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<document id="46B043ADC84DAE928A552AD6D4C5E028" ID-DOI="10.11646/zootaxa.5123.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6399827" ID-ZooBank="036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1648673534065" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard &amp; Janssen, Ronald" docDate="2022" docId="039487D1FFB7FF9FFFBAFAFF699FFB28" docLanguage="en" docName="zootaxa.5123.1.1.pdf" docOrigin="Zootaxa 5123 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5123.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Granulatocincta pelliscrocodili Harzhauser &amp; Landau &amp; Janssen 2022, nov. sp." docType="treatment" docVersion="4" lastPageNumber="56" masterDocId="FFADFFA9FF82FFA8FF2DFFAB6872FFC0" masterDocTitle="The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera" masterLastPageNumber="172" masterPageNumber="1" pageNumber="54" updateTime="1709053659529" updateUser="karina">
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<mods:title id="06FD44182B126E442F28CC60CE023331">The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera</mods:title>
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<mods:namePart id="4AE0F80910E5FFBC9B420EA7CF5F8040">Landau, Bernard</mods:namePart>
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<taxonomicName id="4C3D4D44FFB7FF9DFFBAFAFF6A7CFAAF" ID-CoL="8YMMV" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[151,526,1364,1391]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="pelliscrocodili" status="sp. nov.">
<emphasis id="B949EAD5FFB7FF9DFFBAFAFF6A7CFAAF" bold="true" box="[151,526,1364,1391]" italics="true" pageId="53" pageNumber="54">Granulatocincta pelliscrocodili</emphasis>
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<figureCitation id="13062A42FFB7FF9DFFBAFAD3688DFA53" box="[151,255,1400,1427]" captionStart="FIGURE 17" captionStartId="52.[151,250,1723,1747]" captionTargetBox="[248,1339,181,1698]" captionTargetId="figure-26@52.[248,1339,181,1698]" captionTargetPageId="52" captionText="FIGURE 17 A1A3. Granulatocincta callim nov. sp., holotype, NHMW 1866/0001/0549, Wiesen (Austria). B1B3. Granulatocincta callim nov.sp.,paralectotype,NHMW 1846/0037/0351b, Wiesen (Austria).C1C3. Granulatocincta callim nov. sp., paralectotype, NHMW 1846/0037/0351, Wiesen (Austria). D1D2. Granulatocincta callim nov. sp., paralectotype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). E. Granulatocincta carasiensis (Jekelius, 1944), MNG P 4965, Soceni (Romania). F1F2. Granulatocincta contorta (Švagrovský, 1958), holotype, Z 3076, Kuzmice (Slovakia). G1G2. Granulatocincta contorta (Švagrovský, 1958), paratype, SNMB Z 3074, Kuzmice (Slovakia). H1H3. Granulatocincta pelliscrocodili nov. sp., holotype, RGM 776 907, Gödet Creek near Tilkikaya (Turkey). I1I2. Granulatocincta pelliscrocodili nov. sp., paratype, RGM 776 906, southwest bank of Gödet Creek near Tilkikaya (Turkey). J1J3. Granulatocincta nataliae (Hoernes &amp; Auinger, 1891), holotype, NHMW 1855/0035/0021, Belgrade/Rakovica (Serbia)." figureDoi="http://doi.org/10.5281/zenodo.6399871" httpUri="https://zenodo.org/record/6399871/files/figure.png" pageId="53" pageNumber="54">Figs 17H</figureCitation>
<figureCitation id="13062A42FFB7FF9DFFD2FA20695BFA53" box="[255,297,1401,1433]" captionStart="FIGURE 1" captionStartId="6.[151,250,1903,1927]" captionTargetBox="[171,1417,176,1880]" captionTargetId="figure-26@6.[171,1417,176,1880]" captionTargetPageId="6" captionText="FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: EisenstadtSopron Basin)." figureDoi="http://doi.org/10.5281/zenodo.6399829" httpUri="https://zenodo.org/record/6399829/files/figure.png" pageId="53" pageNumber="54">
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H
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<figureCitation id="13062A42FFB7FF9DFE04FA206943FA59" box="[297,305,1419,1433]" captionStart="FIGURE 3" captionStartId="12.[151,250,1619,1643]" captionTargetBox="[151,1436,181,1595]" captionTargetId="figure-26@12.[151,1436,181,1595]" captionTargetPageId="12" captionText="FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. Clavatula diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. Clavatula mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. Clavatula bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. Clavatula pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. Clavatula romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. Clavatula interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. Clavatula hirmetzli Kovács &amp; Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. Clavatula irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. Clavatula sophiae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. Clavatula veronicae (Hoernes &amp;Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. Clavatula santhai Kovács &amp; Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. Clavatula camillae (Hoernes &amp;Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes &amp; Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &amp;Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes &amp; Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes &amp; Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici." figureDoi="http://doi.org/10.5281/zenodo.6399835" httpUri="https://zenodo.org/record/6399835/files/figure.png" pageId="53" pageNumber="54">
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, I
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<subScript id="17B93482FFB7FF9DFE64FA206923FA59" attach="right" box="[329,337,1419,1433]" fontSize="6" pageId="53" pageNumber="54">1</subScript>
I
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<figureCitation id="13062A42FFB7FF9DFE45FA206902FA59" box="[360,368,1419,1433]" captionStart="FIGURE 2" captionStartId="7.[151,250,1264,1288]" captionTargetBox="[151,1436,597,1241]" captionTargetId="figure-187@7.[151,1436,597,1241]" captionTargetPageId="7" captionText="FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width." figureDoi="http://doi.org/10.5281/zenodo.6399831" httpUri="https://zenodo.org/record/6399831/files/figure.png" pageId="53" pageNumber="54">
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,
<figureCitation id="13062A42FFB7FF9DFE53FAD369FCFA53" box="[382,398,1400,1427]" captionStart="FIGURE 5" captionStartId="15.[151,250,1999,2023]" captionTargetBox="[151,1436,631,1976]" captionTargetId="figure-214@15.[151,1436,631,1976]" captionTargetPageId="15" captionText="FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae." figureDoi="http://doi.org/10.5281/zenodo.6399839" httpUri="https://zenodo.org/record/6399839/files/figure.png" pageId="53" pageNumber="54">5</figureCitation>
