345 lines
49 KiB
XML
345 lines
49 KiB
XML
<document id="71AACC022A0A268947301094A83F322A" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3727F5FFF67FD8A8F05DA6C" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) gupchiensis Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="335" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="332" updateTime="1698468866275" updateUser="ExternalLinkService">
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<mods:titleInfo id="D2F4B610121A5830F56E8AF47EC76A6C">
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<mods:title id="CA12BAB88B47C1E829425D9F3AB0FF0F">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:name id="7F878F4E02BA0C43792F891D78A6505C" type="personal">
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<mods:roleTerm id="FC6094280A1C85AA257E64334CE00C76">Author</mods:roleTerm>
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<mods:namePart id="3F488320CD3D8C368D0E566C00B939E7">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="68838B8A1BC1AEF3C4D635990C694DE3">text</mods:typeOfResource>
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<mods:title id="BB8B6385F4F3CA5B588AFF92DB9B807F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F40922660ADC08DC6E72E790A6DB5857">
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<mods:date id="E621CBD019B11725706117BB22F53520">2017</mods:date>
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<mods:number id="481429CCBC8C7331F64265D27AFF1CA5">2017-09-22</mods:number>
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<mods:detail id="BA4D37F05A90ADE798D68740A671A0E5" type="volume">
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<mods:number id="66BD54BB15E95B97BCF558C61CF0A8B1">4323</mods:number>
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<mods:number id="FAE2722454F19779BDA4980BE1C8F42D">3</mods:number>
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<mods:classification id="40EC2A54C8511F2F6E4CD5272EB181EE">journal article</mods:classification>
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<mods:identifier id="A5A7A6D10DD6140BB1182215A126E4DD" type="DOI">10.11646/zootaxa.4323.3.1</mods:identifier>
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<treatment id="03956A03A3727F5FFF67FD8A8F05DA6C" ID-DOI="http://doi.org/10.5281/zenodo.6015761" ID-GBIF-Taxon="134839418" ID-Zenodo-Dep="6015761" LSID="urn:lsid:plazi:treatment:03956A03A3727F5FFF67FD8A8F05DA6C" httpUri="http://treatment.plazi.org/id/03956A03A3727F5FFF67FD8A8F05DA6C" lastPageId="34" lastPageNumber="335" pageId="31" pageNumber="332">
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<subSubSection id="C326889EA3727F62FF67FD8A8AEEDB6F" pageId="31" pageNumber="332" type="nomenclature">
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<paragraph id="8B83DB15A3727F62FF67FD8A89EEDB0E" blockId="31.[151,690,548,607]" box="[151,690,548,574]" pageId="31" pageNumber="332">
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<heading id="D0CB6C79A3727F62FF67FD8A89EEDB0E" bold="true" box="[151,690,548,574]" fontSize="11" level="1" pageId="31" pageNumber="332" reason="1">
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<emphasis id="B9480707A3727F62FF67FD8A89EEDB0E" bold="true" box="[151,690,548,574]" pageId="31" pageNumber="332">
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<taxonomicName id="4C3CA096A3727F62FF67FD8A8935DB0E" ID-CoL="58LGR" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,617,548,574]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="gupchiensis" status="sp. nov." subGenus="Bottchrus">
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<emphasis id="B9480707A3727F62FF67FD8A8A1DDB0E" bold="true" box="[151,321,548,574]" italics="true" pageId="31" pageNumber="332">Trichotichnus</emphasis>
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(
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<emphasis id="B9480707A3727F62FEA2FD8A8A95DB0E" bold="true" box="[338,457,548,574]" italics="true" pageId="31" pageNumber="332">Bottchrus</emphasis>
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)
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<emphasis id="B9480707A3727F62FE2AFD8A8935DB0E" bold="true" box="[474,617,548,574]" italics="true" pageId="31" pageNumber="332">gupchiensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BBA7CA3727F62FD80FD8A89EEDB0E" box="[624,690,548,574]" pageId="31" pageNumber="332" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B83DB15A3727F62FF67FDE88AEEDB6F" blockId="31.[151,690,548,607]" box="[151,434,582,607]" pageId="31" pageNumber="332">
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(
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<figureCitation id="1307C790A3727F62FF6FFDE88BA9DB6F" box="[159,245,582,607]" captionStart="FIGURES 91 – 94" captionStartId="32.[151,264,1940,1963]" captionTargetBox="[265,1303,196,1917]" captionTargetId="figure@32.[261,1305,193,1921]" captionTargetPageId="32" captionText="FIGURES 91 – 94. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 91, T. gupchiensis sp. n., paratype, male. 92, T. brancuccii sp. n., paratype, male. 93, T. manasluensis sp. n., paratype, male. 94, T. cyanescens Ito, 1998, male from Dhabeli." httpUri="https://zenodo.org/record/919654/files/figure.png" pageId="31" pageNumber="332">Figs 91</figureCitation>
