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<document id="A0D549DA9FF487683324AB9997DE4750" ID-DOI="10.1080/0022293021000030844" ID-ISSN="1464-5262" ID-Zenodo-Dep="5258656" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629911445027" checkinUser="felipe" docAuthor="George, Robert Y." docDate="2004" docId="03998778D051FFE4FDE6FDC8348A05AB" docLanguage="en" docName="JNatHist.38.3.337-373.pdf" docOrigin="Journal of Natural History 38 (3)" docSource="http://www.tandfonline.com/doi/abs/10.1080/0022293021000030844" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Haploniscidae Hansen 1916" docType="treatment" docVersion="3" lastPageNumber="341" masterDocId="FFA0FF00D055FFE1FFC0FFFA307E036E" masterDocTitle="Deep-sea asellote isopods (Crustacea, Eumalacostraca) of the north-west Atlantic: the family Haploniscidae" masterLastPageNumber="373" masterPageNumber="337" pageNumber="340" updateTime="1699127404719" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4D2A5C0EA3E9DC9012643160C1AF49FD">Deep-sea asellote isopods (Crustacea, Eumalacostraca) of the north-west Atlantic: the family Haploniscidae</mods:title>
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<mods:namePart id="355A2145F591CB3C26FE82A1BEB6EA0A">George, Robert Y.</mods:namePart>
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<mods:title id="3EC874014CB01E274C235B9283FC1E7C">Journal of Natural History</mods:title>
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<mods:date id="F9FF64F45AD0FF4789440FEFE8691B67">2004</mods:date>
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<mods:number id="7A335BBFD585D099E2F105C0599073AA">2004-10-10</mods:number>
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<mods:number id="32C0EDC5E52CD15965D09F8305D7634D">38</mods:number>
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<mods:number id="9F7922107052BA97390A6B0129DA5409">3</mods:number>
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<mods:classification id="A6978D012BC947D39F16954B3A6D1A06">journal article</mods:classification>
<mods:identifier id="D8235D77F10774325D1C55A42E541B6F" type="DOI">10.1080/0022293021000030844</mods:identifier>
<mods:identifier id="60604071AD48CC69148B9DBB3E797F9C" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03998778D051FFE4FDE6FDC8348A05AB" LSID="urn:lsid:plazi:treatment:03998778D051FFE4FDE6FDC8348A05AB" httpUri="http://treatment.plazi.org/id/03998778D051FFE4FDE6FDC8348A05AB" lastPageId="5" lastPageNumber="341" pageId="4" pageNumber="340">
<subSubSection id="C32A65E5D051FFE5FDE6FDC832950107" pageId="4" pageNumber="340" type="nomenclature">
<paragraph id="8B8F366ED051FFE5FDE6FDC834790124" blockId="4.[550,1031,562,586]" box="[550,1031,562,586]" pageId="4" pageNumber="340">
<heading id="D0C78102D051FFE5FDE6FDC834790124" box="[550,1031,562,586]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="340" reason="2">
<emphasis id="B944EA7CD051FFE5FDE6FDC8320C0124" bold="true" box="[550,626,562,586]" pageId="4" pageNumber="340">Family</emphasis>
<taxonomicName id="4C304DEDD051FFE5FDBBFDC834790124" ID-CoL="7NHRW" authority="Hansen, 1916" authorityName="Hansen" authorityYear="1916" box="[635,1031,562,586]" class="Malacostraca" family="Haploniscidae" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="family">
<emphasis id="B944EA7CD051FFE5FDBBFDC8331C0124" bold="true" box="[635,866,562,586]" pageId="4" pageNumber="340">HAPLONISCIDAE</emphasis>
Hansen, 1916
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9FDAE32950107" blockId="4.[313,1268,596,671]" box="[313,747,596,618]" pageId="4" pageNumber="340">Synonym: Haploniscini Hansen, 1916: 28.</paragraph>
</subSubSection>
<subSubSection id="C32A65E5D051FFE4FEF9FD94348A05AB" lastPageId="5" lastPageNumber="341" pageId="4" pageNumber="340" type="description">
<paragraph id="8B8F366ED051FFE5FEF9FD9432DB01F1" blockId="4.[313,1268,596,671]" pageId="4" pageNumber="340">
<taxonomicName id="4C304DEDD051FFE5FEF9FD9431B001EA" authorityName="Hansen" authorityYear="1916" box="[313,462,622,644]" class="Malacostraca" family="Haploniscidae" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="family">Haploniscidae</taxonomicName>
: Gurjanova, 1933: 402; Menzies, 1962: 94; Wolf, 1962: 49; Menzies and George, 1972: 107; Lincoln, 1985b: 659.