,
<figureCitation id="13062A42FFB7FF9DFEB7FAD369DAFA52" box="[410,424,1400,1426]" captionStart="FIGURE 6" captionStartId="16.[151,250,1938,1962]" captionTargetBox="[151,1436,345,1915]" captionTargetId="figure-106@16.[151,1436,345,1915]" captionTargetPageId="16" captionText="FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, C. = Clavatula, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia)." figureDoi="http://doi.org/10.5281/zenodo.6399841" httpUri="https://zenodo.org/record/6399841/files/figure.png" pageId="53" pageNumber="54">6</figureCitation>
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<taxonomicName id="4C3D4D44FFB7FF9DFFBAFA166A7CFA15" ID-CoL="8743R" authority="(Michelotti, 1847)" baseAuthorityName="Michelotti" baseAuthorityYear="1847" box="[151,526,1469,1493]" class="Gastropoda" family="Clavatulidae" genus="Clavatula" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB7FF9DFFBAFA16693EFA15" box="[151,332,1469,1493]" italics="true" pageId="53" pageNumber="54">Clavatula sotterii</emphasis>
(
<bibRefCitation id="EFAC4B36FFB7FF9DFE71FA166A75FA15" author="Michelotti, G." box="[348,519,1469,1493]" pageId="53" pageNumber="54" pagination="1 - 408" refId="ref138690" refString="Michelotti, G. (1847) Description des fossiles des terrains miocenes de l'Italie septentrionale. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem, Series 2, 3 (2), 1 - 408. [https: // books. google. be / books? id = aQVqpL 58 HrsC &amp; ots = rhEPJ _ C 6 hy &amp; dq = Michelotti % 2 C % 20 G. % 091847 % 09 Description % 20 des % 20 fossiles % 20 de s % 20 terrains % 20 mioc % C 3 % A 8 nes % 20 de % 20 l'Italie % 20 septentrionale. &amp; lr &amp; pg = PP 5 # v = onepage &amp; q = Michelotti, % 20 G. % 091847 % 09 Description % 20 des % 2]" type="journal article" year="1847">Michelotti, 1847</bibRefCitation>
)
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<treatmentCitation id="0A9C10D6FFB7FF9DFD05FA166B5AFA15" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[552,808,1469,1493]" page="287" pageId="53" pageNumber="54" year="2013">
<bibRefCitation id="EFAC4B36FFB7FF9DFD05FA166B5AFA15" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[552,808,1469,1493]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFD52FA156ADDFA15" box="[639,687,1469,1493]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. 2013: 287
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, pl. 48, figs 712 [
<emphasis id="B949EAD5FFB7FF9DFCD9FA156C68FA15" box="[1012,1050,1470,1493]" italics="true" pageId="53" pageNumber="54">non</emphasis>
<taxonomicName id="4C3D4D44FFB7FF9DFB0EFA166975FA34" ID-CoL="8YMMX" authority="(Michelotti, 1847)" baseAuthorityName="Michelotti" baseAuthorityYear="1847" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB7FF9DFB0EFA166D69FA15" box="[1059,1307,1469,1493]" italics="true" pageId="53" pageNumber="54">Granulatocincta sotterii</emphasis>
(
<bibRefCitation id="EFAC4B36FFB7FF9DFA06FA16688DFA34" author="Michelotti, G." pageId="53" pageNumber="54" pagination="1 - 408" refId="ref138690" refString="Michelotti, G. (1847) Description des fossiles des terrains miocenes de l'Italie septentrionale. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem, Series 2, 3 (2), 1 - 408. [https: // books. google. be / books? id = aQVqpL 58 HrsC &amp; ots = rhEPJ _ C 6 hy &amp; dq = Michelotti % 2 C % 20 G. % 091847 % 09 Description % 20 des % 20 fossiles % 20 de s % 20 terrains % 20 mioc % C 3 % A 8 nes % 20 de % 20 l'Italie % 20 septentrionale. &amp; lr &amp; pg = PP 5 # v = onepage &amp; q = Michelotti, % 20 G. % 091847 % 09 Description % 20 des % 2]" type="journal article" year="1847">Michelotti, 1847</bibRefCitation>
)
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB7FF9DFFBAF9896AF8F864" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8B8236C7FFB7FF9DFFBAF9896D19F8F8" blockId="53.[151,1437,1569,1993]" pageId="53" pageNumber="54">
<emphasis id="B949EAD5FFB7FF9DFFBAF9896934F9FB" bold="true" box="[151,326,1569,1596]" pageId="53" pageNumber="54">Type material.</emphasis>
<materialsCitation id="3B553C9AFFB7FF9DFE63F9896B89F9A6" collectionCode="MD" country="Turkey" location="Godet Creek" municipality="Tilkikaya" pageId="53" pageNumber="54" specimenCode="RGM 776" specimenCount="1" typeStatus="holotype">
<typeStatus id="54868865FFB7FF9DFE63F98969C9F9FC" box="[334,443,1570,1596]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB9B9EBCFFB7FF9DFEEBF9896A4AF9FC" box="[454,568,1570,1596]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="53" pageNumber="54" type="Museum">RGM 776</specimenCode>
907, southwest bank of
<location id="8EE2601CFFB7FF9DFC60F9896BAFF9FC" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFC60F9896BAFF9FC" box="[845,989,1570,1596]" country="Turkey" municipality="Tilkikaya" name="Godet Creek" pageId="53" pageNumber="54">Gödet Creek</location>
near
<collectingMunicipality id="6BE6ACBDFFB7FF9DFB31F9896CF4F9FC" box="[1052,1158,1570,1596]" pageId="53" pageNumber="54">Tilkikaya</collectingMunicipality>
(
<collectingCountry id="F32A7657FFB7FF9DFBBBF9896C9EF9FD" box="[1174,1260,1570,1597]" name="Turkey" pageId="53" pageNumber="54">Turkey</collectingCountry>
), SL:
<quantity id="4CC59B22FFB7FF9DFA1CF9896DEAF9FC" box="[1329,1432,1570,1596]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="53" pageNumber="54" unit="mm" value="17.9">17.9 mm</quantity>
,
<collectionCode id="ED2CAE02FFB7FF9DFFBAF9ED68BAF9A0" box="[151,200,1606,1632]" country="Germany" name="Museum Donaueschingen" pageId="53" pageNumber="54">MD</collectionCode>
:
<quantity id="4CC59B22FFB7FF9DFFFCF9ED695AF9A0" box="[209,296,1606,1632]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="53" pageNumber="54" unit="mm" value="8.6">8.6 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FFB7FF9DFEEFF9ED6AE9F9A0" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[450,667,1606,1632]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFD36F9EC6A20F9A0" box="[539,594,1606,1632]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 8ac); figs 17H
<subScript id="17B93482FFB7FF9DFCE4F9F36BA3F9A6" attach="left" box="[969,977,1624,1638]" fontSize="6" pageId="53" pageNumber="54">1</subScript>
H
<subScript id="17B93482FFB7FF9DFCDEF9F36B89F9A6" attach="left" box="[1011,1019,1624,1638]" fontSize="6" pageId="53" pageNumber="54">3</subScript>
</materialsCitation>
.
