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<mods:title id="5407206EE3BAFFDDD44E863449F0C922">Euscorpius sulfur sp n (Scorpiones Euscorpiidae) a new cave scorpion from Albania and northwestern Greece</mods:title>
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Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratopotype in dorsal (5) and ventral (6) views. Scale bars: 10 mm." captionText-2="Figures 7–14: E. sulfur sp. n. Figures 7, 9, 11–14. Male holotype, carapace and tergites I–III (7), coxosternal area and sternites (9), left legs I–IV, retrolateral aspect (11–14 respectively). Figures 8, 10. Female paratopotype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10)." captionText-3="Figures 15–22: E. sulfur sp. n. Figures 15. Male paratype, telson lateral. Figures 17–19. Male holotype, metasoma and telson lateral (17), dorsal (18), and ventral with sternite VII (19) views. Figures 16, 20–22. Female paratopotype, telson lateral (16), metasoma and telson lateral (20), dorsal (21), and ventral (22) views. Scale bars: 10 mm (17–22)." captionText-4="Figures 23–32. E. sulfur sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable (31) and fixed (32) finger dentition." captionText-5="Figures 33–41. E. sulfur sp. n., female paratopotype, pedipalp segments. Chela dorsal (33), external (34) and ventral (35) views. Patella dorsal (36), external (37) and ventral (38) views. Trochanter and femur dorsal (39) and ventral (40) views. Movable finger dentition (41) Trichobothrial pattern is indicated by white circles (33–39)." captionText-6="Figures 42–49. E. sulfur sp. n., female juvenile paratype from Turtle Cave, pedipalp segments.Chela dorsal (42), external (43) and ventral (44) views. Patella dorsal (45), external (46) and ventral (47) views. Trochanter and femur dorsal (48) and ventral (49) views." captionText-7="Figures 50–51. Figure 50. Map of distribution of E. sulfur sp. n. Figure 51. A map of Sulfur Cave. Surveyes by M. Audy, R. Bouda, drawn by M. Audy (Czech Speleological S ociety 2021)." captionText-8="Figures 52–53. Figure 52. The Sarandaporo Valley at the border between Albania and Greece. The massive Vromoner hypogene sinkhole, visible on the limestone hill in the foreground, communicates through vertical chimneys with Sulfur Cave. Photograph by M. Audy. Figure 53. The spacious VesmÍr (Universe) Dome, situated under the sinkhole (in Figure 51). Several hydrogene sulfide-rich thermal springs (26°C) are located on the passage floor under the dome. The gypsum accumulations are covered with sulfur. Photograph by M. Audy &amp; R. Bouda." figureDoi-0="http://doi.org/10.5281/zenodo.10833728" figureDoi-1="http://doi.org/10.5281/zenodo.10833730" figureDoi-2="http://doi.org/10.5281/zenodo.10833734" figureDoi-3="http://doi.org/10.5281/zenodo.10833736" figureDoi-4="http://doi.org/10.5281/zenodo.10833740" figureDoi-5="http://doi.org/10.5281/zenodo.10833742" figureDoi-6="http://doi.org/10.5281/zenodo.10833744" figureDoi-7="http://doi.org/10.5281/zenodo.10833746" figureDoi-8="http://doi.org/10.5281/zenodo.10833748" httpUri-0="https://zenodo.org/record/10833728/files/figure.png" httpUri-1="https://zenodo.org/record/10833730/files/figure.png" httpUri-2="https://zenodo.org/record/10833734/files/figure.png" httpUri-3="https://zenodo.org/record/10833736/files/figure.png" httpUri-4="https://zenodo.org/record/10833740/files/figure.png" httpUri-5="https://zenodo.org/record/10833742/files/figure.png" httpUri-6="https://zenodo.org/record/10833744/files/figure.png" httpUri-7="https://zenodo.org/record/10833746/files/figure.png" httpUri-8="https://zenodo.org/record/10833748/files/figure.png" pageId="4" pageNumber="3">Figures 1–53</figureCitation>
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<geoCoordinate id="EE195CBAFFE9A458FEF6F9B5FE2FF9F7" box="[274,418,1621,1645]" degrees="20.666667" direction="east" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="20.666668">20.666667°E</geoCoordinate>
, 
<figureCitation id="131626F8FFE9A458FE49F9B5FD89F9F7" box="[429,516,1621,1645]" captionStart="Figures 50–51" captionStartId="12.[115,192,1992,2014]" captionTargetBox="[113,1474,227,1958]" captionTargetId="figure-59@12.[113,1474,224,1977]" captionTargetPageId="12" captionText="Figures 50–51. Figure 50. Map of distribution of E. sulfur sp. n. Figure 51. A map of Sulfur Cave. Surveyes by M. Audy, R. Bouda, drawn by M. Audy (Czech Speleological S ociety 2021)." figureDoi="http://doi.org/10.5281/zenodo.10833746" httpUri="https://zenodo.org/record/10833746/files/figure.png" pageId="4" pageNumber="3">Figs. 51</figureCitation>
and 
<figureCitation id="131626F8FFE9A458FDDEF9B5FDDAF9F7" box="[570,599,1621,1645]" captionStart="Figures 52–53" captionStartId="13.[116,193,1724,1746]" captionTargetBox="[113,1474,238,1708]" captionTargetId="figure-172@13.[113,1474,238,1708]" captionTargetPageId="13" captionText="Figures 52–53. Figure 52. The Sarandaporo Valley at the border between Albania and Greece. The massive Vromoner hypogene sinkhole, visible on the limestone hill in the foreground, communicates through vertical chimneys with Sulfur Cave. Photograph by M. Audy. Figure 53. The spacious VesmÍr (Universe) Dome, situated under the sinkhole (in Figure 51). Several hydrogene sulfide-rich thermal springs (26°C) are located on the passage floor under the dome. The gypsum accumulations are covered with sulfur. Photograph by M. Audy &amp; R. Bouda." figureDoi="http://doi.org/10.5281/zenodo.10833748" httpUri="https://zenodo.org/record/10833748/files/figure.png" pageId="4" pageNumber="3">53</figureCitation>
), 
<specimenCount id="9D2BF1F4FFE9A458FD88F9B5FD00F9F7" box="[620,653,1621,1645]" count="1" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
(
<typeStatus id="549684DFFFE9A458FD78F9B5FD70F9F7" box="[668,765,1621,1645]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
) 
<specimenCount id="9D2BF1F4FFE9A458FF96F995FF19F917" box="[114,148,1653,1677]" count="3" pageId="4" pageNumber="3" type="male">3♂</specimenCount>
<specimenCount id="9D2BF1F4FFE9A458FF70F995FF3DF917" box="[148,176,1653,1677]" count="5" pageId="4" pageNumber="3" type="female">5♀</specimenCount>