</paragraph>
<paragraph id="8B8F366ED051FFE5FEA1FD3232E900D1" blockId="4.[313,1268,711,959]" pageId="4" pageNumber="340">
<emphasis id="B944EA7CD051FFE5FEA1FD3231B301B1" box="[353,461,712,735]" italics="true" pageId="4" pageNumber="340">Diagnosis</emphasis>
.
<taxonomicName id="4C304DEDD051FFE5FE20FD3D322801B1" authorityName="G. O. Sars" authorityYear="1897" box="[480,598,711,735]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="339" phylum="Arthropoda" rank="superFamily" superFamily="Janiroidea">Janiroidea</taxonomicName>
with free cephalon; blind; oval or subrectangular body, dorsoventrally flattened; all pereonites wider than long, free or fused. Pleotelson composed of a single somite, fused with last pereonite in some genera. Anus separated from branchial chamber by robust pre-anal bar. Antenna shorter than body, antenna 1 shorter than 2. Mandible with incisor and lacinia only on left mandible, molar elongate and truncated, palp three-segmented with article 3 reflexed. Pereopods 1–7 ambulatory, dactylus with single claw. Uropods small, uniramous, ventral, subterminal, one- or two-segmented.
</paragraph>
<paragraph id="8B8F366ED051FFE5FD8AFBF23321072E" blockId="4.[313,1268,1031,1146]" pageId="4" pageNumber="340">
Key to the genera of
<taxonomicName id="4C304DEDD051FFE5FC81FBFD339D0771" authorityName="Hansen" authorityYear="1916" box="[833,995,1031,1055]" class="Malacostraca" family="Haploniscidae" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="family">Haploniscidae</taxonomicName>
(figures 2, 3)
</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9FBB3348A0730" blockId="4.[313,1268,1031,1146]" box="[313,1268,1097,1119]" pageId="4" pageNumber="340">1 Pleotelson with posterior part tapering to a blunt point........ 2</paragraph>
<paragraph id="8B8F366ED051FFE5FEFAFB9E348A0717" blockId="4.[313,1268,1031,1146]" box="[314,1268,1124,1146]" pageId="4" pageNumber="340">– Pleotelson truncated or broadly rounded posteriorly......... 3</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9FB6334BA07C1" blockId="4.[313,1268,1177,1279]" box="[313,1220,1177,1199]" pageId="4" pageNumber="340">2 Uropods not concealed, set close together, ventral to the anal valves; female opercular</paragraph>
<paragraph id="8B8F366ED051FFE5FE91FB4E34BA078A" blockId="4.[313,1268,1177,1279]" pageId="4" pageNumber="340">
pleopod fringed with long plumose distal setae.
<taxonomicName id="4C304DEDD051FFE5FC99FB4E348A07A4" authority="Menzies and Schultz, 1968" authorityName="Menzies and Schultz" authorityYear="1968" box="[857,1268,1204,1226]" class="Malacostraca" family="Haploniscidae" genus="Aspidoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD051FFE5FC99FB4E33A707A7" box="[857,985,1204,1225]" italics="true" pageId="4" pageNumber="340">Aspidoniscus</emphasis>
Menzies and Schultz, 1968
</taxonomicName>
– Uropods concealed; female opercular pleopod fringed with simple short marginal setae
</paragraph>
<paragraph id="8B8F366ED051FFE5FE96FB13348A0790" blockId="4.[313,1268,1177,1279]" box="[342,1268,1257,1279]" pageId="4" pageNumber="340">
.................