<materialsCitation id="3B553C9AFFB7FF9DFB25F9ED6ACCF944" collectionCode="RGM" collectorName="Tilkikaya, SL" country="Turkey" location="Godet Creek" pageId="53" pageNumber="54" specimenCode="RGM 794" specimenCount="1" typeStatus="paratype">
<typeStatus id="54868865FFB7FF9DFB25F9ED6C0BF9A0" box="[1032,1145,1606,1632]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
:
<specimenCode id="DB9B9EBCFFB7FF9DFBAFF9ED6C83F9A0" box="[1154,1265,1606,1632]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="53" pageNumber="54" type="Museum">RGM 794</specimenCode>
567, southwest bank of
<location id="8EE2601CFFB7FF9DFFDCF9C1690CF944" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFFDCF9C1690CF944" box="[241,382,1642,1668]" country="Turkey" name="Godet Creek" pageId="53" pageNumber="54">Gödet Creek</location>
near
<collectorName id="26C85311FFB7FF9DFE94F9C16A26F945" box="[441,596,1642,1669]" pageId="53" pageNumber="54">Tilkikaya, SL</collectorName>
:
<quantity id="4CC59B22FFB7FF9DFD7AF9C16ACCF944" box="[599,702,1642,1669]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="53" pageNumber="54" unit="mm" value="15.7">15.7 mm</quantity>
</materialsCitation>
,
<collectionCode id="ED2CAE02FFB7FF9DFDE5F9C16A8BF944" box="[712,761,1642,1668]" country="China" name="Museum of Zoology" pageId="53" pageNumber="54">MG</collectionCode>
:
<quantity id="4CC59B22FFB7FF9DFC2EF9C16B28F944" box="[771,858,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="53" pageNumber="54" unit="mm" value="7.7">7.7 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FFB7FF9DFCDBF9C16CA3F944" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[1014,1233,1642,1668]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFB7DF9C06CF5F944" box="[1104,1159,1642,1668]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 7ac);
<materialsCitation id="3B553C9AFFB7FF9DFFBAF9256B26F968" box="[151,852,1678,1705]" collectionCode="NHMW" collectorName="Lale River, SL" country="Turkey" location="Akbodazi" pageId="53" pageNumber="54" specimenCode="NHMW 1847" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9B9EBCFFB7FF9DFFBAF9256940F969" box="[151,306,1678,1705]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="53" pageNumber="54" type="Museum">NHMW 1847</specimenCode>
/0058/1503,
<location id="8EE2601CFFB7FF9DFE93F9256A5CF969" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFE93F9256A5CF969" box="[446,558,1678,1705]" country="Turkey" name="Akbodazi" pageId="53" pageNumber="54">Akboðazi</location>
,
<collectorName id="26C85311FFB7FF9DFD14F9256A91F968" box="[569,739,1678,1704]" pageId="53" pageNumber="54">Lale River, SL</collectorName>
:
<quantity id="4CC59B22FFB7FF9DFDC3F9256B26F968" box="[750,852,1678,1704]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="53" pageNumber="54" unit="mm" value="16.9">16.9 mm</quantity>
</materialsCitation>
,
<collectionCode id="ED2CAE02FFB7FF9DFC4DF9256BE3F968" box="[864,913,1678,1704]" country="Germany" name="Museum Donaueschingen" pageId="53" pageNumber="54">MD</collectionCode>
:
<quantity id="4CC59B22FFB7FF9DFCB1F9256B86F968" box="[924,1012,1678,1705]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="53" pageNumber="54" unit="mm" value="8.5">8.5 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FFB7FF9DFBB8F9256D06F968" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[1173,1396,1678,1704]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFBDDF9246D5BF968" box="[1264,1321,1678,1704]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 9ab);
<materialsCitation id="3B553C9AFFB7FF9DFE69F9196C77F90C" box="[324,1029,1714,1741]" collectionCode="NHMW" collectorName="Lale River, SL" country="Turkey" location="Akbodazi" pageId="53" pageNumber="54" specimenCode="NHMW 1847" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9B9EBCFFB7FF9DFE69F91969ACF90D" box="[324,478,1714,1741]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="53" pageNumber="54" type="Museum">NHMW 1847</specimenCode>
/0058/1504,
<location id="8EE2601CFFB7FF9DFD41F9196AAEF90D" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFD41F9196AAEF90D" box="[620,732,1714,1741]" country="Turkey" name="Akbodazi" pageId="53" pageNumber="54">Akboðazi</location>
,
<collectorName id="26C85311FFB7FF9DFDC5F9196BE1F90C" box="[744,915,1714,1740]" pageId="53" pageNumber="54">Lale River, SL</collectorName>
: 13.0 mm
</materialsCitation>
,
