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<document id="52D554D7F127D0C95EE0B07B9C740359" ID-DOI="10.5281/zenodo.5334580" ID-ISSN="0374-1036" ID-Zenodo-Dep="5334580" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630175634131" checkinUser="felipe" docAuthor="Fikáček, Martin, Delgado, Juan A. &amp; Gentili, Elio" docDate="2012" docId="0394878EFFA7FFC44282FE89FE5E854B" docLanguage="en" docName="ActaEntMusNatPra.2012.52.suppl.107-130.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Georissus (Neogeorissus) maritimus Fikáček &amp; Delgado &amp; Gentili 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="112" masterDocId="FFADFFF6FFA5FFC14308FFCAFFAB8018" masterDocTitle="The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae)" masterLastPageNumber="130" masterPageNumber="107" pageNumber="109" updateTime="1699175334189" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="1A6D46B221E7F080333B288C42C4D403">
<mods:title id="5B5E29C248C170F5989C79B367709027">The Hydrophiloid beetles of Socotra Island (Coleoptera: Georissidae, Hydrophilidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="2F33215962AAADD5A0233EFD8192084A">Fikáček, Martin</mods:namePart>
<mods:affiliation id="57ED5631A8393606A10E7D45C746C1EC">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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</mods:role>
<mods:namePart id="974B41B9B329B39F640C3A5B5D034D40">Delgado, Juan A.</mods:namePart>
<mods:affiliation id="FE2AF2B5F76025427D6F0670A34A34A3">Departamento de Zoología, Facultad de Biología, Universidad de Murcia, 30100, Murcia, Spain; e-mail: jdelgado @ um. es</mods:affiliation>
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<mods:namePart id="132CDEFA42D9B254FB32A40B8722F6DA">Gentili, Elio</mods:namePart>
<mods:affiliation id="A8B5297618EA8BE6B03C81154AD68733">Via San Gottardo 37, I- 21030 Varese-Rasa, Italy; e-mail: elio. gentili. 32 @ alice. it</mods:affiliation>
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<mods:title id="1FEC799C57E485A07226FD63A4908C23">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="8D00B91A8C0EFD0AA39C7400A40C3622">2012</mods:date>
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<treatment id="0394878EFFA7FFC44282FE89FE5E854B" ID-DOI="http://doi.org/10.5281/zenodo.5413313" ID-Zenodo-Dep="5413313" LSID="urn:lsid:plazi:treatment:0394878EFFA7FFC44282FE89FE5E854B" httpUri="http://treatment.plazi.org/id/0394878EFFA7FFC44282FE89FE5E854B" lastPageId="5" lastPageNumber="112" pageId="2" pageNumber="109">
<subSubSection id="C3276513FFA7FFC34282FE89FCDF8143" box="[394,884,322,347]" pageId="2" pageNumber="109" type="nomenclature">
<paragraph id="8B823698FFA7FFC34282FE89FCDF8143" blockId="2.[394,884,322,347]" box="[394,884,322,347]" pageId="2" pageNumber="109">
<heading id="D0CA81F4FFA7FFC34282FE89FCDF8143" bold="true" box="[394,884,322,347]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="109" reason="3">
<emphasis id="B949EA8AFFA7FFC34282FE89FCDF8143" bold="true" box="[394,884,322,347]" pageId="2" pageNumber="109">
<taxonomicName id="4C3D4D1BFFA7FFC34282FE89FCB28143" ID-CoL="3FS2M" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[394,793,322,347]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov." subGenus="Neogeorissus">
<emphasis id="B949EA8AFFA7FFC34282FE89FE5D8143" bold="true" box="[394,502,323,347]" italics="true" pageId="2" pageNumber="109">Georissus</emphasis>
(
<emphasis id="B949EA8AFFA7FFC3410DFE89FD338143" bold="true" box="[517,664,323,347]" italics="true" pageId="2" pageNumber="109">Neogeorissus</emphasis>
)
<emphasis id="B949EA8AFFA7FFC341A0FE89FCB28143" bold="true" box="[680,793,323,347]" italics="true" pageId="2" pageNumber="109">maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA7FFC34017FE89FCDF8143" box="[799,884,323,347]" pageId="2" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA7FFC34118FEABFD45816C" box="[528,750,353,372]" pageId="2" pageNumber="109" type="description">
<paragraph id="8B823698FFA7FFC34118FEABFD45816C" blockId="2.[528,750,353,372]" box="[528,750,353,372]" pageId="2" pageNumber="109">
(
<figureCitation id="13062A1DFFA7FFC3411EFEABFDC3816C" box="[534,616,353,372]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="3.[113,152,1540,1559]" captionStartId-1="4.[153,192,1033,1052]" captionTargetBox-0="[113,1086,219,1451]" captionTargetBox-1="[266,1016,226,1012]" captionTargetId-0="figure-13@3.[113,1086,219,1452]" captionTargetId-1="figure-178@4.[262,1016,219,1015]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figs. 16. Georissus maritimus sp. nov. (paratypes). 1 pronotum and elytra in dorsal view; 2 head, dorsal view; 34 detail of pronotum (3 dorsal view; 4 dorsolateral view); 5 elytron, dorsolateral view; 6 abdomen." captionText-1="Figs. 78. Genitalia of Socotran endemic Georissus species. 7 G. maritimus sp. nov. (a, b: two different ventral views of the same aedeagus; c: lateral view); 8 G. nemo sp. nov., ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.5334584" figureDoi-1="http://doi.org/10.5281/zenodo.5334586" httpUri-0="https://zenodo.org/record/5334584/files/figure.png" httpUri-1="https://zenodo.org/record/5334586/files/figure.png" pageId="2" pageNumber="109">Figs. 17</figureCitation>
,
