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<document id="78C69535E3E73EC39568A5217B855E04" ID-CLB-Dataset="204092" ID-DOI="10.11646/zootaxa.5219.2.2" ID-GBIF-Dataset="e46fc946-ff26-440a-9b11-6bd5ed10db29" ID-ISSN="1175-5326" ID-Zenodo-Dep="7413348" ID-ZooBank="0745DAF5-B623-401E-BA16-981F8883701B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670487986157" checkinUser="plazi" docAuthor="Telnov, Dmitry" docDate="2022" docId="038E8D48617EA846FF73FAE2F4E6F8F8" docLanguage="en" docName="zootaxa.5219.2.2.pdf" docOrigin="Zootaxa 5219 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lacconotopedilus elongatus Pic 1936, syn. nov." docType="treatment" docVersion="7" lastPageNumber="126" masterDocId="FFB7F530617CA843FFE4FFB7F617FF89" masterDocTitle="Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key" masterLastPageNumber="138" masterPageNumber="121" pageNumber="123" updateTime="1699413211887" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="400341138D7CB16859E791ACB2FE8671">Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key</mods:title>
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<mods:namePart id="BEDEA77048A8B09D65F9F3C0B5BF1AB4">Telnov, Dmitry</mods:namePart>
<mods:affiliation id="82AB1FD79F8DD90623C2DB1190245931">Department of Life Sciences, Natural History Museum, SW 7 5 BD, London, United Kingdom. &amp; Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV- 5401, Daugavpils, Latvia &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV- 1004, Rīga, Latvia</mods:affiliation>
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<treatment id="038E8D48617EA846FF73FAE2F4E6F8F8" ID-DOI="http://doi.org/10.5281/zenodo.7415028" ID-GBIF-Taxon="205926625" ID-Zenodo-Dep="7415028" LSID="urn:lsid:plazi:treatment:038E8D48617EA846FF73FAE2F4E6F8F8" httpUri="http://treatment.plazi.org/id/038E8D48617EA846FF73FAE2F4E6F8F8" lastPageId="5" lastPageNumber="126" pageId="2" pageNumber="123">
<subSubSection id="C33D6FD5617EA841FF73FAE2F4FAFAE6" box="[151,749,1364,1391]" pageId="2" pageNumber="123" type="nomenclature">
<paragraph id="8B983C5E617EA841FF73FAE2F4FAFAE6" blockId="2.[151,749,1364,1391]" box="[151,749,1364,1391]" pageId="2" pageNumber="123">
<heading id="D0D08B32617EA841FF73FAE2F4FAFAE6" bold="true" box="[151,749,1364,1391]" fontSize="11" level="1" pageId="2" pageNumber="123" reason="1">
<emphasis id="B953E04C617EA841FF73FAE2F4FAFAE6" bold="true" box="[151,749,1364,1391]" pageId="2" pageNumber="123">
=
<taxonomicName id="4C2747DD617EA841FF4BFAE3F46EFAE6" authority="Pic, 1936" authorityName="Pic" authorityYear="1936" box="[175,633,1364,1391]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B953E04C617EA841FF4BFAE3F416FAE6" bold="true" box="[175,513,1364,1391]" italics="true" pageId="2" pageNumber="123">Lacconotopedilus elongatus</emphasis>
<bibRefCitation id="EFB641AF617EA841FDEDFAE2F46EFAE6" author="Pic, M." box="[521,633,1365,1391]" pageId="2" pageNumber="123" pagination="10 - 36" refId="ref8326" refString="Pic, M. (1936) Nouveautes diverses. Melanges exotico-entomologiques 68, 10 - 36." type="journal article" year="1936">Pic, 1936</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A2605D37617EA841FD64FAE2F4FAFAE6" box="[640,749,1365,1391]" pageId="2" pageNumber="123">syn. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD5617EA840FF73FA2AF4DDFF70" lastPageId="3" lastPageNumber="124" pageId="2" pageNumber="123" type="materials_examined">
<paragraph id="8B983C5E617EA841FF73FA2AF222F9CE" blockId="2.[151,1437,1437,2040]" pageId="2" pageNumber="123">
<emphasis id="B953E04C617EA841FF73FA2AF75DFA3E" bold="true" box="[151,330,1437,1463]" pageId="2" pageNumber="123">Type material.</emphasis>
<taxonomicName id="4C2747DD617EA841FEB2FA29F7F1FA3E" authorityName="Pic" authorityYear="1936" box="[342,486,1437,1463]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B953E04C617EA841FEB2FA29F7F1FA3E" box="[342,486,1437,1463]" italics="true" pageId="2" pageNumber="123">L. elongatus</emphasis>
</taxonomicName>
,
<materialsCitation id="3B4F3603617EA841FE1DFA2AF222F9CE" ID-GBIF-Occurrence="3988003302" collectionCode="MNHN, TYPE" pageId="2" pageNumber="123" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="549C82FC617EA841FE1DFA2AF44CFA3E" box="[505,603,1437,1463]" pageId="2" pageNumber="123" type="holotype">holotype</typeStatus>
<specimenCount id="9D21F7D7617EA841FD8CFA2AF46AFA3E" box="[616,637,1437,1463]" pageId="2" pageNumber="123" type="female"></specimenCount>
[
<collectionCode id="ED36A49B617EA841FD6BFA2AF4FDFA3E" box="[655,746,1437,1463]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="123">MNHN</collectionCode>
] (
<figureCitation id="131C20DB617EA841FCE1FA2AF566FA31" box="[773,881,1437,1464]" captionStart="FIGURES 58" captionStartId="4.[152,270,1743,1767]" captionTargetBox="[165,1423,198,1719]" captionTargetId="figure-33@4.[150,1438,181,1720]" captionTargetPageId="4" captionText="FIGURES 58. Physciolagria liturata Pic, 1930, holotype ♀ Lacconotopedilus elongatus Pic, 1936. 5—Habitus, dorsal view; 6—ditto, lateral view; 7—ditto, ventral view; 8—Labels [images courtesy Christophe Rivier, MNHN]." figureDoi="http://doi.org/10.5281/zenodo.7413352" httpUri="https://zenodo.org/record/7413352/files/figure.png" pageId="2" pageNumber="123">Figs 58</figureCitation>
): U. 4/IV.
