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<document id="BCD03FA28873FE4FE385C1535BB01A7F" ID-CLB-Dataset="30378" ID-DOI="10.11646/zootaxa.4402.3.4" ID-GBIF-Dataset="6b1f8c1f-1ebb-46ff-a749-c773c5426015" ID-ISSN="1175-5326" ID-Zenodo-Dep="1209708" ID-ZooBank="AFBEB273-0057-4F25-B702-CA2CB4568361" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522317935364" checkinUser="plazi" docAuthor="Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G. &amp; Desutter-Grandcolas, Laure" docDate="2018" docId="039987AF8039FFDCFF4B6DA4FCD934D1" docLanguage="en" docName="zootaxa.4402.3.4.pdf" docOrigin="Zootaxa 4402 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Titanogryllus Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas 2018, n. gen." docType="treatment" docVersion="7" lastPageNumber="489" masterDocId="FFA0FFD7803BFFDEFFDC6D7AFFE73345" masterDocTitle="Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae)" masterLastPageNumber="507" masterPageNumber="487" pageNumber="489" updateTime="1698534362614" updateUser="ExternalLinkService">
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<mods:title id="FA6897A42B24AD55F960874AECC8AC2A">Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae)</mods:title>
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<mods:namePart id="733CD5588BDA402118097A9FCD36F593">Jaiswara, Ranjana</mods:namePart>
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<mods:namePart id="BE760FEC57316DD53DAEA1F128648DD1">Souza-Dias, Pedro G. B.</mods:namePart>
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<mods:namePart id="113D834D6FD9E7C2FC270456A9E9426B">Campos, Lucas Denadai De</mods:namePart>
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<mods:namePart id="B3A093C606962ADC12790519DB294DEB">Redü, Darlan R.</mods:namePart>
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<mods:namePart id="327F4AA53734F22C9361C87D55EA3F70">De Mello, Francisco De A. G.</mods:namePart>
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<mods:namePart id="CE77CBDBAA86CA4018E1217F4BF2133F">Desutter-Grandcolas, Laure</mods:namePart>
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<mods:typeOfResource id="532F78838585BF79C0B6F7E1265E7A5B">text</mods:typeOfResource>
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<mods:date id="3AF02D96E2FB4C019690A063139679DA">2018</mods:date>
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<treatment id="039987AF8039FFDCFF4B6DA4FCD934D1" ID-DOI="http://doi.org/10.5281/zenodo.5987495" ID-GBIF-Taxon="142669833" ID-Zenodo-Dep="5987495" LSID="urn:lsid:plazi:treatment:039987AF8039FFDCFF4B6DA4FCD934D1" httpUri="http://treatment.plazi.org/id/039987AF8039FFDCFF4B6DA4FCD934D1" lastPageNumber="489" pageId="2" pageNumber="489">
<subSubSection id="C32A65328039FFDCFF4B6DA4FBBF3224" pageId="2" pageNumber="489" type="nomenclature">
<paragraph id="8B8F36B98039FFDCFF4B6DA4FBAC33BD" blockId="2.[151,1099,222,248]" box="[151,1099,222,248]" pageId="2" pageNumber="489">
<heading id="D0C781D58039FFDCFF4B6DA4FBAC33BD" bold="true" box="[151,1099,222,248]" fontSize="11" level="1" pageId="2" pageNumber="489" reason="1">
<emphasis id="B944EAAB8039FFDCFF4B6DA4FBAC33BD" bold="true" box="[151,1099,222,248]" pageId="2" pageNumber="489">
<taxonomicName id="4C304D3A8039FFDCFF4B6DA4FC1533BD" ID-CoL="8P27V" authority="Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello" authorityName="Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas" authorityYear="2018" box="[151,1010,222,248]" class="Insecta" family="Gryllidae" genus="Titanogryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944EAAB8039FFDCFF4B6DA4FEDF33BD" bold="true" box="[151,312,222,248]" italics="true" pageId="2" pageNumber="489">Titanogryllus</emphasis>
Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello
</taxonomicName>
<taxonomicNameLabel id="A27757D08039FFDCFC256DA4FBAC33BD" box="[1017,1099,222,248]" pageId="2" pageNumber="489" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B98039FFDCFF4B6C5FFBBF3224" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
Included in the molecular phylogeny of
<taxonomicName id="4C304D3A8039FFDCFDA76C5FFD02327B" box="[635,741,293,318]" class="Insecta" family="Gryllidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="subFamily" subFamily="Gryllinae">Gryllinae</taxonomicName>
produced by Jaiswara
<emphasis id="B944EAAB8039FFDCFC206C5CFBD5327B" box="[1020,1074,293,318]" italics="true" pageId="2" pageNumber="489">et al</emphasis>
. (in preparation) as LDG 281 (
<taxonomicName id="4C304D3A8039FFDCFF7C6C33FBEE3227" authority="Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello" authorityName="Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas" authorityYear="2018" box="[160,1033,329,354]" class="Insecta" family="Gryllidae" genus="Titanogryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="salgado" status="sp. nov.">
<emphasis id="B944EAAB8039FFDCFF7C6C33FE713227" box="[160,406,329,354]" italics="true" pageId="2" pageNumber="489">Titanogryllus salgado</emphasis>
Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello
</taxonomicName>
<taxonomicNameLabel id="A27757D08039FFDCFBCC6C33FBAA3224" box="[1040,1101,329,354]" pageId="2" pageNumber="489" rank="species">
<emphasis id="B944EAAB8039FFDCFBCC6C33FBA03227" bold="true" box="[1040,1095,329,354]" pageId="2" pageNumber="489">n. sp</emphasis>
.