<taxonomicName id="4C304DEDD051FFE5FC1DFB13348A0790" authority="Birstein, 1971" authorityName="Birstein" authorityYear="1971" box="[989,1268,1257,1279]" class="Malacostraca" family="Haploniscidae" genus="Abyssoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD051FFE5FC1DFB13341E0790" box="[989,1120,1257,1278]" italics="true" pageId="4" pageNumber="340">Abyssoniscus</emphasis>
Birstein, 1971
</taxonomicName>
</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9FAE534BA065A" blockId="4.[313,1268,1311,1412]" box="[313,1220,1311,1333]" pageId="4" pageNumber="340">3 Antenna 2 with peduncular articles flattened, article 3 elongate and grooved, article 4</paragraph>
<paragraph id="8B8F366ED051FFE5FE91FAC3348A0604" blockId="4.[313,1268,1311,1412]" pageId="4" pageNumber="340">
short, articles 5 and 6 fused and apically prolonged, flagellum small and slender, subterminal (figure 2C)..........
<taxonomicName id="4C304DEDD051FFE5FC79FAAE348A0604" authority="Menzies, 1962" authorityName="Menzies" authorityYear="1962" box="[953,1268,1364,1386]" class="Malacostraca" family="Haploniscidae" genus="Antennuloniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD051FFE5FC79FAAE34240607" box="[953,1114,1364,1385]" italics="true" pageId="4" pageNumber="340">Antennuloniscus</emphasis>
Menzies, 1962
</taxonomicName>
</paragraph>
<paragraph id="8B8F366ED051FFE5FEFAFA95348A06ED" blockId="4.[313,1268,1311,1412]" box="[314,1268,1390,1412]" pageId="4" pageNumber="340">– Antenna 2 unmodified................. 4</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9FA5E34BA06BB" blockId="4.[313,1269,1443,1519]" pageId="4" pageNumber="340">
4 Pereonites 5–7 free (figure 3D).........
<taxonomicName id="4C304DEDD051FFE5FBD6FA5E34D906D7" authority="George, 2004" authorityName="George" authorityYear="2004" box="[1046,1191,1444,1465]" class="Malacostraca" family="Haploniscidae" genus="Chandraniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944EA7CD051FFE5FBD6FA5E34D906D7" box="[1046,1191,1444,1465]" italics="true" pageId="4" pageNumber="340">Chandraniscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775707D051FFE5FB6EFA5E348B06D7" box="[1198,1269,1444,1465]" pageId="4" pageNumber="340" rank="genus">n. gen.</taxonomicNameLabel>
– Pereonites 5 and 6 fused, or 6 and 7 fused, or 7 fused with pleotelson, or 5–7 and
</paragraph>
<paragraph id="8B8F366ED051FFE5FE91FA23348A0681" blockId="4.[313,1269,1443,1519]" box="[337,1268,1496,1519]" pageId="4" pageNumber="340">pleotelson fused together................. 5</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9F9F4348A054A" blockId="4.[313,1268,1550,1626]" box="[313,1268,1550,1573]" pageId="4" pageNumber="340">5 Pereonites 5–7 fused with the pleotelson to form a single segment (figure 2B)....</paragraph>
<paragraph id="8B8F366ED051FFE5FE96F9D0348A0534" blockId="4.[313,1268,1550,1626]" pageId="4" pageNumber="340">
.................
<taxonomicName id="4C304DEDD051FFE5FC25F9D3348A0551" authority="Hansen, 1916" authorityName="Hansen" authorityYear="1916" box="[997,1268,1577,1599]" class="Malacostraca" family="Haploniscidae" genus="Hydroniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD051FFE5FC25F9D3341F0550" box="[997,1121,1577,1598]" italics="true" pageId="4" pageNumber="340">Hydroniscus</emphasis>
Hansen, 1916
</taxonomicName>
– Pereonites 5 and 6 fused, or 6 and 7 fused, with pleotelson........ 6
</paragraph>
<paragraph id="8B8F366ED051FFE5FEF9F980348A05E1" blockId="4.[313,1268,1657,1733]" box="[313,1268,1657,1680]" pageId="4" pageNumber="340">6 Pereonites 5–7 without any pronounced anterolateral angles in anterior pereonites. 7</paragraph>
<paragraph id="8B8F366ED051FFE5FEFAF96F34BA05C4" blockId="4.[313,1268,1657,1733]" box="[314,1220,1683,1706]" pageId="4" pageNumber="340">– Pereonites 5–7 free, with one or more pereonites 2–4 having prolonged anterolateral</paragraph>
<paragraph id="8B8F366ED051FFE5FE92F955348A05AB" blockId="4.[313,1268,1657,1733]" box="[338,1268,1710,1733]" pageId="4" pageNumber="340">
angles (figure 3B)...........