<figureCitation id="13062A1DFFA7FFC34179FEABFD4C816C" box="[625,743,353,372]" captionStart="Figs" captionStartId="10.[151,190,1491,1510]" captionTargetBox="[153,1125,221,1476]" captionTargetId="figure-12@10.[147,1128,216,1478]" captionTargetPageId="10" captionText="Figs. 1522.Socotran endemic Georissus species and their habitats. 1516 G. maritimus sp. nov.(15 dorsal view; 16 lateral view). 1718 G. nemo sp. nov. (17 dorsal view; 18 lateral view). 1920 the sand tongue 3 km NE of Shuab, type locality of G. maritimus sp. nov. (19 general view of the habitat; 20 satellite view showing the position of the locality); 2122 the stream at Arhar, type locality of G. nemo sp. nov. (21 general view of the habitat; 22 satellite view showing the position of the locality)." figureDoi="http://doi.org/10.5281/zenodo.5334592" httpUri="https://zenodo.org/record/5334592/files/figure.png" pageId="2" pageNumber="109">1516, 1920</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA7FFC3439FFE41FBB18210" pageId="2" pageNumber="109" type="materials_examined">
<paragraph id="8B823698FFA7FFC3439FFE41FC198186" blockId="2.[151,1127,394,521]" box="[151,946,394,414]" pageId="2" pageNumber="109">
<emphasis id="B949EA8AFFA7FFC3439FFE41FEBB8186" bold="true" box="[151,272,394,414]" pageId="2" pageNumber="109">
<typeStatus id="5486883AFFA7FFC3439FFE41FF6F8186" box="[151,196,395,414]" pageId="2" pageNumber="109">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32A7608FFA7FFC3421DFE41FEF88186" box="[277,339,395,414]" name="Yemen" pageId="2" pageNumber="109">Yemen</collectingCountry>
,
<collectingCountry id="F32A7608FFA7FFC34253FE41FE738186" box="[347,472,395,414]" name="Socotra Island" pageId="2" pageNumber="109">Socotra Island</collectingCountry>
, ca.
<quantity id="4CC59B7DFFA7FFC342F6FE41FD828186" box="[510,553,395,414]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="109" unit="km" value="3.0">3 km</quantity>
NE of Shuab,
<geoCoordinate id="EE09505FFFA7FFC341A0FE41FD558186" box="[680,766,394,414]" degrees="12" direction="north" minutes="34.1" orientation="latitude" pageId="2" pageNumber="109" precision="92" value="12.568334">12°34.1N</geoCoordinate>
<geoCoordinate id="EE09505FFFA7FFC3400CFE41FCF08186" box="[772,859,394,414]" degrees="53" direction="east" minutes="23.9" orientation="longitude" pageId="2" pageNumber="109" precision="92" value="53.398335">53°23.9E</geoCoordinate>
,
<quantity id="4CC59B7DFFA7FFC3406CFE41FC2E8186" box="[868,901,395,414]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="109" unit="m" value="3.0">3 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8B823698FFA7FFC3439FFE6CFBB18210" blockId="2.[151,1127,394,521]" pageId="2" pageNumber="109">
<emphasis id="B949EA8AFFA7FFC3439FFE6CFEB681A1" bold="true" box="[151,285,421,441]" pageId="2" pageNumber="109">Type material.</emphasis>
<materialsCitation id="3B553CC5FFA7FFC3422BFE6CFD5C81F6" collectingDate="2012-06-20" collectingDateMax="2012-06-21" collectingDateMin="2012-06-20" collectionCode="NMPC" collectorName="Socotra Expedition &amp; J. Bezdek &amp; J. Hajek &amp; V. Hula &amp; P. Kment &amp; I. Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="3" latitude="12.568334" location="Socotra island" longLatPrecision="129" longitude="53.398335" pageId="2" pageNumber="109" specimenCount="1" typeStatus="holotype">
<typeStatus id="5486883AFFA7FFC3422BFE6CFEDD81AF" box="[291,374,422,441]" pageId="2" pageNumber="109" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED2CAE5DFFA7FFC34297FE6CFE4A81A1" box="[415,481,422,441]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="109">NMPC</collectionCode>
):
<collectingCountry id="F32A7608FFA7FFC342FBFE6CFDED81A1" box="[499,582,422,441]" name="Yemen" pageId="2" pageNumber="109">YEMEN</collectingCountry>
:
<location id="8EE26043FFA7FFC34147FE6CFD6181A0" LSID="urn:lsid:plazi:treatment:0394878EFFA7FFC44282FE89FE5E854B:8EE26043FFA7FFC34147FE6CFD6181A0" box="[591,714,421,441]" country="Yemen" latitude="12.568334" longLatPrecision="129" longitude="53.398335" name="Socotra island" pageId="2" pageNumber="109">Socotra island</location>
/ ca.
<locationDeviation id="9A302FBAFFA7FFC341F3FE6CFC3581A0" box="[763,926,421,441]" location="Shuab" pageId="2" pageNumber="109">
<quantity id="4CC59B7DFFA7FFC341F3FE6CFC8C81A0" box="[763,807,421,441]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="109" unit="km" value="3.0">3 km</quantity>
NE of Shuab
</locationDeviation>
/ mangrove,
<taxonomicName id="4C3D4D1BFFA7FFC3471AFE6CFF7E81CB" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="109" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B949EA8AFFA7FFC3471AFE6CFF7E81CB" italics="true" pageId="2" pageNumber="109">Avicennia marina</emphasis>
</taxonomicName>
/
<geoCoordinate id="EE09505FFFA7FFC343EFFE0AFE9581CC" box="[231,318,447,468]" degrees="12" direction="north" minutes="34.1" orientation="latitude" pageId="2" pageNumber="109" precision="92" value="12.568334">12°34.1N</geoCoordinate>
<geoCoordinate id="EE09505FFFA7FFC3424CFE0AFE3081CC" box="[324,411,447,468]" degrees="53" direction="east" minutes="23.9" orientation="longitude" pageId="2" pageNumber="109" precision="92" value="53.398335">53°23.9E</geoCoordinate>
;
<elevation id="0010D1ABFFA7FFC342ADFE0AFE5E81CB" box="[421,501,448,468]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="109" unit="m" value="3.0">
<quantity id="4CC59B7DFFA7FFC342ADFE0AFE6C81CC" box="[421,455,448,468]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="109" unit="m" value="3.0">3 m</quantity>
a.s.l.