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,
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<figureCitation id="1307C790A3727F62FEF1FDE88A04DB6F" box="[257,344,582,607]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="31" pageNumber="332">95–102</figureCitation>
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,
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<figureCitation id="1307C790A3727F62FE93FDE88AD3DB6F" box="[355,399,582,607]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="31" pageNumber="332">152</figureCitation>
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,
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<emphasis id="B9480707A3727F62FE6BFDE98AF5DB6F" box="[411,425,583,607]" italics="true" pageId="31" pageNumber="332">1</emphasis>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C326889EA3727F62FF67FD208932DA07" pageId="31" pageNumber="332" type="materials_examined">
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<paragraph id="8B83DB15A3727F62FF67FD2089C6DBFB" blockId="31.[151,1437,654,1975]" pageId="31" pageNumber="332">
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<materialsCitation id="3B54D148A3727F62FF67FD2089C6DBFB" ID-GBIF-Occurrence="1655986839" collectingDate="2006-05-18" collectorName="Manaslu Mts. SE & W Gupchi Danda & J. Schmidt" country="Nepal" elevation="2250" latitude="28.143612" location="Manaslu Mts." longLatPrecision="20" longitude="84.744995" pageId="31" pageNumber="332" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B9480707A3727F62FF67FD208A1EDB97" bold="true" box="[151,322,654,679]" pageId="31" pageNumber="332">
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<typeStatus id="548765B7A3727F62FF67FD208B8EDB97" box="[151,210,654,679]" pageId="31" pageNumber="332">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="548765B7A3727F62FEA5FD208A9EDB97" box="[341,450,654,679]" pageId="31" pageNumber="332" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D3A109CA3727F62FE21FD218AB4DB96" box="[465,488,655,678]" pageId="31" pageNumber="332" type="male">♂</specimenCount>
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||
, “
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||
<collectingCountry id="F32B9B85A3727F62FDF2FD218901DB97" box="[514,605,655,679]" name="Nepal" pageId="31" pageNumber="332">NEPAL</collectingCountry>
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||
<collectorName id="26C9BEC3A3727F62FD98FD208868DB97" box="[616,820,654,679]" pageId="31" pageNumber="332">
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||
<location id="8EE38DCEA3727F62FD98FD20885BDB96" LSID="urn:lsid:plazi:treatment:03956A03A3727F5FFF67FD8A8F05DA6C:8EE38DCEA3727F62FD98FD20885BDB96" box="[616,775,654,679]" country="Nepal" latitude="28.143612" longLatPrecision="20" longitude="84.744995" name="Manaslu Mts." pageId="31" pageNumber="332">Manaslu Mts.</location>
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SE
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</collectorName>
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slope,
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<collectorName id="26C9BEC3A3727F62FC7DFD218F06DB97" box="[909,1114,654,679]" pageId="31" pageNumber="332">W Gupchi Danda</collectorName>
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||
<elevation id="00113C26A3727F62FB96FD218E54DB96" box="[1126,1288,655,679]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="31" pageNumber="332" unit="m" value="2250.0" valueMax="2300.0" valueMin="2200.0">
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||
<quantity id="4CC476F0A3727F62FB96FD218E54DB96" box="[1126,1288,655,679]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="31" pageNumber="332" unit="m" value="2250.0" valueMax="2300.0" valueMin="2200.0">2200–2300 m</quantity>
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||
</elevation>
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||
,
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||
<geoCoordinate id="EE08BDD2A3727F62FAE8FD218EC0DB97" box="[1304,1436,655,679]" direction="north" orientation="latitude" pageId="31" pageNumber="332" precision="15" value="28.143612">28°08'37"N</geoCoordinate>
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||
<geoCoordinate id="EE08BDD2A3727F62FF67FD1D8A46DBFB" box="[151,282,691,715]" direction="east" orientation="longitude" pageId="31" pageNumber="332" precision="15" value="84.744995">84°44'42"E</geoCoordinate>
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||
,
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||
<collectingDate id="EFC6043DA3727F62FED5FD1C8AC4DBFB" box="[293,408,690,715]" pageId="31" pageNumber="332" value="2006-05-18">
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||
<date id="FF82FDD5A3727F62FED5FD1C8AC4DBFB" box="[293,408,690,715]" pageId="31" pageNumber="332" value="2006-05-18">18.V.2006</date>
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</collectingDate>
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leg.
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<collectorName id="26C9BEC3A3727F62FE3FFD1D8914DBFB" box="[463,584,690,715]" pageId="31" pageNumber="332">J. Schmidt</collectorName>
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” (cJS).