</taxonomicNameLabel>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A65328039FFDCFF1B6C17FC83328B" pageId="2" pageNumber="489" type="etymology">
<paragraph id="8B8F36B98039FFDCFF1B6C17FC83328B" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6C17FEA932C3" bold="true" box="[199,334,365,390]" pageId="2" pageNumber="489">Etymology.</emphasis>
The word
<emphasis id="B944EAAB8039FFDCFE1A6C14FE1D32C0" box="[454,506,366,389]" italics="true" pageId="2" pageNumber="489">titan</emphasis>
has its origin in the Greek mythology. The titans were six elder, giant gods, sons of Uranus and Gaia. Titan is also a noun related to one that is gigantic in some attribute. Here, we use the prefix
<emphasis id="B944EAAB8039FFDCFA836CE8FF4A3288" italics="true" pageId="2" pageNumber="489">titanto</emphasis>
refer to the large size of the species of this new gryllid genus.
</paragraph>
</subSubSection>
<subSubSection id="C32A65328039FFDCFF1B6CA3FAFE317C" pageId="2" pageNumber="489" type="materials_examined">
<paragraph id="8B8F36B98039FFDCFF1B6CA3FAFF32B7" blockId="2.[151,1437,293,1941]" box="[199,1304,473,498]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6CA3FE8532B7" bold="true" box="[199,354,473,498]" pageId="2" pageNumber="489">Type species.</emphasis>
<taxonomicName id="4C304D3A8039FFDCFEB56CA3FB3532B7" authority="Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello" authorityName="Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas" authorityYear="2018" box="[361,1234,473,498]" class="Insecta" family="Gryllidae" genus="Titanogryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="salgado" status="sp. nov.">
<emphasis id="B944EAAB8039FFDCFEB56CA3FDB932B7" box="[361,606,473,498]" italics="true" pageId="2" pageNumber="489">Titanogryllus salgado</emphasis>
Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello
</taxonomicName>
<taxonomicNameLabel id="A27757D08039FFDCFB046CA3FAFF32B7" box="[1240,1304,473,498]" pageId="2" pageNumber="489" rank="species">
<emphasis id="B944EAAB8039FFDCFB046CA3FAFF32B7" bold="true" box="[1240,1304,473,498]" pageId="2" pageNumber="489">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8B8F36B98039FFDCFF1B6C87FAFE317C" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6C87FE753153" bold="true" box="[199,402,509,534]" pageId="2" pageNumber="489">Species included</emphasis>
.
<taxonomicName id="4C304D3A8039FFDCFE7B6C87FAA63153" authority="Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello" authorityName="Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas" authorityYear="2018" box="[423,1345,509,534]" class="Insecta" family="Gryllidae" genus="Titanogryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="salgado" status="sp. nov.">
<emphasis id="B944EAAB8039FFDCFE7B6C87FD433153" box="[423,676,509,534]" italics="true" pageId="2" pageNumber="489">Titanogryllus salgado</emphasis>
Jaiswara, Souza-Dias, Desutter-Grandcolas &amp; de Mello
</taxonomicName>
<taxonomicNameLabel id="A27757D08039FFDCFA936C87FA733150" box="[1359,1428,509,534]" pageId="2" pageNumber="489" rank="species">
<emphasis id="B944EAAB8039FFDCFA936C87FA6A3153" bold="true" box="[1359,1421,509,534]" pageId="2" pageNumber="489">n. sp</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C304D3A8039FFDCFF4B6F5BFD6E317F" authority="Souza-Dias &amp; de Mello" authorityName="Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas" authorityYear="2018" box="[151,649,545,570]" class="Insecta" family="Gryllidae" genus="Titanogryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="oxossi" status="sp. nov.">
<emphasis id="B944EAAB8039FFDCFF4B6F5BFE9E317C" box="[151,377,545,570]" italics="true" pageId="2" pageNumber="489">Titanogryllus oxossi</emphasis>
Souza-Dias &amp; de Mello
</taxonomicName>
<taxonomicNameLabel id="A27757D08039FFDCFD4C6F5BFD29317C" box="[656,718,545,570]" pageId="2" pageNumber="489" rank="species">
<emphasis id="B944EAAB8039FFDCFD4C6F5BFD2E317F" bold="true" box="[656,713,545,570]" pageId="2" pageNumber="489">n. sp</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="4C304D3A8039FFDCFD016F5BFB34317F" authority="Souza-Dias &amp; de Mello" authorityName="Jaiswara &amp; Souza-Dias &amp; Campos &amp; Redü &amp; De Mello &amp; Desutter-Grandcolas" authorityYear="2018" box="[733,1235,545,570]" class="Insecta" family="Gryllidae" genus="Titanogryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="489" phylum="Arthropoda" rank="species" species="oxente" status="sp. nov.">
<emphasis id="B944EAAB8039FFDCFD016F5BFC25317C" box="[733,962,545,570]" italics="true" pageId="2" pageNumber="489">Titanogryllus oxente</emphasis>
Souza-Dias &amp; de Mello
</taxonomicName>
<taxonomicNameLabel id="A27757D08039FFDCFB066F5BFAFE317C" box="[1242,1305,545,570]" pageId="2" pageNumber="489" rank="species">
<emphasis id="B944EAAB8039FFDCFB066F5BFAF6317F" bold="true" box="[1242,1297,545,570]" pageId="2" pageNumber="489">n. sp</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C32A65328039FFDCFF1B6F3FFBB5311B" box="[199,1106,581,606]" pageId="2" pageNumber="489" type="distribution">