</elevation>
/ saline,
<date id="FF831058FFA7FFC3414CFE0AFD1081CB" box="[580,699,448,467]" pageId="2" pageNumber="109" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">
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</date>
//
<collectorName id="26C8534EFFA7FFC341DCFE0AFCD281CB" box="[724,889,448,467]" pageId="2" pageNumber="109">Socotra Expedition</collectorName>
2012 /
<collectorName id="26C8534EFFA7FFC340B3FE0BFBBA81CB" box="[955,1041,448,468]" pageId="2" pageNumber="109">J. Bezděk</collectorName>
,
<collectorName id="26C8534EFFA7FFC34712FE0BFBC881CB" box="[1050,1123,448,468]" pageId="2" pageNumber="109">J. Hájek</collectorName>
,
<collectorName id="26C8534EFFA7FFC3439FFE11FF7081F6" box="[151,219,475,494]" pageId="2" pageNumber="109">V. Hula</collectorName>
, /
<collectorName id="26C8534EFFA7FFC343F8FE11FEEA81F6" box="[240,321,475,494]" pageId="2" pageNumber="109">P. Kment</collectorName>
,
<collectorName id="26C8534EFFA7FFC34242FE11FE6381F6" box="[330,456,475,494]" pageId="2" pageNumber="109">I. Malenovský</collectorName>
, /
<collectorName id="26C8534EFFA7FFC342D5FE11FDE481F6" box="[477,591,475,494]" pageId="2" pageNumber="109">J. Niedobová</collectorName>
&amp;
<collectorName id="26C8534EFFA7FFC34163FE11FD6081F6" box="[619,715,475,494]" pageId="2" pageNumber="109">L. Purchart</collectorName>
leg.
</materialsCitation>
.
<typeStatus id="5486883AFFA7FFC3400BFE11FCF081F5" box="[771,859,475,494]" pageId="2" pageNumber="109" type="paratype">PARATYPES</typeStatus>
: 4 JJ,
<specimenCount id="9D3BFD11FFA7FFC340AAFE11FC6981F6" box="[930,962,469,494]" pageId="2" pageNumber="109" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D3BFD11FFA7FFC340C5FE11FBBE81F6" box="[973,1045,475,494]" pageId="2" pageNumber="109" type="generic">41 spec.</specimenCount>
(
<collectionCode id="ED2CAE5DFFA7FFC34728FE11FBC881F6" box="[1056,1123,475,494]" pageId="2" pageNumber="109">BMNH</collectionCode>
,
<collectionCode id="ED2CAE5DFFA7FFC3439FFE3CFF738211" box="[151,216,502,521]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="2" pageNumber="109">IRSNB</collectionCode>
,
<collectionCode id="ED2CAE5DFFA7FFC343E9FE3CFE8C8211" box="[225,295,502,521]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="2" pageNumber="109" type="Museum">NHMW</collectionCode>
, NMPC, KSEM,
<specimenCount id="9D3BFD11FFA7FFC342B4FE3FFE528211" box="[444,505,501,521]" pageId="2" pageNumber="109" type="generic">5 spec.</specimenCount>
in DNA grade kept in
<collectionCode id="ED2CAE5DFFA7FFC341B2FE3CFD508211" box="[698,763,502,521]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="109">NMPC</collectionCode>
): same label data as the
<typeStatus id="5486883AFFA7FFC340C0FE3FFBBE8210" box="[968,1045,501,520]" pageId="2" pageNumber="109" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA7FFC4439FFDEBFEDB814B" lastPageId="5" lastPageNumber="112" pageId="2" pageNumber="109" type="description">
<paragraph id="8B823698FFA7FFC3439FFDEBFDDE82A1" blockId="2.[151,1127,545,1561]" pageId="2" pageNumber="109">
<emphasis id="B949EA8AFFA7FFC3439FFDEBFE898221" bold="true" box="[151,290,545,569]" pageId="2" pageNumber="109">Description.</emphasis>
Body narrowly elongate, weakly convex in lateral view. Body length
<quantity id="4CC59B7DFFA7FFC34722FDEBFF5E8240" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="2" pageNumber="109" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.20 1.60 mm</quantity>
(
<typeStatus id="5486883AFFA7FFC3420CFD8BFEC88241" box="[260,355,577,601]" pageId="2" pageNumber="109" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA7FFC34261FD8BFE678241" box="[361,460,577,601]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="109" unit="mm" value="1.4">1.40 mm</quantity>
), width of head
<quantity id="4CC59B7DFFA7FFC3417FFD8BFCB98240" box="[631,786,577,601]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="2" pageNumber="109" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.300.40 mm</quantity>
(
<typeStatus id="5486883AFFA7FFC34028FD8BFCD48241" box="[800,895,577,601]" pageId="2" pageNumber="109" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA7FFC3408DFD8BFC438241" box="[901,1000,577,601]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="109" unit="mm" value="0.35">0.35 mm</quantity>
), maximum width of pronotum
<quantity id="4CC59B7DFFA7FFC3426DFDABFE548260" box="[357,511,609,633]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="2" pageNumber="109" unit="mm" value="0.525" valueMax="0.6" valueMin="0.45">0.450.60 mm</quantity>
(
<typeStatus id="5486883AFFA7FFC34105FDABFDC78261" box="[525,620,609,633]" pageId="2" pageNumber="109" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA7FFC3417AFDABFD7F8261" box="[626,724,609,633]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="109" unit="mm" value="0.55">0.55 mm</quantity>
), maximum width of elytra
<quantity id="4CC59B7DFFA7FFC340F4FDABFF6B8280" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.5" metricValueMin="6.5" pageId="2" pageNumber="109" unit="mm" value="0.75" valueMax="0.85" valueMin="0.65">0.650.85 mm</quantity>
(
<typeStatus id="5486883AFFA7FFC343C7FD4BFE858281" box="[207,302,641,665]" pageId="2" pageNumber="109" type="holotype">holotype</typeStatus>
<quantity id="4CC59B7DFFA7FFC3423CFD4BFE338281" box="[308,408,641,665]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="109" unit="mm" value="0.75">0.75 mm</quantity>
). Coloration of elytra brown to piceous, pronotum dark brown with paler anterior portion, head piceous to black.