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8B83DB15A3727F62FF37FD798932DA07" blockId="31.[151,1437,654,1975]" pageId="31" pageNumber="332">
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||
<materialsCitation id="3B54D148A3727F62FF37FD79881FDBDF" ID-GBIF-Occurrence="1655986825" box="[199,835,726,751]" collectingDate="2006-05-18" collectionCode="ZIN" collectorName="Manaslu Mts. SE & W Gupchi Danda & J. Schmidt" country="Nepal" elevation="2250" latitude="28.143612" location="Manaslu Mts." longLatPrecision="20" longitude="84.744995" pageId="31" pageNumber="332" specimenCount="34" specimenCount-female="15" specimenCount-male="19" typeStatus="paratype">
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<typeStatus id="548765B7A3727F62FF37FD798A65DBDF" box="[199,313,727,751]" pageId="31" pageNumber="332" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="9D3A109CA3727F62FEB2FD798A26DBDE" box="[322,378,727,751]" pageId="31" pageNumber="332" type="male">19 ♂</specimenCount>
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||
,
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||
<specimenCount id="9D3A109CA3727F62FE78FD788A9CDBDE" box="[392,448,726,751]" pageId="31" pageNumber="332" type="female">15 ♀</specimenCount>
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, same data as holotype (cJS,
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<collectionCode id="ED2D43D0A3727F62FCF8FD798867DBDF" box="[776,827,727,751]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="332">ZIN</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B54D148A3727F62FCBFFD7989A0DA23" ID-GBIF-Occurrence="1655986845" collectingDate="2006-05-23" collectorName="E Gupchi Danda & J. Schmidt" country="Nepal" elevation="2400" location="Manaslu Mts." municipality="Gumda" pageId="31" pageNumber="332" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<specimenCount id="9D3A109CA3727F62FCBFFD798824DBDE" box="[847,888,727,751]" pageId="31" pageNumber="332" type="male">1 ♂</specimenCount>
|
||
,
|
||
<location id="8EE38DCEA3727F62FC76FD788F7FDBDE" LSID="urn:lsid:plazi:treatment:03956A03A3727F5FFF67FD8A8F05DA6C:8EE38DCEA3727F62FC76FD788F7FDBDE" box="[902,1059,726,751]" country="Nepal" municipality="Gumda" name="Manaslu Mts." pageId="31" pageNumber="332">Manaslu Mts.</location>
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||
, E-slope,
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||
<collectorName id="26C9BEC3A3727F62FB61FD798E12DBDF" box="[1169,1358,726,751]" pageId="31" pageNumber="332">E Gupchi Danda</collectorName>
|
||
, above
|
||
<collectingMunicipality id="6BE7416FA3727F62FF67FD548BB6DA23" box="[151,234,762,787]" pageId="31" pageNumber="332">Gumda</collectingMunicipality>
|
||
vill.,
|
||
<elevation id="00113C26A3727F62FEDCFD558ADFDA22" box="[300,387,763,787]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="332" unit="m" value="2400.0">
|
||
<quantity id="4CC476F0A3727F62FEDCFD558ADFDA22" box="[300,387,763,787]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="332" unit="m" value="2400.0">2400 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3727F62FE7FFD548959DA23" box="[399,517,762,787]" pageId="31" pageNumber="332" value="2006-05-23">
|
||
<date id="FF82FDD5A3727F62FE7FFD548959DA23" box="[399,517,762,787]" pageId="31" pageNumber="332" value="2006-05-23">23.V.2006</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3727F62FDE0FD5589DADA23" box="[528,646,762,787]" pageId="31" pageNumber="332">J. Schmidt</collectorName>
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||
leg. (cJS)
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||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B54D148A3727F62FCF8FD558936DA07" ID-GBIF-Occurrence="1655986853" collectingDate="2006-05-20" collectionCode="ZIN" collectorName="J. Schmidt" country="Nepal" elevation="2650" latitude="28.149721" location="Manaslu Mts." longLatPrecision="20" longitude="84.76833" municipality="Gumda" pageId="31" pageNumber="332" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="paratype">
|
||
<specimenCount id="9D3A109CA3727F62FCF8FD558868DA22" box="[776,820,763,787]" pageId="31" pageNumber="332" type="male">2 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3727F62FCB0FD548837DA22" box="[832,875,762,787]" pageId="31" pageNumber="332" type="female">5 ♀</specimenCount>
|
||
, ditto but
|
||