<collectionCode id="ED2CAE02FFB7FF9DFB3FF9196C31F90C" box="[1042,1091,1714,1740]" country="Germany" name="Museum Donaueschingen" pageId="53" pageNumber="54">MD</collectionCode>
:
<quantity id="4CC59B22FFB7FF9DFB62F9196CDAF90C" box="[1103,1192,1714,1741]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="53" pageNumber="54" unit="mm" value="6.5">6.5 mm</quantity>
, illustrated
<bibRefCitation id="EFAC4B36FFB7FF9DFA01F919688CF930" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFAA5F91868C6F930" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 10ac);
<materialsCitation id="3B553C9AFFB7FF9DFEF3F97D6DEAF930" box="[478,1432,1750,1777]" collectionCode="RGM" country="Turkey" location="Godet Creek" municipality="Tilkikaya" pageId="53" pageNumber="54" specimenCode="RGM 776" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9B9EBCFFB7FF9DFEF3F97D6A3CF930" box="[478,590,1750,1776]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="53" pageNumber="54" type="Museum">RGM 776</specimenCode>
906, southwest bank of
<location id="8EE2601CFFB7FF9DFC71F97D6B98F930" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFC71F97D6B98F930" box="[860,1002,1750,1776]" country="Turkey" municipality="Tilkikaya" name="Godet Creek" pageId="53" pageNumber="54">Gödet Creek</location>
near
<collectingMunicipality id="6BE6ACBDFFB7FF9DFB08F97D6CFDF930" box="[1061,1167,1750,1776]" pageId="53" pageNumber="54">Tilkikaya</collectingMunicipality>
(
<collectingCountry id="F32A7657FFB7FF9DFBB0F97D6C81F931" box="[1181,1267,1750,1777]" name="Turkey" pageId="53" pageNumber="54">Turkey</collectingCountry>
), SL:
<quantity id="4CC59B22FFB7FF9DFA19F97D6DEAF930" box="[1332,1432,1750,1776]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="53" pageNumber="54" unit="mm" value="11.3">11.3 mm</quantity>
</materialsCitation>
,
<collectionCode id="ED2CAE02FFB7FF9DFFBAF95168BAF8D4" box="[151,200,1786,1812]" country="Germany" name="Museum Donaueschingen" pageId="53" pageNumber="54">MD</collectionCode>
:
<quantity id="4CC59B22FFB7FF9DFFF9F951695EF8D4" box="[212,300,1786,1813]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="53" pageNumber="54" unit="mm" value="5.6">5.6 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FFB7FF9DFEE0F9516ADFF8D4" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[461,685,1786,1812]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFD04F9506A10F8D4" box="[553,610,1786,1812]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 11ac), figs 16I
<subScript id="17B93482FFB7FF9DFCC7F8A76B80F8DA" attach="left" box="[1002,1010,1804,1818]" fontSize="6" pageId="53" pageNumber="54">1</subScript>
I
<subScript id="17B93482FFB7FF9DFB27F8A76C60F8DA" attach="left" box="[1034,1042,1804,1818]" fontSize="6" pageId="53" pageNumber="54">2</subScript>
;
<materialsCitation id="3B553C9AFFB7FF9DFB33F9516A03F8F8" collectionCode="RGM" collectorName="Tilkikaya, SL" country="Turkey" location="Godet Creek" pageId="53" pageNumber="54" specimenCode="RGM 776" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9B9EBCFFB7FF9DFB33F9516CE2F8D4" box="[1054,1168,1786,1812]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="53" pageNumber="54" type="Museum">RGM 776</specimenCode>
905, southwest bank of
<location id="8EE2601CFFB7FF9DFFBAF8B56954F8F8" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFFBAF8B56954F8F8" box="[151,294,1822,1848]" country="Turkey" name="Godet Creek" pageId="53" pageNumber="54">Gödet Creek</location>
near
<collectorName id="26C85311FFB7FF9DFE4EF8B56A72F8F8" box="[355,512,1822,1848]" pageId="53" pageNumber="54">Tilkikaya, SL</collectorName>
:
<quantity id="4CC59B22FFB7FF9DFD26F8B56A03F8F8" box="[523,625,1822,1848]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="53" pageNumber="54" unit="mm" value="21.6">21.6 mm</quantity>
</materialsCitation>
,
<collectionCode id="ED2CAE02FFB7FF9DFD50F8B56ADCF8F8" box="[637,686,1822,1848]" country="Germany" name="Museum Donaueschingen" pageId="53" pageNumber="54">MD</collectionCode>
:
<quantity id="4CC59B22FFB7FF9DFD95F8B56B62F8F8" box="[696,784,1822,1849]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="53" pageNumber="54" unit="mm" value="9.5">9.5 mm</quantity>
, illustrated in
<bibRefCitation id="EFAC4B36FFB7FF9DFC9DF8B56CFCF8F8" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[944,1166,1822,1848]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB7FF9DFB26F8B46C31F8F8" box="[1035,1091,1822,1848]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013
</bibRefCitation>
, pl. 48, figs 12ac).