<collectorName id="26D25988617EA841FC0DFA2AF21EFA3E" box="[1001,1033,1437,1463]" pageId="2" pageNumber="123">Co</collectorName>
2280 [handwritten] //
<collectorName id="26D25988617EA841FAFEFA2AF38BFA3E" box="[1306,1436,1437,1463]" pageId="2" pageNumber="123">Tanganyika</collectorName>
(Conrads) [handwritten] // elongatus n sp [handwritten] // type [handwritten] //
<collectorName id="26D25988617EA841FBECFA76F321FA52" box="[1032,1334,1473,1499]" pageId="2" pageNumber="123">Museum Paris Coll. M. Pic</collectorName>
[printed, label bluish] //
<collectionCode id="ED36A49B617EA841FEA1FA52F79CFA76" box="[325,395,1509,1535]" pageId="2" pageNumber="123">TYPE</collectionCode>
[printed, label red] //
<typeStatus id="549C82FC617EA841FD6FFA52F508FA76" box="[651,799,1509,1535]" pageId="2" pageNumber="123" type="holotype">HOLOTYPE</typeStatus>
[printed]
<taxonomicName id="4C2747DD617EA841FC77FA52F2ECFA76" authority="Pic" authorityName="Pic" authorityYear="1936" box="[915,1275,1509,1535]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="elongatus">Lacconotopedilus elongatus Pic</taxonomicName>
[handwritten]
<collectorName id="26D25988617EA841FF73F9BEF71EF9AA" box="[151,265,1545,1571]" pageId="2" pageNumber="123">Examined</collectorName>
by
<collectorName id="26D25988617EA841FED1F9BEF7DDF9AA" box="[309,458,1545,1571]" pageId="2" pageNumber="123">D.A. Pollock</collectorName>
19 [printed] 97 [handwritten] [label red] //
<collectionCode id="ED36A49B617EA841FC5EF9BEF205F9AA" box="[954,1042,1545,1571]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="123">MNHN</collectionCode>
,
<collectorName id="26D25988617EA841FBFBF9BEF294F9AD" box="[1055,1155,1545,1572]" pageId="2" pageNumber="123">Paris EC</collectorName>
15939 [printed, provided with QR code].
<collectorName id="26D25988617EA841FEAEF99AF761F9CE" box="[330,374,1581,1607]" pageId="2" pageNumber="123">The</collectorName>
type is missing both terminal antennomeres and both forelegs.
</materialsCitation>
</paragraph>
<paragraph id="8B983C5E617EA841FF23F9E6F2CCF975" blockId="2.[151,1437,1437,2040]" pageId="2" pageNumber="123">
<bibRefCitation id="EFB641AF617EA841FF23F9E6F7D6F9E5" author="Pollock, D. A." box="[199,449,1617,1644]" pageId="2" pageNumber="123" pagination="51 - 56" refId="ref8344" refString="Pollock, D. A. (1997) Review and systematic placement of the Afrotropical genus Lacconotopedilus Pic (Coleoptera: Mycteridae: Lacconotinae). African Entomology, 5 (1), 51 - 56." type="journal article" year="1997">Pollock (1997: 5556)</bibRefCitation>
redescribed
<taxonomicName id="4C2747DD617EA841FDA9F9E6F503F9E2" authorityName="Telnov" authorityYear="2022" box="[589,788,1617,1643]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis id="B953E04C617EA841FDA9F9E6F503F9E2" box="[589,788,1617,1643]" italics="true" pageId="2" pageNumber="123">Lacconotopedilus</emphasis>
</taxonomicName>
from the type of
<taxonomicName id="4C2747DD617EA841FC34F9E5F280F9E2" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1930" box="[976,1175,1617,1643]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C617EA841FC34F9E5F280F9E2" box="[976,1175,1617,1643]" italics="true" pageId="2" pageNumber="123">L. singularicornis</emphasis>
</taxonomicName>
and additional material from
<collectingCountry id="F3307CCE617EA841FF30F9C2F752F906" box="[212,325,1653,1679]" name="Botswana" pageId="2" pageNumber="123">Botswana</collectingCountry>
, south-eastern
<collectingCountry id="F3307CCE617EA841FE09F9C2F47FF906" box="[493,616,1653,1679]" name="Democratic Republic of the Congo" pageId="2" pageNumber="123">DR Congo</collectingCountry>
, and
<collectingCountry id="F3307CCE617EA841FD46F9C2F523F906" box="[674,820,1653,1679]" name="South Africa" pageId="2" pageNumber="123">South Africa</collectingCountry>
. Similar as in my case, Pollock was also unable to read in full the locality label of the
<typeStatus id="549C82FC617EA841FDE4F92EF475F93A" box="[512,610,1689,1715]" pageId="2" pageNumber="123" type="holotype">holotype</typeStatus>
<taxonomicName id="4C2747DD617EA841FD88F92DF52FF93A" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1930" box="[620,824,1689,1715]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C617EA841FD88F92DF52FF93A" box="[620,824,1689,1715]" italics="true" pageId="2" pageNumber="123">L. singularicornis</emphasis>
</taxonomicName>
: “Afrique [with other illegible name, something like Lrimave]” (Pollock 1995: 55) and therefore suggested possible conspecifity of