<geoCoordinate id="EE08BDD2A3727F62FC2FFD558F38DA23" box="[991,1124,762,787]" direction="north" orientation="latitude" pageId="31" pageNumber="332" precision="15" value="28.149721">28°08'59"N</geoCoordinate>
|
||
<geoCoordinate id="EE08BDD2A3727F62FB9BFD548FB2DA23" box="[1131,1262,762,787]" direction="east" orientation="longitude" pageId="31" pageNumber="332" precision="15" value="84.76833">84°46'06"E</geoCoordinate>
|
||
,
|
||
<elevation id="00113C26A3727F62FB0AFD548ECBDA22" box="[1274,1431,762,787]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="31" pageNumber="332" unit="m" value="2650.0" valueMax="2800.0" valueMin="2500.0">
|
||
<quantity id="4CC476F0A3727F62FB0AFD548ECBDA22" box="[1274,1431,762,787]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="31" pageNumber="332" unit="m" value="2650.0" valueMax="2800.0" valueMin="2500.0">2500–2800 m</quantity>
|
||
</elevation>
|
||
, 19+
|
||
<collectingDate id="EFC6043DA3727F62FF31FCB08A64DA07" box="[193,312,798,823]" pageId="31" pageNumber="332" value="2006-05-20">
|
||
<date id="FF82FDD5A3727F62FF31FCB08A64DA07" box="[193,312,798,823]" pageId="31" pageNumber="332" value="2006-05-20">20.V.2006</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="26C9BEC3A3727F62FE84FCB18AB5DA07" box="[372,489,798,823]" pageId="31" pageNumber="332">J. Schmidt</collectorName>
|
||
(cJS,
|
||
<collectionCode id="ED2D43D0A3727F62FDDFFCB1893FDA07" box="[559,611,799,823]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="31" pageNumber="332">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3727F5FFF37FCEC8F96D855" lastPageId="34" lastPageNumber="335" pageId="31" pageNumber="332" type="description">
|
||
<paragraph id="8B83DB15A3727F62FF37FCEC8F0CDA6A" blockId="31.[151,1437,654,1975]" box="[199,1104,834,859]" pageId="31" pageNumber="332">
|
||
<emphasis id="B9480707A3727F62FF37FCEC8A0DDA6B" bold="true" box="[199,337,834,859]" pageId="31" pageNumber="332">Description</emphasis>
|
||
. Habitus as in
|
||
<figureCitation id="1307C790A3727F62FE08FCEC8917DA6B" box="[504,587,834,859]" captionStart="FIGURES 91 – 94" captionStartId="32.[151,264,1940,1963]" captionTargetBox="[265,1303,196,1917]" captionTargetId="figure@32.[261,1305,193,1921]" captionTargetPageId="32" captionText="FIGURES 91 – 94. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 91, T. gupchiensis sp. n., paratype, male. 92, T. brancuccii sp. n., paratype, male. 93, T. manasluensis sp. n., paratype, male. 94, T. cyanescens Ito, 1998, male from Dhabeli." httpUri="https://zenodo.org/record/919654/files/figure.png" pageId="31" pageNumber="332">Fig. 91</figureCitation>
|
||
. Body length
|
||
<quantity id="4CC476F0A3727F62FD17FCEC8833DA6A" box="[743,879,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="5.9" metricValueMin="5.2" pageId="31" pageNumber="332" unit="mm" value="5.55" valueMax="5.9" valueMin="5.2">5.2–5.9 mm</quantity>
|
||
, width
|
||
<quantity id="4CC476F0A3727F62FC32FCED8F10DA6A" box="[962,1100,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="31" pageNumber="332" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3727F62FF37FCC88FD5DAF7" blockId="31.[151,1437,654,1975]" pageId="31" pageNumber="332">Colour: Body piceous, shiny on dorsum, with more or less distinct bluish or greenish tinge on elytra and pronotum; labrum, in some specimens also very narrow margins of pronotum and elytra within lateral gutter paler, reddish brown; palpi, antennae and legs brownish yellow, rarely femora slightly infuscate.</paragraph>
|
||
<paragraph id="8B83DB15A3727F62FF37FC7C8A1ADCD3" blockId="31.[151,1437,654,1975]" pageId="31" pageNumber="332">
|
||
Head: Large, with wide neck (in males, HWmax/PWmax = 0.68–0.71, HWmin/PWmax = 0.55–0.58; in females, these indices 0.71–0.73 and 0.59–0.60, respectively). Eyes markedly convex (in males, HWmax/HWmin = 1.22–1.25, and in females, 1.18–1.25), in lateral view somewhat wide oval, occasionally slightly more elongate. Tempora comparatively long, about third to half as long as eye, flat or slightly convex, somewhat abrupt to neck. Genae wide, slightly less than twice as wide as width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye slightly protruded laterally, forming an obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye approximately as wide as width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow by width of antennomere 3 apically. Labrum moderately concave anteriorly. Clypeus slightly emarginate along anterior margin, moderately convex basally, flatly sloped to apex. Frons and vertex convex.
