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<document id="C2809DB8DCEA423BBE32983F40B62C47" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A1385AFF842B15FAB4FD29FECC" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium amazonicum Darby 2020, sp. nov." docType="treatment" docUuid="D685D95C-622F-4042-875B-F74E26A8DB42" docUuidSource="ZooBank" docVersion="7" lastPageNumber="75" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="73" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="1BC16222F33C09357C006B2474621CA4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DED0AEE37F9966EDA9AE57C9DB9DB044">
<mods:title id="3FFB15FA41462AE55C511373BF791958">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
</mods:titleInfo>
<mods:name id="FF12163E6AF9E55603283BBBA37F8A08" type="personal">
<mods:role id="149CA608C6F9D4E7BBFF5A6751F5E887">
<mods:roleTerm id="6646A52B9BE32E37885D19E1FEC8F4E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1155B4D7E696F7637B1A829F0D12E837">Darby, Michael</mods:namePart>
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<mods:typeOfResource id="B1244A64CE048FD34C021590BC8E899D">text</mods:typeOfResource>
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<mods:title id="88EA91A453F412A156AAF43EE1AACCCD">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="4E218EF1513F161336DC63C02C251E74">
<mods:date id="5D34EF91938D9A2D52666FEBE1BBA61A">2020</mods:date>
<mods:detail id="5F337FF05597137C4E3B64231122DF32" type="pubDate">
<mods:number id="323575525D67E82D4858088F804F355E">2020-04-02</mods:number>
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<mods:detail id="1228FCC27CFF2633106E0A37CDA28897" type="volume">
<mods:number id="14EDDDDDB119AE7426698538A6DF88F4">622</mods:number>
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<mods:classification id="4AC28142A391BC4A2C6B8DBEF0726785">journal article</mods:classification>
<mods:identifier id="C64366D6D66CBCCF608371AEFA663DD3" type="CLB-Dataset">22458</mods:identifier>
<mods:identifier id="8C08A7AE246D72AE5B0929FCD82DF69A" type="DOI">10.5852/ejt.2020.622</mods:identifier>
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<treatment id="039987A1385AFF842B15FAB4FD29FECC" ID-DOI="http://doi.org/10.5281/zenodo.3795620" ID-GBIF-Taxon="163877826" ID-Zenodo-Dep="3795620" LSID="urn:lsid:zoobank.org:act:D685D95C-622F-4042-875B-F74E26A8DB42" httpUri="http://treatment.plazi.org/id/039987A1385AFF842B15FAB4FD29FECC" lastPageId="74" lastPageNumber="75" pageId="72" pageNumber="73">
<subSubSection id="C32A653C385AFF862B15FAB4FB5FFA03" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="8B8F36B7385AFF862B15FAB4FC44FAED" blockId="72.[369,1219,1370,1471]" box="[604,984,1370,1397]" pageId="72" pageNumber="73">
<heading id="D0C781DB385AFF862B15FAB4FC44FAED" box="[604,984,1370,1397]" centered="true" fontSize="11" level="2" pageId="72" pageNumber="73" reason="2">
<taxonomicName id="4C304D34385AFF862B15FAB4FCEBFAED" ID-CoL="8XXP5" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[604,887,1370,1397]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="amazonicum" status="sp. nov.">
<emphasis id="B944EAA5385AFF862B15FAB4FCEBFAED" bold="true" box="[604,887,1370,1397]" italics="true" pageId="72" pageNumber="73">Cissidium amazonicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE385AFF862A37FAB5FC44FAED" box="[894,984,1371,1397]" pageId="72" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B8F36B7385AFF862838FA6EFB5FFA03" blockId="72.[369,1219,1370,1471]" box="[369,1219,1408,1435]" pageId="72" pageNumber="73">
<uri id="FFA13AB5385AFF862838FA6EFB5FFA03" box="[369,1219,1408,1435]" pageId="72" pageNumber="73">
urn:lsid:zoobank.org:act:
<uuid id="FF960C62385AFF862BC7FA6EFB5FFA03" box="[654,1219,1408,1435]" pageId="72" pageNumber="73">D685D95C-622F-4042-875B-F74E26A8DB42</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C385AFF862BB9FA4BFCDFFA27" box="[752,835,1445,1471]" pageId="72" pageNumber="73" type="description">
<paragraph id="8B8F36B7385AFF862BB9FA4BFCDFFA27" blockId="72.[369,1219,1370,1471]" box="[752,835,1445,1471]" pageId="72" pageNumber="73">
<figureCitation id="130B2A32385AFF862BB9FA4BFCDFFA27" box="[752,835,1445,1471]" captionStart="Fig" captionStartId="73.[189,232,1962,1988]" captionTargetBox="[190,1386,1061,1920]" captionTargetId="figure@73.[189,1386,1061,1920]" captionTargetPageId="73" captionText="Fig. 46. Cissidium amazonicum sp. nov. A. Habitus. B. Pronotum, × 365. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 480." figureDoi="http://doi.org/10.5281/zenodo.3777376" httpUri="https://zenodo.org/record/3777376/files/figure.png" pageId="72" pageNumber="73">Fig. 46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C385AFF8629F4FA06FCE6F9AA" box="[189,890,1512,1587]" pageId="72" pageNumber="73" type="etymology">
<paragraph id="8B8F36B7385AFF8629F4FA06FED8F99A" blockId="72.[189,324,1512,1538]" box="[189,324,1512,1538]" pageId="72" pageNumber="73">
<heading id="D0C781DB385AFF8629F4FA06FED8F99A" bold="true" box="[189,324,1512,1538]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B944EAA5385AFF8629F4FA06FED8F99A" bold="true" box="[189,324,1512,1538]" pageId="72" pageNumber="73">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4F9F6FCE6F9AA" blockId="72.[189,890,1560,1587]" box="[189,890,1560,1587]" pageId="72" pageNumber="73">Named after the region in which the specimens were found.</paragraph>
</subSubSection>
<subSubSection id="C32A653C385AFF8629F4F9B2FB7FF8AC" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="8B8F36B7385AFF8629F4F9B2FE36F9EE" blockId="72.[189,426,1628,1654]" box="[189,426,1628,1654]" pageId="72" pageNumber="73">
<heading id="D0C781DB385AFF8629F4F9B2FE36F9EE" bold="true" box="[189,426,1628,1654]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B944EAA5385AFF8629F4F9B2FE36F9EE" bold="true" box="[189,426,1628,1654]" pageId="72" pageNumber="73">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B583CEA385AFF8629A3F962FB80F954" ID-GBIF-Occurrence="2609493724" collectionCode="MMUE" collectorName="L. Beck" country="Brazil" location="Amazonas" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Amazonas" typeStatus="holotype">
<paragraph id="8B8F36B7385AFF8629A3F962FEC0F93E" blockId="72.[189,1052,1676,1740]" box="[234,348,1676,1702]" pageId="72" pageNumber="73">
<emphasis id="B944EAA5385AFF8629A3F962FEC0F93E" bold="true" box="[234,348,1676,1702]" pageId="72" pageNumber="73">
<typeStatus id="548B8815385AFF8629A3F962FEC0F93E" box="[234,348,1676,1702]" pageId="72" pageNumber="73" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4F95CFB80F954" blockId="72.[189,1052,1676,1740]" box="[189,1052,1713,1740]" pageId="72" pageNumber="73">
<collectingCountry id="F3277627385AFF8629F4F95CFEBAF954" box="[189,294,1714,1740]" name="Brazil" pageId="72" pageNumber="73">BRAZIL</collectingCountry>
•
<specimenCount id="9D36FD3E385AFF862876F95CFECAF954" box="[319,342,1714,1740]" pageId="72" pageNumber="73" type="female">♀</specimenCount>
;
<locationDeviation id="9A3D2F95385AFF86282DF95FFD19F954" box="[356,645,1713,1740]" pageId="72" pageNumber="73">surroundings of Manaus</locationDeviation>
,
<collectingRegion id="49F4F855385AFF862BC7F95CFC92F954" box="[654,782,1714,1740]" country="Brazil" name="Amazonas" pageId="72" pageNumber="73">Amazonas</collectingRegion>
;
