahassee</location>
,
<collectingRegion id="49C422BFFFEF5020F013FD0C6B859FF8" box="[568,647,671,692]" country="United States of America" name="Florida" pageId="5" pageNumber="520">Florida</collectingRegion>
;
<collectingRegion id="49C422BFFFEF5020F0BFFD0C6A1E9FF8" box="[660,796,671,692]" country="United States of America" name="Florida" pageId="5" pageNumber="520">Florida State</collectingRegion>
<location id="8EDFBA86FFEF5020F10EFD0C6D319FF8" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F10EFD0C6D319FF8" box="[805,1075,671,692]" country="United States of America" name="Collection of Arthropods" pageId="5" pageNumber="520" stateProvince="California">Collection of Arthropods</location>
, Gainesville;
<collectingRegion id="49C422BFFFEF5020F354FD2A68EB9F82" box="[383,489,697,718]" country="United States of America" name="California" pageId="5" pageNumber="520">California</collectingRegion>
<collectingMunicipality id="6BDB7627FFEF5020F3D8FD2A6BE09F82" box="[499,738,697,718]" pageId="5" pageNumber="520">Academy of Sciences</collectingMunicipality>
;
<collectorName id="26F5898BFFEF5020F0DBFD2A6A9F9F82" box="[752,925,697,718]" pageId="5" pageNumber="520">British Museum</collectorName>
(
<collectorName id="26F5898BFFEF5020F185FD2A68279FA5" pageId="5" pageNumber="520">Natural History</collectorName>
);
<collectorName id="26F5898BFFEF5020F311FD476B679FA5" box="[314,613,724,745]" pageId="5" pageNumber="520">Kirby W. Brown Collection</collectorName>
,
<location id="8EDFBA86FFEF5020F05AFD476BCE9FA5" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F05AFD476BCE9FA5" box="[625,716,724,745]" country="United States of America" municipality="Academy of Sciences" name="Paradise" pageId="5" pageNumber="520" stateProvince="California">Paradise</location>
,
<collectingRegion id="49C422BFFFEF5020F0F2FD476A449FA5" box="[729,838,724,745]" country="United States of America" name="California" pageId="5" pageNumber="520">California</collectingRegion>
;
<collectorName id="26F5898BFFEF5020F178FD4768239E48" pageId="5" pageNumber="520">G. H. Nelson Collection</collectorName>
,
<collectingRegion id="49C422BFFFEF5020F306FD7C687A9E48" box="[301,376,751,772]" country="United States of America" name="Kansas" pageId="5" pageNumber="520">Kansas</collectingRegion>
<collectorName id="26F5898BFFEF5020F3ABFD7C68AD9E48" box="[384,431,751,772]" pageId="5" pageNumber="520">City</collectorName>
,
<collectingRegion id="49C422BFFFEF5020F390FD7C6B199E48" box="[443,539,751,772]" country="United States of America" name="Missouri" pageId="5" pageNumber="520">Missouri</collectingRegion>
;
<collectorName id="26F5898BFFEF5020F00CFD7C6A1E9E48" box="[551,796,751,772]" pageId="5" pageNumber="520">R. Turnbow Collection</collectorName>
,
<location id="8EDFBA86FFEF5020F10CFD7C6AA49E48" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F10CFD7C6AA49E48" box="[807,934,751,772]" country="United States of America" name="Fort Rucker" pageId="5" pageNumber="520" stateProvince="Alabama">Fort Rucker</location>
,
<collectingRegion id="49C422BFFFEF5020F19AFD7C6D119E48" box="[945,1043,751,772]" country="United States of America" name="Alabama" pageId="5" pageNumber="520">Alabama</collectingRegion>
; C. B, Barr Collection,
<location id="8EDFBA86FFEF5020F3E5FC9A6B2C9E52" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F3E5FC9A6B2C9E52" box="[462,558,777,798]" country="United States of America" name="Berkeley" pageId="5" pageNumber="520" stateProvince="California">Berkeley</location>
,
<collectingRegion id="49C422BFFFEF5020F016FC9A6BA89E52" box="[573,682,777,798]" country="United States of America" name="California" pageId="5" pageNumber="520">California</collectingRegion>
, and
<collectorName id="26F5898BFFEF5020F0CCFC9A6AF89E52" box="[743,1018,777,798]" pageId="5" pageNumber="520">Rolf L. Aalbu Collection</collectorName>
,
<location id="8EDFBA86FFEF5020F623FC9A684C9E75" LSID="urn:lsid:plazi:treatment:03A95D4BFFEE5023F372FDDC6A769C88:8EDFBA86FFEF5020F623FC9A684C9E75" country="United States of America" name="Sacramento" pageId="5" pageNumber="520" stateProvince="California">Sacramento</location>
,
<collectingRegion id="49C422BFFFEF5020F370FCB768CA9E75" box="[347,456,804,825]" country="United States of America" name="California" pageId="5" pageNumber="520">California</collectingRegion>
.