|
||
<collectingRegion id="49F815F7A3727F62FF67FA958BAADC62" box="[151,246,1339,1362]" country="Myanmar" name="Chin" pageId="31" pageNumber="332">Mentum</collectingRegion>
|
||
and submentum completely fused, submentum on each side with one long seta and ocasionally one short seta laterally of long seta. Left mandible not truncate, blunted at apex. Dorsal microsculpture strongly obliterate, more or less distinct transverse and almost isodiametric meshes evident only laterally behind eyes. Antennae in males extending to the basal edge of pronotum, with antennomeres 4–8 markedly longer than wide, in females slightly shorter.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3727F62FF37FA418A35DE87" blockId="31.[151,1437,654,1975]" pageId="31" pageNumber="332">Pronotum: Slightly transverse (PWmax/PL = 1.33–1.42), widest slightly before anterior third, markedly narrowed posteriad (PWmax/PWmin = 1.20–1.28), with one lateral seta inserted slightly before widest point. Sides rounded anteriorly, nearly straight in basal third and slightly sinuate in short distance before basal angles. Apical margin slightly emarginate, bordered only laterally or almost throughout. Apical angles more or less widely rounded, slightly protruded anteriad. Basal margin nearly straight, in some specimens slightly oblique laterally, vaguely, occasionally very indistinctly, bordered almost throughout, somewhat longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles obtuse, each with a small denticle at apex. Pronotal disc convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally. Lateral gutter narrow, very slightly widened in apical third. Median line fine, interrupted, ended before apical and basal margins, deepened anteriorly. Lateral depressions absent; basal foveae poorly delimited, very small and superficial. Surface comparatively coarsely and irregularly punctate along base and in latero-basal area, with finer obliterate punctation at apical margin. Microsculpture present only at apical angles, consisting of very fine transverse meshes.</paragraph>
|
||
<caption id="DF438B9DA34D7F5DFF67F83A8A01DED8" httpUri="https://zenodo.org/record/919654/files/figure.png" pageId="32" pageNumber="333" targetBox="[265,1303,196,1917]" targetPageId="32">
|
||
<paragraph id="8B83DB15A34D7F5DFF67F83A8A01DED8" blockId="32.[151,1436,1940,2024]" pageId="32" pageNumber="333">
|
||
<emphasis id="B9480707A34D7F5DFF67F83A8A09DE9A" bold="true" box="[151,341,1940,1963]" pageId="32" pageNumber="333">FIGURES 91–94.</emphasis>
|
||
Himalayan species of
|
||
<taxonomicName id="4C3CA096A34D7F5DFDCEF83B8995DE9B" authorityName="Morawitz" authorityYear="1863" box="[574,713,1941,1963]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A34D7F5DFDCEF83B8995DE9B" box="[574,713,1941,1963]" italics="true" pageId="32" pageNumber="333">Trichotichnus</emphasis>
|
||
</taxonomicName>
|
||
subgenus
|
||
<taxonomicName id="4C3CA096A34D7F5DFCC6F83B88C4DE9B" authorityName="Jedlicka" authorityYear="1935" box="[822,920,1941,1963]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A34D7F5DFCC6F83B88C4DE9B" box="[822,920,1941,1963]" italics="true" pageId="32" pageNumber="333">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
, photographs of habitus. 91,
|
||
<taxonomicName id="4C3CA096A34D7F5DFB34F8388E09DE9B" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1220,1365,1941,1963]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="gupchiensis" status="sp. nov.">
|
||
<emphasis id="B9480707A34D7F5DFB34F8388E09DE9B" box="[1220,1365,1941,1963]" italics="true" pageId="32" pageNumber="333">T. gupchiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34D7F5DFAACF83B8ECADE9A" box="[1372,1430,1941,1962]" pageId="32" pageNumber="333" rank="species">
|
||
<emphasis id="B9480707A34D7F5DFAACF83B8ECADE9A" bold="true" box="[1372,1430,1941,1962]" pageId="32" pageNumber="333">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male. 92,
|
||
<taxonomicName id="4C3CA096A34D7F5DFEAEF81B8ABFDEFA" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[350,483,1972,1994]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="brancuccii" status="sp. nov.">
|
||
<emphasis id="B9480707A34D7F5DFEAEF81B8ABFDEFA" box="[350,483,1972,1994]" italics="true" pageId="32" pageNumber="333">T. brancuccii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34D7F5DFE19F81A897DDEF9" box="[489,545,1972,1993]" pageId="32" pageNumber="333" rank="species">
|
||
<emphasis id="B9480707A34D7F5DFE19F81A897DDEF9" bold="true" box="[489,545,1972,1993]" pageId="32" pageNumber="333">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male. 93,
|
||
<taxonomicName id="4C3CA096A34D7F5DFD06F81B88CBDEFA" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[758,919,1972,1994]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov.">
|
||
<emphasis id="B9480707A34D7F5DFD06F81B88CBDEFA" box="[758,919,1972,1994]" italics="true" pageId="32" pageNumber="333">T. manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34D7F5DFC6DF81A8889DEF9" box="[925,981,1972,1993]" pageId="32" pageNumber="333" rank="species">
|
||
<emphasis id="B9480707A34D7F5DFC6DF81A8889DEF9" bold="true" box="[925,981,1972,1993]" pageId="32" pageNumber="333">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male. 94,