<taxonomicName id="4C304DEDD051FFE5FC7EF955348A05AB" authority="Lincoln, 1985" authorityName="Lincoln" authorityYear="1985" box="[958,1268,1710,1733]" class="Malacostraca" family="Haploniscidae" genus="Chauliodoniscus" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD051FFE5FC7EF955342105AA" box="[958,1119,1711,1732]" italics="true" pageId="4" pageNumber="340">Chauliodoniscus</emphasis>
Lincoln, 1985
</taxonomicName>
</paragraph>
<caption id="DF4F66E6D050FFE4FEF9FCCD34960009" ID-DOI="http://doi.org/10.5281/zenodo.5258660" ID-Zenodo-Dep="5258660" httpUri="https://zenodo.org/record/5258660/files/figure.png" pageId="5" pageNumber="341" startId="5.[313,329,823,844]" targetBox="[315,1272,172,802]" targetPageId="5">
<paragraph id="8B8F366ED050FFE4FEF9FCCD34960009" blockId="5.[313,1268,823,872]" pageId="5" pageNumber="341">
F. 2. Four Haploniscid genera: (A)
<taxonomicName id="4C304DEDD050FFE4FD2DFCCD344D0022" authority="Richardson, 1908" authorityName="Richardson" authorityYear="1908" box="[749,1075,823,845]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FD2DFCCD33180022" box="[749,870,823,844]" italics="true" pageId="5" pageNumber="341">Haploniscus</emphasis>
Richardson, 1908
</taxonomicName>
; (B)
<taxonomicName id="4C304DEDD050FFE4FBB8FCCD31E90009" authority="Hansen" authorityName="Hansen" authorityYear="1916" class="Malacostraca" family="Haploniscidae" genus="Hydroniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FBB8FCCD348A0022" box="[1144,1268,823,844]" italics="true" pageId="5" pageNumber="341">Hydroniscus</emphasis>
Hansen
</taxonomicName>
; (C)
<taxonomicName id="4C304DEDD050FFE4FE0FFCAB33720009" authority="Menzies, 1962" authorityName="Menzies" authorityYear="1962" box="[463,780,849,871]" class="Malacostraca" family="Haploniscidae" genus="Antennuloniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FE0FFCAB320E0008" box="[463,624,849,870]" italics="true" pageId="5" pageNumber="341">Antennuloniscus</emphasis>
Menzies, 1962
</taxonomicName>
; (D)
<taxonomicName id="4C304DEDD050FFE4FC85FCAB349D0009" authority="Menzies and Schultz, 1968" authorityName="Menzies and Schultz" authorityYear="1968" box="[837,1251,849,871]" class="Malacostraca" family="Haploniscidae" genus="Aspidoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FC85FCAB33BB0008" box="[837,965,849,870]" italics="true" pageId="5" pageNumber="341">Aspidoniscus</emphasis>
Menzies and Schultz, 1968
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF4F66E6D050FFE4FEF9FA2834E4056C" ID-DOI="http://doi.org/10.5281/zenodo.5258662" ID-Zenodo-Dep="5258662" httpUri="https://zenodo.org/record/5258662/files/figure.png" pageId="5" pageNumber="341" startId="5.[313,329,1490,1511]" targetBox="[314,1272,909,1471]" targetPageId="5">
<paragraph id="8B8F366ED050FFE4FEF9FA2834E4056C" blockId="5.[313,1268,1490,1539]" pageId="5" pageNumber="341">
F. 3. Four Haploniscid genera: (A)
<taxonomicName id="4C304DEDD050FFE4FD2AFA28346E0689" authority="Birstein, 1971" authorityName="Birstein" authorityYear="1971" box="[746,1040,1490,1512]" class="Malacostraca" family="Haploniscidae" genus="Abyssoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FD2AFA2833130689" box="[746,877,1490,1511]" italics="true" pageId="5" pageNumber="341">Abyssoniscus</emphasis>
Birstein, 1971