<paragraph id="8B8F36B98039FFDCFF1B6F3FFBB5311B" blockId="2.[151,1437,293,1941]" box="[199,1106,581,606]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6F3FFE87311B" bold="true" box="[199,352,581,606]" pageId="2" pageNumber="489">Distribution.</emphasis>
Brazilian Atlantic Forest, in the states of
<collectingRegion id="49F4F85B8039FFDCFCEF6F3FFC34311B" box="[819,979,581,606]" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="489">Espírito Santo</collectingRegion>
and
<collectingRegion id="49F4F85B8039FFDCFBD56F3FFBAD311B" box="[1033,1098,581,606]" country="Brazil" name="Bahia" pageId="2" pageNumber="489">Bahia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A65328039FFDCFF1B6F13FED63083" pageId="2" pageNumber="489" type="diagnosis">
<paragraph id="8B8F36B98039FFDCFF1B6F13FED63083" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6F13FEA631C7" bold="true" box="[199,321,617,642]" pageId="2" pageNumber="489">Diagnosis.</emphasis>
Species large, robust, more than
<quantity id="4CC89B5C8039FFDCFD1D6F10FD1631C7" box="[705,753,618,642]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="489" unit="cm" value="3.0">3cm</quantity>
of body length. Head wide, glabrous. Ocelli almost in line. Pronotum DD saddle-shaped, glabrous, wider than long. TIII subapical spurs 3/3; serrulation only anterior to subapical spurs.
<emphasis id="B944EAAB8039FFDCFE856FCBFE72318F" bold="true" box="[345,405,689,714]" pageId="2" pageNumber="489">Male</emphasis>
. FWs reaching half of abdomen; anal vein area bulged dorsally; harp crossed with 46 almost transverse veins; mirror undivided, distinct from apical venation; apical field short, reduced; CuPb short; lateral field with 35 longitudinal veins.
<emphasis id="B944EAAB8039FFDCFDB56F83FCF23057" bold="true" box="[617,789,761,786]" pageId="2" pageNumber="489">Male genitalia</emphasis>
. Pseudepiphallic sclerite elongated; apex of lateral lophi bifurcated; median lophi short; rami elongated, bent towards ventral side, almost connected; ectophallic arc dorsal, thin, almost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold elongated, strongly sclerotized; dorsal cavity short.
<emphasis id="B944EAAB8039FFDCFEA26E1FFE32303B" bold="true" box="[382,469,869,894]" pageId="2" pageNumber="489">Female</emphasis>
. Larger than males. FWs small, very reduced. Ovipositor short; apex of ovipositor modified, with conspicuous ventral concavity; apex pointed and curved downwards in lateral view, with ventral protuberance.
</paragraph>
</subSubSection>
<subSubSection id="C32A65328039FFDCFF1B6EABFCD934D1" pageId="2" pageNumber="489" type="description">
<paragraph id="8B8F36B98039FFDCFF1B6EABFED43610" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6EABFEBE30AF" bold="true" box="[199,345,977,1002]" pageId="2" pageNumber="489">Description.</emphasis>
Species large, robust, more than
<quantity id="4CC89B5C8039FFDCFD126EA8FD1930AF" box="[718,766,978,1002]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="489" unit="cm" value="3.0">3cm</quantity>
of body length (
<figureCitation id="130B2A3C8039FFDCFC656EABFBDA30AF" box="[953,1085,977,1002]" captionStart="FIGURE 1" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[189,1386,224,1901]" captionTargetId="figure@3.[168,1404,193,1929]" captionTargetPageId="3" captionText="FIGURE 1. Titanogryllus n. gen. Male habitus of (A) T. salgado n. sp.; (B) T. oxossi n. sp.; (C) T. oxente n. sp. Scale bar: 10mm" httpUri="https://zenodo.org/record/1209710/files/figure.png" pageId="2" pageNumber="489">Figs. 1AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB956EA8FB8930AF" box="[1097,1134,978,1002]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">2A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFBA66EA8FB7930AF" box="[1146,1182,978,1002]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">3A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB766EA8FB2830AF" box="[1194,1231,978,1002]" captionStart="FIGURE 4" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[197,1351,531,1923]" captionTargetId="figure@7.[197,1360,519,1935]" captionTargetPageId="7" captionText="FIGURE 4. Titanogryllus salgado n. gen. n. sp. General morphology of the male paratype (MNHN). (A) habitus, dorsal; (B) habitus, lateral; (C) head, frontal;(D) FW, dorsal. Scale bar: 10mm." httpUri="https://zenodo.org/record/1209716/files/figure.png" pageId="2" pageNumber="489">4A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB076EA8FAC730AC" box="[1243,1312,978,1002]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7A, B</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFAF06EA8FAB730AF" box="[1324,1360,978,1002]" captionStart="FIGURE 8" captionStartId="11.[151,250,1804,1826]" captionTargetBox="[190,1391,221,1771]" captionTargetId="figure@11.[190,1397,193,1783]" captionTargetPageId="11" captionText="FIGURE 8. Titanogryllus oxossi n. gen. n. sp. General morphology of the allotype (MZSP). (A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal; (F) ovipositor, ventral; (G) ovipositor, lateral. Scale bar:A, 10mm; BG, 1mm" httpUri="https://zenodo.org/record/1209724/files/figure.png" pageId="2" pageNumber="489">8A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFA806EA8FA7730AF" box="[1372,1424,978,1002]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11A</figureCitation>