<taxonomicName id="4C2747DD617EA841FBF5F909F28BF95E" authorityName="Pic" authorityYear="1936" box="[1041,1180,1725,1751]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B953E04C617EA841FBF5F909F28BF95E" box="[1041,1180,1725,1751]" italics="true" pageId="2" pageNumber="123">L. elongatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2747DD617EA841FB37F909F38BF95E" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1930" box="[1235,1436,1725,1751]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C617EA841FB37F909F38BF95E" box="[1235,1436,1725,1751]" italics="true" pageId="2" pageNumber="123">L. singularicornis</emphasis>
</taxonomicName>
but did not propose the formal synonymy due to lack of comparative material (
<bibRefCitation id="EFB641AF617EA841FBEDF956F2D9F975" author="Pollock, D. A." box="[1033,1230,1761,1788]" pageId="2" pageNumber="123" pagination="51 - 56" refId="ref8344" refString="Pollock, D. A. (1997) Review and systematic placement of the Afrotropical genus Lacconotopedilus Pic (Coleoptera: Mycteridae: Lacconotinae). African Entomology, 5 (1), 51 - 56." type="journal article" year="1997">Pollock 1997: 53</bibRefCitation>
).
</paragraph>
<paragraph id="8B983C5E617EA841FF23F8B2F2BBF8CA" blockId="2.[151,1437,1437,2040]" pageId="2" pageNumber="123">
The
<typeStatus id="549C82FC617EA841FF1FF8B2F74AF896" box="[251,349,1797,1823]" pageId="2" pageNumber="123" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2747DD617EA841FE62F8B1F446F896" authorityName="Telnov" authorityYear="2022" baseAuthorityName="Pic" baseAuthorityYear="1930" box="[390,593,1797,1823]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="singularicornis">
<emphasis id="B953E04C617EA841FE62F8B1F446F896" box="[390,593,1797,1823]" italics="true" pageId="2" pageNumber="123">L. singularicornis</emphasis>
</taxonomicName>
was obtained from unknown locality, since I was not able to attribute the type locality (spelled as Limawe, Limowe, Limare or Limare) to any existing place in Africa.
</paragraph>
<paragraph id="8B983C5E617EA840FF23F8FAF4DDFF70" blockId="2.[151,1437,1437,2040]" lastBlockId="3.[151,1437,151,393]" lastPageId="3" lastPageNumber="124" pageId="2" pageNumber="123">
The oldest label of the
<typeStatus id="549C82FC617EA841FE33F8FAF42EF8EE" box="[471,569,1869,1895]" pageId="2" pageNumber="123" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2747DD617EA841FD80F8F9F4E6F8EE" authorityName="Pic" authorityYear="1936" box="[612,753,1869,1895]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B953E04C617EA841FD80F8F9F4E6F8EE" box="[612,753,1869,1895]" italics="true" pageId="2" pageNumber="123">L. elongatus</emphasis>
</taxonomicName>
is handwritten by an unknown person, while the next one, claiming the specimen is from Conrads material from “Tanganyika”, was undoubtedly added later by Pic (judging from handwriting on both labels). Had the type specimen really been collected by P. Conrads, the letter “U.” on the initial label likely refers to Ukerewe Island, Lake Victoria,
<collectingCountry id="F3307CCE617EA841FC9AF80EF5F1F85A" box="[894,998,1977,2003]" name="Tanzania" pageId="2" pageNumber="123">Tanzania</collectingCountry>
, where Conrads extensively collected 19121931 (
<bibRefCitation id="EFB641AF617EA841FEC7F86AF42CF871" author="Lanjouw, J. &amp; Stafleu, F. A." box="[291,571,2013,2040]" pageId="2" pageNumber="123" refId="ref8218" refString="Lanjouw, J. &amp; Stafleu, F. A. (1954) Collectors, first instalment, A - D. In: Index Herbariorum. Part II. Regnum Vegetable. Vol. 2. International Bureau for Plant Taxonomy and Nomenclature of the Association for Plant Taxonomy, Utrecht, pp. 1 - 174." type="book" year="1954">Lanjouw &amp; Stafleu 1954</bibRefCitation>
). In support of this hypothesis, there are
<specimenCount id="9D21F7D7617EA841FC18F86AF283F87E" box="[1020,1172,2013,2039]" pageId="2" pageNumber="123" type="generic" typeStatus="syntypes">nine syntypes</specimenCount>
of
<taxonomicName id="4C2747DD617EA840FB5CF86AF755FF38" authority="Marshall, 1939" authorityName="Marshall" authorityYear="1939" class="Insecta" family="Brentidae" genus="Nanophyes" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="3" lastPageNumber="124" order="Coleoptera" pageId="2" pageNumber="123" phylum="Arthropoda" rank="species" species="conradsi">
<emphasis id="B953E04C617EA841FB5CF86AF38BF87E" box="[1208,1436,2013,2039]" italics="true" pageId="2" pageNumber="123">Nanophyes conradsi</emphasis>