</paragraph>
<paragraph id="8B823698FFA7FFC343BDFD0BFDE983E0" blockId="2.[151,1127,545,1561]" pageId="2" pageNumber="109">
Head (
<figureCitation id="13062A1DFFA7FFC343F4FD0BFE9682C1" box="[252,317,705,729]" captionStart="Figs" captionStartId="3.[113,152,1540,1559]" captionTargetBox="[113,1086,219,1451]" captionTargetId="figure-13@3.[113,1086,219,1452]" captionTargetPageId="3" captionText="Figs. 16. Georissus maritimus sp. nov. (paratypes). 1 pronotum and elytra in dorsal view; 2 head, dorsal view; 34 detail of pronotum (3 dorsal view; 4 dorsolateral view); 5 elytron, dorsolateral view; 6 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334584" httpUri="https://zenodo.org/record/5334584/files/figure.png" pageId="2" pageNumber="109">Fig. 2</figureCitation>
). Clypeus weakly convex, with few weakly developed granules anteromedially, anterior margin with elevated bead without distinct granules; each side of clypeus with deep pit anterolaterally; lateral portion of clypeus posteriorly divided from mesal portion by high ridge without granules, ridges bent laterad and bearing more or less distinct granules more anteriorly. Frons elevated above clypeus, bearing large elongate rhomboid plate mesally reaching to posterior part of frons; laterally of mesal plate with high and wide ridge on each side; posterior part of median rhomboid and longitudinal ridges connected by group or large and low granules posteriorly; frons with two pairs of deep pits, one situated anteromesally, second laterally. Clypeus and median portion of frons with weak mesh-like microsculpture. Antennal club with three antennomeres.
</paragraph>
<paragraph id="8B823698FFA7FFC343BDFBCBFCA28521" blockId="2.[151,1127,545,1561]" pageId="2" pageNumber="109">
Prothorax (
<figureCitation id="13062A1DFFA7FFC34222FBCBFE3B8401" box="[298,400,1025,1049]" captionStart="Figs" captionStartId="3.[113,152,1540,1559]" captionTargetBox="[113,1086,219,1451]" captionTargetId="figure-13@3.[113,1086,219,1452]" captionTargetPageId="3" captionText="Figs. 16. Georissus maritimus sp. nov. (paratypes). 1 pronotum and elytra in dorsal view; 2 head, dorsal view; 34 detail of pronotum (3 dorsal view; 4 dorsolateral view); 5 elytron, dorsolateral view; 6 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334584" httpUri="https://zenodo.org/record/5334584/files/figure.png" pageId="2" pageNumber="109">Figs. 34</figureCitation>
). Pronotum 1.15× as wide as long, with maximum width at midlength; lateral portions rounded, without distinct projections. Median portion with two closely situated submedian ridges throughout, delimiting shallow median groove; each lateral lobe divided from mesal portion by longitudinal ridge falling into separate granules posteriorly. Anterior half of pronotum without granules, only bearing small pits intermixed with sparsely arranged punctures. Posterior half of pronotum without distinct bulges, with sparsely arranged large but low granules throughout the surface. Pronotum with two pairs of deep pits, one anteriorly and the other posteriorly of lateral lobe. Whole surface between granules with weak meashlike microsculpture. Ventral portion of prothorax with extremelly large antennal grooves, otherwise corresponding with
<taxonomicName id="4C3D4D1BFFA7FFC342D3FAE8FCAE8521" authority="(Rossi, 1794)" baseAuthorityName="Rossi" baseAuthorityYear="1794" box="[475,773,1313,1337]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="109" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B949EA8AFFA7FFC342D3FAE8FDC08521" box="[475,619,1314,1337]" italics="true" pageId="2" pageNumber="109">G. crenulatus</emphasis>
(Rossi, 1794)
</taxonomicName>
.