</paragraph>
<paragraph id="8B8236C7FFB7FF9DFFEAF8E96CFDF89C" blockId="53.[151,1437,1569,1993]" box="[199,1167,1857,1885]" pageId="53" pageNumber="54">
<materialsCitation id="3B553C9AFFB7FF9DFFEAF8E96CF9F89C" box="[199,1163,1857,1885]" country="Turkey" county="South-west" location="Tilkikaya" municipality="Godet River" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Karaman" typeStatus="holotype">
<emphasis id="B949EAD5FFB7FF9DFFEAF8E96914F89B" bold="true" box="[199,358,1857,1884]" pageId="53" pageNumber="54">
<typeStatus id="54868865FFB7FF9DFFEAF8E96970F89C" box="[199,258,1858,1884]" pageId="53" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
<collectingCounty id="62E34E4BFFB7FF9DFE41F8E96998F89C" box="[364,490,1858,1884]" pageId="53" pageNumber="54">South-west</collectingCounty>
bank of
<collectingMunicipality id="6BE6ACBDFFB7FF9DFD60F8E96AA5F89C" box="[589,727,1858,1884]" pageId="53" pageNumber="54">Gödet River</collectingMunicipality>
at
<location id="8EE2601CFFB7FF9DFDD5F8E96B10F89C" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFDD5F8E96B10F89C" box="[760,866,1858,1884]" country="Turkey" county="South-west" municipality="Godet River" name="Tilkikaya" pageId="53" pageNumber="54" stateProvince="Karaman">Tilkikaya</location>
(
<collectingCountry id="F32A7657FFB7FF9DFC5CF8E96BB5F89D" box="[881,967,1858,1885]" name="Turkey" pageId="53" pageNumber="54">Turkey</collectingCountry>
),
<collectingRegion id="49F9F825FFB7FF9DFCF7F8E96C33F89C" box="[986,1089,1858,1884]" country="Turkey" name="Karaman" pageId="53" pageNumber="54">Karaman</collectingRegion>
<location id="8EE2601CFFB7FF9DFB65F8E96CF9F89C" LSID="urn:lsid:plazi:treatment:039487D1FFB7FF9FFFBAFAFF699FFB28:8EE2601CFFB7FF9DFB65F8E96CF9F89C" box="[1096,1163,1858,1884]" country="Turkey" county="South-west" municipality="Godet River" name="Basin" pageId="53" pageNumber="54" stateProvince="Karaman">Basin</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8236C7FFB7FF9DFFEAF8CD6B6BF840" blockId="53.[151,1437,1569,1993]" box="[199,793,1894,1920]" pageId="53" pageNumber="54">
<emphasis id="B949EAD5FFB7FF9DFFEAF8CD691DF840" bold="true" box="[199,367,1894,1920]" pageId="53" pageNumber="54">
<typeStatus id="54868865FFB7FF9DFFEAF8CD6970F840" box="[199,258,1894,1920]" pageId="53" pageNumber="54">Type</typeStatus>
stratum.
</emphasis>
Silt and clay of the Týrtar Formation.
</paragraph>
<paragraph id="8B8236C7FFB7FF9DFFEAF8226AF8F864" blockId="53.[151,1437,1569,1993]" box="[199,650,1929,1956]" pageId="53" pageNumber="54">
<emphasis id="B949EAD5FFB7FF9DFFEAF822688FF863" bold="true" box="[199,253,1929,1955]" pageId="53" pageNumber="54">Age.</emphasis>
Middle Miocene, late Serravallian.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB7FF9DFFEAF8056D19F808" box="[199,1387,1966,1993]" pageId="53" pageNumber="54" type="etymology">
<paragraph id="8B8236C7FFB7FF9DFFEAF8056D19F808" blockId="53.[151,1437,1569,1993]" box="[199,1387,1966,1993]" pageId="53" pageNumber="54">
<emphasis id="B949EAD5FFB7FF9DFFEAF805693CF808" bold="true" box="[199,334,1966,1992]" pageId="53" pageNumber="54">Etymology.</emphasis>
From
<emphasis id="B949EAD5FFB7FF9DFEB5F80569A7F808" box="[408,469,1966,1992]" italics="true" pageId="53" pageNumber="54">pellis</emphasis>
(Latin: skin) and
<emphasis id="B949EAD5FFB7FF9DFDB3F8056B61F808" box="[670,787,1966,1992]" italics="true" pageId="53" pageNumber="54">crocodilus</emphasis>
(Latin: crocodile), referring to the peculiar sculpture.
</paragraph>
</subSubSection>
<caption id="DF42664FFFB4FF9EFFBAF91C6D7BF80B" ID-DOI="http://doi.org/10.5281/zenodo.6399873" ID-Zenodo-Dep="6399873" httpUri="https://zenodo.org/record/6399873/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1719,1743]" targetBox="[151,1436,181,1695]" targetPageId="54">
<paragraph id="8B8236C7FFB4FF9EFFBAF91C6D7BF80B" blockId="54.[151,1437,1719,1995]" pageId="54" pageNumber="55">
<emphasis id="B949EAD5FFB4FF9EFFBAF91C694DF90F" bold="true" box="[151,319,1719,1743]" pageId="54" pageNumber="55">FIGURE 18 A.</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFE64F91C6AEEF90F" authority="granulatocincta" box="[329,668,1719,1743]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFE64F91C6AEEF90F" box="[329,668,1719,1743]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), paralectotype, SNSB-BSPG AS VII 2057, Enzesfeld (Austria).
<emphasis id="B949EAD5FFB4FF9EFE40F97069F6F933" bold="true" box="[365,388,1755,1779]" pageId="54" pageNumber="55">B.</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFEA1F9706AAFF933" authority="granulatocincta" box="[396,733,1755,1779]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFEA1F9706AAFF933" box="[396,733,1755,1779]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), lectotype, SNSB-BSPG AS VII 2056, Enzesfeld (Austria).
<emphasis id="B949EAD5FFB4FF9EFE44F95469DDF8D7" bold="true" box="[361,431,1791,1821]" pageId="54" pageNumber="55">
C
<subScript id="17B93482FFB4FF9EFE56F8A469F0F8DD" attach="left" box="[379,386,1807,1821]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
C
<subScript id="17B93482FFB4FF9EFE8FF8A469DBF8DD" attach="right" box="[418,425,1807,1821]" fontSize="6" pageId="54" pageNumber="55">2</subScript>
.