Marshall, 1939
</taxonomicName>
(
<taxonomicName id="4C2747DD617FA840FEB6FF20F47AFF38" authority="Billberg, 1820" authorityName="Billberg" authorityYear="1820" box="[338,621,151,177]" class="Insecta" family="Brentidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="family">Brentidae Billberg, 1820</taxonomicName>
) present at
<collectionCode id="ED36A49B617FA840FD10FF20F55CFF38" box="[756,843,151,177]" pageId="3" pageNumber="124">BMNH</collectionCode>
, all described from Conrads collected material from Ukerewe Island, each bearing handwritten collecting locality code label “2215” which is close to the “2280” code as on the initial label of the
<taxonomicName id="4C2747DD617FA840FE2BFF68F44DFF70" authorityName="Pic" authorityYear="1936" box="[463,602,223,249]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B953E04C617FA840FE2BFF68F44DFF70" box="[463,602,223,249]" italics="true" pageId="3" pageNumber="124">L. elongatus</emphasis>
</taxonomicName>
<typeStatus id="549C82FC617FA840FD84FF68F4D1FF70" box="[608,710,223,249]" pageId="3" pageNumber="124" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD5617FA846FF23FEB4F764F8C4" lastPageId="5" lastPageNumber="126" pageId="3" pageNumber="124" type="description">
<paragraph id="8B983C5E617FA840FF23FEB4F420FE00" blockId="3.[151,1437,151,393]" pageId="3" pageNumber="124">
<emphasis id="B953E04C617FA840FF23FEB4F769FE94" bold="true" box="[199,382,259,285]" pageId="3" pageNumber="124">Measurements.</emphasis>
<typeStatus id="549C82FC617FA840FE60FEB4F7FBFE94" box="[388,492,259,285]" pageId="3" pageNumber="124" type="holotype">Holotype</typeStatus>
<specimenCount id="9D21F7D7617FA840FE16FEB4F410FE94" box="[498,519,259,285]" pageId="3" pageNumber="124" type="female"></specimenCount>
(
<figureCitation id="131C20DB617FA840FDF1FEB4F46EFE94" box="[533,633,259,285]" captionStart="FIGURES 14" captionStartId="3.[152,270,1959,1983]" captionTargetBox="[164,1406,433,1933]" captionTargetId="figure-116@3.[162,1426,417,1936]" captionTargetPageId="3" captionText="FIGURES 14. Physciolagria liturata Pic, 1930, holotype ♀. 1—Habitus, dorsal view; 2—ditto, lateral view; 3—Forebody, anterodorsal view; 4—Labels [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413350" httpUri="https://zenodo.org/record/7413350/files/figure.png" pageId="3" pageNumber="124">Figs 14</figureCitation>
), total body length
<quantity id="4CDF91BB617FA840FCB4FEB4F5BEFE94" box="[848,937,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="3" pageNumber="124" unit="mm" value="8.3">8.3 mm</quantity>
; head length
<quantity id="4CDF91BB617FA840FBDFFEB4F285FE94" box="[1083,1170,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="124" unit="mm" value="1.3">1.3 mm</quantity>
, head width across eyes
<quantity id="4CDF91BB617FA840FF73FE90F6E7FEC8" box="[151,240,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="124" unit="mm" value="1.4">1.4 mm</quantity>
, pronotal length
<quantity id="4CDF91BB617FA840FE4BFE90F410FEC8" box="[431,519,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="124" unit="mm" value="1.2">1.2 mm</quantity>
, maximum pronotal width
<quantity id="4CDF91BB617FA840FCDDFE91F585FEC8" box="[825,914,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="124" unit="mm" value="1.5">1.5 mm</quantity>
, minimum pronotal
<quantity id="4CDF91BB617FA840FB9CFE90F2C7FEC8" box="[1144,1232,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="124" unit="mm" value="1.2">1.2 mm</quantity>
, elytral length
<quantity id="4CDF91BB617FA840FA9DFE91F6D2FEEC" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="124" unit="mm" value="5.8">5.8 mm</quantity>
, combined maximum elytral width in apical third
<quantity id="4CDF91BB617FA840FD12FEFCF52DFEEC" box="[758,826,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="124" unit="mm" value="3.0">3 mm</quantity>
. Selected comparative
<specimenCount id="9D21F7D7617FA840FBD8FEFCF271FEEC" box="[1084,1126,331,357]" pageId="3" pageNumber="124" type="male">♂♂</specimenCount>
from
<collectingCountry id="F3307CCE617FA840FB4EFEFCF318FEEC" box="[1194,1295,331,357]" name="Tanzania" pageId="3" pageNumber="124">Tanzania</collectingCountry>
<quantity id="4CDF91BB617FA840FAF2FEFDF38AFEEC" box="[1302,1437,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="5.8" metricValueMin="5.7" pageId="3" pageNumber="124" unit="mm" value="5.75" valueMax="5.8" valueMin="5.7">5.75.8 mm</quantity>
long (see Checklist for locality data).