</paragraph>
<paragraph id="8B823698FFA7FFC543BDFA8BFC3884B2" blockId="2.[151,1127,545,1561]" lastBlockId="4.[151,1128,1138,1578]" lastPageId="4" lastPageNumber="111" pageId="2" pageNumber="109">
Mesothorax. Elytra (
<figureCitation id="13062A1DFFA7FFC3429AFA8BFE5C8541" box="[402,503,1345,1369]" captionStart="Figs" captionStartId="3.[113,152,1540,1559]" captionTargetBox="[113,1086,219,1451]" captionTargetId="figure-13@3.[113,1086,219,1452]" captionTargetPageId="3" captionText="Figs. 16. Georissus maritimus sp. nov. (paratypes). 1 pronotum and elytra in dorsal view; 2 head, dorsal view; 34 detail of pronotum (3 dorsal view; 4 dorsolateral view); 5 elytron, dorsolateral view; 6 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334584" httpUri="https://zenodo.org/record/5334584/files/figure.png" pageId="2" pageNumber="109">Figs. 1, 5</figureCitation>
) combined 1.25× as long as wide, 1.75× as long as pronotum; base of elytra ca. as wide as maximum width of pronotum, maximum width of elytra in anterior third, then weakly narrowing posteriad to posterior fifth, apex strongly narrowed. Elytra completely devoid of granules, interstices without microsculpture. Elytral suture and intervals 17 evently convex, interval 8 reduced, interval 9 slightly elevated; lateral portion with suboval depression at anterior third across intervals 89; all intervals more convex posteriorly than anteriorly. Elytral series 111 regular, serial punctures small but sharply impressed. Lateral-most portion of elytron declined, hence elytra laterally without clear projections. Median pentagonal protuberance of mesoventrite flat, with distinct pits.
</paragraph>
<caption id="DF426610FFA6FFC24379F9CEFB8A862A" ID-DOI="http://doi.org/10.5281/zenodo.5334584" ID-Zenodo-Dep="5334584" httpUri="https://zenodo.org/record/5334584/files/figure.png" pageId="3" pageNumber="110" startId="3.[113,152,1540,1559]" targetBox="[113,1086,219,1451]" targetPageId="3">
<paragraph id="8B823698FFA6FFC24379F9CEFB8A862A" blockId="3.[113,1089,1540,1587]" pageId="3" pageNumber="110">
Figs. 16.
<taxonomicName id="4C3D4D1BFFA6FFC243C3F9CFFED7860F" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[203,380,1541,1559]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="110" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA6FFC243C3F9CFFED7860F" box="[203,380,1541,1559]" italics="true" pageId="3" pageNumber="110">Georissus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA6FFC24289F9CFFE6F8600" box="[385,452,1541,1560]" pageId="3" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
(paratypes). 1 pronotum and elytra in dorsal view; 2 head, dorsal view; 34 detail of pronotum (3 dorsal view; 4 dorsolateral view); 5 elytron, dorsolateral view; 6 abdomen.
</paragraph>
</caption>
<caption id="DF426610FFA1FFC54391FBC3FCEF842E" ID-DOI="http://doi.org/10.5281/zenodo.5334586" ID-Zenodo-Dep="5334586" httpUri="https://zenodo.org/record/5334586/files/figure.png" pageId="4" pageNumber="111" startId="4.[153,192,1033,1052]" targetBox="[266,1016,226,1012]" targetPageId="4">
<paragraph id="8B823698FFA1FFC54391FBC3FCEF842E" blockId="4.[151,1126,1033,1079]" pageId="4" pageNumber="111">
Figs. 78. Genitalia of Socotran endemic
<taxonomicName id="4C3D4D1BFFA1FFC542F7FBC0FD058404" box="[511,686,1033,1052]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="undefined-7">
<emphasis id="B949EA8AFFA1FFC542F7FBC0FDFF8404" box="[511,596,1034,1052]" italics="true" pageId="4" pageNumber="111">Georissus</emphasis>
species. 7
</taxonomicName>
<taxonomicName id="4C3D4D1BFFA1FFC541CCFBC0FC9D8404" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[708,822,1034,1052]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA1FFC541CCFBC0FC9D8404" box="[708,822,1034,1052]" italics="true" pageId="4" pageNumber="111">G. maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA1FFC54033FBC3FCD48404" box="[827,895,1033,1052]" pageId="4" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
(a, b: two different ventral views of the same aedeagus; c: lateral view); 8
<taxonomicName id="4C3D4D1BFFA1FFC54133FBEEFD28842E" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[571,643,1060,1078]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="111" phylum="Arthropoda" rank="species" species="nemo" status="sp. nov.">
<emphasis id="B949EA8AFFA1FFC54133FBEEFD28842E" box="[571,643,1060,1078]" italics="true" pageId="4" pageNumber="111">G. nemo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA1FFC54180FBEEFD66842F" box="[648,717,1060,1079]" pageId="4" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
, ventral view.
</paragraph>
</caption>
<paragraph id="8B823698FFA1FFC543BDFB78FDA88532" blockId="4.[151,1128,1138,1578]" pageId="4" pageNumber="111">Metathorax. Metaventrite ca. 1.2× as long as mesoventrite, flat, with few scattered indistinct granules along posterior margin and four deep pits along anterior margin; lateral portions divided from mesal part by longitudinal ridges. Metathoracic wings absent in specimens examined for this character (n=5).</paragraph>
<paragraph id="8B823698FFA1FFC543BDFAF8FBA385B2" blockId="4.[151,1128,1138,1578]" pageId="4" pageNumber="111">
Abdomen (
<figureCitation id="13062A1DFFA1FFC54225FAF8FEC78552" box="[301,364,1330,1354]" captionStart="Figs" captionStartId="3.[113,152,1540,1559]" captionTargetBox="[113,1086,219,1451]" captionTargetId="figure-13@3.[113,1086,219,1452]" captionTargetPageId="3" captionText="Figs. 16. Georissus maritimus sp. nov. (paratypes). 1 pronotum and elytra in dorsal view; 2 head, dorsal view; 34 detail of pronotum (3 dorsal view; 4 dorsolateral view); 5 elytron, dorsolateral view; 6 abdomen." figureDoi="http://doi.org/10.5281/zenodo.5334584" httpUri="https://zenodo.org/record/5334584/files/figure.png" pageId="4" pageNumber="111">Fig. 6</figureCitation>
) gradually narrowing posteriad. Ventrite 1 flat mesally, with lateral portion declines and divided by ridge; median portion with sparsely arranged punctures, without granules; posterior margin of ventrite 1 with pair of large tubercles facing enlarged granules of ventrite 2. Ventrites 34 with weakly developed granules along anterior margin.