</paragraph>
<paragraph id="8BBFEC5DFFEF5020F33BFCAD6A879908" blockId="5.[246,1079,270,1652]" pageId="5" pageNumber="520">
<emphasis id="B974304FFFEF5020F33BFCAD68DE9E1F" bold="true" box="[272,476,830,851]" pageId="5" pageNumber="520">Other Specimens.</emphasis>
<collectingRegion id="49C422BFFFEF5020F3C2FCAC6B429E18" box="[489,576,831,852]" country="United States of America" name="Texas" pageId="5" pageNumber="520">TEXAS</collectingRegion>
, Culberson County: (1)
<quantity id="4CF841B8FFEF5020F17EFCAC6AAA9E18" box="[853,936,831,852]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="5" pageNumber="520" unit="mi" value="5.0">5 miles</quantity>
east of Van Horn,
<date id="FFBECA9DFFEF5020F310FCCA68D19E22" box="[315,467,857,878]" pageId="5" pageNumber="520" value="1970-09-12">12 Sept. 1970</date>
, A. S. Menke. Reeves County: (4) Pecos,
<date id="FFBECA9DFFEF5020F1B0FCCA6D309E22" box="[923,1074,857,878]" pageId="5" pageNumber="520" value="1938-09-23">23 Sept. 1938</date>
, D. J. and J. N. Knull. Winkler County: (91)
<quantity id="4CF841B8FFEF5020F0F4FCE76A2D9EC5" box="[735,815,884,905]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="5" pageNumber="520" unit="mi" value="4.0">4 miles</quantity>
northeast of Kermit,
<date id="FFBECA9DFFEF5020F637FCE7687E9EE8" pageId="5" pageNumber="520" value="1971-08-25">25 August 1971</date>
(on tassels of tall grass), K. W. Brown, C. OBrien and P. Petrulis; (25)
<quantity id="4CF841B8FFEF5020F301FC3A68859EF2" box="[298,391,937,958]" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="5" pageNumber="520" unit="mi" value="11.0">11 miles</quantity>
northeast of Kermit,
<date id="FFBECA9DFFEF5020F05BFC3A6A079EF2" box="[624,773,937,958]" pageId="5" pageNumber="520" value="1971-09-12">12 Sept. 1971</date>
(on tassels of tall grass), K. W. Brown, (1)
<quantity id="4CF841B8FFEF5020F38BFC5768FC9E95" box="[416,510,964,985]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="5" pageNumber="520" unit="mi" value="10.0">10 miles</quantity>
northeast of Kermit on Hwy 115, 3,200,
<date id="FFBECA9DFFEF5020F1F7FC57682F9EB8" pageId="5" pageNumber="520" value="1971-09-12">12 Sept. 1971</date>
, K. W. Brown.
<collectingRegion id="49C422BFFFEF5020F3F3FC4C6B859EB8" box="[472,647,991,1012]" country="United States of America" name="New Mexico" pageId="5" pageNumber="520">NEW MEXICO</collectingRegion>
, Chaves County: (10) Mescalero Sands, Caprock,
<date id="FFBECA9DFFEF5020F376FC6A68DF9942" box="[349,477,1017,1038]" pageId="5" pageNumber="520" value="1954-07-09">9 July 1954</date>
, Cazier and Gertsch; (3) Mescalero Sands,
<quantity id="4CF841B8FFEF5020F180FC6A6D049942" box="[939,1030,1017,1038]" metricMagnitude="4" metricUnit="m" metricValue="7.242048" pageId="5" pageNumber="520" unit="mi" value="45.0">45 miles</quantity>
east of Roswell,
<date id="FFBECA9DFFEF5020F351FB876B3B9965" box="[378,569,1044,1065]" pageId="5" pageNumber="520" value="1959-09-12" valueMax="1959-09-13" valueMin="1959-09-12">1213 Sept. 1959</date>
, E. R. Tinkham. Lea County: (8)
<quantity id="4CF841B8FFEF5020F183FB876AF59965" box="[936,1015,1044,1065]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="5" pageNumber="520" unit="mi" value="4.0">4 miles</quantity>
north of Maljamar,
<date id="FFBECA9DFFEF5020F3A3FBBC6B129908" box="[392,528,1071,1092]" pageId="5" pageNumber="520" value="1962-09-01">1 Sept. 1962</date>
(on
<taxonomicName id="4C0097DEFFEF5020F06EFBBC6BBA9908" box="[581,696,1071,1092]" class="Magnoliopsida" family="Asteraceae" genus="Helianthus" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="520" phylum="Tracheophyta" rank="genus">
<emphasis id="B974304FFFEF5020F06EFBBC6BBA9908" box="[581,696,1071,1092]" italics="true" pageId="5" pageNumber="520">Helianthus</emphasis>
</taxonomicName>
), H. V. Weems, Jr.