</paragraph>
<caption id="DF586CD6617FA840FF7CF810F442F86D" ID-DOI="http://doi.org/10.5281/zenodo.7413350" ID-Zenodo-Dep="7413350" httpUri="https://zenodo.org/record/7413350/files/figure.png" pageId="3" pageNumber="124" startId="3.[152,270,1959,1983]" targetBox="[164,1406,433,1933]" targetPageId="3">
<paragraph id="8B983C5E617FA840FF7CF810F442F86D" blockId="3.[151,1437,1959,2020]" pageId="3" pageNumber="124">
<emphasis id="B953E04C617FA840FF7CF810F755F849" bold="true" box="[152,322,1959,1984]" pageId="3" pageNumber="124">FIGURES 14.</emphasis>
<taxonomicName id="4C2747DD617FA840FEADF81FF482F849" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[329,661,1960,1984]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C617FA840FEADF81FF43DF849" box="[329,554,1960,1984]" italics="true" pageId="3" pageNumber="124">Physciolagria liturata</emphasis>
<bibRefCitation id="EFB641AF617FA840FDD6F81FF482F849" author="Pic, M." box="[562,661,1960,1984]" pageId="3" pageNumber="124" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
, holotype ♀. 1—Habitus, dorsal view; 2—ditto, lateral view; 3—Forebody, anterodorsal view; 4—Labels [not to scale].
</paragraph>
</caption>
<caption id="DF586CD66178A847FF7CF978F2BEF882" ID-DOI="http://doi.org/10.5281/zenodo.7413352" ID-Zenodo-Dep="7413352" httpUri="https://zenodo.org/record/7413352/files/figure.png" pageId="4" pageNumber="125" startId="4.[152,270,1743,1767]" targetBox="[165,1423,198,1719]" targetPageId="4">
<paragraph id="8B983C5E6178A847FF7CF978F2BEF882" blockId="4.[151,1436,1743,1804]" pageId="4" pageNumber="125">
<emphasis id="B953E04C6178A847FF7CF978F756F96E" bold="true" box="[152,321,1743,1767]" pageId="4" pageNumber="125">FIGURES 58.</emphasis>
<taxonomicName id="4C2747DD6178A847FEA2F967F499F961" authority="Pic, 1930" authorityName="Pic" authorityYear="1930" box="[326,654,1744,1768]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C6178A847FEA2F967F430F961" box="[326,551,1744,1768]" italics="true" pageId="4" pageNumber="125">Physciolagria liturata</emphasis>
<bibRefCitation id="EFB641AF6178A847FDC9F967F499F961" author="Pic, M." box="[557,654,1744,1768]" pageId="4" pageNumber="125" pagination="7 - 8" refId="ref8278" refString="Pic, M. (1930) Coleopteres exotiques en partie nouveaux (suite). L'Echange, Revue Linneenne, 46 (440), 7 - 8." type="journal article" year="1930">Pic, 1930</bibRefCitation>
</taxonomicName>
, holotype ♀
<taxonomicName id="4C2747DD6178A847FCF4F967F281F96E" authority="Pic, 1936" authorityName="Pic" authorityYear="1936" box="[784,1174,1743,1768]" class="Insecta" family="Mycteridae" genus="Lacconotopedilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B953E04C6178A847FCF4F967F23AF961" box="[784,1069,1744,1768]" italics="true" pageId="4" pageNumber="125">Lacconotopedilus elongatus</emphasis>
<bibRefCitation id="EFB641AF6178A847FBD7F967F281F96E" author="Pic, M." box="[1075,1174,1743,1768]" pageId="4" pageNumber="125" pagination="10 - 36" refId="ref8326" refString="Pic, M. (1936) Nouveautes diverses. Melanges exotico-entomologiques 68, 10 - 36." type="journal article" year="1936">Pic, 1936</bibRefCitation>
</taxonomicName>
. 5—Habitus, dorsal view; 6—ditto, lateral view; 7—ditto, ventral view; 8—Labels [images courtesy Christophe Rivier, MNHN].
</paragraph>
</caption>
<paragraph id="8B983C5E6178A846FF23F88EF49DFA48" blockId="4.[151,1437,1849,2020]" lastBlockId="5.[151,1437,151,1905]" lastPageId="5" lastPageNumber="126" pageId="4" pageNumber="125">
<emphasis id="B953E04C6178A847FF23F88EF77AF8DA" bold="true" box="[199,365,1849,1875]" pageId="4" pageNumber="125">Redescription</emphasis>
,
<specimenCount id="9D21F7D76178A847FE9AF88DF784F8DD" box="[382,403,1850,1876]" pageId="4" pageNumber="125" type="male"></specimenCount>
from Ukiriguru,
<collectingCountry id="F3307CCE6178A847FDBFF88DF4D7F8DD" box="[603,704,1850,1876]" name="Tanzania" pageId="4" pageNumber="125">Tanzania</collectingCountry>
(
<figureCitation id="131C20DB6178A847FD35F88DF550F8DD" box="[721,839,1850,1876]" captionStart="FIGURES 911" captionStartId="6.[152,270,1387,1411]" captionTargetBox="[212,1387,181,1363]" captionTargetId="figure-33@6.[193,1394,181,1363]" captionTargetPageId="6" captionText="FIGURES 911. Physciolagria liturata Pic, 1930, ♂ from Homa, Kenya. 9—Habitus, dorsal view; 10—Head, dorsal view; 11—Left antenna, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413356" httpUri="https://zenodo.org/record/7413356/files/figure.png" pageId="4" pageNumber="125">Figs 911</figureCitation>
; consider Checklist below for specimen label data): Dorsal forebody generally dark brown, pronotal basal area narrowly paler, clypeus and labrum dirty yellow to yellowish brown. Elytra ochre with black, strongly elongate sutural spot in apical third leaving suture and apex of elytra narrowly yellow. Antennae black. Maxillary palps pale brown. Legs black-brown to brown, tarsi yellowish brown, trochanters yellowish brown. Venter generally brown, ventral and posterolateral parts of prothoracic hypomeron yellow to yellowish brown. Head (
<figureCitation id="131C20DB6179A846FD4BFF20F533FF38" box="[687,804,151,177]" captionStart="FIGURES 911" captionStartId="6.[152,270,1387,1411]" captionTargetBox="[212,1387,181,1363]" captionTargetId="figure-33@6.[193,1394,181,1363]" captionTargetPageId="6" captionText="FIGURES 911. Physciolagria liturata Pic, 1930, ♂ from Homa, Kenya. 9—Habitus, dorsal view; 10—Head, dorsal view; 11—Left antenna, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413356" httpUri="https://zenodo.org/record/7413356/files/figure.png" pageId="5" pageNumber="126">Figs 910</figureCitation>
