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<document id="5D2F9135BB6B062ADB4127B2307FDEBD" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13886FF5B2B3BFDA8FBCDFEE5" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium lisae Darby 2015" docType="treatment" docVersion="7" lastPageNumber="150" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="149" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="39A7EF6447B12C3B5E5CB0EE25FAD3EF">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<mods:namePart id="A73FC41F1629AD0D1839C210EDCB17A9">Darby, Michael</mods:namePart>
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<mods:title id="B1E0D20E79A7E685C88B0797287FB4AB">European Journal of Taxonomy</mods:title>
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<mods:date id="9A7330C3B63FF8B8BCCD2161F9A352BC">2020</mods:date>
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<mods:number id="A10043CDEC95BF73D3361A09BE59695E">2020-04-02</mods:number>
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<mods:classification id="8E7C58EC9BEBC5F2E176E6462C9E1B1D">journal article</mods:classification>
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<mods:identifier id="F78C63584EB7E7649B274F6057371D7D" type="DOI">10.5852/ejt.2020.622</mods:identifier>
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<treatment id="039987A13886FF5B2B3BFDA8FBCDFEE5" ID-DOI="http://doi.org/10.5281/zenodo.3795678" ID-GBIF-Taxon="163877804" ID-Zenodo-Dep="3795678" LSID="urn:lsid:plazi:treatment:039987A13886FF5B2B3BFDA8FBCDFEE5" httpUri="http://treatment.plazi.org/id/039987A13886FF5B2B3BFDA8FBCDFEE5" lastPageId="149" lastPageNumber="150" pageId="148" pageNumber="149">
<subSubSection id="C32A653C3886FF5A2B3BFDA8FC5EFDF9" box="[626,962,582,609]" pageId="148" pageNumber="149" type="nomenclature">
<paragraph id="8B8F36B73886FF5A2B3BFDA8FC5EFDF9" blockId="148.[626,962,582,647]" box="[626,962,582,609]" pageId="148" pageNumber="149">
<heading id="D0C781DB3886FF5A2B3BFDA8FC5EFDF9" box="[626,962,582,609]" centered="true" fontSize="11" level="2" pageId="148" pageNumber="149" reason="2">
<taxonomicName id="4C304D343886FF5A2B3BFDA8FC5EFDF9" ID-CoL="VK8F" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[626,962,582,609]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="148" pageNumber="149" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B944EAA53886FF5A2B3BFDA8FCB5FDF9" bold="true" box="[626,809,582,609]" italics="true" pageId="148" pageNumber="149">Cissidium lisae</emphasis>
<bibRefCitation id="EFA14B463886FF5A2A78FDA8FC5EFDF9" author="Darby M." box="[817,962,582,609]" pageId="148" pageNumber="149" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3886FF5A2B8AFD82FCECFD1E" box="[707,880,620,647]" pageId="148" pageNumber="149" type="description">
<paragraph id="8B8F36B73886FF5A2B8AFD82FCECFD1E" blockId="148.[626,962,582,647]" box="[707,880,620,647]" pageId="148" pageNumber="149">
<figureCitation id="130B2A323886FF5A2B8AFD82FC80FD1E" box="[707,796,620,646]" captionStart="Fig" captionStartId="4.[189,232,1717,1743]" captionTargetBox="[190,1397,393,1672]" captionTargetId="figure@4.[189,1398,393,1676]" captionTargetPageId="4" captionText="Fig. 2. A–B. Mesoventra of two species of Dacrysoma Grebennikov, 2009 to show form of mid-keel and perforation fringed by setae. C–E. Figures to explain use of the terms foveae, distinct foveolae and shallow foveolae in distinguishing the main species groups. C. Distinct foveolae, showing setae arising from the anterior margin of the depressions. D. Foveae and foveolae, showing setae randomly associated with foveae. E. Shallow foveolae, many setae rubbed off, but insertion points marked by punctures with slightly raised circular rims. F–I. Figures to show different forms of antennae. F. Elongate antenna of Cissidium lisae Darby, 2015. G. Globular antennomere of Cissidium globosum Darby, 2015. H. Female and male antennae of Cissidium petri Darby, 2015. I. Abnormal antennomere XI of Cissidium tigrum Darby, 2015." figureDoi="http://doi.org/10.5281/zenodo.3777286" httpUri="https://zenodo.org/record/3777286/files/figure.png" pageId="148" pageNumber="149">Figs 2F</figureCitation>
,
<figureCitation id="130B2A323886FF5A2A61FD82FCDBFD1F" box="[808,839,620,647]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[195,1398,757,1880]" captionTargetId="figure@6.[189,1398,756,1884]" captionTargetPageId="6" captionText="Fig. 5. Figures to show keel shapes. A. Cissidium bifoveolatum sp. nov., × 515. B. Cissidium lucidulum sp. nov., × 780. C. Cissidium scutellaris (Deane, 1931), × 1480. D. Cissidium tigrum Darby, 2015, × 840. E. Cissidium sawadai Darby, 2015, × 680. F. Cissidium lisae Darby, 2015, × 823." figureDoi="http://doi.org/10.5281/zenodo.3777292" httpUri="https://zenodo.org/record/3777292/files/figure.png" pageId="148" pageNumber="149">5F</figureCitation>
,
<figureCitation id="130B2A323886FF5A2A1AFD82FCECFD1E" box="[851,880,620,646]" captionStart="Fig" captionStartId="149.[189,232,1435,1461]" captionTargetBox="[212,1376,453,1383]" captionTargetId="figure@149.[208,1376,441,1389]" captionTargetPageId="149" captionText="Fig. 94. Cissidium lisae Darby, 2015. A. Habitus. B. Pronotum, C. Mesoventrite showing median process of collar, mid-keel and keel, × 550." figureDoi="http://doi.org/10.5281/zenodo.3777472" httpUri="https://zenodo.org/record/3777472/files/figure.png" pageId="148" pageNumber="149">94</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C3886FF5A29F4FD5EFCE5F873" pageId="148" pageNumber="149" type="materials_examined">
<paragraph id="8B8F36B73886FF5A29F4FD5EFE36FD52" blockId="148.[189,426,688,714]" box="[189,426,688,714]" pageId="148" pageNumber="149">
<heading id="D0C781DB3886FF5A29F4FD5EFE36FD52" bold="true" box="[189,426,688,714]" fontSize="11" level="3" pageId="148" pageNumber="149" reason="3">
<emphasis id="B944EAA53886FF5A29F4FD5EFE36FD52" bold="true" box="[189,426,688,714]" pageId="148" pageNumber="149">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA3886FF5A29A3FD0CFBBAFCD0" ID-GBIF-Occurrence="2609493601" collectedFrom="sifting litter" collectingDate="2013-11-28" collectionCode="UASC" collectorName="Dep., W &amp; Winkler app. extr. &amp; P. Banar" country="Bolivia" elevation="2450" latitude="-17.836668" location="Siberia Pass" longLatPrecision="21" longitude="-64.70222" municipality="Amboro" pageId="148" pageNumber="149" specimenCount="1" specimenCount-female="1" stateProvince="Santa Cruz" typeStatus="holotype">
<paragraph id="8B8F36B73886FF5A29A3FD0CFEC0FD64" blockId="148.[189,1399,738,840]" box="[234,348,738,764]" pageId="148" pageNumber="149">
<emphasis id="B944EAA53886FF5A29A3FD0CFEC0FD64" bold="true" box="[234,348,738,764]" pageId="148" pageNumber="149">
<typeStatus id="548B88153886FF5A29A3FD0CFEC0FD64" box="[234,348,738,764]" pageId="148" pageNumber="149" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FCE6FBBAFCD0" blockId="148.[189,1399,738,840]" pageId="148" pageNumber="149">
<collectingCountry id="F32776273886FF5A29F4FCE6FEA8FCBA" box="[189,308,776,802]" name="Bolivia" pageId="148" pageNumber="149">BOLIVIA</collectingCountry>
•
<specimenCount id="9D36FD3E3886FF5A2807FCE6FEF9FCBA" box="[334,357,776,802]" pageId="148" pageNumber="149" type="female">♀</specimenCount>
;
<collectingRegion id="49F4F8553886FF5A283DFCE6FE6AFCBA" box="[372,502,776,802]" country="Bolivia" name="Santa Cruz" pageId="148" pageNumber="149">Santa Cruz</collectingRegion>
<collectorName id="26C553613886FF5A28B7FCE6FDC3FCBA" box="[510,607,776,802]" pageId="148" pageNumber="149">Dep., W</collectorName>
<collectingMunicipality id="6BEBACCD3886FF5A2B2FFCE9FD54FCB9" box="[614,712,775,801]" pageId="148" pageNumber="149">Amboro</collectingMunicipality>
NP,
<location id="8EEF606C3886FF5A2A48FCE9FC0CFCBA" LSID="urn:lsid:plazi:treatment:039987A13886FF5B2B3BFDA8FBCDFEE5:8EEF606C3886FF5A2A48FCE9FC0CFCBA" box="[769,912,775,802]" country="Bolivia" latitude="-17.836668" longLatPrecision="21" longitude="-64.70222" municipality="Amboro" name="Siberia Pass" pageId="148" pageNumber="149" stateProvince="Santa Cruz">Siberia Pass</location>
env.;
<geoCoordinate id="EE0450703886FF5A2A9EFCE8FBFBFCBA" box="[983,1127,774,802]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="148" pageNumber="149" precision="15" seconds="12" value="-17.836668">17°50′12″ S</geoCoordinate>
,
<geoCoordinate id="EE0450703886FF5A2D3CFCE8FA8DFCBA" box="[1141,1297,774,802]" degrees="64" direction="west" minutes="42" orientation="longitude" pageId="148" pageNumber="149" precision="15" seconds="08" value="-64.70222">64°42′08″ W</geoCoordinate>
;
<elevation id="001DD1843886FF5A2C54FCE9FF6AFCDF" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="148" pageNumber="149" unit="m" value="2450.0">
<quantity id="4CC89B523886FF5A2C54FCE9FAEBFCBA" box="[1309,1399,775,802]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="148" pageNumber="149" unit="m" value="2450.0">2450 m</quantity>
a.s.l.
</elevation>
;
<date id="FF8E10773886FF5A284AFCC0FE39FCD0" box="[259,421,814,840]" pageId="148" pageNumber="149" value="2013-11-28">
<collectingDate id="EFCAE99F3886FF5A284AFCC0FE39FCD0" box="[259,421,814,840]" pageId="148" pageNumber="149" value="2013-11-28">28 Nov. 2013</collectingDate>
</date>
;
<collectedFrom id="382C754A3886FF5A28F8FCC3FDA2FCDF" box="[433,574,813,839]" pageId="148" pageNumber="149">sifting litter</collectedFrom>
;
<collectorName id="26C553613886FF5A2B03FCC3FCBCFCD0" box="[586,800,813,840]" pageId="148" pageNumber="149">Winkler app. extr.</collectorName>
;
<collectorName id="26C553613886FF5A2A65FCC0FC12FCD0" box="[812,910,814,840]" pageId="148" pageNumber="149">P. Baňař</collectorName>
leg.;
<collectionCode id="ED21AE723886FF5A2A99FCC0FBBEFCD0" box="[976,1058,814,840]" country="Bolivia" httpUri="http://grbio.org/cool/kme8-7yd8" name="Museo de Historia Natural &amp;quot;Noel Kempff Mercado&amp;quot;" pageId="148" pageNumber="149">UASC</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B8F36B73886FF5A29A3FC9CFEFBFC14" blockId="148.[189,1399,882,1022]" box="[234,359,882,908]" pageId="148" pageNumber="149">
<materialsCitation id="3B583CEA3886FF5A29A3FC9CFEFBFC14" ID-GBIF-Occurrence="2609493600" box="[234,359,882,908]" collectedFrom="sifting litter" collectingDate="2013-11-28" collectionCode="MMBC" collectorName="Dep., W &amp; Winkler &amp; P. Banar" country="Bolivia" elevation="2450" latitude="-17.836668" location="BOLIVIA" longLatPrecision="21" longitude="-64.70222" municipality="Amboro" pageId="148" pageNumber="149" specimenCount="1" specimenCount-female="1" stateProvince="Santa Cruz" typeStatus="paratype">
<emphasis id="B944EAA53886FF5A29A3FC9CFEFBFC14" bold="true" box="[234,359,882,908]" pageId="148" pageNumber="149">
<typeStatus id="548B88153886FF5A29A3FC9CFEFBFC14" box="[234,359,882,908]" pageId="148" pageNumber="149" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FC76FE5DFC66" blockId="148.[189,1399,882,1022]" pageId="148" pageNumber="149">
<collectingCountry id="F32776273886FF5A29F4FC76FEA8FC2A" box="[189,308,920,946]" name="Bolivia" pageId="148" pageNumber="149">BOLIVIA</collectingCountry>
•
<specimenCount id="9D36FD3E3886FF5A2806FC76FEE1FC2A" box="[335,381,920,946]" pageId="148" pageNumber="149" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED21AE723886FF5A2A03FC76FC39FC2A" box="[842,933,920,946]" collectionName="Czech Republic, Brno, Moravske Muzeum [Moravian Museum]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="148" pageNumber="149">MMBC</collectionCode>
•
<materialsCitation id="3B583CEA3886FF5A2A88FC76FE21FC66" ID-GBIF-Occurrence="2609493605" collectedFrom="sifting litter" collectionCode="BMNH" collectorName="Winkler app. extr. &amp; P. Banar" country="Bolivia" elevation="2481" latitude="-17.840279" location="Siberia Pass" longLatPrecision="21" longitude="-64.69945" pageId="148" pageNumber="149" specimenCount="1" specimenCount-female="1" stateProvince="Santa Cruz" typeStatus="paratype">
<specimenCount id="9D36FD3E3886FF5A2A88FC76FC73FC2A" box="[961,1007,920,946]" pageId="148" pageNumber="149" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49F4F8553886FF5A2AB6FC76FB1EFC2A" box="[1023,1154,920,946]" country="Bolivia" name="Santa Cruz" pageId="148" pageNumber="149">Santa Cruz</collectingRegion>
Dep.,
<locationDeviation id="9A3D2F953886FF5A2D9BFC76FAEBFC2A" box="[1234,1399,920,946]" location="Amboro" pageId="148" pageNumber="149">W of Amboro</locationDeviation>
NP,
<location id="8EEF606C3886FF5A29A6FC50FEE2FC40" LSID="urn:lsid:plazi:treatment:039987A13886FF5B2B3BFDA8FBCDFEE5:8EEF606C3886FF5A29A6FC50FEE2FC40" box="[239,382,958,984]" country="Bolivia" latitude="-17.840279" longLatPrecision="21" longitude="-64.69945" name="Siberia Pass" pageId="148" pageNumber="149" stateProvince="Santa Cruz">Siberia Pass</location>
env.,
<geoCoordinate id="EE0450703886FF5A288CFC52FDCAFC40" box="[453,598,956,984]" degrees="17" direction="south" minutes="50" orientation="latitude" pageId="148" pageNumber="149" precision="15" seconds="25" value="-17.840279">17°50′25″ S</geoCoordinate>
,
<geoCoordinate id="EE0450703886FF5A2B2DFC52FC9CFC40" box="[612,768,956,984]" degrees="64" direction="west" minutes="41" orientation="longitude" pageId="148" pageNumber="149" precision="15" seconds="58" value="-64.69945">64°41′58″ W</geoCoordinate>
;
<elevation id="001DD1843886FF5A2A45FC50FC3BFC40" box="[780,935,958,984]" metricMagnitude="3" metricUnit="m" metricValue="2.481" pageId="148" pageNumber="149" unit="m" value="2481.0">
<quantity id="4CC89B523886FF5A2A45FC50FCFAFC40" box="[780,870,958,984]" metricMagnitude="3" metricUnit="m" metricValue="2.481" pageId="148" pageNumber="149" unit="m" value="2481.0">2481 m</quantity>
a.s.l.
</elevation>
; 2013;
<collectedFrom id="382C754A3886FF5A2D49FC50FB12FC40" box="[1024,1166,958,984]" pageId="148" pageNumber="149">sifting litter</collectedFrom>
;
<collectorName id="26C553613886FF5A2DD3FC50FAEEFC40" box="[1178,1394,958,984]" pageId="148" pageNumber="149">Winkler app. extr.</collectorName>
;
<collectorName id="26C553613886FF5A29F4FC0AFE83FC66" box="[189,287,996,1022]" pageId="148" pageNumber="149">P. Baňař</collectorName>
leg.;
<collectionCode id="ED21AE723886FF5A2828FC0AFE21FC66" box="[353,445,996,1022]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="148" pageNumber="149">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FBC6FD8DFBDA" blockId="148.[189,529,1064,1090]" box="[189,529,1064,1090]" pageId="148" pageNumber="149">
<heading id="D0C781DB3886FF5A29F4FBC6FD8DFBDA" bold="true" box="[189,529,1064,1090]" fontSize="11" level="3" pageId="148" pageNumber="149" reason="3">
<emphasis id="B944EAA53886FF5A29F4FBC6FD8DFBDA" bold="true" box="[189,529,1064,1090]" pageId="148" pageNumber="149">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FBB7FB76FB02" blockId="148.[189,1399,1113,1178]" pageId="148" pageNumber="149">
The following description amplifies the
<typeStatus id="548B88153886FF5A2BDBFBB4FD59FBEC" box="[658,709,1114,1140]" pageId="148" pageNumber="149">type</typeStatus>
description of
<bibRefCitation id="EFA14B463886FF5A2A3DFBB7FB8AFBEC" author="Darby M." box="[884,1046,1113,1140]" pageId="148" pageNumber="149" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby (2015)</bibRefCitation>
, by adding more details of the meso- and metaventral characters, in order to conform with the entries in the present paper.
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FB2DFD36FB46" blockId="148.[189,682,1219,1246]" box="[189,682,1219,1246]" pageId="148" pageNumber="149">
SIZE. Habitus (
<figureCitation id="130B2A323886FF5A282EFB2DFE48FB45" box="[359,468,1219,1245]" captionStart="Fig" captionStartId="149.[189,232,1435,1461]" captionTargetBox="[212,1376,453,1383]" captionTargetId="figure@149.[208,1376,441,1389]" captionTargetPageId="149" captionText="Fig. 94. Cissidium lisae Darby, 2015. A. Habitus. B. Pronotum, C. Mesoventrite showing median process of collar, mid-keel and keel, × 550." figureDoi="http://doi.org/10.5281/zenodo.3777472" httpUri="https://zenodo.org/record/3777472/files/figure.png" pageId="148" pageNumber="149">Fig. 94A</figureCitation>
), length
<quantity id="4CC89B523886FF5A2B73FB2DFD39FB46" box="[570,677,1219,1246]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="148" pageNumber="149" unit="mm" value="0.8">0.80 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FAE6FCC8FAB9" blockId="148.[189,852,1287,1314]" box="[189,852,1287,1314]" pageId="148" pageNumber="149">COLOUR. Yellow brown, antennae and legs dusky yellow.</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FAA2FE80FA14" blockId="148.[189,1398,1356,1420]" pageId="148" pageNumber="149">
HEAD. Width across eyes
<quantity id="4CC89B523886FF5A28BFFAA2FDF9FAFE" box="[502,613,1356,1382]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="148" pageNumber="149" unit="mm" value="0.22">0.22 mm</quantity>
; antennomeres 3–11 length
<quantity id="4CC89B523886FF5A2AF5FAA2FBB5FAFE" box="[956,1065,1356,1383]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="148" pageNumber="149" unit="mm" value="0.45">0.45 mm</quantity>
long (
<figureCitation id="130B2A323886FF5A2D35FAA2FB45FAFE" box="[1148,1241,1356,1382]" captionStart="Fig" captionStartId="4.[189,232,1717,1743]" captionTargetBox="[190,1397,393,1672]" captionTargetId="figure@4.[189,1398,393,1676]" captionTargetPageId="4" captionText="Fig. 2. A–B. Mesoventra of two species of Dacrysoma Grebennikov, 2009 to show form of mid-keel and perforation fringed by setae. C–E. Figures to explain use of the terms foveae, distinct foveolae and shallow foveolae in distinguishing the main species groups. C. Distinct foveolae, showing setae arising from the anterior margin of the depressions. D. Foveae and foveolae, showing setae randomly associated with foveae. E. Shallow foveolae, many setae rubbed off, but insertion points marked by punctures with slightly raised circular rims. F–I. Figures to show different forms of antennae. F. Elongate antenna of Cissidium lisae Darby, 2015. G. Globular antennomere of Cissidium globosum Darby, 2015. H. Female and male antennae of Cissidium petri Darby, 2015. I. Abnormal antennomere XI of Cissidium tigrum Darby, 2015." figureDoi="http://doi.org/10.5281/zenodo.3777286" httpUri="https://zenodo.org/record/3777286/files/figure.png" pageId="148" pageNumber="149">Fig. 2F</figureCitation>
); mentum as
<figureCitation id="130B2A323886FF5A29F4FA9CFE80FA14" box="[189,284,1394,1420]" captionStart="Fig" captionStartId="5.[189,232,1927,1953]" captionTargetBox="[232,1334,1274,1884]" captionTargetId="figure@5.[232,1334,1274,1887]" captionTargetPageId="5" captionText="Fig. 4. Menta and submenta. A. Cissidium petri Darby, 2015, × 1380. B. Cissidium tigrum Darby, 2015, × 1220. C. Cissidium amboroense Darby, 2015, × 1260. D. Cissidium losbanos sp. nov., × 1100. E. Cissidium triangulum Darby, 2019, × 650. F. Cissidium pilosellum sp. nov., × 1420." figureDoi="http://doi.org/10.5281/zenodo.3777290" httpUri="https://zenodo.org/record/3777290/files/figure.png" pageId="148" pageNumber="149">Fig. 4B.</figureCitation>
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4FA58FC9DF984" blockId="148.[189,1399,1462,1564]" pageId="148" pageNumber="149">
PRONOTUM. Length
<quantity id="4CC89B523886FF5A28D7FA58FD96FA48" box="[414,522,1462,1488]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="148" pageNumber="149" unit="mm" value="0.19">0.19 mm</quantity>
, width
<quantity id="4CC89B523886FF5A2B29FA58FD50FA48" box="[608,716,1462,1488]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="148" pageNumber="149" unit="mm" value="0.28">0.28 mm</quantity>
, sparsely pubescent and foveolate; lateral margins sharply angled, strongly concave posteriad to rectangular hind angles, less markedly concave anteriorly to distinct obtusely rounded front angles (
<figureCitation id="130B2A323886FF5A2BC1F9ECFD68F984" box="[648,756,1538,1564]" captionStart="Fig" captionStartId="149.[189,232,1435,1461]" captionTargetBox="[212,1376,453,1383]" captionTargetId="figure@149.[208,1376,441,1389]" captionTargetPageId="149" captionText="Fig. 94. Cissidium lisae Darby, 2015. A. Habitus. B. Pronotum, C. Mesoventrite showing median process of collar, mid-keel and keel, × 550." figureDoi="http://doi.org/10.5281/zenodo.3777472" httpUri="https://zenodo.org/record/3777472/files/figure.png" pageId="148" pageNumber="149">Fig. 94B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4F9A9FEABF91F" blockId="148.[189,1398,1606,1671]" pageId="148" pageNumber="149">
ELYTRA.
<quantity id="4CC89B523886FF5A286AF9A8FE13F9F9" box="[291,399,1606,1633]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="148" pageNumber="149" unit="mm" value="0.55">0.55 mm</quantity>
long,
<quantity id="4CC89B523886FF5A2896F9A8FDD7F9F9" box="[479,587,1606,1633]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="148" pageNumber="149" unit="mm" value="0.41">0.41 mm</quantity>
wide, pubescence a little shorter and more decumbent than on the pronotum.
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4F95FFC2EF8FA" blockId="148.[189,1398,1712,1891]" pageId="148" pageNumber="149">
MESOVENTRITE. Median process of the very narrow collar pointed; mid-keel strongly widened anteriorly, hind angles with carinae to mesocoxal anterior borders, bluntly raised medially before junction with keel; keel with ± five setae, parallel-sided to rounded termination at base of mesocoxae (
<figureCitation id="130B2A323886FF5A2DFEF912FA93F88F" box="[1207,1295,1788,1815]" captionStart="Fig" captionStartId="6.[189,232,1930,1956]" captionTargetBox="[195,1398,757,1880]" captionTargetId="figure@6.[189,1398,756,1884]" captionTargetPageId="6" captionText="Fig. 5. Figures to show keel shapes. A. Cissidium bifoveolatum sp. nov., × 515. B. Cissidium lucidulum sp. nov., × 780. C. Cissidium scutellaris (Deane, 1931), × 1480. D. Cissidium tigrum Darby, 2015, × 840. E. Cissidium sawadai Darby, 2015, × 680. F. Cissidium lisae Darby, 2015, × 823." figureDoi="http://doi.org/10.5281/zenodo.3777292" httpUri="https://zenodo.org/record/3777292/files/figure.png" pageId="148" pageNumber="149">Fig. 5F</figureCitation>
); humeri toothed with flat, sharply pointed carinae reaching posteriorly to mid-point of mesoventrite, mesoventral lateral margins entirely effaced, present as apodemes (
<figureCitation id="130B2A323886FF5A2A70F8A6FC3AF8FA" box="[825,934,1864,1890]" captionStart="Fig" captionStartId="149.[189,232,1435,1461]" captionTargetBox="[212,1376,453,1383]" captionTargetId="figure@149.[208,1376,441,1389]" captionTargetPageId="149" captionText="Fig. 94. Cissidium lisae Darby, 2015. A. Habitus. B. Pronotum, C. Mesoventrite showing median process of collar, mid-keel and keel, × 550." figureDoi="http://doi.org/10.5281/zenodo.3777472" httpUri="https://zenodo.org/record/3777472/files/figure.png" pageId="148" pageNumber="149">Fig. 94C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B73886FF5A29F4F863FDDAF83E" blockId="148.[189,582,1932,1959]" box="[189,582,1932,1959]" pageId="148" pageNumber="149">METAVENTRITE. Spines very small.</paragraph>
<paragraph id="8B8F36B73886FF5A29F4F83FFCE5F873" blockId="148.[189,889,2001,2027]" box="[189,889,2001,2027]" pageId="148" pageNumber="149">GENITALIA. Female spermatheca globular. Males not known.</paragraph>
</subSubSection>
<subSubSection id="C32A653C3887FF5B29F4FEE5FBCDFEE5" pageId="149" pageNumber="150" type="discussion">
<paragraph id="8B8F36B73887FF5B29F4FEE5FEB3FEBD" blockId="149.[189,303,267,293]" box="[189,303,267,293]" pageId="149" pageNumber="150">
<heading id="D0C781DB3887FF5B29F4FEE5FEB3FEBD" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="149" pageNumber="150" reason="3">
<emphasis id="B944EAA53887FF5B29F4FEE5FEB3FEBD" bold="true" box="[189,303,267,293]" pageId="149" pageNumber="150">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73887FF5B29F4FED2FBCDFEE5" blockId="149.[189,1398,316,381]" pageId="149" pageNumber="150">
Differs from the other Bolivian
<taxonomicName id="4C304D343887FF5B2B0AFED3FD24FECF" box="[579,696,317,343]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA53887FF5B2B0AFED3FD24FECF" box="[579,696,317,343]" italics="true" pageId="149" pageNumber="150">Cissidium</emphasis>
</taxonomicName>
, except
<taxonomicName id="4C304D343887FF5B2A68FED3FBD7FECF" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[801,1099,316,343]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="149" pageNumber="150" phylum="Arthropoda" rank="species" species="sawadai">
<emphasis id="B944EAA53887FF5B2A68FED3FC35FECF" box="[801,937,317,343]" italics="true" pageId="149" pageNumber="150">C. sawadai</emphasis>
<bibRefCitation id="EFA14B463887FF5B2AFAFED2FBD7FECF" author="Darby M." box="[947,1099,316,343]" pageId="149" pageNumber="150" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
, by the long and narrow elytra and distinguishable from that species by the strongly tapering mid-keel.
</paragraph>
</subSubSection>
</treatment>
</document>
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