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFEF5020F33BFBDA6B7F99B2" pageId="5" pageNumber="520" type="biology_ecology">
<paragraph id="8BBFEC5DFFEF5020F33BFBDA6B7F99B2" blockId="5.[246,1079,270,1652]" pageId="5" pageNumber="520">
<emphasis id="B974304FFFEF5020F33BFBDA686F9912" bold="true" box="[272,365,1097,1118]" pageId="5" pageNumber="520">Habitat.</emphasis>
Most of the large series taken by us in Ward and Winkler Counties, Texas, were found head down in the leaf axils of dune­stabilizing grasses. Specimens of this grass from Monahans were determined as
<taxonomicName id="4C0097DEFFEF5020F152FBEC683799E2" authority="Vasey" authorityName="Vasey" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="520" phylum="Tracheophyta" rank="species" species="havardii">
<emphasis id="B974304FFFEF5020F152FBEC6D3499D8" box="[889,1078,1151,1172]" italics="true" pageId="5" pageNumber="520">Panicum havardii</emphasis>
Vasey
</taxonomicName>
by Dr. David Keil. This is apparently a native grass in arroyos and sand hills of western Texas, southern New
<collectingRegion id="49C422BFFFEF5020F0ADFB276BD49985" box="[646,726,1204,1225]" country="Mexico" name="Mexico" pageId="5" pageNumber="520">Mexico</collectingRegion>
and northern
<collectingCountry id="F317ACCDFFEF5020F140FB276AB99985" box="[875,955,1204,1225]" name="Mexico" pageId="5" pageNumber="520">Mexico</collectingCountry>
(Hitchcock 1935). A few specimens were taken on the sand around clumps of grass and one was found on a sunflower head.
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFEF5020F33BFA976BAE9898" pageId="5" pageNumber="520" type="diagnosis">
<paragraph id="8BBFEC5DFFEF5020F33BFA976BAE9898" blockId="5.[246,1079,270,1652]" pageId="5" pageNumber="520">
<emphasis id="B974304FFFEF5020F33BFA9768A39855" bold="true" box="[272,417,1284,1305]" pageId="5" pageNumber="520">Comparison.</emphasis>
Other than the superficial resemblance to
<taxonomicName id="4C0097DEFFEF5020F174FA976D349855" box="[863,1078,1284,1305]" class="Insecta" family="Tenebrionidae" genus="Epitragopsis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="520" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B974304FFFEF5020F174FA976AE79855" box="[863,997,1284,1305]" italics="true" pageId="5" pageNumber="520">Epitragopsis</emphasis>
species
</taxonomicName>
discussed in the generic comparison, this species is most closely approached by
<taxonomicName id="4C0097DEFFEF5020F332FAA96B419802" authority="(Casey)" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[281,579,1337,1359]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="520" phylum="Arthropoda" rank="species" species="ovale">
<emphasis id="B974304FFFEF5020F332FAA968E89803" box="[281,490,1338,1359]" italics="true" pageId="5" pageNumber="520">Lobometopon ovale</emphasis>
(Casey)
</taxonomicName>
found in eastern
<collectingRegion id="49C422BFFFEF5020F12AFAAA6A3F9802" box="[769,829,1337,1358]" country="United States of America" name="Texas" pageId="5" pageNumber="520">Texas</collectingRegion>
(Triplehorn 1980). Besides characters already discussed at the generic level,
<taxonomicName id="4C0097DEFFEF5020F16BFAC76A949825" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[832,918,1364,1385]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="520" phylum="Arthropoda" rank="species" species="ovale">
<emphasis id="B974304FFFEF5020F16BFAC76A949825" box="[832,918,1364,1385]" italics="true" pageId="5" pageNumber="520">L. ovale</emphasis>
</taxonomicName>
is less densely hirsute, posterior prothoracic angles not or only very weakly everted, and body slightly more robust. Furthermore
<taxonomicName id="4C0097DEFFEF5020F041FA196BC398D3" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[618,705,1418,1439]" class="Insecta" family="Tenebrionidae" genus="Lobometopon" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="520" phylum="Arthropoda" rank="species" species="ovale">
<emphasis id="B974304FFFEF5020F041FA196BC398D3" box="[618,705,1418,1439]" italics="true" pageId="5" pageNumber="520">L. ovale</emphasis>
</taxonomicName>
is associated with the dicotyledon plant
<taxonomicName id="4C0097DEFFEF5020F318FA376BD998F5" box="[307,731,1444,1465]" class="Magnoliopsida" family="Polygonaceae" genus="Eriogonum" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="520" phylum="Tracheophyta" rank="species" species="multiflorum">
<emphasis id="B974304FFFEF5020F318FA376B2E98F5" box="[307,556,1444,1465]" italics="true" pageId="5" pageNumber="520">Eriogonum multiflorum</emphasis>
(Polygonaceae)
</taxonomicName>
, while
<taxonomicName id="4C0097DEFFEF5020F100FA376AA498F5" box="[811,934,1444,1465]" class="Insecta" family="Tenebrionidae" genus="Epitragosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="520" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B974304FFFEF5020F100FA376AA498F5" box="[811,934,1444,1465]" italics="true" pageId="5" pageNumber="520">E. arenaria</emphasis>
</taxonomicName>
is associated with a monocotyledon grass (Graminae).