). Head large, glabrous, wider than long, as wide as pronotum (
<figureCitation id="130B2A3C8039FFDCFC806E8FFC04374B" box="[860,995,1013,1038]" captionStart="FIGURE 1" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[189,1386,224,1901]" captionTargetId="figure@3.[168,1404,193,1929]" captionTargetPageId="3" captionText="FIGURE 1. Titanogryllus n. gen. Male habitus of (A) T. salgado n. sp.; (B) T. oxossi n. sp.; (C) T. oxente n. sp. Scale bar: 10mm" httpUri="https://zenodo.org/record/1209710/files/figure.png" pageId="2" pageNumber="489">Figs. 1AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC2E6E8CFBDB3748" box="[1010,1084,1014,1038]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">2A, D</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB966E8CFB89374B" box="[1098,1134,1014,1038]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">3A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFBA16E8CFB223748" box="[1149,1221,1014,1038]" captionStart="FIGURE 4" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[197,1351,531,1923]" captionTargetId="figure@7.[197,1360,519,1935]" captionTargetPageId="7" captionText="FIGURE 4. Titanogryllus salgado n. gen. n. sp. General morphology of the male paratype (MNHN). (A) habitus, dorsal; (B) habitus, lateral; (C) head, frontal;(D) FW, dorsal. Scale bar: 10mm." httpUri="https://zenodo.org/record/1209716/files/figure.png" pageId="2" pageNumber="489">4A, B</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB086E8CFB1E374B" box="[1236,1273,1014,1038]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFADA6E8CFACC374B" box="[1286,1323,1014,1038]" captionStart="FIGURE 8" captionStartId="11.[151,250,1804,1826]" captionTargetBox="[190,1391,221,1771]" captionTargetId="figure@11.[190,1397,193,1783]" captionTargetPageId="11" captionText="FIGURE 8. Titanogryllus oxossi n. gen. n. sp. General morphology of the allotype (MZSP). (A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal; (F) ovipositor, ventral; (G) ovipositor, lateral. Scale bar:A, 10mm; BG, 1mm" httpUri="https://zenodo.org/record/1209724/files/figure.png" pageId="2" pageNumber="489">8A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFAE56E8CFA69374B" box="[1337,1422,1014,1038]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11A, B</figureCitation>
). Occiput, vertex and fastigium glabrous (
<figureCitation id="130B2A3C8039FFDCFD846963FD103774" box="[600,759,1049,1074]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">Figs. 2B, D, F</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFCDE6960FCC13777" box="[770,806,1050,1074]" captionStart="FIGURE 4" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[197,1351,531,1923]" captionTargetId="figure@7.[197,1360,519,1935]" captionTargetPageId="7" captionText="FIGURE 4. Titanogryllus salgado n. gen. n. sp. General morphology of the male paratype (MNHN). (A) habitus, dorsal; (B) habitus, lateral; (C) head, frontal;(D) FW, dorsal. Scale bar: 10mm." httpUri="https://zenodo.org/record/1209716/files/figure.png" pageId="2" pageNumber="489">4C</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFCEC6960FC6D3774" box="[816,906,1050,1074]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7B, C, I</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC4A6960FC0D3777" box="[918,1002,1050,1074]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11BD</figureCitation>
). Epistomal suture angular. Vertex flat. Eyes small, not protruding. Ocelli arranged almost in straight line; median ocellus very slightly ahead of lateral ocelli, distance between lateral ocelli twice the distance between one lateral and median ocellus. Fastigium twice wider than scape (
<figureCitation id="130B2A3C8039FFDCFEB969FFFE2E37DB" box="[357,457,1157,1182]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">Figs. 2D</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFE0A69FCFE1D37DB" box="[470,506,1158,1182]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">3A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFDDB69FCFDCD37D8" box="[519,554,1158,1181]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7B</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFDEA69FCFD8037DB" box="[566,615,1158,1182]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11B</figureCitation>
). Lower margin of eyes and antennal pit almost at same level. Maxillary palpi short, 3rd and 4th joints subequal, 5th joint longest, regularly widened on its length, apex rounded (