</paragraph>
<paragraph id="8B823698FFA1FFC443BDFA78FEDB814B" blockId="4.[151,1128,1138,1578]" lastBlockId="5.[113,1090,219,1363]" lastPageId="5" lastPageNumber="112" pageId="4" pageNumber="111">
Male genitalia (
<figureCitation id="13062A1DFFA1FFC54268FA78FE0F85D2" box="[352,420,1458,1482]" captionStart="Figs" captionStartId="4.[153,192,1033,1052]" captionTargetBox="[266,1016,226,1012]" captionTargetId="figure-178@4.[262,1016,219,1015]" captionTargetPageId="4" captionText="Figs. 78. Genitalia of Socotran endemic Georissus species. 7 G. maritimus sp. nov. (a, b: two different ventral views of the same aedeagus; c: lateral view); 8 G. nemo sp. nov., ventral view." figureDoi="http://doi.org/10.5281/zenodo.5334586" httpUri="https://zenodo.org/record/5334586/files/figure.png" pageId="4" pageNumber="111">Fig. 7</figureCitation>
). Aedeagus
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long. Parameres 1.6× as long as phallobase, their combined width narrower than maximum width of phallobase; parameres nearly parallel-sided throughout except subbasally, subbasally slightly constricted; apex rounded at outer margin, bluntly pointed mesally; apex with numerous large pores, ventral portion of paramere with numerous sparsely arranged micropores. Median lobe 0.70× length of parameres, narrowly triangular apically, apex sharp, gonopore situated subapically, struts ca. 0.25× as long as apical portion of median lobe. Phallobase widening posteriad, without large posterior opening.
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA0FFC44379FE91FECA836B" pageId="5" pageNumber="112" type="diagnosis">
<paragraph id="8B823698FFA0FFC44379FE91FECA836B" blockId="5.[113,1090,219,1363]" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FE91FEC6816B" bold="true" box="[113,365,347,371]" pageId="5" pageNumber="112">Differential diagnosis.</emphasis>
Based on the evenly convex elytral intervals,
<taxonomicName id="4C3D4D1BFFA0FFC4406AFE97FBEB816C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[866,1088,349,372]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC4406AFE97FBEB816C" box="[866,1088,349,372]" italics="true" pageId="5" pageNumber="112">Georissus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC44379FEB6FF6F818B" box="[113,196,380,403]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
may be easily diagnosed from the species of the
<taxonomicName id="4C3D4D1BFFA0FFC441C2FEB7FCEA818C" authorityName="Laporte de Castelnau" authorityYear="1840" box="[714,833,381,404]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B949EA8AFFA0FFC441C2FEB7FCEA818C" box="[714,833,381,404]" italics="true" pageId="5" pageNumber="112">G. costatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3D4D1BFFA0FFC4407CFEB7FC46818C" baseAuthorityName="Erichson" baseAuthorityYear="1847" box="[884,1005,381,404]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="caelatus">
<emphasis id="B949EA8AFFA0FFC4407CFEB7FC46818C" box="[884,1005,381,404]" italics="true" pageId="5" pageNumber="112">G. caelatus</emphasis>
</taxonomicName>
species groups sensu
<bibRefCitation id="EFAC4B69FFA0FFC4420AFE56FE0D81AB" author="DELEVE J." box="[258,422,411,435]" pageId="5" pageNumber="112" pagination="1 - 23" refId="ref12544" refString="DELEVE J. 1967 a: Notes sur les Georissus d'Afrique et descriptions d'especes nouvelles (Coleoptera Georissidae). Bulletin de l ' Institute Royal des Sciences Naturelles de Belgique 43 (26): 1 - 23." type="journal article" year="1967">DELÈVE (1967a</bibRefCitation>
,b). It may be easily distinguished from the majority of the species with equally elevated elytral intervals by the absence of a median rhomboid impression on the pronotum, which it shares with the African species
<taxonomicName id="4C3D4D1BFFA0FFC441EAFE17FBB881EB" authority="Grouvelle, 1915" authorityName="Grouvelle" authorityYear="1915" box="[738,1043,475,500]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="B949EA8AFFA0FFC441EAFE17FCF581EC" box="[738,862,477,500]" italics="true" pageId="5" pageNumber="112">G. sordidus</emphasis>
Grouvelle, 1915
</taxonomicName>
and
<taxonomicName id="4C3D4D1BFFA0FFC44379FE37FE25820C" authority="Grouvelle, 1909" authorityName="Grouvelle" authorityYear="1909" box="[113,398,507,532]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="B949EA8AFFA0FFC44379FE37FF77820C" box="[113,220,509,532]" italics="true" pageId="5" pageNumber="112">G. bicolor</emphasis>
Grouvelle, 1909
</taxonomicName>
. The latter two species may be however immediately distinguished from