</paragraph>
</subSubSection>
<subSubSection id="C31ABFD6FFEF5023F33BFA4A6A769C88" lastPageId="6" lastPageNumber="521" pageId="5" pageNumber="520" type="description">
<paragraph id="8BBFEC5DFFEF5023F33BFA4A68CB9C38" blockId="5.[246,1079,270,1652]" lastBlockId="6.[249,1081,271,452]" lastPageId="6" lastPageNumber="521" pageId="5" pageNumber="520">
<emphasis id="B974304FFFEF5020F33BFA4A688098A2" bold="true" box="[272,386,1497,1518]" pageId="5" pageNumber="520">Variation.</emphasis>
Three large series are on hand from rather close locations; (1) Monahans Sand Hills State Park, (2)
<quantity id="4CF841B8FFEF5020F0A0FA676BD89B45" box="[651,730,1524,1545]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="5" pageNumber="520" unit="mi" value="4.0">4 miles</quantity>
northeast of Kermit, and (3)
<quantity id="4CF841B8FFEF5020F637FA67682D9B68" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="5" pageNumber="520" unit="mi" value="11.0">11 miles</quantity>
northeast of Kermit. Kermit is approximately
<quantity id="4CF841B8FFEF5020F16BF99C6AA29B68" box="[832,928,1551,1572]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="5" pageNumber="520" unit="mi" value="20.0">20 miles</quantity>
northwest of Monahans Sand Hills. When placed side by side, series of each population are recognizably distinct. Most notable are size differences (the Monahans specimens are larger than the Kermit specimens) but also there are subtle morphological differences. This is apparently a case of microgeographic variation similar to
<taxonomicName id="4C0097DEFFEC5023F347FEB96A819C72" authority="(Doyen and Slobodchikoff 1984)" baseAuthorityName="Doyen and Slobodchikoff" baseAuthorityYear="1984" box="[364,899,297,319]" class="Insecta" family="Tenebrionidae" genus="Coelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="521" phylum="Arthropoda" rank="species" species="ciliatus">
<emphasis id="B974304FFFEC5023F347FEB96B0F9C73" box="[364,525,298,319]" italics="true" pageId="6" pageNumber="521">Coelus ciliatus</emphasis>
(Doyen and Slobodchikoff 1984)
</taxonomicName>
. Because of this variability we have restricted the
<typeStatus id="54BB52FFFFEC5023F076FED76B889C15" box="[605,650,324,345]" pageId="6" pageNumber="521">type</typeStatus>
series to those specimens collected only at the
<typeStatus id="54BB52FFFFEC5023F315FECC68699C38" box="[318,363,351,372]" pageId="6" pageNumber="521">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BBFEC5DFFEC5023F338FEEA6A769C88" blockId="6.[249,1081,271,452]" pageId="6" pageNumber="521">
<emphasis id="B974304FFFEC5023F338FEEA68F89CC2" bold="true" box="[275,506,377,398]" pageId="6" pageNumber="521">Sexual Dimorphism.</emphasis>
Other than the structure of abdominal tergites 7 and 8, there appears to be little sexual dimorphism in this species. There is a weak tendency to a slightly smaller and narrower body in males.
</paragraph>
</subSubSection>
</treatment>
</document>