) transverse, moderately glossy dorsally, flattened, with extremely large and globose hemispherical compound eyes occupying most of head. Compound eye touching antennal insertion, with inconspicuous notch at point of contact. Interfacetal setae dense, rather long. Frontoclypeal suture or impression not indicated. Labrum broadly emarginate at anterior margin. Frons extremely narrow between compound eyes (at its narrowest point frons is narrower than length of basal antennomere), strongly widens posteriad. Antennal insertion free, not concealed by anterolateral margin of frons. Tempora not indicated, head base straight, not narrowed to a neck, hidden beneath anterior margin of pronotum. Dorsal cranial punctures dense and rather coarse, irregular (especially anteriad eyes). Intervening spaces moderately glossy on frons. Head dorsal setation yellowish, inconspicuous, suberect. Antenna (
<figureCitation id="131C20DB6179A846FD4AFE00F515FE58" box="[686,770,439,465]" captionStart="FIGURES 911" captionStartId="6.[152,270,1387,1411]" captionTargetBox="[212,1387,181,1363]" captionTargetId="figure-33@6.[193,1394,181,1363]" captionTargetPageId="6" captionText="FIGURES 911. Physciolagria liturata Pic, 1930, ♂ from Homa, Kenya. 9—Habitus, dorsal view; 10—Head, dorsal view; 11—Left antenna, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413356" httpUri="https://zenodo.org/record/7413356/files/figure.png" pageId="5" pageNumber="126">Fig. 11</figureCitation>
) extending beyond mesocoxa when directed backwards. Basal antennomere short, about 1.2× as long as wide. Antennomere three about 1.21.3× as long as antennomere two. Antennomeres 48 asymmetrical, 410 strongly transverse. Terminal antennomere greatly elongated, flattened dorso-ventrally and somewhat shallowly impressed on whole ventral surface, densely rugulose, rounded apically, with an inconspicuous, obtuse angulation about midlength of anterior margin, significantly longer than combined length of antennomeres 110. Terminal maxillary palpomere slightly fusiform, narrow. Pronotum (
<figureCitation id="131C20DB6179A846FA8EFDDCF6C6FD20" captionStart="FIGURES 911" captionStartId="6.[152,270,1387,1411]" captionTargetBox="[212,1387,181,1363]" captionTargetId="figure-33@6.[193,1394,181,1363]" captionTargetPageId="6" captionText="FIGURES 911. Physciolagria liturata Pic, 1930, ♂ from Homa, Kenya. 9—Habitus, dorsal view; 10—Head, dorsal view; 11—Left antenna, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413356" httpUri="https://zenodo.org/record/7413356/files/figure.png" pageId="5" pageNumber="126">Figs 910</figureCitation>
) moderately glossy dorsally, transverse, markedly narrower than head across compound eyes, subtruncate at anterior margin. Lateral margins subparallel. Anterolateral angles obsolete, posterolateral angles broadly obtuse to rounded. Pronotal base broadly rounded, with narrow bead. Pronotal disc flattened dorsally. Punctures circular, regular, intervening spaces glossy, narrower than to twice as wide as punctures. Pronotal dorsal pubescence yellow, moderately long, rather dense, suberect. Scutellar shield triangular, rounded apically. Elytra (
<figureCitation id="131C20DB6179A846FB7EFCA8F2CBFCB0" box="[1178,1244,799,825]" captionStart="FIGURES 911" captionStartId="6.[152,270,1387,1411]" captionTargetBox="[212,1387,181,1363]" captionTargetId="figure-33@6.[193,1394,181,1363]" captionTargetPageId="6" captionText="FIGURES 911. Physciolagria liturata Pic, 1930, ♂ from Homa, Kenya. 9—Habitus, dorsal view; 10—Head, dorsal view; 11—Left antenna, ventral view [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413356" httpUri="https://zenodo.org/record/7413356/files/figure.png" pageId="5" pageNumber="126">Fig. 9</figureCitation>
) elongate, nearly parallel-sided, slightly glossy, flattened dorsally, somewhat malacoderm-like thin and flimsy. Humerus broadly rounded. Punctures on elytral disc coarser and denser than those on pronotum, with irregular intervening spaces which are generally narrower than punctures; punctures not or hardly becoming more delicate in apical third. Elytral pubescence yellow, dense, suberect, moderately long, directed posteriad. Sutural striae not indicated. Epipleuron short, moderately broad, present from humerus towards about posterior margin of mesocoxal cavity. Metathoracic wings fully developed. All legs of similar shape, sparsely setose. Femora not clavate, tibiae nearly straight, rugulose. Pro- and mesothoracic tibia about as long as corresponding femur, metathoracic tibia longer than metathoracic femur. Tibial terminal spurs paired, inconspicuously short. Penultimate tarsomeres not bilobed, with truncate ventral pad. Basal metathoracic tarsomere shorter than the combined length of all remaining metathoracic tarsomeres. Pretarsal claws short, curved, distinctly cleft. Tergite VII subtruncate at posterior margin (
<figureCitation id="131C20DB6179A846FB6BFB30F2F3FB28" box="[1167,1252,1159,1185]" captionStart="FIGURES 2027" captionStartId="11.[152,270,1776,1800]" captionTargetBox="[159,1428,199,1738]" captionTargetId="figure-15@11.[151,1436,181,1752]" captionTargetPageId="11" captionText="FIGURES 2027. Male terminalia of Physciolagria Pic, 1930. 20—P. liturata Pic, 1930, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—P. smithi sp. nov., paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, spiculum gastrale; 27—P. smithi sp. nov., paratype, spiculum gastrale [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="5" pageNumber="126">Fig. 20</figureCitation>