<taxonomicName id="4C3D4D1BFFA0FFC443A1FDD7FE98822C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[169,307,541,564]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC443A1FDD7FE98822C" box="[169,307,541,564]" italics="true" pageId="5" pageNumber="112">G. maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC44230FDD6FE23822B" box="[312,392,540,563]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
by much larger and wider pale-coloured body, morphology of male genitalia and the sculpture of the abdomen (see
<bibRefCitation id="EFAC4B69FFA0FFC4417EFDF6FCBB824B" author="DELEVE J." box="[630,784,571,595]" pageId="5" pageNumber="112" pagination="1 - 23" refId="ref12544" refString="DELEVE J. 1967 a: Notes sur les Georissus d'Afrique et descriptions d'especes nouvelles (Coleoptera Georissidae). Bulletin de l ' Institute Royal des Sciences Naturelles de Belgique 43 (26): 1 - 23." type="journal article" year="1967">DELÈVE 1967a</bibRefCitation>
), as well as by the sculpture of the pronotum which lacks any bulges in the posterior half in
<taxonomicName id="4C3D4D1BFFA0FFC44005FD97FC32826C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[781,921,605,628]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC44005FD97FC32826C" box="[781,921,605,628]" italics="true" pageId="5" pageNumber="112">G. maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC44097FD96FC5A826B" box="[927,1009,604,627]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
In fact,
<taxonomicName id="4C3D4D1BFFA0FFC44379FDB7FEFB828C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[113,336,637,660]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC44379FDB7FEFB828C" box="[113,336,637,660]" italics="true" pageId="5" pageNumber="112">Georissus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC4425FFDB6FE00828B" box="[343,427,636,659]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
seems to be unique among all described species of the genus by its pronotal sculpture, i.e. the combination of the absence of a median rhomboid depression, absence of granules in anterior half of the pronotum and absence of elevated granulate bulges posteriorly. By the extremely long and narrow parameres and posteriorly widening phallobase, The new species resembles
<taxonomicName id="4C3D4D1BFFA0FFC4411BFD37FCEB830C" authority="Grouvelle, 1909" authorityName="Grouvelle" authorityYear="1909" box="[531,832,763,788]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="alticosta">
<emphasis id="B949EA8AFFA0FFC4411BFD37FD24830C" box="[531,655,765,788]" italics="true" pageId="5" pageNumber="112">G. alticosta</emphasis>
Grouvelle, 1909
</taxonomicName>
(which has costate even elytral intervals),
<taxonomicName id="4C3D4D1BFFA0FFC44223FCD7FD9E832B" authority="Deleve, 1967" authorityName="Deleve" authorityYear="1967" box="[299,565,795,820]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B949EA8AFFA0FFC44223FCD7FE08832C" box="[299,419,797,820]" italics="true" pageId="5" pageNumber="112">G. alluaudi</emphasis>
Delève, 1967
</taxonomicName>
(which has the pronotum with a rhomboid central depression),
<taxonomicName id="4C3D4D1BFFA0FFC443F2FCF7FDF6834C" authority="Fairmaire, 1898" authorityName="Fairmaire" authorityYear="1898" box="[250,605,827,852]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="acutecostatus">
<emphasis id="B949EA8AFFA0FFC443F2FCF7FE07834C" box="[250,428,829,852]" italics="true" pageId="5" pageNumber="112">G. acutecostatus</emphasis>
Fairmaire, 1898
</taxonomicName>
and
<taxonomicName id="4C3D4D1BFFA0FFC44199FCF7FCD1834B" authority="Deleve, 1969" authorityName="Deleve" authorityYear="1969" box="[657,890,827,852]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="biroi">
<emphasis id="B949EA8AFFA0FFC44199FCF7FD4E834C" box="[657,741,829,852]" italics="true" pageId="5" pageNumber="112">G. biroi</emphasis>
Delève, 1969
</taxonomicName>
(both with costate even elytral intervals).
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA0FFC44379FCB6FC5084AB" pageId="5" pageNumber="112" type="etymology">
<paragraph id="8B823698FFA0FFC44379FCB6FC5783AC" blockId="5.[113,1090,219,1363]" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FCB6FF59838C" bold="true" box="[113,242,892,916]" pageId="5" pageNumber="112">Etymology.</emphasis>
The species name refers to the
<typeStatus id="5486883AFFA0FFC44134FCB6FDC1838C" box="[572,618,892,916]" pageId="5" pageNumber="112">type</typeStatus>
locality of this species, which is on the bank of a brackish lagoon situated at the estuary of a temporary stream to the Arabian Sea.