</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFE98F9546B77F8D7" authority="granulatocincta" box="[437,773,1791,1815]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFE98F9546B77F8D7" box="[437,773,1791,1815]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), NHMW 2021/0126/0006, Gainfarn (Austria).
<emphasis id="B949EAD5FFB4FF9EFE29F8886938F8FB" bold="true" box="[260,330,1827,1857]" pageId="54" pageNumber="55">
D
<subScript id="17B93482FFB4FF9EFE3BF898696FF881" attach="left" box="[278,285,1843,1857]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
D
<subScript id="17B93482FFB4FF9EFE10F8986936F881" attach="right" box="[317,324,1843,1857]" fontSize="6" pageId="54" pageNumber="55">2</subScript>
.
</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFE7CF8886AD4F8FB" authority="granulatocincta" box="[337,678,1827,1851]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFE7CF8886AD4F8FB" box="[337,678,1827,1851]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), 3T; M.61.4366, Buda-hegy, Sámsonháza (Hungary).
<emphasis id="B949EAD5FFB4FF9EFE3BF8EC692BF89F" bold="true" box="[278,345,1863,1893]" pageId="54" pageNumber="55">
E
<subScript id="17B93482FFB4FF9EFE0BF8FC695FF8A5" attach="left" box="[294,301,1879,1893]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
E
<subScript id="17B93482FFB4FF9EFE66F8FC6920F8A5" attach="left" box="[331,338,1879,1893]" fontSize="6" pageId="54" pageNumber="55">3</subScript>
.
</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFE45F8EC6AB2F89F" authority="granulatocincta" box="[360,704,1863,1887]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFE45F8EC6AB2F89F" box="[360,704,1863,1887]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), NHMW 1846/0028/0008, Enzesfeld (Austria).
<emphasis id="B949EAD5FFB4FF9EFFD2F8C0694DF843" bold="true" box="[255,319,1899,1929]" pageId="54" pageNumber="55">
F
<subScript id="17B93482FFB4FF9EFE23F8D06967F849" attach="left" box="[270,277,1915,1929]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
F
<subScript id="17B93482FFB4FF9EFE1FF8D0694BF849" attach="right" box="[306,313,1915,1929]" fontSize="6" pageId="54" pageNumber="55">3</subScript>
.
</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFE68F8C06AE7F843" authority="granulatocincta" box="[325,661,1899,1923]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFE68F8C06AE7F843" box="[325,661,1899,1923]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria).
<emphasis id="B949EAD5FFB4FF9EFFBAF8246892F867" bold="true" box="[151,224,1935,1965]" pageId="54" pageNumber="55">
G
<subScript id="17B93482FFB4FF9EFF86F83468C0F86D" attach="left" box="[171,178,1951,1965]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
G
<subScript id="17B93482FFB4FF9EFFFEF83468A8F86D" attach="left" box="[211,218,1951,1965]" fontSize="6" pageId="54" pageNumber="55">3</subScript>
.
</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFFC4F8246A48F867" authority="granulatocincta" box="[233,570,1935,1959]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFFC4F8246A48F867" box="[233,570,1935,1959]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), NHMW 1855/0045/0486, Gainfarn (Austria).
<emphasis id="B949EAD5FFB4FF9EFA7EF8246DEEF867" bold="true" box="[1363,1436,1935,1965]" pageId="54" pageNumber="55">
H
<subScript id="17B93482FFB4FF9EFA4AF8346D1CF86D" attach="left" box="[1383,1390,1951,1965]" fontSize="6" pageId="54" pageNumber="55">1</subScript>
H
<subScript id="17B93482FFB4FF9EFAA2F8346DE4F86D" attach="left" box="[1423,1430,1951,1965]" fontSize="6" pageId="54" pageNumber="55">3</subScript>
.
</emphasis>
<taxonomicName id="4C3D4D44FFB4FF9EFFBAF8186995F80B" authority="granulatocincta" box="[151,487,1971,1995]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="granulatocincta">
<emphasis id="B949EAD5FFB4FF9EFFBAF8186995F80B" box="[151,487,1971,1995]" italics="true" pageId="54" pageNumber="55">Granulatocincta granulatocincta</emphasis>
</taxonomicName>
(Münster in Goldfuss, 1841), NHMW 1855/0045/0136, Steinebrunn (Austria).
</paragraph>
</caption>
<subSubSection id="C327654CFFB5FF9FFFEAFF3D6B94FEDC" pageId="55" pageNumber="56" type="diagnosis">
<paragraph id="8B8236C7FFB5FF9FFFEAFF3D6B94FEDC" blockId="55.[151,1437,150,1257]" pageId="55" pageNumber="56">
<emphasis id="B949EAD5FFB5FF9FFFEAFF3D6930FF70" bold="true" box="[199,322,150,176]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
<taxonomicName id="4C3D4D44FFB5FF9FFE64FF3D6A29FF70" box="[329,603,150,176]" pageId="55" pageNumber="56">
<emphasis id="B949EAD5FFB5FF9FFE64FF3D6A70FF70" box="[329,514,150,176]" italics="true" pageId="55" pageNumber="56">Granulatocincta</emphasis>
species
</taxonomicName>
of small size, paucispiral protoconch, spire coeloconoid, broad buccinoid profile and barrel-shaped last whorl. Finely sculptured, early whorls with sinuous axial riblets crossed by spiral cords. Later whorls with low cords of primary to tertiary strength bearing vaguely quadratic tubercles axially arranged to form sinuous riblets. Siphonal fasciole not developed. Short siphonal canal.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB5FF9FFFEAFE8D6ACCFCF8" pageId="55" pageNumber="56" type="description">
<paragraph id="8B8236C7FFB5FF9FFFEAFE8D6ACCFCF8" blockId="55.[151,1437,150,1257]" pageId="55" pageNumber="56">
<emphasis id="B949EAD5FFB5FF9FFFEAFE8D692BFE80" bold="true" box="[199,345,294,320]" pageId="55" pageNumber="56">Description.</emphasis>
Shell small, solid, broad buccinoid weakly coeloconoid spire; apical angle ~4550°. Protoconch worn, but paucispiral. Teleoconch of up to eight whorls initially tall and flat-sided, later rather depressed. Early teleoconch whorls bearing close-set, sinuous axial riblets, crossed by weak spiral cords. Fourth teleoconch whorl rapidly broadening, becoming squatter, tripartite, with broad subsutural collar covered in fine spirals. In some specimens subsutural collar swollen, making spire weakly gradate. On penultimate whorl secondary weakly beaded spiral cords intercalated above and below suprasutural cord. Suture narrowly impressed, linear. Last whorl 6570% of total height, barrel-shaped. Subsutural collar flattened or swollen, bearing close-set, narrow cords. Subsutural ramp weakly concave, bearing close-set, narrow spiral cords. One cord placed at mid-ramp, beaded, comma-shaped riblets. Shoulder hardly developed. Sculpture at mid-whorl and base consisting of flattened cords of alternating strength, tertiary threads present in some specimens, separated by narrow grooves, bearing low subquadrate tubercles. Tubercles aligned axially forming vague sinuous axial riblets; siphonal fasciole not developed. Aperture ovate; anal sinus moderately wide and deep, symmetrically U-shaped, with apex at beaded cord on ramp; siphonal canal moderately short, wide, shallowly notched at tip. Columella strongly excavated in upper third, straight below, moderately twisted to the left, smooth. Columellar and parietal callus strongly thickened, sharply delimited, forming broad callus rim and weak pseudumbilical chink.