), morphological sternite VII short and broad, subtruncate at posterior margin (
<figureCitation id="131C20DB6179A846FCA2FB1CF58EFB4C" box="[838,921,1195,1221]" captionStart="FIGURES 2027" captionStartId="11.[152,270,1776,1800]" captionTargetBox="[159,1428,199,1738]" captionTargetId="figure-15@11.[151,1436,181,1752]" captionTargetPageId="11" captionText="FIGURES 2027. Male terminalia of Physciolagria Pic, 1930. 20—P. liturata Pic, 1930, tergite VII, dorsal view; 21—ditto, morphological sternite VII, ventral view; 22—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, tergite VII, dorsal view; 23—ditto, morphological sternite VII, ventral view; 24—P. smithi sp. nov., paratype, tergite VII, dorsal view; 25—ditto, morphological sternite VII, ventral view; 26—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana, spiculum gastrale; 27—P. smithi sp. nov., paratype, spiculum gastrale [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413364" httpUri="https://zenodo.org/record/7413364/files/figure.png" pageId="5" pageNumber="126">Fig. 21</figureCitation>
). Tergite VIII and morphological sternite VIII short and broad, subtruncate at posterior margin. Aedeagus with median lobe longer than tegmen. Median lobe somewhat funnel-shaped, wide basally in dorsal and lateral view, strongly narrowed towards rounded apex (
<figureCitation id="131C20DB6179A846FA8EFB44F6F5FAB8" captionStart="FIGURES 3644" captionStartId="13.[152,270,1906,1930]" captionTargetBox="[187,1403,196,1860]" captionTargetId="figure-15@13.[166,1422,181,1883]" captionTargetPageId="13" captionText="FIGURES 3644. Median lobe of Physciolagria Pic, 1930. 36—P. liturata Pic, 1930; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—P. smithi sp. nov., paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="5" pageNumber="126">Figs 3638</figureCitation>
). Gonopore armature as in
<figureCitation id="131C20DB6179A846FDEBFAA0F476FAB8" box="[527,609,1303,1329]" captionStart="FIGURES 3644" captionStartId="13.[152,270,1906,1930]" captionTargetBox="[187,1403,196,1860]" captionTargetId="figure-15@13.[166,1422,181,1883]" captionTargetPageId="13" captionText="FIGURES 3644. Median lobe of Physciolagria Pic, 1930. 36—P. liturata Pic, 1930; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—P. smithi sp. nov., paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="5" pageNumber="126">Fig. 38</figureCitation>
. Apical portion with several pores, apical margin with 56 radial channels (
<figureCitation id="131C20DB6179A846FF7BFA8CF736FADC" box="[159,289,1339,1365]" captionStart="FIGURES 3644" captionStartId="13.[152,270,1906,1930]" captionTargetBox="[187,1403,196,1860]" captionTargetId="figure-15@13.[166,1422,181,1883]" captionTargetPageId="13" captionText="FIGURES 3644. Median lobe of Physciolagria Pic, 1930. 36—P. liturata Pic, 1930; 37—ditto, apical portion (note apical pores); 38—ditto, gonopore armature; 39—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 40—ditto, apical portion (note apical spines); 41—ditto, gonopore armature; 42—P. smithi sp. nov., paratype; 43—ditto, apical portion (note apical spines); 44—gonopore armature [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413368" httpUri="https://zenodo.org/record/7413368/files/figure.png" pageId="5" pageNumber="126">Figs 3637</figureCitation>
). Tegmen stout, with parameres externally evenly, broadly curved; paramere apex rounded, with several moderately long setae raising from pores (
<figureCitation id="131C20DB6179A846FD9CFAE8F4EAFAF0" box="[632,765,1375,1401]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="5" pageNumber="126">Figs 2829</figureCitation>
). Outer margin of paramere narrow (
<figureCitation id="131C20DB6179A846FB44FAE8F2E2FAF0" box="[1184,1269,1375,1401]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="5" pageNumber="126">Fig. 28</figureCitation>
). Inner margin of paramere nearly straight except at apex where it strongly curves inwards (
<figureCitation id="131C20DB6179A846FC12FA34F261FA14" box="[1014,1142,1411,1437]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="5" pageNumber="126">Fig. 2829</figureCitation>
). Ventral side of paramere with sparse, moderately long setae (
<figureCitation id="131C20DB6179A846FDCDFA10F46AFA48" box="[553,637,1447,1473]" captionStart="FIGURES 2835" captionStartId="12.[152,270,1855,1879]" captionTargetBox="[170,1426,197,1817]" captionTargetId="figure-33@12.[162,1426,181,1831]" captionTargetPageId="12" captionText="FIGURES 2835. Tegmen of Physciolagria Pic, 1930. 28—P. liturata Pic, 1930; 29—ditto, parameres, apical portion (note moderately long apical setation); 30—P. singularicornis (Pic, 1935), ♂ from Sebele, Botswana; 31—ditto, paramere, apical portion, apical setation removed; 32—ditto, parameres, apical portion (note long apical setation); 33—P. smithi sp. nov., paratype; 34—ditto, paramere, apical portion, apical setation removed; 35—ditto, parameres, apical portion (note long apical setation) [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413366" httpUri="https://zenodo.org/record/7413366/files/figure.png" pageId="5" pageNumber="126">Fig. 29</figureCitation>
).