|
||
<taxonomicName id="4C3CA096A34D7F5DFB5AF81B8EC4DEFA" authority="Ito, 1998" authorityName="Ito" authorityYear="1998" box="[1194,1432,1972,1994]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="cyanescens">
|
||
<emphasis id="B9480707A34D7F5DFB5AF81B8E68DEFA" box="[1194,1332,1973,1994]" italics="true" pageId="32" pageNumber="333">T. cyanescens</emphasis>
|
||
Ito, 1998
|
||
</taxonomicName>
|
||
, male from Dhabeli.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF438B9DA34C7F5CFF67FBB7886DDDBC" httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="33" pageNumber="334" targetBox="[172,1414,205,1019]" targetPageId="33">
|
||
<paragraph id="8B83DB15A34C7F5CFF67FBB7886DDDBC" blockId="33.[151,1436,1049,1164]" pageId="33" pageNumber="334">
|
||
<emphasis id="B9480707A34C7F5CFF67FBB78A3CDD1F" bold="true" box="[151,352,1049,1072]" pageId="33" pageNumber="334">FIGURES 95–102.</emphasis>
|
||
<taxonomicName id="4C3CA096A34C7F5CFE95FBB489BBDD00" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[357,743,1050,1072]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="gupchiensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34C7F5CFE95FBB48AACDD00" box="[357,496,1050,1072]" italics="true" pageId="33" pageNumber="334">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34C7F5CFE0EFBB4893DDD00" box="[510,609,1050,1072]" italics="true" pageId="33" pageNumber="334">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A34C7F5CFD9FFBB489BBDD00" box="[623,743,1050,1072]" italics="true" pageId="33" pageNumber="334">gupchiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34C7F5CFD1CFBB48879DD1F" box="[748,805,1050,1071]" pageId="33" pageNumber="334" rank="species">
|
||
<emphasis id="B9480707A34C7F5CFD1CFBB48879DD1F" bold="true" box="[748,805,1050,1071]" pageId="33" pageNumber="334">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(95–98, paratypes; 99–102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98–99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95–97, ventral view; 98–99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95–99, 102; B: Figs 100–101, 89).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83DB15A34C7F5CFF37FB1D8AB2DF04" blockId="33.[151,1437,1203,2020]" pageId="33" pageNumber="334">Elytra: In lateral and caudal view convex, in dorsal view oval, elongate, moderately wide (in males, EL/EW = 1.32–1.35; EL/PL = 2.21–2.33; EW/PWmax = 1.19–1.27; in females, these indices 1.31–1.36, 2.27–2.38, 1.23– 1.27, respectively), widest slightly behind middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, usually with a tiny denticle visible from behind. Subapical sinuation very shallow. Sutural angle almost rectangular at apex, with small acute tooth projected posteriad. Basal edge slightly sinuate laterally, forming an obtuse angle with lateral margin. Striae smooth or finely crenulate anteriorly, slightly impressed along entire length. Parascutellar setigerous pore large. Parascutellar striole short, at most as long as distance from parascutellar pore to suture, with apex free. Intervals flat. Interval 3 with a discal setigerous pore at stria 2 at or behind middle. Marginal umbilicate series widely interrupted at middle, with 5–7 pores in anterior group and 8–9 pores in posterior group. Microsculpture strongly obliterate, indistinct on disc, fine transverse meshes evident at apex and on one or two lateral intervals.</paragraph>
|
||
<paragraph id="8B83DB15A34C7F5CFF37F991890FDF68" blockId="33.[151,1437,1203,2020]" box="[199,595,1599,1624]" pageId="33" pageNumber="334">Hindwings: Reduced to tiny scales.</paragraph>
|
||
<paragraph id="8B83DB15A34C7F5CFF37F9CD8EC7DF90" blockId="33.[151,1437,1203,2020]" pageId="33" pageNumber="334">
|
||
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
|
||
<figureCitation id="1307C790A34C7F5CFBA1F9CD8FFFDF4C" box="[1105,1187,1635,1660]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="33" pageNumber="334">Fig. 95</figureCitation>
|
||
) markedly wider than long, narrowed posteriad. Apex of last visible (VII) abdominal sternite in both sexes widely rounded (
|
||
<figureCitation id="1307C790A34C7F5CFAFCF9298ED3DF90" box="[1292,1423,1671,1696]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="33" pageNumber="334">Figs 96–97</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A34C7F5CFF37F9028ADEDE48" blockId="33.[151,1437,1203,2020]" pageId="33" pageNumber="334">Legs: Protibia not sulcate. Profemur with four to seven setae on anterio-ventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) moderately widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Metatarsomeres glabrous on dorsal side (at most with few scattered very shot setae on some tarsomeres), pro- and mesotarsomeres each with few somewhat long setae at their apical margin. Tarsomere 5 with two pairs of ventro-lateral setae. In male, pro- and mesotarsi very moderately widened.</paragraph>
|
||
<paragraph id="8B83DB15A34C7F5CFF37F82D893FDED4" blockId="33.[151,1437,1203,2020]" pageId="33" pageNumber="334">Female genitalia: Laterotergite (hemisternite) symmetrical, with two thick setae apically. Basal stylomere with one strong preapical spine on external margin. Apical stylomere moderately curved, with a peg-like spine at both ventral and dorsal edges of outer margin.</paragraph>