</paragraph>
</subSubSection>
<subSubSection id="C327654CFFB5FF9FFFEAFCE9699FFB28" pageId="55" pageNumber="56" type="discussion">
<paragraph id="8B8236C7FFB5FF9FFFEAFCE96BC3FBBC" blockId="55.[151,1437,150,1257]" pageId="55" pageNumber="56">
<emphasis id="B949EAD5FFB5FF9FFFEAFCE9693EFC9C" bold="true" box="[199,332,834,860]" pageId="55" pageNumber="56">Discussion.</emphasis>
This species was tentatively treated as
<taxonomicName id="4C3D4D44FFB5FF9FFC2DFCE96C90FC9C" authority="(Michelotti, 1847)" baseAuthorityName="Michelotti" baseAuthorityYear="1847" box="[768,1250,834,860]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB5FF9FFC2DFCE96C7CFC9C" box="[768,1038,834,860]" italics="true" pageId="55" pageNumber="56">Granulatocincta sotterii</emphasis>
(
<bibRefCitation id="EFAC4B36FFB5FF9FFB30FCE96CA8FC9C" author="Michelotti, G." box="[1053,1242,834,860]" pageId="55" pageNumber="56" pagination="1 - 408" refId="ref138690" refString="Michelotti, G. (1847) Description des fossiles des terrains miocenes de l'Italie septentrionale. Natuurkundige Verhandelingen van de Hollandsche Maatschappij der Wetenschappen te Haarlem, Series 2, 3 (2), 1 - 408. [https: // books. google. be / books? id = aQVqpL 58 HrsC &amp; ots = rhEPJ _ C 6 hy &amp; dq = Michelotti % 2 C % 20 G. % 091847 % 09 Description % 20 des % 20 fossiles % 20 de s % 20 terrains % 20 mioc % C 3 % A 8 nes % 20 de % 20 l'Italie % 20 septentrionale. &amp; lr &amp; pg = PP 5 # v = onepage &amp; q = Michelotti, % 20 G. % 091847 % 09 Description % 20 des % 2]" type="journal article" year="1847">Michelotti, 1847</bibRefCitation>
)
</taxonomicName>
by
<bibRefCitation id="EFAC4B36FFB5FF9FFA26FCE96897FC40" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." pageId="55" pageNumber="56" pagination="3 - 576" refId="ref137659" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576" type="journal article" year="2013">
Landau
<emphasis id="B949EAD5FFB5FF9FFA48FCE86DEFFC9C" box="[1381,1437,834,860]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
(2013)
</bibRefCitation>
.
<taxonomicName id="4C3D4D44FFB5FF9FFFDCFCCD6A70FC40" baseAuthorityName="Michelotti" baseAuthorityYear="1847" box="[241,514,870,896]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB5FF9FFFDCFCCD6A70FC40" box="[241,514,870,896]" italics="true" pageId="55" pageNumber="56">Granulatocincta sotterii</emphasis>
</taxonomicName>
is a common species in the Tortonian of
<collectingCountry id="F32A7657FFB5FF9FFCF0FCCD6C62FC40" box="[989,1040,870,896]" name="Italy" pageId="55" pageNumber="56">Italy</collectingCountry>
and has been frequently described and illustrated (
<bibRefCitation id="EFAC4B36FFB5FF9FFE66FC216A34FC64" author="Montanaro, E." box="[331,582,906,932]" pageId="55" pageNumber="56" pagination="115 - 191" refId="ref139628" refString="Montanaro, E. (1937) Studi monografici sulla malacologia miocenica modenese. Parte I. I mollusci tortoniani di Montegibbio. Palaeontographia Italica, New Series 7, 37, 115 - 191." type="journal article" year="1937">Montanaro 1937: 117</bibRefCitation>
, pl. 5, figs 218;
<bibRefCitation id="EFAC4B36FFB5FF9FFC3AFC216C46FC64" author="Ruggieri, C. &amp; Davoli, F." box="[791,1076,906,932]" pageId="55" pageNumber="56" pagination="41 - 85" refId="ref142914" refString="Ruggieri, C. &amp; Davoli, F. (1984) Malacofauna di Casa Nova Calisese (Sogliano, Forli), Palaeontographia Italica, 73, 41 - 85." type="journal article" year="1984">Ruggieri &amp; Davoli 1984</bibRefCitation>
, pl. 4, figs 1, 6, 1314, 1819,
<bibRefCitation id="EFAC4B36FFB5FF9FFFBAFC056954FC08" author="Davoli, F." box="[151,294,942,968]" pageId="55" pageNumber="56" pagination="27 - 109" refId="ref131183" refString="Davoli, F. (1990) La collezione di &quot; Fossili Miocenici di Sogliano &quot; di Ludovico Foresti: revisione ed illustrazione. Atti della Societa dei Naturalisti e Matematici di Modena, 121, 27 - 109." type="journal article" year="1990">Davoli 1990</bibRefCitation>
, pl. 8, figs 28). These Italian specimens of
<taxonomicName id="4C3D4D44FFB5FF9FFC30FC046BFDFC08" baseAuthorityName="Michelotti" baseAuthorityYear="1847" box="[797,911,943,969]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB5FF9FFC30FC046BFDFC08" box="[797,911,943,969]" italics="true" pageId="55" pageNumber="56">G. sotterii</emphasis>
</taxonomicName>