</taxonomicName>
; (B)
<taxonomicName id="4C304DEDD050FFE4FB93FA28325D056D" authority="Lincoln, 1985" authorityName="Lincoln" authorityYear="1985" class="Malacostraca" family="Haploniscidae" genus="Chauliodoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FB93FA28348A0689" box="[1107,1268,1490,1511]" italics="true" pageId="5" pageNumber="341">Chauliodoniscus</emphasis>
Lincoln, 1985
</taxonomicName>
; (C)
<taxonomicName id="4C304DEDD050FFE4FD9CFA1733FF056D" authority="Lincoln, 1985" authorityName="Lincoln" authorityYear="1985" box="[604,897,1516,1539]" class="Malacostraca" family="Haploniscidae" genus="Mastigoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FD9CFA173294056C" box="[604,746,1517,1538]" italics="true" pageId="5" pageNumber="341">Mastigoniscus</emphasis>
Lincoln, 1985
</taxonomicName>
; (D)
<taxonomicName id="4C304DEDD050FFE4FC7BFA163432056F" authority="George, 2004" authorityName="George" authorityYear="2004" box="[955,1100,1516,1537]" class="Malacostraca" family="Haploniscidae" genus="Chandraniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944EA7CD050FFE4FC7BFA163432056F" box="[955,1100,1516,1537]" italics="true" pageId="5" pageNumber="341">Chandraniscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775707D050FFE4FB93FA1734E4056C" box="[1107,1178,1517,1538]" pageId="5" pageNumber="341" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8F366ED050FFE4FEF9F9D034BB052E" blockId="5.[313,1268,1577,1733]" box="[313,1221,1577,1600]" pageId="5" pageNumber="341">7 Pereonites 5–7 fused between each other, and pereonite 7 fused or not fused with</paragraph>
<paragraph id="8B8F366ED050FFE4FE91F9BE34BB05FE" blockId="5.[313,1268,1577,1733]" pageId="5" pageNumber="341">
pleotelson but pereonite 7 not reduced (figure 2A)..
<taxonomicName id="4C304DEDD050FFE4FC7EF9BE348A0534" authority="Richardson, 1908" authorityName="Richardson" authorityYear="1908" box="[958,1268,1604,1626]" class="Malacostraca" family="Haploniscidae" genus="Haploniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FC7EF9BE34490537" box="[958,1079,1604,1625]" italics="true" pageId="5" pageNumber="341">Haploniscus</emphasis>
Richardson, 1908
</taxonomicName>
– Posterior pereonites fused mid-dorsally between each other and with pleotelson, and pereonite 7 much reduced in adult, conspicuously shorter and narrower than pereonite
</paragraph>
<paragraph id="8B8F366ED050FFE4FE92F96E348A05C4" blockId="5.[313,1268,1577,1733]" box="[338,1268,1684,1706]" pageId="5" pageNumber="341">6, and pleotelson with posterolateral angles conspicuously produced (figure 3C)...</paragraph>
<paragraph id="8B8F366ED050FFE4FE96F955348A05AB" blockId="5.[313,1268,1577,1733]" box="[342,1268,1710,1733]" pageId="5" pageNumber="341">
.................
<taxonomicName id="4C304DEDD050FFE4FC11F955348A05AB" authority="Lincoln, 1985" authorityName="Lincoln" authorityYear="1985" box="[977,1268,1710,1733]" class="Malacostraca" family="Haploniscidae" genus="Mastigoniscus" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B944EA7CD050FFE4FC11F955342105AA" box="[977,1119,1711,1732]" italics="true" pageId="5" pageNumber="341">Mastigoniscus</emphasis>
Lincoln, 1985
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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