<figureCitation id="130B2A3C8039FFDCFACD69D6FA943781" box="[1297,1395,1196,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">Figs. 2B</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFAA269D7FA703781" box="[1406,1431,1197,1220]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">
<date id="FF8E10798039FFDCFAA269D7FA703781" box="[1406,1431,1197,1220]" pageId="2" pageNumber="489">7I</date>
</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFF4B69AAFF2037AD" box="[151,199,1232,1256]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11C</figureCitation>
). Pronotum DD wider than long, glabrous (
<figureCitation id="130B2A3C8039FFDCFD6669AAFCA737AD" box="[698,832,1232,1257]" captionStart="FIGURE 1" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[189,1386,224,1901]" captionTargetId="figure@3.[168,1404,193,1929]" captionTargetPageId="3" captionText="FIGURE 1. Titanogryllus n. gen. Male habitus of (A) T. salgado n. sp.; (B) T. oxossi n. sp.; (C) T. oxente n. sp. Scale bar: 10mm" httpUri="https://zenodo.org/record/1209710/files/figure.png" pageId="2" pageNumber="489">Figs. 1AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC9269AAFC7037AD" box="[846,919,1232,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">2A, D</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC7869AAFC2F37AD" box="[932,968,1232,1256]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">3A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC0A69AAFC1D37AD" box="[982,1018,1232,1256]" captionStart="FIGURE 4" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[197,1351,531,1923]" captionTargetId="figure@7.[197,1360,519,1935]" captionTargetPageId="7" captionText="FIGURE 4. Titanogryllus salgado n. gen. n. sp. General morphology of the male paratype (MNHN). (A) habitus, dorsal; (B) habitus, lateral; (C) head, frontal;(D) FW, dorsal. Scale bar: 10mm." httpUri="https://zenodo.org/record/1209716/files/figure.png" pageId="2" pageNumber="489">4A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFBD469ABFBA837AD" box="[1032,1103,1233,1257]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7A, B</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB8069AAFB6737AD" box="[1116,1152,1232,1256]" captionStart="FIGURE 8" captionStartId="11.[151,250,1804,1826]" captionTargetBox="[190,1391,221,1771]" captionTargetId="figure@11.[190,1397,193,1783]" captionTargetPageId="11" captionText="FIGURE 8. Titanogryllus oxossi n. gen. n. sp. General morphology of the allotype (MZSP). (A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal; (F) ovipositor, ventral; (G) ovipositor, lateral. Scale bar:A, 10mm; BG, 1mm" httpUri="https://zenodo.org/record/1209724/files/figure.png" pageId="2" pageNumber="489">8A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB5269AAFB0537AD" box="[1166,1250,1232,1256]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11A, B</figureCitation>
). Tympanum on both sides of TI. TI with 2 ventral spurs and 1 dorsal (inner). TII with 4 apical spurs. Legs III not specially elongated; TIII with spines before subapical spurs; subapical spurs 3/3 (
<figureCitation id="130B2A3C8039FFDCFC3E6862FB843675" box="[994,1123,1304,1329]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">
Figs.
<date id="FF8E10798039FFDCFBF96862FBDA3675" box="[1061,1085,1304,1328]" pageId="2" pageNumber="489">2I</date>
K
</figureCitation>
). Double row of spines on basitarsus-III.
</paragraph>
<paragraph id="8B8F36B98039FFDCFF1B681AFBFF354D" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B681AFEEC363C" bold="true" box="[199,267,1376,1401]" pageId="2" pageNumber="489">Male.</emphasis>
Metanotum without metanotal structures. FWs covering half of abdomen; stridulatory file present; harp crossed by 4-6 almost transverse veins; mirror undivided, wider than long, clearly distinct from surrounding veins; chords 1 and 2 elongated, close to each other, well-separated from chord 3; apical field reduced; lateral field with 35 longitudinal, parallel veins (
<figureCitation id="130B2A3C8039FFDCFDCC68B6FD7F36A1" box="[528,664,1484,1509]" captionStart="FIGURE 1" captionStartId="3.[151,250,1954,1976]" captionTargetBox="[189,1386,224,1901]" captionTargetId="figure@3.[168,1404,193,1929]" captionTargetPageId="3" captionText="FIGURE 1. Titanogryllus n. gen. Male habitus of (A) T. salgado n. sp.; (B) T. oxossi n. sp.; (C) T. oxente n. sp. Scale bar: 10mm" httpUri="https://zenodo.org/record/1209710/files/figure.png" pageId="2" pageNumber="489">Figs. 1AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFD7B68B6FCF536A1" box="[679,786,1484,1508]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">2A, C, E</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFCFD68B6FC8B36A1" box="[801,876,1484,1508]" captionStart="FIGURE 4" captionStartId="7.[151,250,1966,1988]" captionTargetBox="[197,1351,531,1923]" captionTargetId="figure@7.[197,1360,519,1935]" captionTargetPageId="7" captionText="FIGURE 4. Titanogryllus salgado n. gen. n. sp. General morphology of the male paratype (MNHN). (A) habitus, dorsal; (B) habitus, lateral; (C) head, frontal;(D) FW, dorsal. Scale bar: 10mm." httpUri="https://zenodo.org/record/1209716/files/figure.png" pageId="2" pageNumber="489">4A, D</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFCA768B6FC0636A1" box="[891,993,1484,1508]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7A, DF</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC3368B6FBA336A1" box="[1007,1092,1484,1508]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11A, E</figureCitation>