</paragraph>
<paragraph id="8B823698FFA0FFC44379FC71FC5084AB" blockId="5.[113,1090,219,1363]" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FC71FE2583CB" bold="true" box="[113,398,955,979]" pageId="5" pageNumber="112">Collection circumstances.</emphasis>
The
<typeStatus id="5486883AFFA0FFC442CEFC76FE5883CC" box="[454,499,956,980]" pageId="5" pageNumber="112">type</typeStatus>
series was collected at night on a sandbar between the sea and a brackish lake in an estuary of a temporary stream (
<figureCitation id="13062A1DFFA0FFC441D5FC11FCF683EC" box="[733,861,987,1012]" captionStart="Figs" captionStartId="10.[151,190,1491,1510]" captionTargetBox="[153,1125,221,1476]" captionTargetId="figure-12@10.[147,1128,216,1478]" captionTargetPageId="10" captionText="Figs. 1522.Socotran endemic Georissus species and their habitats. 1516 G. maritimus sp. nov.(15 dorsal view; 16 lateral view). 1718 G. nemo sp. nov. (17 dorsal view; 18 lateral view). 1920 the sand tongue 3 km NE of Shuab, type locality of G. maritimus sp. nov. (19 general view of the habitat; 20 satellite view showing the position of the locality); 2122 the stream at Arhar, type locality of G. nemo sp. nov. (21 general view of the habitat; 22 satellite view showing the position of the locality)." figureDoi="http://doi.org/10.5281/zenodo.5334592" httpUri="https://zenodo.org/record/5334592/files/figure.png" pageId="5" pageNumber="112">Figs. 1920</figureCitation>
). The specimens were collected on places without vegetation which were wet due to the seepage of the subsurface water. The surrounding vegetation (which usually began within ca. 2 meters of the sites with
<taxonomicName id="4C3D4D1BFFA0FFC440D0FBD7FF70844C" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC440D0FBD7FF70844C" italics="true" pageId="5" pageNumber="112">Georissus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC443EBFBF6FE9C844C" box="[227,311,1084,1108]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
) consisted of the low succulent shrubs of
<taxonomicName id="4C3D4D1BFFA0FFC441F0FBF7FE6F846B" authority="(Moric.) Moris" authorityName="Moris" baseAuthorityName="Moric." class="Magnoliopsida" family="Amaranthaceae" genus="Arthrocnemum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="112" phylum="Tracheophyta" rank="species" species="macrostachyum">
<emphasis id="B949EA8AFFA0FFC441F0FBF7FBEA844C" box="[760,1089,1085,1108]" italics="true" pageId="5" pageNumber="112">Arthrocnemum macrostachyum</emphasis>
(Moric.) Moris (Amaranthaceae)
</taxonomicName>
,
<taxonomicName id="4C3D4D1BFFA0FFC442C4FB97FCF8846B" authority="(Vierh.) Radcl." authorityName="Radcl." baseAuthorityName="Vierh." box="[460,851,1115,1140]" class="Magnoliopsida" family="Plumbaginaceae" genus="Limonium" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="112" phylum="Tracheophyta" rank="species" species="socotranum">
<emphasis id="B949EA8AFFA0FFC442C4FB97FD18846C" box="[460,691,1117,1140]" italics="true" pageId="5" pageNumber="112">Limonium socotranum</emphasis>
(Vierh.) Radcl.
</taxonomicName>
-Sm. (
<taxonomicName id="4C3D4D1BFFA0FFC44085FB91FB91846B" box="[909,1082,1115,1139]" class="Magnoliopsida" family="Plumbaginaceae" kingdom="Plantae" order="Caryophyllales" pageId="5" pageNumber="112" phylum="Tracheophyta" rank="family">Plumbaginaceae</taxonomicName>
) and isolated groups of mangrove trees
<taxonomicName id="4C3D4D1BFFA0FFC4410BFBB7FBBC848B" authority="(Forssk.) Vierh." authorityName="Vierh." baseAuthorityName="Forssk." box="[515,1047,1147,1172]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="5" pageNumber="112" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B949EA8AFFA0FFC4410BFBB7FD17848C" box="[515,700,1149,1172]" italics="true" pageId="5" pageNumber="112">Avicennia marina</emphasis>
(Forssk.) Vierh. (Avicenniaceae)
</taxonomicName>
. No specimen of
<taxonomicName id="4C3D4D1BFFA0FFC443FEFB57FE6484AC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[246,463,1181,1204]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC443FEFB57FE6484AC" box="[246,463,1181,1204]" italics="true" pageId="5" pageNumber="112">Georissus maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC442DDFB56FD8D84AB" box="[469,550,1180,1203]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
bore the substrate layer on the body dorsum.
</paragraph>
</subSubSection>
<subSubSection id="C3276513FFA0FFC44379FB71FE5E854B" pageId="5" pageNumber="112" type="distribution">
<paragraph id="8B823698FFA0FFC44379FB71FD2A84CB" blockId="5.[113,1090,219,1363]" box="[113,641,1211,1236]" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FB71FEAF84CB" bold="true" box="[113,260,1211,1235]" pageId="5" pageNumber="112">Distribution.</emphasis>
Known only from the
<typeStatus id="5486883AFFA0FFC442FFFB76FD8E84CC" box="[503,549,1212,1236]" pageId="5" pageNumber="112">type</typeStatus>
locality.
</paragraph>
<paragraph id="8B823698FFA0FFC44379FB16FE5E854B" blockId="5.[113,1090,219,1363]" pageId="5" pageNumber="112">
<emphasis id="B949EA8AFFA0FFC44379FB16FF0584EC" bold="true" box="[113,174,1244,1268]" pageId="5" pageNumber="112">Note.</emphasis>
The unique morphology of
<taxonomicName id="4C3D4D1BFFA0FFC442E9FB17FDC484EC" authority="Fikáček &amp; Delgado &amp; Gentili, 2012" authorityName="Fikáček &amp; Delgado &amp; Gentili" authorityYear="2012" box="[481,623,1245,1268]" class="Insecta" family="Georissidae" genus="Georissus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="112" phylum="Arthropoda" rank="species" species="maritimus" status="sp. nov.">
<emphasis id="B949EA8AFFA0FFC442E9FB17FDC484EC" box="[481,623,1245,1268]" italics="true" pageId="5" pageNumber="112">G. maritimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27A57F1FFA0FFC4417FFB16FD6784EB" box="[631,716,1244,1267]" pageId="5" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
makes it impossible to assign this species to any of the species groups proposed by
<bibRefCitation id="EFAC4B69FFA0FFC44174FB36FCB6850B" author="DELEVE J." box="[636,797,1275,1299]" pageId="5" pageNumber="112" pagination="1 - 23" refId="ref12544" refString="DELEVE J. 1967 a: Notes sur les Georissus d'Afrique et descriptions d'especes nouvelles (Coleoptera Georissidae). Bulletin de l ' Institute Royal des Sciences Naturelles de Belgique 43 (26): 1 - 23." type="journal article" year="1967">DELÈVE (1967a</bibRefCitation>
,b) and it seems to be rather isolated from the known species of the genus. Further studies are thus necessary to understand the geographic origin of the species.
</paragraph>
</subSubSection>
</treatment>
</document>