show some variability in sculpture, but are all characterized by a prominent subsutural collar with large beads, a prominent central spiral cord of strong beads and a prominently beaded shoulder, but lack secondary and tertiary spiral cords. Thus,
<taxonomicName id="4C3D4D44FFB5FF9FFB16FC5D6DEFFBD0" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1083,1437,1014,1040]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="pelliscrocodili" status="sp. nov.">
<emphasis id="B949EAD5FFB5FF9FFB16FC5D6DEFFBD0" box="[1083,1437,1014,1040]" italics="true" pageId="55" pageNumber="56">Granulatocincta pelliscrocodili</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFB5FF9FFFBAFBB06882FBF5" box="[151,240,1051,1077]" pageId="55" pageNumber="56" rank="species">nov. sp.</taxonomicNameLabel>
differs significantly from
<taxonomicName id="4C3D4D44FFB5FF9FFD0EFBB06AE5FBF4" baseAuthorityName="Michelotti" baseAuthorityYear="1847" box="[547,663,1051,1077]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="sotterii">
<emphasis id="B949EAD5FFB5FF9FFD0EFBB06AE5FBF4" box="[547,663,1051,1077]" italics="true" pageId="55" pageNumber="56">G. sotterii</emphasis>
</taxonomicName>
in its delicate and complex sculpture.
<taxonomicName id="4C3D4D44FFB5FF9FFB77FBB16D11FBF4" authority="Harzhauser &amp; Landau &amp; Janssen, 2022" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[1114,1379,1050,1076]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="callim" status="sp. nov.">
<emphasis id="B949EAD5FFB5FF9FFB77FBB16D11FBF4" box="[1114,1379,1050,1076]" italics="true" pageId="55" pageNumber="56">Granulatocincta callim</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57AEFFB5FF9FFA40FBB068C5FB99" pageId="55" pageNumber="56" rank="species">nov. sp.</taxonomicNameLabel>
differs in the same features from
<taxonomicName id="4C3D4D44FFB5FF9FFD11FB946B72FB98" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[572,768,1086,1113]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="pelliscrocodili">
<emphasis id="B949EAD5FFB5FF9FFD11FB946B72FB98" box="[572,768,1086,1113]" italics="true" pageId="55" pageNumber="56">G. pelliscrocodili</emphasis>
</taxonomicName>
. Within
<taxonomicName id="4C3D4D44FFB5FF9FFC49FB956C6FFB98" authorityName="Harzhauser &amp; Landau &amp; Janssen" authorityYear="2022" box="[868,1053,1086,1112]" class="Gastropoda" family="Clavatulidae" genus="Granulatocincta" kingdom="Animalia" order="Neogastropoda" pageId="55" pageNumber="56" phylum="Mollusca" rank="genus">
<emphasis id="B949EAD5FFB5FF9FFC49FB956C6FFB98" box="[868,1053,1086,1112]" italics="true" pageId="55" pageNumber="56">Granulatocincta</emphasis>
</taxonomicName>
, this species is outstanding by its fragmented primary sculpture and reduced central spiral cord of beads.
</paragraph>
<paragraph id="8B8236C7FFB5FF9FFFEAFB2D6C04FB60" blockId="55.[151,1437,150,1257]" box="[199,1142,1158,1185]" pageId="55" pageNumber="56">
<emphasis id="B949EAD5FFB5FF9FFFEAFB2D69DAFB60" bold="true" box="[199,424,1158,1184]" pageId="55" pageNumber="56">Paleoenvironment.</emphasis>
Inner neritic brackish water environments (
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Landau
<emphasis id="B949EAD5FFB5FF9FFCC3FB2C6C54FB60" box="[1006,1062,1158,1184]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
<paragraph id="8B8236C7FFB5FF9FFFEAFB01699FFB28" blockId="55.[151,1437,150,1257]" pageId="55" pageNumber="56">
<emphasis id="B949EAD5FFB5FF9FFFEAFB016A70FB04" bold="true" box="[199,514,1194,1220]" pageId="55" pageNumber="56">Proto-Mediterranean Sea:</emphasis>
Serravallian (middle Miocene):
<emphasis id="B949EAD5FFB5FF9FFC5CFB006C50FB04" box="[881,1058,1195,1220]" italics="true" pageId="55" pageNumber="56">
<collectingRegion id="49F9F825FFB5FF9FFC5CFB006BA9FB04" box="[881,987,1195,1220]" country="Turkey" name="Karaman" pageId="55" pageNumber="56">Karaman</collectingRegion>
Basin
</emphasis>
: Akboðazi, Seyithasan, Tilkikaya (
<collectingCountry id="F32A7657FFB5FF9FFFB2FB656881FB29" box="[159,243,1230,1257]" name="Turkey" pageId="55" pageNumber="56">Turkey</collectingCountry>
) (
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Landau
<emphasis id="B949EAD5FFB5FF9FFE48FB6469EFFB28" box="[357,413,1230,1256]" italics="true" pageId="55" pageNumber="56">et al.</emphasis>
2013
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>