|
||
<paragraph id="8B83DB15A34F7F5FFF37FF398F96D855" blockId="34.[151,1437,151,861]" pageId="34" pageNumber="335">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A34F7F5FFD51FF39886CD980" box="[673,816,151,176]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Figs 98–102</figureCitation>
|
||
) in lateral view arcuate, C-shaped, with ventral margin evenly emarginate in middle portion and with comparatively long and narrow terminal lamella bent ventrad; ventral margin of terminal lamella convex; in dorsal view median lobe almost straight, strongly narrowed to apex in distal half. Terminal lamella (
|
||
<figureCitation id="1307C790A34F7F5FFDFAFEAA8930D82C" box="[522,620,260,285]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Fig. 100</figureCitation>
|
||
) in dorsal view narrow, almost parallel-sided, hardly widened apically. Apical capitulum in lateral view slightly oblique, more strongly prominent ventrally than dorsally (
|
||
<figureCitation id="1307C790A34F7F5FFAFBFE898ED3D870" box="[1291,1423,295,320]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Figs 98–99</figureCitation>
|
||
), oval in caudal view (
|
||
<figureCitation id="1307C790A34F7F5FFE71FEE28ABFD854" box="[385,483,332,357]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Fig. 101</figureCitation>
|
||
). Internal sac without individual spines and distinct spiny patches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA34F7F5FFF37FEC18A08DA25" pageId="34" pageNumber="335" type="etymology">
|
||
<paragraph id="8B83DB15A34F7F5FFF37FEC18E34D8B7" blockId="34.[151,1437,151,861]" box="[199,1384,367,392]" pageId="34" pageNumber="335">
|
||
<emphasis id="B9480707A34F7F5FFF37FEC18A1AD8B8" bold="true" box="[199,326,367,392]" pageId="34" pageNumber="335">Etymology</emphasis>
|
||
. The specific name refers to the Gupchi Danda mountain range where the new species occurs.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A34F7F5FFF37FE3A8A08DA25" blockId="34.[151,1437,151,861]" pageId="34" pageNumber="335">
|
||
<emphasis id="B9480707A34F7F5FFF37FE3A8A06D89D" bold="true" box="[199,346,404,429]" pageId="34" pageNumber="335">Comparison</emphasis>
|
||
. This new species is particularly similar in size and gestalt to
|
||
<taxonomicName id="4C3CA096A34F7F5FFBB3FE3B8FADD89C" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1091,1265,404,428]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34F7F5FFBB3FE3B8F0AD89C" box="[1091,1110,405,428]" italics="true" pageId="34" pageNumber="335">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34F7F5FFB9CFE3B8F21D89C" box="[1132,1149,405,428]" italics="true" pageId="34" pageNumber="335">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34F7F5FFB68FE3B8FADD89C" box="[1176,1265,405,428]" italics="true" pageId="34" pageNumber="335">minutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34F7F5FFB0CFE3A8E1CD89D" box="[1276,1344,404,429]" pageId="34" pageNumber="335" rank="species">
|
||
<emphasis id="B9480707A34F7F5FFB0CFE3A8E1CD89D" bold="true" box="[1276,1344,404,429]" pageId="34" pageNumber="335">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3CA096A34F7F5FFAA1FE3B8A42D8E0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34F7F5FFAA1FE3B8E3AD89C" box="[1361,1382,405,428]" italics="true" pageId="34" pageNumber="335">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34F7F5FFA8AFE3B8ED7D89C" box="[1402,1419,405,428]" italics="true" pageId="34" pageNumber="335">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34F7F5FFF67FE198A42D8E0" box="[151,286,439,464]" italics="true" pageId="34" pageNumber="335">ganeshensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9480707A34F7F5FFED5FE198A30D8E0" bold="true" box="[293,364,439,464]" pageId="34" pageNumber="335">
|
||
<taxonomicNameLabel id="A27BBA7CA34F7F5FFED5FE198A3AD8E0" box="[293,358,439,464]" pageId="34" pageNumber="335" rank="species">sp. n.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3CA096A34F7F5FFE54FE178915D8E0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[420,585,439,464]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34F7F5FFE54FE178AEBD8E0" box="[420,439,441,464]" italics="true" pageId="34" pageNumber="335">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34F7F5FFE39FE178A86D8E0" box="[457,474,441,464]" italics="true" pageId="34" pageNumber="335">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34F7F5FFE02FE198915D8E0" box="[498,585,439,464]" italics="true" pageId="34" pageNumber="335">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34F7F5FFDA0FE1989CCD8E0" box="[592,656,439,464]" pageId="34" pageNumber="335" rank="species">
|
||