<specimenCount id="9D2BF1F4FFE9A458FF54F995FE8FF916" box="[176,258,1653,1677]" pageId="4" pageNumber="3" type="female">1juv. ♀</specimenCount>
(
<typeStatus id="549684DFFFE9A458FEEAF995FE2AF917" box="[270,423,1653,1677]" pageId="4" pageNumber="3" type="paratopotype">paratopotypes</typeStatus>
) and 
<collectingCounty id="62F342F1FFE9A458FE3BF995FDD0F917" box="[479,605,1653,1677]" pageId="4" pageNumber="3">Turtle Cave</collectingCounty>
, 
<geoCoordinate id="EE195CBAFFE9A458FD82F995FC89F916" box="[614,772,1653,1677]" degrees="40" direction="north" minutes="05" orientation="latitude" pageId="4" pageNumber="3" precision="1" seconds="53.59" value="40.098217">40°05'53.59''N</geoCoordinate>
<geoCoordinate id="EE195CBAFFE9A458FF96F975FE9DF936" box="[114,272,1685,1709]" degrees="20" direction="east" minutes="40" orientation="longitude" pageId="4" pageNumber="3" precision="1" seconds="33.86" value="20.676071">20°40'33.86''E</geoCoordinate>
, 
<specimenCount id="9D2BF1F4FFE9A458FEFBF975FEE6F937" box="[287,363,1685,1709]" pageId="4" pageNumber="3" type="male">1juv.♂</specimenCount>
<specimenCount id="9D2BF1F4FFE9A458FE8FF975FE3DF937" box="[363,432,1685,1709]" pageId="4" pageNumber="3" type="female">1juv.♀</specimenCount>
(
<typeStatus id="549684DFFFE9A458FE27F975FDA0F937" box="[451,557,1685,1709]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
, 
<figureCitation id="131626F8FFE9A458FDDAF975FD48F937" box="[574,709,1685,1709]" captionStart="Figures 42–49" captionStartId="10.[113,190,2000,2022]" captionTargetBox="[144,1411,215,1976]" captionTargetId="figure-66@10.[113,1474,209,1987]" captionTargetPageId="10" captionText="Figures 42–49. E. sulfur sp. n., female juvenile paratype from Turtle Cave, pedipalp segments.Chela dorsal (42), external (43) and ventral (44) views. Patella dorsal (45), external (46) and ventral (47) views. Trochanter and femur dorsal (48) and ventral (49) views." figureDoi="http://doi.org/10.5281/zenodo.10833744" httpUri="https://zenodo.org/record/10833744/files/figure.png" pageId="4" pageNumber="3">Figs. 42–49</figureCitation>
), 3– 
<date id="FF931CBDFFE9A458FD12F975FED2F957" pageId="4" pageNumber="3" value="2023-05-04" valueMax="2023-06-09" valueMin="2023-05-04">
<collectingDate id="EFD7E555FFE9A458FD12F975FED2F957" pageId="4" pageNumber="3" value="2023-05-04" valueMax="2023-06-09" valueMin="2023-05-04">4 May and 9 June 2023</collectingDate>
</date>
, leg. 
<collectorName id="26D85FABFFE9A458FE7EF955FE76F957" box="[410,507,1717,1741]" pageId="4" pageNumber="3">M. Audy</collectorName>
, S. M. Sarbu
</materialsCitation>
, 
<materialsCitation id="3B453020FFE9A458FD73F955FF64F976" collectionCode="T" collectorName="M. Kenesz" country="Albania" pageId="4" pageNumber="3" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED3CA2B8FFE9A458FD73F955FD24F956" box="[663,681,1717,1740]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="3">T</collectionCode>
. Brad &amp; 
<collectorName id="26D85FABFFE9A458FF96F935FF64F976" box="[114,233,1749,1772]" pageId="4" pageNumber="3">M. Kenesz</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33769F6FFE9A458FF96F8F5FD07F8D7" pageId="4" pageNumber="3" type="etymology">
<paragraph id="8B923A7DFFE9A458FF96F8F5FD07F8D7" blockId="4.[114,772,1813,1869]" pageId="4" pageNumber="3">
ETYMOLOGY. Named after the Sulfur Cave, the 
<typeStatus id="549684DFFFE9A458FD85F8F5FD02F8B7" box="[609,655,1813,1837]" pageId="4" pageNumber="3">type</typeStatus>
locality, to indicate an exceptional habitat of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C33769F6FFE9A458FF96F895FBB8FA10" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="8B923A7DFFE9A458FF96F895FBB8FA10" blockId="4.[114,772,1909,2029]" lastBlockId="4.[817,1476,1074,1418]" pageId="4" pageNumber="3">
DIAGNOSIS (
<specimenCount id="9D2BF1F4FFE9A458FF15F895FE85F817" box="[241,264,1909,1933]" count="1" pageId="4" pageNumber="3" type="male">♂</specimenCount>
<specimenCount id="9D2BF1F4FFE9A458FEECF895FE95F817" box="[264,280,1909,1933]" count="1" pageId="4" pageNumber="3" type="female">♀</specimenCount>
). Total length 
<quantity id="4CD59798FFE9A458FE49F895FDADF816" box="[429,544,1909,1933]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="4" pageNumber="3" unit="mm" value="31.5" valueMax="35.0" valueMin="28.0">28–35 mm</quantity>
. Color reddish yellow to brown, telson yellow. Pedipalp patella external trichobothria numbers: 4 
<emphasis id="B959E66FFFE9A458FF0FF855FE89F857" box="[235,260,1973,1997]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
, 4 
<emphasis id="B959E66FFFE9A458FEC5F855FEB7F857" box="[289,314,1973,1997]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
<subScript id="17A93838FFE9A458FEDEF825FECCF849" attach="left" box="[314,321,1989,2003]" fontSize="6" pageId="4" pageNumber="3">a</subScript>
, 2 
<emphasis id="B959E66FFFE9A458FEBAF855FE0FF857" box="[350,386,1973,1997]" italics="true" pageId="4" pageNumber="3">esb</emphasis>
, 4 
<emphasis id="B959E66FFFE9A458FE7AF856FE30F857" box="[414,445,1974,1997]" italics="true" pageId="4" pageNumber="3">em</emphasis>
, 4 
<emphasis id="B959E66FFFE9A458FE3DF856FE7AF857" box="[473,503,1974,1997]" italics="true" pageId="4" pageNumber="3">est</emphasis>
, 5–6 
<emphasis id="B959E66FFFE9A458FDCAF856FDCCF857" box="[558,577,1974,1997]" italics="true" pageId="4" pageNumber="3">et</emphasis>
, ventral aspect of patella with 8 trichobothria. Pectinal teeth number 
<quantity id="4CD59798FFE9A458FD5FF835FC89F877" box="[699,772,2005,2029]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="4" pageNumber="3" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6–8 in</quantity>
males, 
<quantity id="4CD59798FFE9A458FC98FBD2FC32FBD0" box="[892,959,1074,1098]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="4" pageNumber="3" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
females. Chelicerae yellow without reticulation. Male with pedipalp finger marginal profile 
<typeStatus id="549684DFFFE9A458FAF4FBB2FAB3FBF0" box="[1296,1342,1106,1130]" pageId="4" pageNumber="3">type</typeStatus>
C (KovařÍk &amp; ŠťÁhlavský, 2020: 2), female with fingers undulate. Dorsal metasomal carinae on segments I– IV irregularly granulated, mainly in male. Ventrolateral carinae on metasomal segments II–IV present and smooth. Metasoma V ventrally with median carina present. Metasoma finely granulated dorsally in male and smooth in female. Chela length/width ratio, 
<quantity id="4CD59798FFE9A458FAD8FAF2FA4EFAB0" box="[1340,1475,1298,1322]" metricMagnitude="-2" metricUnit="m" metricValue="6.0833" metricValueMax="6.1214" metricValueMin="6.0451999999999995" pageId="4" pageNumber="3" unit="in" value="2.395" valueMax="2.41" valueMin="2.38">2.38–2.41 in</quantity>
male, 
<quantity id="4CD59798FFE9A458FC92FAD2FC74FAD0" box="[886,1017,1330,1354]" metricMagnitude="0" metricUnit="m" metricValue="3.46202" metricValueMax="6.858" metricValueMin="0.06604" pageId="4" pageNumber="3" unit="in" value="136.3" valueMax="270.0" valueMin="2.6">2.60–270 in</quantity>
female. Metasoma IV length/width ratio, 2.0– 
<quantity id="4CD59798FFE9A458FC86FAB2FC2CFAF0" box="[866,929,1362,1386]" metricMagnitude="-2" metricUnit="m" metricValue="5.334" pageId="4" pageNumber="3" unit="in" value="2.1">2.1 in</quantity>
both sexes. Telson length/depth ratio, 
<quantity id="4CD59798FFE9A458FAB0FAB2FA4EFAF0" box="[1364,1475,1362,1386]" metricMagnitude="-2" metricUnit="m" metricValue="6.096" metricValueMax="6.35" metricValueMin="5.842" pageId="4" pageNumber="3" unit="in" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 in</quantity>
male, 
<quantity id="4CD59798FFE9A458FC97FA92FC53FA10" box="[883,990,1394,1418]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" metricValueMax="8.89" metricValueMin="8.382" pageId="4" pageNumber="3" unit="in" value="3.4" valueMax="3.5" valueMin="3.3">3.3–3.5 in</quantity>
female.
</paragraph>
</subSubSection>
<subSubSection id="C33769F6FFE9A457FCD5FA52FD04FB19" lastPageId="11" lastPageNumber="10" pageId="4" pageNumber="3" type="description">
<paragraph id="8B923A7DFFE9A458FCD5FA52FC22F9D0" blockId="4.[817,1476,1458,2026]" pageId="4" pageNumber="3">
DESCRIPTION (
<specimenCount id="9D2BF1F4FFE9A458FC2FFA52FC56FA50" box="[971,987,1458,1482]" count="1" pageId="4" pageNumber="3" type="female">♀</specimenCount>
<specimenCount id="9D2BF1F4FFE9A458FC3FFA52FC7FFA50" box="[987,1010,1458,1482]" count="1" pageId="4" pageNumber="3" type="male">♂</specimenCount>
). Total length 
<quantity id="4CD59798FFE9A458FB68FA52FB73FA53" box="[1164,1278,1458,1482]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="4" pageNumber="3" unit="mm" value="31.5" valueMax="35.0" valueMin="28.0">28–35 mm</quantity>
in both sexes. The habitus is shown in 
<figureCitation id="131626F8FFE9A458FBF6FA32FBF1FA70" box="[1042,1148,1490,1514]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStartId-0="3.[116,193,2003,2025]" captionStartId-1="5.[115,192,2058,2080]" captionTargetBox-0="[116,1477,229,1992]" captionTargetBox-1="[139,1427,206,2035]" captionTargetId-0="figure-39@3.[116,1477,226,1995]" captionTargetId-1="figure-52@5.[113,1474,191,2050]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–2: Euscorpius sulfur sp. n., females paratopotypes in vivo habitus under UV (1, photograph by Ruxandra Nitescu) and white (2, photograph by Traian Brad) light." captionText-1="Figures 3–6: Euscorpius sulfur sp. n. Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratopotype in dorsal (5) and ventral (6) views. Scale bars: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10833728" figureDoi-1="http://doi.org/10.5281/zenodo.10833730" httpUri-0="https://zenodo.org/record/10833728/files/figure.png" httpUri-1="https://zenodo.org/record/10833730/files/figure.png" pageId="4" pageNumber="3">Figs. 2–6</figureCitation>
. For position and distribution of trichobothria on pedipalps, see 
<figureCitation id="131626F8FFE9A458FB53FA12FAB2F990" box="[1207,1343,1522,1546]" captionStart="Figures 33–41" captionStartId="9.[116,193,1957,1979]" captionTargetBox="[137,1419,266,1906]" captionTargetId="figure-63@9.[113,1474,238,1937]" captionTargetPageId="9" captionText="Figures 33–41. E. sulfur sp. n., female paratopotype, pedipalp segments. Chela dorsal (33), external (34) and ventral (35) views. Patella dorsal (36), external (37) and ventral (38) views. Trochanter and femur dorsal (39) and ventral (40) views. Movable finger dentition (41) Trichobothrial pattern is indicated by white circles (33–39)." figureDoi="http://doi.org/10.5281/zenodo.10833742" httpUri="https://zenodo.org/record/10833742/files/figure.png" pageId="4" pageNumber="3">Figs. 33–38</figureCitation>
. For sexual dimorphism, see below in the description. For measurements, see 
<tableCitation id="C6AF0FC6FFE9A458FCBEF9D2FC24F9D0" box="[858,937,1586,1610]" captionStart="Table 1" captionStartId="4.[114,170,994,1015]" captionText="Table 1. Comparative measurements of types of Euscorpius sulfur sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="4" pageNumber="3">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B923A7DFFE9A458FCD5F9B1FAA9F930" blockId="4.[817,1476,1458,2026]" pageId="4" pageNumber="3">
<emphasis id="B959E66FFFE9A458FCD5F9B1FC21F9F3" bold="true" box="[817,940,1617,1641]" pageId="4" pageNumber="3">Coloration</emphasis>
(
<figureCitation id="131626F8FFE9A458FC58F9B2FBA8F9F0" box="[956,1061,1618,1642]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStartId-0="3.[116,193,2003,2025]" captionStartId-1="5.[115,192,2058,2080]" captionTargetBox-0="[116,1477,229,1992]" captionTargetBox-1="[139,1427,206,2035]" captionTargetId-0="figure-39@3.[116,1477,226,1995]" captionTargetId-1="figure-52@5.[113,1474,191,2050]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–2: Euscorpius sulfur sp. n., females paratopotypes in vivo habitus under UV (1, photograph by Ruxandra Nitescu) and white (2, photograph by Traian Brad) light." captionText-1="Figures 3–6: Euscorpius sulfur sp. n. Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratopotype in dorsal (5) and ventral (6) views. Scale bars: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10833728" figureDoi-1="http://doi.org/10.5281/zenodo.10833730" httpUri-0="https://zenodo.org/record/10833728/files/figure.png" httpUri-1="https://zenodo.org/record/10833730/files/figure.png" pageId="4" pageNumber="3">Figs. 2–6</figureCitation>
). Base color uniformly reddish yellow to brown including sternites, telson yellow, pedipalps reddish brown. Chelicerae yellow without reticulation.
</paragraph>
<paragraph id="8B923A7DFFE9A458FCD5F951FA22F870" blockId="4.[817,1476,1458,2026]" pageId="4" pageNumber="3">
<emphasis id="B959E66FFFE9A458FCD5F951FBC0F950" bold="true" box="[817,1101,1713,1738]" pageId="4" pageNumber="3">Carapace and mesosoma</emphasis>
(
<figureCitation id="131626F8FFE9A458FBB8F952FB5EF950" box="[1116,1235,1714,1738]" captionStart="Figures 7–14" captionStartId="6.[116,193,1958,1980]" captionTargetBox="[113,1474,268,1924]" captionTargetId="figure-75@6.[113,1474,268,1924]" captionTargetPageId="6" captionText="Figures 7–14: E. sulfur sp. n. Figures 7, 9, 11–14. Male holotype, carapace and tergites I–III (7), coxosternal area and sternites (9), left legs I–IV, retrolateral aspect (11–14 respectively). Figures 8, 10. Female paratopotype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10)." figureDoi="http://doi.org/10.5281/zenodo.10833734" httpUri="https://zenodo.org/record/10833734/files/figure.png" pageId="4" pageNumber="3">Figs. 7–10</figureCitation>
). Carapace very finely granulated with several smooth areas; carinae absent. Anterior margin of carapace slightly concave. Carapace with two lateral eyes. Tergites very finely granulated, more so in male, without carinae developed. Tergite VII lacking median and paired lateral carinae. Sternites III–VII smooth and lustrous; VII lacking median and paired lateral carinae. Stigmata small, narrow ellipical. Pectinal teeth number 6–8 (1 x 6, 2 x 7, 9 x 8) in males and 6–7 (9 x 6, 5 x 7) in females, fulcra present. Pectines with 3 marginal lamellae and 5–6 middle lamellae.
</paragraph>
<caption id="DF526AF5FFE8A459FF97F7EAFDEBF7A7" ID-DOI="http://doi.org/10.5281/zenodo.10833730" ID-Zenodo-Dep="10833730" httpUri="https://zenodo.org/record/10833730/files/figure.png" pageId="5" pageNumber="4" startId="5.[115,192,2058,2080]" targetBox="[139,1427,206,2035]" targetPageId="5" targetType="figure">
<paragraph id="8B923A7DFFE8A459FF97F7EAFDEBF7A7" blockId="5.[115,1473,2058,2109]" pageId="5" pageNumber="4">
<emphasis id="B959E66FFFE8A459FF97F7EAFF67F785" bold="true" box="[115,234,2058,2080]" pageId="5" pageNumber="4">Figures 3–6</emphasis>
: 
<taxonomicName id="4C2D41FEFFE8A459FF1CF7EBFE2EF785" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[248,419,2058,2080]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="sulfur" status="sp. nov.">
<emphasis id="B959E66FFFE8A459FF1CF7EBFE2EF785" box="[248,419,2058,2080]" italics="true" pageId="5" pageNumber="4">Euscorpius sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE8A459FE4EF7EAFE6CF7BA" box="[426,481,2058,2080]" pageId="5" pageNumber="4" rank="species">
<emphasis id="B959E66FFFE8A459FE4EF7EAFE4CF785" bold="true" box="[426,449,2058,2079]" pageId="5" pageNumber="4">sp</emphasis>
. 
<emphasis id="B959E66FFFE8A459FE29F7EAFE57F785" bold="true" box="[461,474,2058,2079]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B959E66FFFE8A459FE03F7EAFDD2F785" bold="true" box="[487,607,2058,2080]" pageId="5" pageNumber="4">Figures 3–4</emphasis>
. Male holotype in dorsal (3) and ventral (4) views. 
<emphasis id="B959E66FFFE8A459FBBCF7EAFB5DF785" bold="true" box="[1112,1232,2058,2080]" pageId="5" pageNumber="4">Figures 5–6</emphasis>
. Female paratopotype in dorsal (5) and ventral (6) views. Scale bars: 10 mm.
</paragraph>
</caption>
<caption id="DF526AF5FFEBA45AFF90F846FF68F86F" ID-DOI="http://doi.org/10.5281/zenodo.10833734" ID-Zenodo-Dep="10833734" httpUri="https://zenodo.org/record/10833734/files/figure.png" pageId="6" pageNumber="5" startId="6.[116,193,1958,1980]" targetBox="[113,1474,268,1924]" targetPageId="6" targetType="figure">
<paragraph id="8B923A7DFFEBA45AFF90F846FF68F86F" blockId="6.[116,1474,1958,2037]" pageId="6" pageNumber="5">
<emphasis id="B959E66FFFEBA45AFF90F846FF7AF821" bold="true" box="[116,247,1958,1980]" pageId="6" pageNumber="5">Figures 7–14</emphasis>
: 
<taxonomicName id="4C2D41FEFFEBA45AFEE0F847FEDAF821" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[260,343,1958,1980]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFEBA45AFEE0F847FEDAF821" box="[260,343,1958,1980]" italics="true" pageId="6" pageNumber="5">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFEBA45AFEBAF846FE19F826" box="[350,404,1958,1980]" pageId="6" pageNumber="5" rank="species">
<emphasis id="B959E66FFFEBA45AFEBAF846FEF8F821" bold="true" box="[350,373,1958,1979]" pageId="6" pageNumber="5">sp</emphasis>
. 
<emphasis id="B959E66FFFEBA45AFE65F846FE03F821" bold="true" box="[385,398,1958,1979]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B959E66FFFEBA45AFE7FF846FE77F821" bold="true" box="[411,506,1958,1980]" pageId="6" pageNumber="5">Figures 7</emphasis>
, 
<emphasis id="B959E66FFFEBA45AFDE3F846FD9EF821" bold="true" box="[519,531,1958,1979]" pageId="6" pageNumber="5">9</emphasis>
, 
<emphasis id="B959E66FFFEBA45AFDFBF846FDD7F821" bold="true" box="[543,602,1958,1980]" pageId="6" pageNumber="5">11–14</emphasis>
. Male holotype, carapace and tergites I–III (7), coxosternal area and sternites (9), left legs I–IV, retrolateral aspect (11–14 respectively). 
<emphasis id="B959E66FFFEBA45AFDC3F823FD08F842" bold="true" box="[551,645,1987,2009]" pageId="6" pageNumber="5">Figures 8</emphasis>
, 
<emphasis id="B959E66FFFEBA45AFD74F823FD25F842" bold="true" box="[656,680,1987,2008]" pageId="6" pageNumber="5">10</emphasis>
. Female paratopotype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10).
</paragraph>
</caption>
<caption id="DF526AF5FFEAA45BFF95F833FC85F7B8" ID-DOI="http://doi.org/10.5281/zenodo.10833736" ID-Zenodo-Dep="10833736" httpUri="https://zenodo.org/record/10833736/files/figure.png" pageId="7" pageNumber="6" startId="7.[113,190,2003,2025]" targetBox="[135,1474,256,1989]" targetPageId="7" targetType="figure">
<paragraph id="8B923A7DFFEAA45BFF95F833FC85F7B8" blockId="7.[113,1472,2003,2083]" pageId="7" pageNumber="6">
<emphasis id="B959E66FFFEAA45BFF95F833FE8FF872" bold="true" box="[113,258,2003,2025]" pageId="7" pageNumber="6">Figures 15–22</emphasis>
: 
<taxonomicName id="4C2D41FEFFEAA45BFEEBF834FEEEF872" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[271,355,2003,2025]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFEAA45BFEEBF834FEEEF872" box="[271,355,2003,2025]" italics="true" pageId="7" pageNumber="6">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFEAA45BFE8FF834FE2FF873" box="[363,418,2004,2025]" pageId="7" pageNumber="6" rank="species">
<emphasis id="B959E66FFFEAA45BFE8FF834FE0FF873" bold="true" box="[363,386,2004,2025]" pageId="7" pageNumber="6">sp</emphasis>
. 
<emphasis id="B959E66FFFEAA45BFE6AF834FE16F873" bold="true" box="[398,411,2004,2025]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B959E66FFFEAA45BFE4DF833FD98F872" bold="true" box="[425,533,2003,2025]" pageId="7" pageNumber="6">Figures 15</emphasis>
. Male paratype, telson lateral. 
<emphasis id="B959E66FFFEAA45BFCA0F833FC59F872" bold="true" box="[836,980,2003,2025]" pageId="7" pageNumber="6">Figures 17–19</emphasis>
. Male holotype, metasoma and telson lateral (17), dorsal (18), and ventral with sternite VII (19) views. 
<emphasis id="B959E66FFFEAA45BFD88F810FD5AF79F" bold="true" box="[620,727,2032,2054]" pageId="7" pageNumber="6">Figures 16</emphasis>
, 
<emphasis id="B959E66FFFEAA45BFD07F810FC92F79F" bold="true" box="[739,799,2032,2054]" pageId="7" pageNumber="6">20–22</emphasis>
. Female paratopotype, telson lateral (16), metasoma and telson lateral (20), dorsal (21), and ventral (22) views. Scale bars: 10 mm (17–22).
</paragraph>
</caption>
<caption id="DF526AF5FFE5A454FF90F9BEFA3CF90B" ID-DOI="http://doi.org/10.5281/zenodo.10833740" ID-Zenodo-Dep="10833740" httpUri="https://zenodo.org/record/10833740/files/figure.png" pageId="8" pageNumber="7" startId="8.[116,193,1630,1652]" targetBox="[123,1437,256,1600]" targetPageId="8" targetType="figure">
<paragraph id="8B923A7DFFE5A454FF90F9BEFA3CF90B" blockId="8.[116,1474,1630,1681]" pageId="8" pageNumber="7">
<emphasis id="B959E66FFFE5A454FF90F9BEFE8FF9E9" bold="true" box="[116,258,1630,1652]" pageId="8" pageNumber="7">Figures 23–32</emphasis>
. 
<taxonomicName id="4C2D41FEFFE5A454FEEAF9BFFEEDF9EE" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[270,352,1631,1652]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFE5A454FEEAF9BFFEEDF9EE" box="[270,352,1631,1652]" italics="true" pageId="8" pageNumber="7">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE5A454FE82F9BFFE16F9EE" box="[358,411,1631,1652]" pageId="8" pageNumber="7" rank="species">
<emphasis id="B959E66FFFE5A454FE82F9BFFEF0F9EE" bold="true" box="[358,381,1631,1652]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="B959E66FFFE5A454FE6CF9BFFE18F9EE" bold="true" box="[392,405,1631,1652]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable (31) and fixed (32) finger dentition.
</paragraph>
</caption>
<paragraph id="8B923A7DFFE5A454FF95F95AFC5BF888" blockId="8.[113,771,1722,2034]" lastBlockId="8.[816,1474,1722,2034]" pageId="8" pageNumber="7">
<emphasis id="B959E66FFFE5A454FF95F95AFEFFF948" bold="true" box="[113,370,1722,1746]" pageId="8" pageNumber="7">Metasoma and telson</emphasis>
(
<figureCitation id="131626F8FFE5A454FE6CF95AFD9EF948" box="[392,531,1722,1746]" captionStart="Figures 15–22" captionStartId="7.[113,190,2003,2025]" captionTargetBox="[135,1474,256,1989]" captionTargetId="figure-76@7.[113,1474,234,1995]" captionTargetPageId="7" captionText="Figures 15–22: E. sulfur sp. n. Figures 15. Male paratype, telson lateral. Figures 17–19. Male holotype, metasoma and telson lateral (17), dorsal (18), and ventral with sternite VII (19) views. Figures 16, 20–22. Female paratopotype, telson lateral (16), metasoma and telson lateral (20), dorsal (21), and ventral (22) views. Scale bars: 10 mm (17–22)." figureDoi="http://doi.org/10.5281/zenodo.10833736" httpUri="https://zenodo.org/record/10833736/files/figure.png" pageId="8" pageNumber="7">Figs. 15–22</figureCitation>
). Metasoma sparsely hirsute. Metasoma I–V very finely granulated in male, several fine granules also present on lateral surfaces of metasoma I and V in both sexes. Dorsal carinae on metasomal segments I–V irregularly granulated in both sexes, more in males; dorsolateral carinae absent; ventrolateral carinae present or indicated and smooth on segments I–IV, granulated on segment V. Metasoma V ventrally granulated with median carina present, metasoma I–IV with ventral median carinae absent. Anal arch with small pigmented granules. Telson rather smooth, elongate in female and swollen in male, with annular ring indicated in both sexes. Aculeus short, more curved in male.
</paragraph>
<paragraph id="8B923A7DFFE5A457FCD4F8FAFD4FFE39" blockId="8.[816,1474,1722,2034]" lastBlockId="11.[113,771,234,547]" lastPageId="11" lastPageNumber="10" pageId="8" pageNumber="7">
<emphasis id="B959E66FFFE5A454FCD4F8FAFC12F8A8" bold="true" box="[816,927,1818,1842]" pageId="8" pageNumber="7">Pedipalps</emphasis>
(
<figureCitation id="131626F8FFE5A454FC51F8FAFBCDF8A8" box="[949,1088,1818,1842]" captionStart-0="Figures 23–32" captionStart-1="Figures 33–41" captionStart-2="Figures 42–49" captionStartId-0="8.[116,193,1630,1652]" captionStartId-1="9.[116,193,1957,1979]" captionStartId-2="10.[113,190,2000,2022]" captionTargetBox-0="[123,1437,256,1600]" captionTargetBox-1="[137,1419,266,1906]" captionTargetBox-2="[144,1411,215,1976]" captionTargetId-0="figure-250@8.[113,1474,234,1623]" captionTargetId-1="figure-63@9.[113,1474,238,1937]" captionTargetId-2="figure-66@10.[113,1474,209,1987]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="Figures 23–32. E. sulfur sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable (31) and fixed (32) finger dentition." captionText-1="Figures 33–41. E. sulfur sp. n., female paratopotype, pedipalp segments. Chela dorsal (33), external (34) and ventral (35) views. Patella dorsal (36), external (37) and ventral (38) views. Trochanter and femur dorsal (39) and ventral (40) views. Movable finger dentition (41) Trichobothrial pattern is indicated by white circles (33–39)." captionText-2="Figures 42–49. E. sulfur sp. n., female juvenile paratype from Turtle Cave, pedipalp segments.Chela dorsal (42), external (43) and ventral (44) views. Patella dorsal (45), external (46) and ventral (47) views. Trochanter and femur dorsal (48) and ventral (49) views." figureDoi-0="http://doi.org/10.5281/zenodo.10833740" figureDoi-1="http://doi.org/10.5281/zenodo.10833742" figureDoi-2="http://doi.org/10.5281/zenodo.10833744" httpUri-0="https://zenodo.org/record/10833740/files/figure.png" httpUri-1="https://zenodo.org/record/10833742/files/figure.png" httpUri-2="https://zenodo.org/record/10833744/files/figure.png" pageId="8" pageNumber="7">Figs. 23–49</figureCitation>
). Pedipalps very sparsely hirsute. Patella with 23–24 (4 
<emphasis id="B959E66FFFE5A454FBF7F8DAFBA1F8C8" box="[1043,1068,1850,1874]" italics="true" pageId="8" pageNumber="7">eb</emphasis>
, 4 
<emphasis id="B959E66FFFE5A454FBA2F8DAFBD2F8C8" box="[1094,1119,1850,1874]" italics="true" pageId="8" pageNumber="7">eb</emphasis>
<subScript id="17A93838FFE5A454FBBBF8AAFBEBF8C2" attach="left" box="[1119,1126,1866,1880]" fontSize="6" pageId="8" pageNumber="7">a</subScript>
, 2 
<emphasis id="B959E66FFFE5A454FB64F8DAFB29F8C8" box="[1152,1188,1850,1874]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
, 4 
<emphasis id="B959E66FFFE5A454FB5AF8DBFB50F8C8" box="[1214,1245,1851,1874]" italics="true" pageId="8" pageNumber="7">em</emphasis>
, 4 
<emphasis id="B959E66FFFE5A454FB13F8DBFA98F8C8" box="[1271,1301,1851,1874]" italics="true" pageId="8" pageNumber="7">est</emphasis>
, 5–6 
<emphasis id="B959E66FFFE5A454FAADF8DBFAD1F8C8" box="[1353,1372,1851,1874]" italics="true" pageId="8" pageNumber="7">et</emphasis>
) external and 8 ventral trichobothria.Chela with 4 trichobothria in ventral series, of which 
<emphasis id="B959E66FFFE5A454FC0DF89BFC74F808" box="[1001,1017,1915,1938]" italics="true" pageId="8" pageNumber="7">V</emphasis>
<subScript id="17A93838FFE5A454FC1DF86AFB8CF802" attach="left" box="[1017,1025,1930,1944]" fontSize="6" pageId="8" pageNumber="7">4</subScript>
is located external to the ventroexternal carina, on the external surface near 
<emphasis id="B959E66FFFE5A454FB58F87AFB57F828" box="[1212,1242,1946,1970]" italics="true" pageId="8" pageNumber="7">Eb</emphasis>
<subScript id="17A93838FFE5A454FB3DF84AFB6CF822" attach="left" box="[1241,1249,1962,1976]" fontSize="6" pageId="8" pageNumber="7">1</subScript>
. Entire femur finely granulated, strongly ventrally and patella fine granulated irregularly. Femur with granulated developed carinae; ventroexternal carina incomplete. Patella with 5 complete carinae including irregular wide externomedian carina. Dorsal patella spur well developed in both sexes. Entire manus with fine, rounded granules, which do not form a median carina; only five chelal carinae developed. Male with pedipalp finger undulation profile 
<typeStatus id="549684DFFFE6A457FED2FE6BFEE9FE39" box="[310,356,395,419]" pageId="11" pageNumber="10">type</typeStatus>
C, female with fingers undulate.
</paragraph>
<caption id="DF526AF5FFE4A455FF90F845FD3CF86E" ID-DOI="http://doi.org/10.5281/zenodo.10833742" ID-Zenodo-Dep="10833742" httpUri="https://zenodo.org/record/10833742/files/figure.png" pageId="9" pageNumber="8" startId="9.[116,193,1957,1979]" targetBox="[137,1419,266,1906]" targetPageId="9" targetType="figure">
<paragraph id="8B923A7DFFE4A455FF90F845FD3CF86E" blockId="9.[116,1474,1957,2036]" pageId="9" pageNumber="8">
<emphasis id="B959E66FFFE4A455FF90F845FE88F820" bold="true" box="[116,261,1957,1979]" pageId="9" pageNumber="8">Figures 33–41</emphasis>
. 
<taxonomicName id="4C2D41FEFFE4A455FEF0F845FEE4F820" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[276,361,1957,1978]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFE4A455FEF0F845FEE4F820" box="[276,361,1957,1978]" italics="true" pageId="9" pageNumber="8">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE4A455FE96F845FE27F820" box="[370,426,1957,1978]" pageId="9" pageNumber="8" rank="species">
<emphasis id="B959E66FFFE4A455FE96F845FE04F820" bold="true" box="[370,393,1957,1978]" pageId="9" pageNumber="8">sp</emphasis>
. 
<emphasis id="B959E66FFFE4A455FE73F845FE29F820" bold="true" box="[407,420,1957,1978]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, female paratopotype, pedipalp segments. Chela dorsal (33), external (34) and ventral (35) views. Patella dorsal (36), external (37) and ventral (38) views. Trochanter and femur dorsal (39) and ventral (40) views. Movable finger dentition (41) Trichobothrial pattern is indicated by white circles (33–39).
</paragraph>
</caption>
<caption id="DF526AF5FFE7A456FF95F830FAAAF799" ID-DOI="http://doi.org/10.5281/zenodo.10833744" ID-Zenodo-Dep="10833744" httpUri="https://zenodo.org/record/10833744/files/figure.png" pageId="10" pageNumber="9" startId="10.[113,190,2000,2022]" targetBox="[144,1411,215,1976]" targetPageId="10" targetType="figure">
<paragraph id="8B923A7DFFE7A456FF95F830FAAAF799" blockId="10.[113,1433,2000,2051]" pageId="10" pageNumber="9">
<emphasis id="B959E66FFFE7A456FF95F830FF72F87F" bold="true" box="[113,255,2000,2022]" pageId="10" pageNumber="9">Figures 42–49</emphasis>
. 
<taxonomicName id="4C2D41FEFFE7A456FEEEF831FED1F87F" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[266,348,2000,2022]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFE7A456FEEEF831FED1F87F" box="[266,348,2000,2022]" italics="true" pageId="10" pageNumber="9">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE7A456FE85F830FE1BF87C" box="[353,406,2000,2022]" pageId="10" pageNumber="9" rank="species">
<emphasis id="B959E66FFFE7A456FE85F830FEF5F87F" bold="true" box="[353,376,2000,2021]" pageId="10" pageNumber="9">sp</emphasis>
. 
<emphasis id="B959E66FFFE7A456FE67F830FE1DF87F" bold="true" box="[387,400,2000,2021]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, female juvenile paratype from Turtle Cave, pedipalp segments. Chela dorsal (42), external (43) and ventral (44) views. Patella dorsal (45), external (46) and ventral (47) views. Trochanter and femur dorsal (48) and ventral (49) views.
</paragraph>
</caption>
<paragraph id="8B923A7DFFE6A457FF95FE4BFE2AFDB9" blockId="11.[113,771,234,547]" pageId="11" pageNumber="10">
<emphasis id="B959E66FFFE6A457FF95FE4BFF2BFE59" bold="true" box="[113,166,427,451]" pageId="11" pageNumber="10">Legs</emphasis>
(
<figureCitation id="131626F8FFE6A457FF5FFE4BFECEFE59" box="[187,323,427,451]" captionStart="Figures 7–14" captionStartId="6.[116,193,1958,1980]" captionTargetBox="[113,1474,268,1924]" captionTargetId="figure-75@6.[113,1474,268,1924]" captionTargetPageId="6" captionText="Figures 7–14: E. sulfur sp. n. Figures 7, 9, 11–14. Male holotype, carapace and tergites I–III (7), coxosternal area and sternites (9), left legs I–IV, retrolateral aspect (11–14 respectively). Figures 8, 10. Female paratopotype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10)." figureDoi="http://doi.org/10.5281/zenodo.10833734" httpUri="https://zenodo.org/record/10833734/files/figure.png" pageId="11" pageNumber="10">Figs. 11–14</figureCitation>
). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules.
</paragraph>
<paragraph id="8B923A7DFFE6A457FF95FDABFDA9FD59" blockId="11.[113,771,587,1157]" pageId="11" pageNumber="10">
AFFINITIES. 
<taxonomicName id="4C2D41FEFFE6A457FF15FDABFE22FDF9" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[241,431,587,611]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="sulfur" status="sp. nov.">
<emphasis id="B959E66FFFE6A457FF15FDABFE22FDF9" box="[241,431,587,611]" italics="true" pageId="11" pageNumber="10">Euscorpius sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE6A457FE53FDABFE78FDF8" box="[439,501,587,611]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B959E66FFFE6A457FE53FDABFE5DFDF9" bold="true" box="[439,464,587,611]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="B959E66FFFE6A457FE3BFDABFE63FDF9" bold="true" box="[479,494,587,611]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
is morphologically most similar to 
<taxonomicName id="4C2D41FEFFE6A457FF08FD8BFD85FD19" authority="Birula, 1903" authorityName="Birula" authorityYear="1903" box="[236,520,619,643]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="candiota">
<emphasis id="B959E66FFFE6A457FF08FD8BFEE3FD19" box="[236,366,619,643]" italics="true" pageId="11" pageNumber="10">E. candiota</emphasis>
Birula, 1903
</taxonomicName>
from 
<collectingCountry id="F33A7AEDFFE6A457FDBCFD8BFD2EFD19" box="[600,675,619,643]" name="Greece" pageId="11" pageNumber="10">Greece</collectingCountry>
(
<collectingRegion id="49E9F49FFFE6A457FD5CFD8BFD7AFD19" box="[696,759,619,643]" country="Greece" name="Kriti" pageId="11" pageNumber="10">Crete</collectingRegion>
), which differs by having ventral aspect of pedipalp patella with 9–10 trichobothria (
<quantity id="4CD59798FFE6A457FEA3FD4BFEFDFD59" box="[327,368,683,707]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="11" pageNumber="10" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C2D41FEFFE6A457FE92FD4BFE5FFD59" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[374,466,683,707]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFE6A457FE92FD4BFE5FFD59" box="[374,466,683,707]" italics="true" pageId="11" pageNumber="10">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE6A457FE3DFD4BFD9EFD59" box="[473,531,683,707]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B959E66FFFE6A457FE3DFD4BFE7FFD59" bold="true" box="[473,498,683,707]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="B959E66FFFE6A457FE1BFD4BFD83FD59" bold="true" box="[511,526,683,707]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8B923A7DFFE6A457FF45FD2BFD90FC19" blockId="11.[113,771,587,1157]" pageId="11" pageNumber="10">
Geographically the closest cave species, 
<taxonomicName id="4C2D41FEFFE6A457FD82FD2BFEF6FC99" authority="Tropea &amp; Ozimec, 2020" authorityName="Tropea &amp; Ozimec" authorityYear="2020" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="biokovensis">
<emphasis id="B959E66FFFE6A457FD82FD2BFC8EFD79" box="[614,771,715,739]" italics="true" pageId="11" pageNumber="10">E. biokovensis</emphasis>
Tropea &amp; Ozimec, 2020
</taxonomicName>
described from caves from 
<collectingCountry id="F33A7AEDFFE6A457FD50FD0BFC8EFC99" box="[692,771,747,771]" name="Croatia" pageId="11" pageNumber="10">Croatia</collectingCountry>
and 
<collectingCountry id="F33A7AEDFFE6A457FF4EFCEBFE08FCB9" box="[170,389,779,803]" name="Bosnia and Herzegovina" pageId="11" pageNumber="10">Bosnia-Herzegovina</collectingCountry>
differs by having metasoma V ventral with median carina obsolete to absent and male with pedipalp finger marginal profile 
<typeStatus id="549684DFFFE6A457FE23FCABFE78FCF9" box="[455,501,843,867]" pageId="11" pageNumber="10">type</typeStatus>
B (
<typeStatus id="549684DFFFE6A457FDFEFCABFDC4FCF9" box="[538,585,843,867]" pageId="11" pageNumber="10">type</typeStatus>
C in 
<taxonomicName id="4C2D41FEFFE6A457FD66FCABFD50FCF9" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" box="[642,733,843,867]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFE6A457FD66FCABFD50FCF9" box="[642,733,843,867]" italics="true" pageId="11" pageNumber="10">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE6A457FD06FCABFF0BFC18" pageId="11" pageNumber="10" rank="species">
<emphasis id="B959E66FFFE6A457FD06FCABFD76FCF9" bold="true" box="[738,763,843,867]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="B959E66FFFE6A457FF95FC8BFF0DFC19" bold="true" box="[113,128,875,899]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
, see KovařÍk &amp; ŠťÁhlavský, 2020: 2).
</paragraph>
<paragraph id="8B923A7DFFE6A457FF45FC6BFD04FB19" blockId="11.[113,771,587,1157]" pageId="11" pageNumber="10">
Some other species described recently from the region differ in having 5–6 trichobothria in 
<emphasis id="B959E66FFFE6A457FDE9FC4BFDABFC59" box="[525,550,939,963]" italics="true" pageId="11" pageNumber="10">eb</emphasis>
series on pedipalp patella external (
<taxonomicName id="4C2D41FEFFE6A457FEC8FC2BFC8EFC79" authority="KovarIk &amp; StAhlavsky, 2020" authorityName="KovarIk &amp; StAhlavsky" authorityYear="2020" box="[300,771,971,997]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="bonacinai">
<emphasis id="B959E66FFFE6A457FEC8FC2BFE35FC79" box="[300,440,971,995]" italics="true" pageId="11" pageNumber="10">E. bonacinai</emphasis>
KovařÍk &amp; ŠťÁhlavský, 2020
</taxonomicName>
from 
<collectingCountry id="F33A7AEDFFE6A457FF54FC0BFE89FB99" box="[176,260,1003,1027]" name="Albania" pageId="11" pageNumber="10">Albania</collectingCountry>
or 
<taxonomicName id="4C2D41FEFFE6A457FED7FC0BFC8EFB99" authority="KovarIk &amp; StAhlavsky, 2020" authorityName="KovarIk &amp; StAhlavsky" authorityYear="2020" box="[307,771,1003,1029]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="kabateki">
<emphasis id="B959E66FFFE6A457FED7FC0BFE3DFB99" box="[307,432,1003,1027]" italics="true" pageId="11" pageNumber="10">E. kabateki</emphasis>
KovařÍk &amp; ŠťÁhlavský, 2020
</taxonomicName>
from 
<collectingCountry id="F33A7AEDFFE6A457FF4BFBEBFF72FBB9" box="[175,255,1035,1059]" name="Greece" pageId="11" pageNumber="10">Greece</collectingCountry>
). 
<taxonomicName id="4C2D41FEFFE6A457FEF0FBEBFD48FBB9" authority="KovarIk &amp; StAhlavsky, 2020" authorityName="KovarIk &amp; StAhlavsky" authorityYear="2020" box="[276,709,1035,1061]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="janstai">
<emphasis id="B959E66FFFE6A457FEF0FBEBFEF3FBB8" box="[276,382,1035,1058]" italics="true" pageId="11" pageNumber="10">E. janstai</emphasis>
KovařÍk &amp; ŠťÁhlavský, 2020
</taxonomicName>
from North 
<collectingCountry id="F33A7AEDFFE6A457FF51FBCBFEA6FBD9" box="[181,299,1067,1091]" name="Macedonia" pageId="11" pageNumber="10">Macedonia</collectingCountry>
differs by darker colored telson and legs and male with pedipalp finger marginal profile 
<typeStatus id="549684DFFFE6A457FDD0FBABFDEFFBF9" box="[564,610,1099,1123]" pageId="11" pageNumber="10">type</typeStatus>
A (
<typeStatus id="549684DFFFE6A457FD60FBABFD3EFBF9" box="[644,691,1099,1123]" pageId="11" pageNumber="10">type</typeStatus>
C in 
<taxonomicName id="4C2D41FEFFE6A457FD0FFBABFF22FB19" authority="Kovařík &amp; Audy &amp; Sarbu &amp; Fet, 2023" authorityName="Kovařík &amp; Audy &amp; Sarbu &amp; Fet" authorityYear="2023" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="sulfur" status="sp. n.">
<emphasis id="B959E66FFFE6A457FD0FFBABFF22FB19" italics="true" pageId="11" pageNumber="10">E. sulfur</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5B14FFE6A457FF52FB8BFF7FFB18" box="[182,242,1131,1155]" pageId="11" pageNumber="10" rank="species">
<emphasis id="B959E66FFFE6A457FF52FB8BFF42FB19" bold="true" box="[182,207,1131,1155]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="B959E66FFFE6A457FF39FB8BFF61FB19" bold="true" box="[221,236,1131,1155]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
, see KovařÍk &amp; ŠťÁhlavský, 2020: 2).
</paragraph>
</subSubSection>
<subSubSection id="C33769F6FFE6A457FF95FB4BFDFBFAD9" pageId="11" pageNumber="10" type="distribution">
<paragraph id="8B923A7DFFE6A457FF95FB4BFDFBFAD9" blockId="11.[113,771,1195,1347]" pageId="11" pageNumber="10">
DISTRIBUTION. Sulfidic caves in the Vromoner Canyon, Sarandaporo Valley, on the border between 
<collectingCountry id="F33A7AEDFFE6A457FD92FB2BFD47FB79" box="[630,714,1227,1251]" name="Albania" pageId="11" pageNumber="10">Albania</collectingCountry>
and 
<collectingCountry id="F33A7AEDFFE6A457FF95FB0BFF31FA99" box="[113,188,1259,1283]" name="Greece" pageId="11" pageNumber="10">Greece</collectingCountry>
(
<figureCitation id="131626F8FFE6A457FF32FB0BFEE9FA98" box="[214,356,1258,1283]" captionStart-0="Figures 50–51" captionStart-1="Figures 52–53" captionStartId-0="12.[115,192,1992,2014]" captionStartId-1="13.[116,193,1724,1746]" captionTargetBox-0="[113,1474,227,1958]" captionTargetBox-1="[113,1474,238,1708]" captionTargetId-0="figure-59@12.[113,1474,224,1977]" captionTargetId-1="figure-172@13.[113,1474,238,1708]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figures 50–51. Figure 50. Map of distribution of E. sulfur sp. n. Figure 51. A map of Sulfur Cave. Surveyes by M. Audy, R. Bouda, drawn by M. Audy (Czech Speleological S ociety 2021)." captionText-1="Figures 52–53. Figure 52. The Sarandaporo Valley at the border between Albania and Greece. The massive Vromoner hypogene sinkhole, visible on the limestone hill in the foreground, communicates through vertical chimneys with Sulfur Cave. Photograph by M. Audy. Figure 53. The spacious VesmÍr (Universe) Dome, situated under the sinkhole (in Figure 51). Several hydrogene sulfide-rich thermal springs (26°C) are located on the passage floor under the dome. The gypsum accumulations are covered with sulfur. Photograph by M. Audy &amp; R. Bouda." figureDoi-0="http://doi.org/10.5281/zenodo.10833746" figureDoi-1="http://doi.org/10.5281/zenodo.10833748" httpUri-0="https://zenodo.org/record/10833746/files/figure.png" httpUri-1="https://zenodo.org/record/10833748/files/figure.png" pageId="11" pageNumber="10">Figs. 50–53</figureCitation>
). The majority of specimens were observed within the Albanian territory; the entrance to the Sulfur Cave lies on the Greek side of the border.
</paragraph>
</subSubSection>
</treatment>
</document>