<collectorName id="26C55361385AFF862A53F95CFCEAF953" box="[794,886,1713,1740]" pageId="72" pageNumber="73">L. Beck</collectorName>
leg.;
<collectionCode id="ED21AE72385AFF862AF0F95CFB84F954" box="[953,1048,1714,1740]" collectionName="United Kingdom, Manchester, The University, Manchester Museum" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="72" pageNumber="73">MMUE</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="8B8F36B7385AFF8629A3F91AFE7DF896" blockId="72.[189,1251,1780,1844]" box="[234,481,1780,1806]" pageId="72" pageNumber="73">
<emphasis id="B944EAA5385AFF8629A3F91AFE7DF896" bold="true" box="[234,481,1780,1806]" pageId="72" pageNumber="73">Additional material</emphasis>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4F8F4FB7FF8AC" blockId="72.[189,1251,1780,1844]" box="[189,1251,1818,1844]" pageId="72" pageNumber="73">
A second teneral specimen with the same collection details in
<collectionCode id="ED21AE72385AFF862ADCF8F4FC6CF8AC" box="[917,1008,1818,1844]" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="72" pageNumber="73">MMUE</collectionCode>
may be this species.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C385AFF8729F4F8B3FC6AFC59" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="description">
<paragraph id="8B8F36B7385AFF8629F4F8B3FED2F8EF" blockId="72.[189,334,1885,1911]" box="[189,334,1885,1911]" pageId="72" pageNumber="73">
<heading id="D0C781DB385AFF8629F4F8B3FED2F8EF" bold="true" box="[189,334,1885,1911]" fontSize="11" level="3" pageId="72" pageNumber="73" reason="3">
<emphasis id="B944EAA5385AFF8629F4F8B3FED2F8EF" bold="true" box="[189,334,1885,1911]" pageId="72" pageNumber="73">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7385AFF8629F4F863FD36F830" blockId="72.[189,682,1933,1960]" box="[189,682,1933,1960]" pageId="72" pageNumber="73">
SIZE. Habitus (
<figureCitation id="130B2A32385AFF86282EF863FE49F830" box="[359,469,1933,1960]" captionStart="Fig" captionStartId="73.[189,232,1962,1988]" captionTargetBox="[190,1386,1061,1920]" captionTargetId="figure@73.[189,1386,1061,1920]" captionTargetPageId="73" captionText="Fig. 46. Cissidium amazonicum sp. nov. A. Habitus. B. Pronotum, × 365. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 480." figureDoi="http://doi.org/10.5281/zenodo.3777376" httpUri="https://zenodo.org/record/3777376/files/figure.png" pageId="72" pageNumber="73">Fig. 46A</figureCitation>
), length
<quantity id="4CC89B52385AFF862B73F863FD39F830" box="[570,677,1933,1960]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="72" pageNumber="73" unit="mm" value="0.73">0.73 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B7385AFF8729F4F83FFEB3FEF3" blockId="72.[189,873,2001,2027]" lastBlockId="73.[189,1398,267,363]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
COLOUR. Dark red brown, antennae and legs dusky yellow. HEAD. With a linear depression behind the eyes effaced medially, distance across eyes
<quantity id="4CC89B52385BFF872C49FEE5FAEEFEBE" box="[1280,1394,267,294]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="73" pageNumber="74" unit="mm" value="0.27">0.27 mm</quantity>
; antennomeres III–X1, length
<quantity id="4CC89B52385BFF872B6DFEC0FD08FED1" box="[548,660,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="73" pageNumber="74" unit="mm" value="0.39">0.39 mm</quantity>
, IV–IX
<quantity id="4CC89B52385BFF872BB5FEC0FCF0FED1" box="[764,876,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="73" pageNumber="74" unit="mm" value="0.21">0.21 mm</quantity>
, X–XI
<quantity id="4CC89B52385BFF872A82FEC0FBA7FED1" box="[971,1083,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="73" pageNumber="74" unit="mm" value="0.16">0.16 mm</quantity>
; mentum and submentum obscured.
</paragraph>
<paragraph id="8B8F36B7385BFF8729F4FE7CFD44FE6A" blockId="73.[189,1398,401,498]" pageId="73" pageNumber="74">
PRONOTUM. Length
<quantity id="4CC89B52385BFF8728E9FE7CFD91FE34" box="[416,525,402,428]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="73" pageNumber="74" unit="mm" value="0.22">0.22 mm</quantity>
, width
<quantity id="4CC89B52385BFF872B2DFE7CFD4DFE34" box="[612,721,402,428]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="73" pageNumber="74" unit="mm" value="0.39">0.39 mm</quantity>
, distinctly foveolate but foveolae absent medially; lateral margins bordered and sharply angled, posterior margin bordered including emargination opposite scutellum, hind angles rectangular (
<figureCitation id="130B2A32385BFF872B16FE39FD50FE6A" box="[607,716,471,498]" captionStart="Fig" captionStartId="73.[189,232,1962,1988]" captionTargetBox="[190,1386,1061,1920]" captionTargetId="figure@73.[189,1386,1061,1920]" captionTargetPageId="73" captionText="Fig. 46. Cissidium amazonicum sp. nov. A. Habitus. B. Pronotum, × 365. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 480." figureDoi="http://doi.org/10.5281/zenodo.3777376" httpUri="https://zenodo.org/record/3777376/files/figure.png" pageId="73" pageNumber="74">Fig. 46B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7385BFF8729F4FDF6FADAFDAA" blockId="73.[189,1350,536,563]" box="[189,1350,536,563]" pageId="73" pageNumber="74">
ELYTRA. Length
<quantity id="4CC89B52385BFF872833FDF6FE7AFDAA" box="[378,486,536,563]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="73" pageNumber="74" unit="mm" value="0.57">0.57 mm</quantity>
, width
<quantity id="4CC89B52385BFF872B72FDF6FD3BFDAA" box="[571,679,536,563]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="73" pageNumber="74" unit="mm" value="0.51">0.51 mm</quantity>
. pubescent and more shallowly foveolate than pronotum.
</paragraph>
<paragraph id="8B8F36B7385BFF8729F4FDB7FE02FD44" blockId="73.[189,1398,601,732]" pageId="73" pageNumber="74">
MESOVENTRITE. Medial extension of collar parallel-sided extending almost to middle of the mid-keel; midkeel widest anteriorly tapering strongly to junction with keel; keel setose, tapering to a blunt termination between mesocoxae; mesoventral lateral margins without serrations, sharply angulate; humeri bluntly toothed (
<figureCitation id="130B2A32385BFF87286CFD2FFE0EFD44" box="[293,402,705,732]" captionStart="Fig" captionStartId="73.[189,232,1962,1988]" captionTargetBox="[190,1386,1061,1920]" captionTargetId="figure@73.[189,1386,1061,1920]" captionTargetPageId="73" captionText="Fig. 46. Cissidium amazonicum sp. nov. A. Habitus. B. Pronotum, × 365. C. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 480." figureDoi="http://doi.org/10.5281/zenodo.3777376" httpUri="https://zenodo.org/record/3777376/files/figure.png" pageId="73" pageNumber="74">Fig. 46C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7385BFF8729F4FCECFEE2FCA7" blockId="73.[189,1399,770,831]" pageId="73" pageNumber="74">
METAVENTRITE. With scattered pubescence medially, disc simple, length
<quantity id="4CC89B52385BFF872D61FCECFB05FC84" box="[1064,1177,770,796]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="73" pageNumber="74" unit="mm" value="0.17">0.17 mm</quantity>
, distance between spines
<quantity id="4CC89B52385BFF872847FCCAFEE5FCA7" box="[270,377,804,831]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="73" pageNumber="74" unit="mm" value="0.14">0.14 mm</quantity>
.