). Supra anal plate constricted medially; subgenital plate triangular, apex rounded (
<figureCitation id="130B2A3C8039FFDCFD3C688AFC84354D" box="[736,867,1520,1545]" captionStart="FIGURE 2" captionStartId="5.[151,250,1833,1855]" captionTargetBox="[176,1386,386,1748]" captionTargetId="figure@5.[176,1386,373,1776]" captionTargetPageId="5" captionText="FIGURE 2. Titanogryllus salgado n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, lateral; (C) FW, lateral; (D) head and pronotum, dorsal; (E) FW, dorsal; (F) head, frontal; (G) supra anal plate; (H) subgenital plate;(I) TIIIand basitarsusIII;(J) outer apical spurs of TIII; (K) innerapical spurs of TIII.Scale bar:A, 10mm; BK, 1mm." httpUri="https://zenodo.org/record/1209712/files/figure.png" pageId="2" pageNumber="489">Figs. 2G, H</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFCB2688AFC55354D" box="[878,946,1520,1544]" captionStart="FIGURE7" captionStartId="10.[151,250,1802,1824]" captionTargetBox="[188,1390,210,1765]" captionTargetId="figure@10.[188,1398,193,1780]" captionTargetPageId="10" captionText="FIGURE7. Titanogryllus oxossi n. gen. n. sp. General morphology of the holotype (MZSP).(A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D &amp; E) FW, dorsal; (F) FW, lateral; (G) supra anal plate; (H) subgenital plate; (I) head, frontal. Scale bar:A, 10mm; BI, 1mm." httpUri="https://zenodo.org/record/1209722/files/figure.png" pageId="2" pageNumber="489">7G, H</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC61688AFBEC354D" box="[957,1035,1520,1544]" captionStart="FIGURE 11" captionStartId="15.[151,250,1946,1968]" captionTargetBox="[188,1398,378,1921]" captionTargetId="figure@15.[188,1398,355,1943]" captionTargetPageId="15" captionText="FIGURE 11. Titanogryllus oxente n. gen. n. sp. General morphology of the holotype (MZSP). (A) habitus, dorsal; (B) head and pronotum, dorsal; (C) head and pronotum, lateral; (D) head, frontal; (E) FW, dorsal; (F) supra anal plate; (G) subgenital plate. Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209730/files/figure.png" pageId="2" pageNumber="489">11F, G</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B98039FFDCFF1B6B6EFF5D3441" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6B6EFE9D3568" bold="true" box="[199,378,1556,1581]" pageId="2" pageNumber="489">Male genitalia.</emphasis>
Male genitalia elongated. Pseudepiphallus: lateral lophi wide, apex bifurcated; medial lophi short; rami elongated, bent towards ventral side, ramus almost connected anteriorly (
<figureCitation id="130B2A3C8039FFDCFBB16B42FB123514" box="[1133,1269,1592,1617]" captionStart="FIGURE 5" captionStartId="8.[151,250,1709,1731]" captionTargetBox="[204,1379,217,1668]" captionTargetId="figure@8.[188,1398,193,1687]" captionTargetPageId="8" captionText="FIGURE 5. Titanogryllus salgado n. gen. n. sp. Male holotype and female allotype (MZSP) genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views.Female genitalia in (D) ventral, (E) dorsal and (F) lateral view.Scale bar:1mm." httpUri="https://zenodo.org/record/1209718/files/figure.png" pageId="2" pageNumber="489">Figs. 5AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFAD86B42FAAE3514" box="[1284,1353,1592,1617]" captionStart="FIGURE 6" captionStartId="9.[151,250,1608,1630]" captionTargetBox="[213,1364,539,1543]" captionTargetId="figure@9.[188,1398,519,1586]" captionTargetPageId="9" captionText="FIGURE 6. Titanogryllus salgado n. gen. n. sp. Male genitalia of the paratype (MNHN). Male genitalia in (A) dorsal, (B) ventral and (C) lateral views." httpUri="https://zenodo.org/record/1209720/files/figure.png" pageId="2" pageNumber="489">6AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFA846B42FA7F3515" box="[1368,1432,1592,1616]" captionStart="FIGURE 9" captionStartId="12.[151,250,1848,1870]" captionTargetBox="[169,1428,204,1816]" captionTargetId="figure@12.[151,1436,193,1826]" captionTargetPageId="12" captionText="FIGURE 9. Titanogryllus oxossi n. gen. n. sp. Male genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views. Apex and median part of pseudepiphallus in (D) dorsal, (E) ventral and (F) lateral views. Scale bar:1mm." httpUri="https://zenodo.org/record/1209726/files/figure.png" pageId="2" pageNumber="489">9AF</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFF4B6B26FF0B3531" box="[151,236,1628,1652]" captionStart="FIGURE12" captionStartId="16.[151,250,1362,1384]" captionTargetBox="[203,1369,225,1296]" captionTargetId="figure@16.[188,1398,193,1319]" captionTargetPageId="16" captionText="FIGURE12.Titanogryllus oxente n. gen.n.sp. Male genitalia in(A) dorsal,(B) ventral and (C)lateral views.Scale bar:1mm." httpUri="https://zenodo.org/record/1209732/files/figure.png" pageId="2" pageNumber="489">12AC</figureCitation>
). Pseudepiphallic apodemes absent. Pseudepiphallic parameres well sclerotized. Ectophallic invagination: ectophallic apodemes elongated; ectophallic arc dorsal, thin, almost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold elongated, strongly sclerotized (
<figureCitation id="130B2A3C8039FFDCFCBD6BDEFC0F35F8" box="[865,1000,1700,1725]" captionStart="FIGURE 5" captionStartId="8.[151,250,1709,1731]" captionTargetBox="[204,1379,217,1668]" captionTargetId="figure@8.[188,1398,193,1687]" captionTargetPageId="8" captionText="FIGURE 5. Titanogryllus salgado n. gen. n. sp. Male holotype and female allotype (MZSP) genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views.Female genitalia in (D) ventral, (E) dorsal and (F) lateral view.Scale bar:1mm." httpUri="https://zenodo.org/record/1209718/files/figure.png" pageId="2" pageNumber="489">Figs. 5AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFC2B6BDEFBDB35F8" box="[1015,1084,1700,1725]" captionStart="FIGURE 6" captionStartId="9.[151,250,1608,1630]" captionTargetBox="[213,1364,539,1543]" captionTargetId="figure@9.[188,1398,519,1586]" captionTargetPageId="9" captionText="FIGURE 6. Titanogryllus salgado n. gen. n. sp. Male genitalia of the paratype (MNHN). Male genitalia in (A) dorsal, (B) ventral and (C) lateral views." httpUri="https://zenodo.org/record/1209720/files/figure.png" pageId="2" pageNumber="489">6AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB966BDEFB7735F9" box="[1098,1168,1700,1724]" captionStart="FIGURE 9" captionStartId="12.[151,250,1848,1870]" captionTargetBox="[169,1428,204,1816]" captionTargetId="figure@12.[151,1436,193,1826]" captionTargetPageId="12" captionText="FIGURE 9. Titanogryllus oxossi n. gen. n. sp. Male genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views. Apex and median part of pseudepiphallus in (D) dorsal, (E) ventral and (F) lateral views. Scale bar:1mm." httpUri="https://zenodo.org/record/1209726/files/figure.png" pageId="2" pageNumber="489">9AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB426BDEFB1435F9" box="[1182,1267,1700,1724]" captionStart="FIGURE12" captionStartId="16.[151,250,1362,1384]" captionTargetBox="[203,1369,225,1296]" captionTargetId="figure@16.[188,1398,193,1319]" captionTargetPageId="16" captionText="FIGURE12.Titanogryllus oxente n. gen.n.sp. Male genitalia in(A) dorsal,(B) ventral and (C)lateral views.Scale bar:1mm." httpUri="https://zenodo.org/record/1209732/files/figure.png" pageId="2" pageNumber="489">12AC</figureCitation>
). Endophallus: endophallic sclerite small, related to ectophallic fold; endophallic cavity reduced (
<figureCitation id="130B2A3C8039FFDCFBF06BB2FB5635A4" box="[1068,1201,1736,1761]" captionStart="FIGURE 5" captionStartId="8.[151,250,1709,1731]" captionTargetBox="[204,1379,217,1668]" captionTargetId="figure@8.[188,1398,193,1687]" captionTargetPageId="8" captionText="FIGURE 5. Titanogryllus salgado n. gen. n. sp. Male holotype and female allotype (MZSP) genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views.Female genitalia in (D) ventral, (E) dorsal and (F) lateral view.Scale bar:1mm." httpUri="https://zenodo.org/record/1209718/files/figure.png" pageId="2" pageNumber="489">Figs. 5AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB606BB2FAE535A4" box="[1212,1282,1736,1761]" captionStart="FIGURE 6" captionStartId="9.[151,250,1608,1630]" captionTargetBox="[213,1364,539,1543]" captionTargetId="figure@9.[188,1398,519,1586]" captionTargetPageId="9" captionText="FIGURE 6. Titanogryllus salgado n. gen. n. sp. Male genitalia of the paratype (MNHN). Male genitalia in (A) dorsal, (B) ventral and (C) lateral views." httpUri="https://zenodo.org/record/1209720/files/figure.png" pageId="2" pageNumber="489">6AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFAD16BB2FAB435A5" box="[1293,1363,1736,1760]" captionStart="FIGURE 9" captionStartId="12.[151,250,1848,1870]" captionTargetBox="[169,1428,204,1816]" captionTargetId="figure@12.[151,1436,193,1826]" captionTargetPageId="12" captionText="FIGURE 9. Titanogryllus oxossi n. gen. n. sp. Male genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views. Apex and median part of pseudepiphallus in (D) dorsal, (E) ventral and (F) lateral views. Scale bar:1mm." httpUri="https://zenodo.org/record/1209726/files/figure.png" pageId="2" pageNumber="489">9AC</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFA826BB2FF4A3441" captionStart="FIGURE12" captionStartId="16.[151,250,1362,1384]" captionTargetBox="[203,1369,225,1296]" captionTargetId="figure@16.[188,1398,193,1319]" captionTargetPageId="16" captionText="FIGURE12.Titanogryllus oxente n. gen.n.sp. Male genitalia in(A) dorsal,(B) ventral and (C)lateral views.Scale bar:1mm." httpUri="https://zenodo.org/record/1209732/files/figure.png" pageId="2" pageNumber="489">12A C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B98039FFDCFF1B6A6AFCD934D1" blockId="2.[151,1437,293,1941]" pageId="2" pageNumber="489">
<emphasis id="B944EAAB8039FFDCFF1B6A6AFEF9346C" bold="true" box="[199,286,1808,1833]" pageId="2" pageNumber="489">Female</emphasis>
. Larger than males (
<figureCitation id="130B2A3C8039FFDCFDD56A6AFD89346D" box="[521,622,1808,1833]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">Figs. 3A</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFDA76A6AFD46346D" box="[635,673,1808,1832]" captionStart="FIGURE 8" captionStartId="11.[151,250,1804,1826]" captionTargetBox="[190,1391,221,1771]" captionTargetId="figure@11.[190,1397,193,1783]" captionTargetPageId="11" captionText="FIGURE 8. Titanogryllus oxossi n. gen. n. sp. General morphology of the allotype (MZSP). (A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal; (F) ovipositor, ventral; (G) ovipositor, lateral. Scale bar:A, 10mm; BG, 1mm" httpUri="https://zenodo.org/record/1209724/files/figure.png" pageId="2" pageNumber="489">8A</figureCitation>
). FW small, very reduced (
<figureCitation id="130B2A3C8039FFDCFC0B6A6AFBDD346D" box="[983,1082,1808,1833]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">Figs. 3B</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFB9B6A6AFB8D346D" box="[1095,1130,1808,1832]" captionStart="FIGURE 8" captionStartId="11.[151,250,1804,1826]" captionTargetBox="[190,1391,221,1771]" captionTargetId="figure@11.[190,1397,193,1783]" captionTargetPageId="11" captionText="FIGURE 8. Titanogryllus oxossi n. gen. n. sp. General morphology of the allotype (MZSP). (A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal; (F) ovipositor, ventral; (G) ovipositor, lateral. Scale bar:A, 10mm; BG, 1mm" httpUri="https://zenodo.org/record/1209724/files/figure.png" pageId="2" pageNumber="489">8B</figureCitation>
). Ovipositor short; apex of ovipositor modified, with conspicuous concavity; apex pointed and curved in lateral view, with ventral protuberance (
<figureCitation id="130B2A3C8039FFDCFEE16A22FE0D3435" box="[317,490,1880,1905]" captionStart="FIGURE 3" captionStartId="6.[151,250,1745,1767]" captionTargetBox="[210,1368,207,1687]" captionTargetId="figure@6.[197,1368,193,1709]" captionTargetPageId="6" captionText="FIGURE 3. Titanogryllus salgado n. gen. n. sp. General morphology of the allotype.(A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal;(F)ovipositor,ventral; (G) ovipositor, lateral.Scale bar:A, 10mm; BG, 1mm." httpUri="https://zenodo.org/record/1209714/files/figure.png" pageId="2" pageNumber="489">Figs. 3A, EG</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFE266A22FD813435" box="[506,614,1880,1904]" captionStart="FIGURE 8" captionStartId="11.[151,250,1804,1826]" captionTargetBox="[190,1391,221,1771]" captionTargetId="figure@11.[190,1397,193,1783]" captionTargetPageId="11" captionText="FIGURE 8. Titanogryllus oxossi n. gen. n. sp. General morphology of the allotype (MZSP). (A) habitus, dorsal; (B) FW, lateral; (C) supra anal plate; (D) subgenital plate; (E) ovipositor, dorsal; (F) ovipositor, ventral; (G) ovipositor, lateral. Scale bar:A, 10mm; BG, 1mm" httpUri="https://zenodo.org/record/1209724/files/figure.png" pageId="2" pageNumber="489">8A, EG</figureCitation>
).
<emphasis id="B944EAAB8039FFDCFDA26A22FCAF3434" bold="true" box="[638,840,1880,1905]" pageId="2" pageNumber="489">Female genitalia</emphasis>
. Copulatory papilla small, rounded; base medially extended ventrally; apex membranous (
<figureCitation id="130B2A3C8039FFDCFD8D6A06FD3734D0" box="[593,720,1916,1941]" captionStart="FIGURE 5" captionStartId="8.[151,250,1709,1731]" captionTargetBox="[204,1379,217,1668]" captionTargetId="figure@8.[188,1398,193,1687]" captionTargetPageId="8" captionText="FIGURE 5. Titanogryllus salgado n. gen. n. sp. Male holotype and female allotype (MZSP) genitalia. Male genitalia in (A) dorsal, (B) ventral and (C) lateral views.Female genitalia in (D) ventral, (E) dorsal and (F) lateral view.Scale bar:1mm." httpUri="https://zenodo.org/record/1209718/files/figure.png" pageId="2" pageNumber="489">Figs. 5DF</figureCitation>
,
<figureCitation id="130B2A3C8039FFDCFD006A06FCD634D1" box="[732,817,1916,1940]" captionStart="FIGURE 10" captionStartId="13.[151,250,640,662]" captionTargetBox="[203,1385,208,574]" captionTargetId="figure@13.[188,1398,193,619]" captionTargetPageId="13" captionText="FIGURE 10. Titanogryllus oxossi n. gen. n. sp. Female genitalia. Copulatory papilla in (A) ventral, (B) dorsal and(C) lateral views." httpUri="https://zenodo.org/record/1209728/files/figure.png" pageId="2" pageNumber="489">10AC</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>