</paragraph>
<paragraph id="8B983C5E6179A846FF23FA7DF272F934" blockId="5.[151,1437,151,1905]" pageId="5" pageNumber="126">
<emphasis id="B953E04C6179A846FF23FA7DF77AFA6D" bold="true" box="[199,365,1482,1508]" pageId="5" pageNumber="126">Redescription</emphasis>
,
<specimenCount id="9D21F7D76179A846FE99FA7CF785FA6C" box="[381,402,1483,1509]" pageId="5" pageNumber="126" type="female"></specimenCount>
<typeStatus id="549C82FC6179A846FE7EFA7CF417FA6C" box="[410,512,1483,1509]" pageId="5" pageNumber="126" type="holotype">holotype</typeStatus>
. Female (
<figureCitation id="131C20DB6179A846FD8AFA7CF4C4FA6C" box="[622,723,1483,1509]" captionStart="FIGURES 14" captionStartId="3.[152,270,1959,1983]" captionTargetBox="[164,1406,433,1933]" captionTargetId="figure-116@3.[162,1426,417,1936]" captionTargetPageId="3" captionText="FIGURES 14. Physciolagria liturata Pic, 1930, holotype ♀. 1—Habitus, dorsal view; 2—ditto, lateral view; 3—Forebody, anterodorsal view; 4—Labels [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413350" httpUri="https://zenodo.org/record/7413350/files/figure.png" pageId="5" pageNumber="126">Figs 13</figureCitation>
) generally longer than male (see measurements, above). Frons comparatively wider between compound eyes. Antenna (
<figureCitation id="131C20DB6179A846FCDCFA58F5B0F980" box="[824,935,1519,1545]" captionStart="FIGURES 14" captionStartId="3.[152,270,1959,1983]" captionTargetBox="[164,1406,433,1933]" captionTargetId="figure-116@3.[162,1426,417,1936]" captionTargetPageId="3" captionText="FIGURES 14. Physciolagria liturata Pic, 1930, holotype ♀. 1—Habitus, dorsal view; 2—ditto, lateral view; 3—Forebody, anterodorsal view; 4—Labels [not to scale]." figureDoi="http://doi.org/10.5281/zenodo.7413350" httpUri="https://zenodo.org/record/7413350/files/figure.png" pageId="5" pageNumber="126">Figs 1, 3</figureCitation>
) comparatively shorter than that of male. Antennomere three triangular, twice as long as antennomere two. Antennomeres 49 slightly asymmetrical, 510 strongly transverse, 910 distinctly broadly emarginate at posterior margin. Terminal antennomere asymmetrically ovoid, truncate apically, about as long as 44.5 preceding antennomeres.Terminal maxillary palpomere subtriangular, slightly widened apically. Basal metathoracic tarsomere longer than combined length of all remaining metathoracic tarsomeres. Tergite VII and morphological sternite VII subtruncate at posterior margin.
</paragraph>
<paragraph id="8B983C5E6179A846FF23F971F764F8C4" blockId="5.[151,1437,151,1905]" pageId="5" pageNumber="126">
<emphasis id="B953E04C6179A846FF23F971F7F2F969" bold="true" box="[199,485,1734,1760]" pageId="5" pageNumber="126">Intraspecific variability.</emphasis>
Specimens significantly vary in body length (see Measurements). Colour of pronotum varies from entirely black to brown. In
<specimenCount id="9D21F7D76179A846FDB7F95CF47FF88C" box="[595,616,1771,1797]" pageId="5" pageNumber="126" type="female"></specimenCount>
<typeStatus id="549C82FC6179A846FD8BF95CF4C6F88C" box="[623,721,1771,1797]" pageId="5" pageNumber="126" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2747DD6179A846FD13F95CF570F88C" authorityName="Pic" authorityYear="1930" box="[759,871,1771,1797]" class="Insecta" family="Mycteridae" genus="Physciolagria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="126" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="B953E04C6179A846FD13F95CF570F88C" box="[759,871,1771,1797]" italics="true" pageId="5" pageNumber="126">P. liturata</emphasis>
</taxonomicName>
the dark apical elytral spot is not touching sutural or apical margin leaving them narrowly yellow. Elytron with or without indistinct tracks of four vaguely raised longitudinal costae.
</paragraph>
</subSubSection>
<subSubSection id="C33D6FD56179A846FF23F8E0F4E6F8F8" box="[199,753,1879,1905]" pageId="5" pageNumber="126" type="biology_ecology">
<paragraph id="8B983C5E6179A846FF23F8E0F4E6F8F8" blockId="5.[151,1437,151,1905]" box="[199,753,1879,1905]" pageId="5" pageNumber="126">
<emphasis id="B953E04C6179A846FF23F8E0F73BF8F8" bold="true" box="[199,300,1879,1905]" pageId="5" pageNumber="126">Ecology.</emphasis>
Some specimens were attracted to light.
</paragraph>
</subSubSection>
</treatment>
</document>