</paragraph>
<paragraph id="8B8F36B7385BFF8729F4FC88FE5EFC18" blockId="73.[189,450,870,896]" box="[189,450,870,896]" pageId="73" pageNumber="74">WINGS. Macropterous.</paragraph>
<paragraph id="8B8F36B7385BFF8729F4FC49FC6AFC59" blockId="73.[189,1014,935,961]" box="[189,1014,935,961]" pageId="73" pageNumber="74">GENITALIA. The female spermatheca was not detected in the dissection.</paragraph>
</subSubSection>
<caption id="DF4F663F385BFF8729F4F844FC54F870" ID-DOI="http://doi.org/10.5281/zenodo.3777376" ID-Zenodo-Dep="3777376" httpUri="https://zenodo.org/record/3777376/files/figure.png" pageId="73" pageNumber="74" startId="73.[189,232,1962,1988]" targetBox="[190,1386,1061,1920]" targetPageId="73">
<paragraph id="8B8F36B7385BFF8729F4F844FC54F870" blockId="73.[189,1398,1962,2024]" pageId="73" pageNumber="74">
<emphasis id="B944EAA5385BFF8729F4F844FE80F85D" bold="true" box="[189,284,1962,1989]" pageId="73" pageNumber="74">Fig. 46.</emphasis>
<taxonomicName id="4C304D34385BFF87286EF844FDA7F85C" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[295,571,1962,1988]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="amazonicum" status="sp. nov.">
<emphasis id="B944EAA5385BFF87286EF844FDA7F85C" box="[295,571,1962,1988]" italics="true" pageId="73" pageNumber="74">Cissidium amazonicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE385BFF872B0FF845FD38F85D" box="[582,676,1963,1989]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B944EAA5385BFF872BE7F844FD5FF85C" bold="true" box="[686,707,1962,1988]" pageId="73" pageNumber="74">A</emphasis>
. Habitus.
<emphasis id="B944EAA5385BFF872A0BF844FCCAF85C" bold="true" box="[834,854,1962,1988]" pageId="73" pageNumber="74">B</emphasis>
. Pronotum, × 365.
<emphasis id="B944EAA5385BFF872D0FF844FBC7F85C" bold="true" box="[1094,1115,1962,1988]" pageId="73" pageNumber="74">C</emphasis>
. Mesoventrite showing median process of collar, mid-keel, keel and lateral margin, × 480.
</paragraph>
</caption>
<subSubSection id="C32A653C3858FF8429F4FEE5FD29FECC" box="[189,693,267,340]" pageId="74" pageNumber="75" type="discussion">
<paragraph id="8B8F36B73858FF8429F4FEE5FEB3FEBD" blockId="74.[189,303,267,293]" box="[189,303,267,293]" pageId="74" pageNumber="75">
<heading id="D0C781DB3858FF8429F4FEE5FEB3FEBD" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="B944EAA53858FF8429F4FEE5FEB3FEBD" bold="true" box="[189,303,267,293]" pageId="74" pageNumber="75">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B73858FF8429F4FED7FD29FECC" blockId="74.[189,693,313,340]" box="[189,693,313,340]" pageId="74" pageNumber="75">
The only species from
<collectingCountry id="F32776273858FF842883FED7FD8EFECB" box="[458,530,313,339]" name="Brazil" pageId="74" pageNumber="75">Brazil</collectingCountry>
in this group.
</paragraph>
</subSubSection>
</treatment>
</document>
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