<emphasis id="B9480707A34F7F5FFDA0FE1989CCD8E0" bold="true" box="[592,656,439,464]" pageId="34" pageNumber="335">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C3CA096A34F7F5FFD67FE1988B3D8E0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[663,1007,439,464]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="gupchiensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34F7F5FFD67FE19886DD8E0" box="[663,817,439,464]" italics="true" pageId="34" pageNumber="335">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34F7F5FFCB2FE17880FD8E0" box="[834,851,441,464]" italics="true" pageId="34" pageNumber="335">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34F7F5FFC9BFE1988B3D8E0" box="[875,1007,439,464]" italics="true" pageId="34" pageNumber="335">gupchiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34F7F5FFC07FE198F6BD8E0" box="[1015,1079,439,464]" pageId="34" pageNumber="335" rank="species">
|
||
<emphasis id="B9480707A34F7F5FFC07FE198F6BD8E0" bold="true" box="[1015,1079,439,464]" pageId="34" pageNumber="335">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shares with these species many morphological characters including mentum and submentum completely fused, metepisternum slightly wider than long, humerus with more or less distinct denticle at apex, microsculpture on pronotal and elytral discs strongly reduced, and median lobe of aedeagus markedly bent ventrad basally and apically. It is distinctly distinguished from all these species by having protibia not sulcate on dorsal side, terminal lamella of aedeagus much longer (
|
||
<figureCitation id="1307C790A34F7F5FFA9AFDE98A7ADBB4" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Figs 98–100, 102</figureCitation>
|
||
), and apical capitulum more prominent ventrad than dorsad in lateral view (
|
||
<figureCitation id="1307C790A34F7F5FFB8EFDC28FA2DBB4" box="[1150,1278,620,645]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Figs 98–99</figureCitation>
|
||
) and elongate oval in caudal view (
|
||
<figureCitation id="1307C790A34F7F5FFE62FD218AA5DB98" box="[402,505,655,680]" captionStart="FIGURES 95 – 102" captionStartId="33.[151,264,1049,1072]" captionTargetBox="[172,1414,205,1019]" captionTargetId="figure@33.[164,1424,193,1029]" captionTargetPageId="33" captionText="FIGURES 95 – 102. Trichotichnus (Bottchrus) gupchiensis sp. n. (95 – 98, paratypes; 99 – 102, holotype). 95, left metepisternum; 96, apex of last (VII) abdominal sternite, male; 97, same, female; 98 – 99, 102, median lobe of aedeagus; 100, terminal lamella and apical capitulum; 101, apical capitulum. 95 – 97, ventral view; 98 – 99, lateral view; 100, 102, dorsal view; 101, caudal view. Scale bars = 0.25 mm (A: Figs 95 – 99, 102; B: Figs 100 – 101, 89)." httpUri="https://zenodo.org/record/919658/files/figure.png" pageId="34" pageNumber="335">Fig. 101</figureCitation>
|
||
). Among other distinctive characters of
|
||
<taxonomicName id="4C3CA096A34F7F5FFC25FD3F8FF3DB98" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[981,1199,655,680]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="335" phylum="Arthropoda" rank="species" species="gupchiensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A34F7F5FFC25FD3F88B4DB98" box="[981,1000,657,680]" italics="true" pageId="34" pageNumber="335">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A34F7F5FFC0EFD3F8F53DB98" box="[1022,1039,657,680]" italics="true" pageId="34" pageNumber="335">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A34F7F5FFBDBFD218FF3DB98" box="[1067,1199,655,680]" italics="true" pageId="34" pageNumber="335">gupchiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA34F7F5FFB4BFD218FA2DB98" box="[1211,1278,655,680]" pageId="34" pageNumber="335" rank="species">
|
||
<emphasis id="B9480707A34F7F5FFB4BFD218FA2DB98" bold="true" box="[1211,1278,655,680]" pageId="34" pageNumber="335">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, it should be mentioned elytral sutural angle with small acute tooth projecting posteriad, and pronotum with lateral gutter narrow, very slightly widened in apical third, and with basal margin usually nearly straight, occasionally slightly oblique laterally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA34F7F5FFF37FCB18F05DA6C" pageId="34" pageNumber="335" type="distribution">
|
||
<paragraph id="8B83DB15A34F7F5FFF37FCB18F05DA6C" blockId="34.[151,1437,151,861]" pageId="34" pageNumber="335">
|
||
<emphasis id="B9480707A34F7F5FFF37FCB18A05DA08" bold="true" box="[199,345,799,824]" pageId="34" pageNumber="335">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A34F7F5FFE9BFCB18A92DA08" box="[363,462,799,824]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="34" pageNumber="335">Fig. 152</figureCitation>
|
||
,
|
||
<emphasis id="B9480707A34F7F5FFE2DFC8E8AB7DA08" box="[477,491,800,824]" italics="true" pageId="34" pageNumber="335">1</emphasis>
|
||
). Up to today only found on Gupchi Danda mountain range on southern slope of Manaslu Himal, Gorkha District, western
|
||
<collectingRegion id="49F815F7A34F7F5FFD9DFCEA899CDA6D" box="[621,704,836,861]" country="Sri Lanka" name="Central" pageId="34" pageNumber="335">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A34F7F5FFD37FCEA8851DA6D" box="[711,781,836,861]" name="Nepal" pageId="34" pageNumber="335">Nepal</collectingCountry>
|
||
, at altitudes of
|
||
<quantity id="4CC476F0A34F7F5FFC47FCEA8F09DA6C" box="[951,1109,836,860]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="34" pageNumber="335" unit="m" value="2500.0" valueMax="2800.0" valueMin="2200.0">2200–2800 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |