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<document id="0CBCCC639584557C95EF88983846CA24" ID-DOI="10.5281/zenodo.178040" ID-GBIF-Dataset="2cbe1502-c057-4a0c-8f15-c1815dcd4d20" ID-ISSN="1175-5326" ID-Zenodo-Dep="178040" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459879787698" checkinUser="plazi" docAuthor="Osawa, Masayuki" docDate="2007" docId="03809477FF90BF31ACF1EB1749839A96" docLanguage="en" docName="zt01548p049.pdf" docOrigin="Zootaxa 1548" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Polyonyx biunguiculatus Dana 1852" docType="treatment" docVersion="14" lastPageNumber="36" masterDocId="FFB9EC0FFF8EBF12AC66EE724A039B34" masterDocTitle="Porcellanidae (Crustacea: Decapoda: Anomura) from New Caledonia and the Loyalty Islands" masterLastPageNumber="49" masterPageNumber="1" pageNumber="31" updateTime="1698227420190" updateUser="plazi">
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<mods:title id="0A48F293E466A127AE7C145958F4C3F4">Porcellanidae (Crustacea: Decapoda: Anomura) from New Caledonia and the Loyalty Islands</mods:title>
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<mods:date id="EC469FF9A64EFDB7FC8C8B13E92A8FD8">2007</mods:date>
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<treatment id="03809477FF90BF31ACF1EB1749839A96" ID-DOI="http://doi.org/10.5281/zenodo.6250066" ID-GBIF-Taxon="119370691" ID-Zenodo-Dep="6250066" LSID="urn:lsid:plazi:treatment:03809477FF90BF31ACF1EB1749839A96" httpUri="http://treatment.plazi.org/id/03809477FF90BF31ACF1EB1749839A96" lastPageId="35" lastPageNumber="36" pageId="30" pageNumber="31">
<subSubSection id="C33376EAFF90BF0CACF1EB174B259E96" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8B962561FF90BF0CACF1EB17486F9E4B" blockId="30.[151,620,1381,1442]" box="[151,620,1381,1407]" pageId="30" pageNumber="31">
<heading id="D0DE920DFF90BF0CACF1EB17486F9E4B" bold="true" box="[151,620,1381,1407]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<taxonomicName id="4C295EE2FF90BF0CACF1EB17486F9E4B" ID-CoL="9ac949af-104f-45ed-9d83-c92284c1138d" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[151,620,1381,1407]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FF90BF0CACF1EB17486F9E4B" bold="true" box="[151,620,1381,1407]" pageId="30" pageNumber="31">
<emphasis id="B95DF973FF90BF0CACF1EB174BC79E4B" bold="true" box="[151,452,1381,1407]" italics="true" pageId="30" pageNumber="31">Polyonyx biunguiculatus</emphasis>
(
<bibRefCitation id="EFB85890FF90BF0CADB2EB1748609E4B" author="Dana" box="[468,611,1381,1407]" pageId="30" pageNumber="45" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U. S. N., 13, i - viii, 1 - 685, Philadelphia." type="journal article" year="1852">Dana, 1852</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B962561FF90BF0CACF1EBFA4B259E96" blockId="30.[151,620,1381,1442]" box="[151,294,1416,1442]" pageId="30" pageNumber="31">
(
<figureCitation id="131239E4FF90BF0CACF9EBFA4AF09E96" box="[159,243,1416,1442]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="30" pageNumber="31">Figs. 9</figureCitation>
,
<figureCitation id="131239E4FF90BF0CAC99EBFA4B1E9E96" box="[255,285,1416,1442]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="30" pageNumber="31">10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33376EAFF90BF0CACF1EBAA49509D5B" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="8B962561FF90BF0CACF1EBAA4B929D3B" blockId="30.[151,1436,1496,1743]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB39024FFF90BF0CACF1EBAA4B929D3B" pageId="30" pageNumber="31">
<taxonomicName id="4C295EE2FF90BF0CACF1EBAA486D9EDB" authority="Dana, 1852: 411" authorityName="Dana" authorityPageNumber="411" authorityYear="1852" box="[151,622,1496,1519]" class="Malacostraca" family="Porcellanidae" genus="Porcellana" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FF90BF0CACF1EBAA4BB19EDB" box="[151,434,1496,1519]" italics="true" pageId="30" pageNumber="31">Porcellana biunguiculatus</emphasis>
<treatmentCitation id="0A880370FF90BF0CADDCEBAA486D9EDB" author="Dana" box="[442,622,1496,1519]" page="411" pageId="30" pageNumber="31" year="1852">
<bibRefCitation id="EFB85890FF90BF0CADDCEBAA483A9EDB" author="Dana" box="[442,569,1496,1519]" pageId="30" pageNumber="45" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U. S. N., 13, i - viii, 1 - 685, Philadelphia." type="journal article" year="1852">Dana, 1852</bibRefCitation>
: 411
</treatmentCitation>
</taxonomicName>
; 1855: pl. 26, fig, 1ad (
<typeStatus id="54929BC3FF90BF0CAF1EEBAA49A49EDB" box="[888,935,1496,1519]" pageId="30" pageNumber="31">type</typeStatus>
locality: Loyalty Islands, restricted by present
<typeStatus id="54929BC3FF90BF0CACA1EB8A4B189D3B" box="[199,283,1528,1551]" pageId="30" pageNumber="31" type="neotype">neotype</typeStatus>
selection).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B962561FF90BF0CACF1E86A4B059D7B" blockId="30.[151,1436,1496,1743]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB39024FFF90BF0CACF1E86A4B059D7B" pageId="30" pageNumber="31">
<taxonomicName id="4C295EE2FF90BF0CACF1E86A48859D1B" ID-CoL="990fb004-ee1f-4efb-b88a-529d5e68436a" authority="Yang 1996: 262" authorityName="Yang" authorityPageNumber="262" authorityYear="1996" box="[151,646,1560,1583]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="parabiunguiculatus">
<emphasis id="B95DF973FF90BF0CACF1E86A4BD19D1B" box="[151,466,1560,1583]" italics="true" pageId="30" pageNumber="31">Polyonyx parabiunguiculatus</emphasis>
<treatmentCitation id="0A880370FF90BF0CADBFE86A48859D1B" author="Yang" box="[473,646,1560,1583]" page="262" pageId="30" pageNumber="31" year="1996">
<bibRefCitation id="EFB85890FF90BF0CADBFE86A48539D1B" author="Yang" box="[473,592,1560,1583]" pageId="30" pageNumber="49" refString="Yang, S. (1996) New species and new records of porcellanids crabs (Crustacea: Decapoda: Anomura) from Nansha Islands, China. Studies on marine fauna and flora and biogeography of the Nansha Islands and neighboring waters, II, 258 - 269. [In Chinese with English summary]" type="book chapter" year="1996">Yang 1996</bibRefCitation>
: 262
</treatmentCitation>
</taxonomicName>
, 268, fig. 5 (
<typeStatus id="54929BC3FF90BF0CAF68E86A493E9D1B" box="[782,829,1560,1583]" pageId="30" pageNumber="31">type</typeStatus>
locality: Nansha Islands, South
<collectingCountry id="F33E65F1FF90BF0CA8FFE86A4EDA9D1B" box="[1177,1241,1560,1583]" name="China" pageId="30" pageNumber="31">China</collectingCountry>
Sea) (new synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B962561FF90BF0CACF1E82A49509D5B" blockId="30.[151,1436,1496,1743]" box="[151,851,1624,1647]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB39024FFF90BF0CACF1E82A49509D5B" box="[151,851,1624,1647]" pageId="30" pageNumber="31">
?
<taxonomicName id="4C295EE2FF90BF0CACCFE82A4BB29D5B" ID-CoL="9ac949af-104f-45ed-9d83-c92284c1138d" box="[169,433,1624,1647]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FF90BF0CACCFE82A4BB29D5B" box="[169,433,1624,1647]" italics="true" pageId="30" pageNumber="31">Polyonyx biunguiculatus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A880370FF90BF0CADB4E82A48819D5B" author="Haig" box="[466,642,1624,1647]" page="286" pageId="30" pageNumber="31" year="1983">
<bibRefCitation id="EFB85890FF90BF0CADB4E82A484E9D5B" author="Haig" box="[466,589,1624,1647]" pageId="30" pageNumber="46" refString="Haig, J. (1983) Porcellanidae (Decapoda, Anomura) from the Seychelles, western Indian Ocean. Crustaceana, 45, 279 - 289." type="journal article" year="1983">Haig, 1983</bibRefCitation>
: 286
</treatmentCitation>
; 1992: 320, fig. 16.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33376EAFF90BF0CACF1E80A4B0F9DFB" pageId="30" pageNumber="31" type="description">
<paragraph id="8B962561FF90BF0CACF1E80A49459DBB" blockId="30.[151,1436,1496,1743]" box="[151,838,1656,1679]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="AB39024FFF90BF0CACF1E80A49459DBB" box="[151,838,1656,1679]" pageId="30" pageNumber="31">
Not
<taxonomicName id="4C295EE2FF90BF0CACA3E80A4BCE9DBB" ID-CoL="9ac949af-104f-45ed-9d83-c92284c1138d" box="[197,461,1656,1679]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FF90BF0CACA3E80A4BCE9DBB" box="[197,461,1656,1679]" italics="true" pageId="30" pageNumber="31">Polyonyx biunguiculatus</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0A880370FF90BF0CAD88E80A48B89DBB" author="Miyake" box="[494,699,1656,1679]" page="371" pageId="30" pageNumber="31" year="1942">
<bibRefCitation id="EFB85890FF90BF0CAD88E80A48849DBB" author="Miyake" box="[494,647,1656,1679]" pageId="30" pageNumber="47" refString="Miyake, S. (1942) Studies on the decapod crustaceans of Micronsia. III. Porcellanidae. The Palau Tropical Biological Station Studies, 2, 329 - 379." type="journal article" year="1942">Miyake, 1942</bibRefCitation>
: 371
</treatmentCitation>
, figs. 3032.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B962561FF90BF0CACF1E8EA4B0F9DFB" blockId="30.[151,1436,1496,1743]" pageId="30" pageNumber="31">
? Not
<taxonomicName id="4C295EE2FF90BF0CACBFE8EA4BE19D9B" box="[217,482,1688,1711]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FF90BF0CACBFE8EA4BE19D9B" box="[217,482,1688,1711]" italics="true" pageId="30" pageNumber="31">Polyonyx biunguiculatus</emphasis>
</taxonomicName>
.—
<bibRefCitation id="EFB85890FF90BF0CAE65E8EA487D9D9B" author="Haig" box="[515,638,1688,1711]" pageId="30" pageNumber="46" refString="Haig, J. (1964) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. 81. Porcellanid crabs from the Indo- West Pacific, Part I. Videnskabelige Meddelelser Dansk Naturhistrisk Forening i Kj g benhavn, 126, 355 - 386." type="journal article" year="1964">Haig, 1964</bibRefCitation>
: 377; 1965: 112; 1979: 130, figs. 1619.—
<bibRefCitation id="EFB85890FF90BF0CA82EE8EA4F529D9B" author="Yang" box="[1096,1361,1688,1711]" pageId="30" pageNumber="49" refString="Yang, S. &amp; Naiyanetr, P. (1997) Thailand's porcellanid crabs (Crustacea: Decapoda: Anomura). Memoirs of Beijing Natural History Museum, 56, 1 - 13." type="journal article" year="1997">Yang &amp; Naiyanetr 1997</bibRefCitation>
: 7, fig. 4AE.
</paragraph>
</subSubSection>
<subSubSection id="C33376EAFF90BF0CACF1E96D49919CD3" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8B962561FF90BF0CACF1E96D4B7A9C4B" blockId="30.[151,1436,1823,2023]" pageId="30" pageNumber="31">
<emphasis id="B95DF973FF90BF0CACF1E96D4E6D9C0D" bold="true" box="[151,1134,1823,1849]" pageId="30" pageNumber="31">
Material examined.
<typeStatus id="54929BC3FF90BF0CADFFE96D4BD59C0D" box="[409,470,1823,1849]" pageId="30" pageNumber="31">Type</typeStatus>
material: Lifou
<collectingCountry id="F33E65F1FF90BF0CAECBE96D48FE9C0D" box="[685,765,1823,1849]" name="Iceland" pageId="30" pageNumber="31">Island</collectingCountry>
. Santal Bay. Atelier
<collectionCode id="ED38BDA4FF90BF0CA86CE96D4E6A9C0D" box="[1034,1129,1823,1849]" pageId="30" pageNumber="31">LIFOU</collectionCode>
.
</emphasis>
St. 1462 (St. 19), Arète, Aimé Martin (= Acadro),
<geoCoordinate id="EE1D43A6FF90BF0CADA3E930483D9C68" box="[453,574,1858,1884]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="92" value="-20.785">20°47.1S</geoCoordinate>
,
<geoCoordinate id="EE1D43A6FF90BF0CAE2FE93048D09C68" box="[585,723,1858,1884]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="92" value="167.05333">167°03.2E</geoCoordinate>
,
<quantity id="4CD18884FF90BF0CAEB8E93049549C68" box="[734,855,1858,1884]" metricMagnitude="1" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="30" pageNumber="31" unit="m" value="95.0" valueMax="120.0" valueMin="70.0">70120 m</quantity>
, dredge,
<date id="FF9703A1FF90BF0CAFA6E9304E629C68" box="[960,1121,1858,1884]" pageId="30" pageNumber="31" value="2000-11-21">21 Nov. 2000</date>
, male CL
<quantity id="4CD18884FF90BF0CA8BEE9304F379C68" box="[1240,1332,1858,1884]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="31" unit="mm" value="2.8">2.8 mm</quantity>
,
<typeStatus id="54929BC3FF90BF0CA959E9304F9F9C68" box="[1343,1436,1858,1884]" pageId="30" pageNumber="31" type="neotype">neotype</typeStatus>
(MNHN-Ga 6466).
</paragraph>
<paragraph id="8B962561FF90BF0CACA1E9F549919CD3" blockId="30.[151,1436,1823,2023]" pageId="30" pageNumber="31">
<emphasis id="B95DF973FF90BF0CACA1E9F54B879C95" bold="true" box="[199,388,1927,1953]" pageId="30" pageNumber="31">
<collectingCountry id="F33E65F1FF90BF0CACA1E9F54B879C95" box="[199,388,1927,1953]" name="New Caledonia" pageId="30" pageNumber="31">New Caledonia</collectingCountry>
</emphasis>
. Koumac, Grand récif, lagoon, outer reef slope,
<quantity id="4CD18884FF90BF0CAFDDE9FA49FA9C96" box="[955,1017,1928,1954]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="31" unit="m" value="12.0">12 m</quantity>
,
<date id="FF9703A1FF90BF0CA862E9FA4E8C9C96" box="[1028,1167,1928,1954]" pageId="30" pageNumber="31" value="1993-10-07">7 Oct. 1993</date>
,
<specimenCount id="9D2FEEE8FF90BF0CA8FDE9FA4F029C96" box="[1179,1281,1928,1954]" pageId="30" pageNumber="31" type="female">1 female</specimenCount>
CL
<quantity id="4CD18884FF90BF0CA952E9FA4F929C96" box="[1332,1425,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="31" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="B95DF973FF90BF0CACA1E9D849CC9CF0" bold="true" box="[199,975,1962,1988]" pageId="30" pageNumber="31">
Loyalty Islands. Lifou
<collectingCountry id="F33E65F1FF90BF0CAD9DE9D848489CF0" box="[507,587,1962,1988]" name="Iceland" pageId="30" pageNumber="31">Island</collectingCountry>
. Santal Bay. Atelier
<collectionCode id="ED38BDA4FF90BF0CAF0CE9D849CA9CF0" box="[874,969,1962,1988]" pageId="30" pageNumber="31">LIFOU</collectionCode>
.
</emphasis>
St. 1459 (St. 6), Ngoni,
<geoCoordinate id="EE1D43A6FF90BF0CA978E9D84F949CF0" box="[1310,1431,1962,1988]" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="92" value="-20.783333">20°47.0S</geoCoordinate>
,
<geoCoordinate id="EE1D43A6FF90BF0CACF1E9BF4B239CD3" box="[151,288,1997,2023]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="92" value="167.05">167°03.0E</geoCoordinate>
,
<quantity id="4CD18884FF90BF0CAD4AE9BF4B969CD3" box="[300,405,1997,2023]" metricMagnitude="1" metricUnit="m" metricValue="6.75" metricValueMax="8.0" metricValueMin="5.5" pageId="30" pageNumber="31" unit="m" value="67.5" valueMax="80.0" valueMin="55.0">5580 m</quantity>
, dredge,
<date id="FF9703A1FF90BF0CAE66E9BF48A29CD3" box="[512,673,1997,2023]" pageId="30" pageNumber="31" value="2000-11-13">13 Nov. 2000</date>
,
<specimenCount id="9D2FEEE8FF90BF0CAECBE9BF48FF9CD3" box="[685,764,1997,2023]" pageId="30" pageNumber="31" type="male">1 male</specimenCount>
CL
<quantity id="4CD18884FF90BF0CAF56E9BF498E9CD3" box="[816,909,1997,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="30" pageNumber="31" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33376EAFF91BF30ACA1EEE54F6D9A96" lastPageId="34" lastPageNumber="35" pageId="31" pageNumber="32" type="description">
<paragraph id="8B962561FF91BF0DACA1EEE54BA99AB6" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
<emphasis id="B95DF973FF91BF0DACA1EEE54B5B9B85" bold="true" box="[199,344,151,177]" pageId="31" pageNumber="32">Description</emphasis>
. Carapace (
<figureCitation id="131239E4FF91BF0DAD8AEEEA48369B86" box="[492,565,152,178]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
A) transversely subrectangular, 1.2 times as broad as long, broadest on anterior branchial margin. Dorsal surface convex, nearly smooth, with scattered, very short striae and small pits. Rostrum (
<figureCitation id="131239E4FF91BF0DAD2BEEAF4B949BC3" box="[333,407,221,247]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
B) broad, bent ventrally, trilobate in frontal view; median lobe triangular, exceeding obtuse lateral lobes, with median longitudinal groove on dorsal surface. Orbits shallow, supra-orbital margins oblique; outer orbital angles rounded. Protogastric ridges and cervical grooves faintly demarcated. Branchial margins somewhat constricted medially; posterior margin and adjacent region with long transverse rugae. Cardiac region smooth.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1EFF84E689ADE" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
Third thoracic sternite (
<figureCitation id="131239E4FF91BF0DADB9EFF848249A90" box="[479,551,394,420]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
C) with anterior margin trilobate, bearing sparse setae; median lobe broad, low subtriangular; lateral lobes distinctly exceeding median lobe, each with rounded apex. Fourth thoracic sternite with transverse series of short setae along weakly concave, anterior margin.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1EF804F2C9938" blockId="31.[151,1437,151,1807]" box="[199,1327,498,524]" pageId="31" pageNumber="32">
Ocular peduncles (
<figureCitation id="131239E4FF91BF0DADC5EF804BEA9938" box="[419,489,498,524]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
A, D) moderately large; dorsal extension onto cornea weakly produced.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1EC674AFC9940" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
Basal article of antennular peduncles (
<figureCitation id="131239E4FF91BF0DAEE8EC6748D6991B" box="[654,725,533,559]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
D) unarmed; anterior surface concave, weakly rugose; anterior, dorsal, and ventral margins weakly tuberculate; ventral surface with short transverse ridge on median lateral part.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1EC0F499599CB" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
Antennal peduncles (
<figureCitation id="131239E4FF91BF0DADA0EC0F480C99A3" box="[454,527,637,663]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
A, D) short, slender, nearly smooth, unarmed. First article largest, strongly produced forward in lateral view, broadly in contact with lower orbital margin, with longitudinal ridge along ventral margin; lateral surface concave; anterior margin narrowly rounded. Second, third, and fourth articles roundly subrectangular, third article elongate, fourth article short.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1ED7A4B2E9F03" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
Third maxilliped (
<figureCitation id="131239E4FF91BF0DADFBED7A4BE19816" box="[413,482,776,802]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="31" pageNumber="32">Fig. 9</figureCitation>
E) with coxa bearing small ronded projection on distoflexor margin; distomedian projection not articulated. Basis articulating with ischium, rounded subtriangular. Ischium broad, ovate, with few delicate striae on ventral surface, with weak longitudinal ridge along extensor margin; anterior extensor projection rounded. Merus with laminate, broad, rounded subrectangular lobe on ventroflexor margin; ventral surface weakly striate. Carpus with subtriangular projection on median part of flexor margin and row of short rugae on extensor surface. Propodus relatively short, somewhat tapering distally. Dactylus short, rounded subtriangular. Merus to dactylus with long setae on flexor margin. Exopod with proximal article small; distal article laminate, robust, nearly reaching distal margin of merus, with distal flagellum (not illustrated), proximal part inflated.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1EA324F489DB0" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
Chelipeds (
<figureCitation id="131239E4FF91BF0DAD2AEA324B9C9F6E" box="[332,415,1088,1114]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
AH) unequal, subcylindrical, inflated. Larger cheliped (
<figureCitation id="131239E4FF91BF0DA85DEA324E8C9F6E" box="[1083,1167,1088,1114]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
AC, F, G) with merus provided with rounded transverse crest submedially on rugose dorsal surface; dorsoflexor margin with round lobe distally. Carpus 2.2 (female)3.0 (male) times as long as broad; dorso-extensor margin rounded; dorsoflexor margin weakly concave, smooth or weakly tuberculate, with rounded proximal angle; dorsodistal margin nearly transverse. Chela relatively broad, ovate, 1.41.5 times as long as carpus, 2.32.4 times as long as high, lying on extensor side; dactylus opening at oblique angle; extensor margin smooth, slightly concave at base of fixed finger; dorsal surface generally smooth, with longitudinal blunt ridge, short rugae, and scattered short setae along extensor margin. Palm with dorsal surface convex, no dorsomedian longitudinal ridge present; dorsoflexor margin with faintly or weakly developed, longitudinal rugose ridge; dorsoflexor distal part with low projection extending onto dactylus; ventral surface with rounded longitudinal ridge proximally along midline, distal flexor margin with few short simple setae. Fixed finger with weakly or moderately curved distal claw; dorsal surface with scattered short simple setae; cutting edge with row of small, rounded and subtriangular teeth (female) or unarmed except for large proximal tooth with marginal tubercles (male); ventral surface with few short simple setae along cutting edge. Dactylus 0.40.5 length of chela, with strongly curved distal claw; dorsal surface nearly smooth; dorso-extensor margin rounded (male) or with rugose ridge (female); cutting edge concave, with row of small, rounded and subtriangular teeth (female) or unarmed except for broad proximal tooth (male); ventral surface with few short simple setae along cutting edge.
</paragraph>
<paragraph id="8B962561FF91BF0DACA1E8FF49F69C3B" blockId="31.[151,1437,151,1807]" pageId="31" pageNumber="32">
Smaller cheliped (
<figureCitation id="131239E4FF91BF0DADC5E8FF4BF89D93" box="[419,507,1677,1703]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="31" pageNumber="32">Fig. 10</figureCitation>
D, E, H) genarally similar to larger cheliped in shape, except for: chela with small protuberances and subacute tubercles on distal 1/22/3 of extensor margin and adjacent dorsal surface; cutting edges of fingers nearly transverse, each with row of smaller teeth; dactylus opening at more stronger angle, with more strongly crested, longitudinal ridge along flexor margin.
</paragraph>
<caption id="DF5675E9FFAEBF32ACF1E8B04F049C9D" httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="32" pageNumber="33" targetBox="[164,1404,209,1692]" targetPageId="32">
<paragraph id="8B962561FFAEBF32ACF1E8B04F049C9D" blockId="32.[151,1436,1730,1961]" pageId="32" pageNumber="33">
<emphasis id="B95DF973FFAEBF32ACF1E8B04B1C9DEE" bold="true" box="[151,287,1730,1754]" pageId="32" pageNumber="33">FIGURE 9.</emphasis>
<taxonomicName id="4C295EE2FFAEBF32AD4CE8B048D49DED" authority="(Dana, 1852)" authorityName="Dana" authorityYear="1852" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[298,727,1730,1753]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFAEBF32AD4CE8B048349DED" box="[298,567,1730,1753]" italics="true" pageId="32" pageNumber="33">Polyonyx biunguiculatus</emphasis>
(Dana, 1852)
</taxonomicName>
, neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (AH); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm.
</paragraph>
</caption>
<caption id="DF5675E9FFAFBF33ACF1E8B04B789C75" httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="33" pageNumber="34" targetBox="[171,1412,184,1678]" targetPageId="33">
<paragraph id="8B962561FFAFBF33ACF1E8B04B789C75" blockId="33.[151,1437,1730,1857]" pageId="33" pageNumber="34">
<emphasis id="B95DF973FFAFBF33ACF1E8B04B289DEE" bold="true" box="[151,299,1730,1754]" pageId="33" pageNumber="34">FIGURE 10.</emphasis>
<taxonomicName id="4C295EE2FFAFBF33AD53E8B048E29DED" authority="(Dana, 1852)" authorityName="Dana" authorityYear="1852" baseAuthorityName="Dana" baseAuthorityYear="1852" box="[309,737,1730,1753]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFAFBF33AD53E8B048419DED" box="[309,578,1730,1753]" italics="true" pageId="33" pageNumber="34">Polyonyx biunguiculatus</emphasis>
(Dana, 1852)
</taxonomicName>
, neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (AE); female, CL 2.5 mm, New Caledonia, Koumac (FH). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B962561FFAFBF30ACA1E90748DC9A4E" blockId="33.[151,1436,1909,2015]" lastBlockId="34.[151,1437,152,2018]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Ambulatory legs (
<figureCitation id="131239E4FFAFBF33ADC1E9074BF29CBB" box="[423,497,1909,1935]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="33" pageNumber="34">Fig. 9</figureCitation>
FI) stout, subcylindrical, with few scattered, simple setae marginally, most numerous on propodi; lateral surfaces nearly smooth. Meri somewhat compressed lateromesially, elongated ovate, decreasing in size posteriorly; extensor margin unarmed but slightly crenulated, weakly convex; distoflexor margins of lateral and mesial surfaces without lobes or spines. Carpi moderately elongate; lateral surface with longitudinal rows of short rugae on extensor half; disto-extensor and distoflexor corners unarmed, narrowly rounded. Propodi 1.6 times as long as dactyli, 3.4 times as long as high; extensor margin slightly crenulated; flexor margin with 4 corneous spines, proximal spine smaller than others, distal paired spines subequal in size. Dactyli each terminating in weakly curved, bifurcate claw, flexor claw stouter but shoter than extensor claw; flexor margin with 2 small corneous spines.
</paragraph>
<paragraph id="8B962561FFACBF30ACA1EFFA4F6D9A96" blockId="34.[151,1437,152,2018]" box="[199,1390,392,418]" pageId="34" pageNumber="35">
<collectingCountry id="F33E65F1FFACBF30ACA1EFFA4B009A96" box="[199,259,392,418]" name="Malta" pageId="34" pageNumber="35">Male</collectingCountry>
with pair of short but developed pleopods modified as gonopods on second abdominal segment.
</paragraph>
</subSubSection>
<subSubSection id="C33376EAFFACBF31ACA1EFDD4E109A66" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8B962561FFACBF30ACA1EFDD4F96992E" blockId="34.[151,1437,152,2018]" pageId="34" pageNumber="35">
<emphasis id="B95DF973FFACBF30ACA1EFDD4B3A9AFD" bold="true" box="[199,313,431,457]" pageId="34" pageNumber="35">Remarks</emphasis>
.
<bibRefCitation id="EFB85890FFACBF30AD2FEFC24BDE9AFE" author="Dana" box="[329,477,432,458]" pageId="34" pageNumber="45" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U. S. N., 13, i - viii, 1 - 685, Philadelphia." type="journal article" year="1852">Dana (1852)</bibRefCitation>
did not specify the
<typeStatus id="54929BC3FFACBF30AEADEFC248FE9AFE" box="[715,765,432,458]" pageId="34" pageNumber="35">type</typeStatus>
locality of
<taxonomicName id="4C295EE2FFACBF30AFE1EFC24EB79AFD" box="[903,1204,432,457]" class="Malacostraca" family="Porcellanidae" genus="Porcellana" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculata">
<emphasis id="B95DF973FFACBF30AFE1EFC24EB79AFD" box="[903,1204,432,457]" italics="true" pageId="34" pageNumber="35">Porcellana biunguiculata</emphasis>
</taxonomicName>
and the
<typeStatus id="54929BC3FFACBF30A97DEFC24F4D9AFE" box="[1307,1358,432,458]" pageId="34" pageNumber="35">type</typeStatus>
specimen is apparently lost (see
<bibRefCitation id="EFB85890FFACBF30ADBBEFAA48FB9AC6" author="Deiss" box="[477,760,472,498]" pageId="34" pageNumber="45" refString="Deiss, W. A. &amp; Manning, R. B. (1981) The fate of the invertebrate collections of the North Pacific Exploring Expedition, 1853 - 1856. In: Wheeler, A. &amp; Price, J. H. (Eds.), History in the Service of Systematics. Society for the Bibliography of Natural History Special Publication, Number 1, London, pp. 79 - 85." type="book chapter" year="1981">Deiss &amp; Manning 1981</bibRefCitation>
). Therefore, the male specimen from the Loyalty Islands (CL
<quantity id="4CD18884FFACBF30ACABEC724B29992E" box="[205,298,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="34" pageNumber="35" unit="mm" value="2.8">2.8 mm</quantity>
, MNHN-Ga 6466) is herein selected as the
<typeStatus id="54929BC3FFACBF30AF4CEC724984992E" box="[810,903,512,538]" pageId="34" pageNumber="35" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C295EE2FFACBF30AFC8EC724EC6992D" box="[942,1221,512,537]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculata">
<emphasis id="B95DF973FFACBF30AFC8EC724EC6992D" box="[942,1221,512,537]" italics="true" pageId="34" pageNumber="35">Polyonyx biunguiculata</emphasis>
</taxonomicName>
to fix its identity.
</paragraph>
<paragraph id="8B962561FFACBF30ACA1EC5A4B68986E" blockId="34.[151,1437,152,2018]" pageId="34" pageNumber="35">
The specimens examined agree well with the original description and figures of
<taxonomicName id="4C295EE2FFACBF30A8C2EC5A4F709975" box="[1188,1395,552,577]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30A8C2EC5A4F709975" box="[1188,1395,552,577]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFB85890FFACBF30ACF1EC224B26995E" author="Dana" box="[151,293,592,618]" pageId="34" pageNumber="45" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U. S. N., 13, i - viii, 1 - 685, Philadelphia." type="journal article" year="1852">Dana (1852</bibRefCitation>
,
<bibRefCitation id="EFB85890FFACBF30AD54EC224B73995E" author="Dana" box="[306,368,592,618]" pageId="34" pageNumber="45" refString="Dana, J. D. (1855) Crustacea. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U. S. N., 14 (Atlas), 1 - 27, 96 pls., Philadelphia." type="journal article" year="1855">1855</bibRefCitation>
), especially on account of the larger chela being rather smooth on the dorsal surface (
<figureCitation id="131239E4FFACBF30A90AEC224AB699A6" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
A, B, F, G). Although
<bibRefCitation id="EFB85890FFACBF30ADD8EC0A485199A6" author="Dana" box="[446,594,632,658]" pageId="34" pageNumber="45" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the command of Charles Wilkes, U. S. N., 13, i - viii, 1 - 685, Philadelphia." type="journal article" year="1852">Dana (1852)</bibRefCitation>
did not indicate the sex of the specimen, it seems to be a male because of the possession of a large proximal tooth on the cutting edge of the fixed finger (
<figureCitation id="131239E4FFACBF30A833ECD24EAF998E" box="[1109,1196,672,698]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
B). The female lacks such a large tooth (
<figureCitation id="131239E4FFACBF30AD1AECBA4BD099D6" box="[380,467,712,738]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
G). The smooth condition of the larger chela in males is also illustrated for
<collectingCountry id="F33E65F1FFACBF30A93DECBA4ADB983E" name="Hong Kong" pageId="34" pageNumber="35">Hong Kong</collectingCountry>
material of
<taxonomicName id="4C295EE2FFACBF30AD03EC82482C983D" box="[357,559,752,777]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AD03EC82482C983D" box="[357,559,752,777]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB85890FFACBF30AE58EC824928983E" author="Haig" box="[574,811,752,778]" pageId="34" pageNumber="46" refString="Haig, J. (1992) Hong Kong's porcellanid crabs. In: Morton, B. (Ed.), The marine flora and fauna of Hong Kong and southern China III. Proceedings of the Fourth International Marine Biological Workshop. Hong Kong University Press, Hong Kong, pp. 303 - 327." type="proceedings paper" year="1992">Haig 1992, fig. 16C</bibRefCitation>
), whereas the chela is figured as rather tuberculate on the fixed finger in the material from the
<collectingCountry id="F33E65F1FFACBF30AE17ED6A48E09806" box="[625,739,792,818]" name="Indonesia" pageId="34" pageNumber="35">Moluccas</collectingCountry>
and Gulf of
<collectingCountry id="F33E65F1FFACBF30AF1CED6A49E19806" box="[890,994,792,818]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
(
<bibRefCitation id="EFB85890FFACBF30AF97ED6A4E709806" author="Haig" box="[1009,1139,792,818]" pageId="34" pageNumber="46" refString="Haig, J. (1979) Expedition Rumphius II (1975) Crustaces parasites, commensaux, etc. (Th. Monod et R. Serene, ed.). V. Porcellanidae (Crustacea, Decapoda, Anomura). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e, section A, 1, 119 - 135." type="journal article" year="1979">Haig 1979</bibRefCitation>
: fig. 19;
<bibRefCitation id="EFB85890FFACBF30A8BDED6A4AFF986E" author="Yang" pageId="34" pageNumber="49" refString="Yang, S. &amp; Naiyanetr, P. (1997) Thailand's porcellanid crabs (Crustacea: Decapoda: Anomura). Memoirs of Beijing Natural History Museum, 56, 1 - 13." type="journal article" year="1997">Yang &amp; Naiyanetr 1997</bibRefCitation>
: fig. 4D).
</paragraph>
<paragraph id="8B962561FFACBF30ACA1ED1A49EE9F7E" blockId="34.[151,1437,152,2018]" pageId="34" pageNumber="35">
<bibRefCitation id="EFB85890FFACBF30ACA1ED1A4B5598B6" author="Haig" box="[199,342,872,898]" pageId="34" pageNumber="46" refString="Haig, J. (1964) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. 81. Porcellanid crabs from the Indo- West Pacific, Part I. Videnskabelige Meddelelser Dansk Naturhistrisk Forening i Kj g benhavn, 126, 355 - 386." type="journal article" year="1964">Haig (1964)</bibRefCitation>
showed that
<taxonomicName id="4C295EE2FFACBF30AD90ED1A48C198B5" box="[502,706,872,897]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AD90ED1A48C198B5" box="[502,706,872,897]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
could be distinguished from
<taxonomicName id="4C295EE2FFACBF30A879ED1A4EA798B5" box="[1055,1188,872,897]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="obesulus">
<emphasis id="B95DF973FFACBF30A879ED1A4EA798B5" box="[1055,1188,872,897]" italics="true" pageId="34" pageNumber="35">P. obesulus</emphasis>
</taxonomicName>
by the dactyli of the ambulatory legs each with two small spines on the flexor margin and the absence of pleopods in males, based on an examination of a good series of
<taxonomicName id="4C295EE2FFACBF30AE35EDCA491E98E5" box="[595,797,952,977]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AE35EDCA491E98E5" box="[595,797,952,977]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
from the Indo-Malayan area. The absence of pleopods in males was also confirmed in the Western Australian and
<collectingCountry id="F33E65F1FFACBF30AF3AED9249CD98CE" box="[860,974,992,1018]" name="Indonesia" pageId="34" pageNumber="35">Moluccas</collectingCountry>
material (
<bibRefCitation id="EFB85890FFACBF30A82EED924ECB98CE" author="Haig" box="[1096,1224,992,1018]" pageId="34" pageNumber="46" refString="Haig, J. (1965) The Porcellanidae (Crustacea, Anomura) of Western Australia with description of four new Australian species. Journal of the Royal Society of Western Australia, 48, 97 - 117." type="journal article" year="1965">Haig 1965</bibRefCitation>
,
<bibRefCitation id="EFB85890FFACBF30A8B0ED924F1698CE" author="Haig" box="[1238,1301,992,1018]" pageId="34" pageNumber="46" refString="Haig, J. (1979) Expedition Rumphius II (1975) Crustaces parasites, commensaux, etc. (Th. Monod et R. Serene, ed.). V. Porcellanidae (Crustacea, Decapoda, Anomura). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e, section A, 1, 119 - 135." type="journal article" year="1979">1979</bibRefCitation>
). However,
<bibRefCitation id="EFB85890FFACBF30ACF1EA7A4B249F16" author="Haig" box="[151,295,1032,1058]" pageId="34" pageNumber="46" refString="Haig, J. (1983) Porcellanidae (Decapoda, Anomura) from the Seychelles, western Indian Ocean. Crustaceana, 45, 279 - 289." type="journal article" year="1983">Haig (1983)</bibRefCitation>
showed that this character was less reliable than she had supposed since the all males from the “Reves 2” collection from the
<collectingCountry id="F33E65F1FFACBF30AD9AEA42487A9F7E" box="[508,633,1072,1098]" name="Seychelles" pageId="34" pageNumber="35">Seychelles</collectingCountry>
have well-developed pleopods.
</paragraph>
<paragraph id="8B962561FFACBF30ACA1EA2A49029E86" blockId="34.[151,1437,152,2018]" pageId="34" pageNumber="35">
The smooth larger chela and presence of male pleopods are species-specific characters for the true
<taxonomicName id="4C295EE2FFACBF30A9E0EA2A4B469FAD" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30A9E0EA2A4B469FAD" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
, and the specimens with tuberculate chelae and no pleopods in males represent a different, undescribed species. It is likely that the material from the
<collectingCountry id="F33E65F1FFACBF30AF5AEADA49BA9FF6" box="[828,953,1192,1218]" name="Seychelles" pageId="34" pageNumber="35">Seychelles</collectingCountry>
and
<collectingCountry id="F33E65F1FFACBF30AF94EADA4E789FF6" box="[1010,1147,1192,1218]" name="Hong Kong" pageId="34" pageNumber="35">Hong Kong</collectingCountry>
reported by
<bibRefCitation id="EFB85890FFACBF30A976EADA4F949FF6" author="Haig" box="[1296,1431,1192,1218]" pageId="34" pageNumber="46" refString="Haig, J. (1983) Porcellanidae (Decapoda, Anomura) from the Seychelles, western Indian Ocean. Crustaceana, 45, 279 - 289." type="journal article" year="1983">Haig (1983</bibRefCitation>
,
<bibRefCitation id="EFB85890FFACBF30ACF1EAA24AD09FDE" author="Haig" box="[151,211,1232,1258]" pageId="34" pageNumber="46" refString="Haig, J. (1992) Hong Kong's porcellanid crabs. In: Morton, B. (Ed.), The marine flora and fauna of Hong Kong and southern China III. Proceedings of the Fourth International Marine Biological Workshop. Hong Kong University Press, Hong Kong, pp. 303 - 327." type="proceedings paper" year="1992">1992</bibRefCitation>
) belongs to true
<taxonomicName id="4C295EE2FFACBF30ADF9EAA2486F9FDD" box="[415,620,1232,1257]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30ADF9EAA2486F9FDD" box="[415,620,1232,1257]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
, whereas the specimens from the the Indo-Malayan area,
<collectingCountry id="F33E65F1FFACBF30A945EAA24F9B9FDE" box="[1315,1432,1232,1258]" name="Indonesia" pageId="34" pageNumber="35">Moluccas</collectingCountry>
, Gulf of
<collectingCountry id="F33E65F1FFACBF30AC90EA8A4B629E26" box="[246,353,1272,1298]" name="Thailand" pageId="34" pageNumber="35">Thailand</collectingCountry>
, and Western
<collectingCountry id="F33E65F1FFACBF30AE6FEA8A48779E26" box="[521,628,1272,1298]" name="Australia" pageId="34" pageNumber="35">Australia</collectingCountry>
reported by
<bibRefCitation id="EFB85890FFACBF30AF6BEA8A49959E26" author="Haig" box="[781,918,1272,1298]" pageId="34" pageNumber="46" refString="Haig, J. (1964) Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. 81. Porcellanid crabs from the Indo- West Pacific, Part I. Videnskabelige Meddelelser Dansk Naturhistrisk Forening i Kj g benhavn, 126, 355 - 386." type="journal article" year="1964">Haig (1964</bibRefCitation>
,
<bibRefCitation id="EFB85890FFACBF30AFC5EA8A49E39E26" author="Haig" box="[931,992,1272,1298]" pageId="34" pageNumber="46" refString="Haig, J. (1965) The Porcellanidae (Crustacea, Anomura) of Western Australia with description of four new Australian species. Journal of the Royal Society of Western Australia, 48, 97 - 117." type="journal article" year="1965">1965</bibRefCitation>
,
<bibRefCitation id="EFB85890FFACBF30AF8BEA8A4E2A9E26" author="Haig" box="[1005,1065,1272,1298]" pageId="34" pageNumber="46" refString="Haig, J. (1979) Expedition Rumphius II (1975) Crustaces parasites, commensaux, etc. (Th. Monod et R. Serene, ed.). V. Porcellanidae (Crustacea, Decapoda, Anomura). Bulletin du Museum National d'Histoire Naturelle, Paris, 4 e, section A, 1, 119 - 135." type="journal article" year="1979">1979</bibRefCitation>
) and
<bibRefCitation id="EFB85890FFACBF30A80BEA8A4F9E9E26" author="Yang" box="[1133,1437,1272,1298]" pageId="34" pageNumber="49" refString="Yang, S. &amp; Naiyanetr, P. (1997) Thailand's porcellanid crabs (Crustacea: Decapoda: Anomura). Memoirs of Beijing Natural History Museum, 56, 1 - 13." type="journal article" year="1997">Yang &amp; Naiyanetr (1997)</bibRefCitation>
are an undescribed species; although the true identities of these specimens need re-examination. On the other hand, re-examination of the specimens from
<collectingCountry id="F33E65F1FFACBF30AECFEB3A48E99E56" box="[681,746,1352,1378]" name="Palau" pageId="34" pageNumber="35">Palau</collectingCountry>
(ZLKU 10318,
<specimenCount id="9D2FEEE8FFACBF30AFCAEB3A4E4F9E56" box="[940,1100,1352,1378]" pageId="34" pageNumber="35" type="generic">11 specimens</specimenCount>
) reported by
<bibRefCitation id="EFB85890FFACBF30A88AEB3A4F9F9E56" author="Miyake" box="[1260,1436,1352,1378]" pageId="34" pageNumber="47" refString="Miyake, S. (1942) Studies on the decapod crustaceans of Micronsia. III. Porcellanidae. The Palau Tropical Biological Station Studies, 2, 329 - 379." type="journal article" year="1942">Miyake (1942)</bibRefCitation>
showed that they had small tubercles on the base of the fixed finger of each cheliped and no pleopods in males, and thus belonged to the undescribed species.
</paragraph>
<paragraph id="8B962561FFACBF30ACA1EBB24F5C9D4E" blockId="34.[151,1437,152,2018]" pageId="34" pageNumber="35">
<bibRefCitation id="EFB85890FFACBF30ACA1EBB24B5F9EEE" author="Baba" box="[199,348,1472,1498]" pageId="34" pageNumber="45" refString="Baba, K. (1989) Anomuran crustaceans obtained by dredging from Oshima Strait, Amami-Oshima of the Ryukyu Islands. Memoirs of the National Science Museum, Tokyo, 22, 127 - 134." type="journal article" year="1989">Baba (1989)</bibRefCitation>
recorded
<taxonomicName id="4C295EE2FFACBF30ADBFEBB248AB9EED" box="[473,680,1472,1497]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30ADBFEBB248AB9EED" box="[473,680,1472,1497]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
from the Ryukyu Islands and noted that the coloration of the specimens is different from that provided by
<bibRefCitation id="EFB85890FFACBF30AEC4EB9A49579D36" author="Miyake" box="[674,852,1512,1538]" pageId="34" pageNumber="47" refString="Miyake, S. (1942) Studies on the decapod crustaceans of Micronsia. III. Porcellanidae. The Palau Tropical Biological Station Studies, 2, 329 - 379." type="journal article" year="1942">Miyake (1942)</bibRefCitation>
. His specimens were totally reddish with irregular reticulation of white, and had the carapace with a rough longitudinal stripe of white color, whereas
<bibRefCitation id="EFB85890FFACBF30A924E8624AE79D66" author="Miyake" pageId="34" pageNumber="47" refString="Miyake, S. (1942) Studies on the decapod crustaceans of Micronsia. III. Porcellanidae. The Palau Tropical Biological Station Studies, 2, 329 - 379." type="journal article" year="1942">Miyake (1942)</bibRefCitation>
noted the dorsal surface (of the carapace) to be light orange yellow, the ventral (surface) whitish. New material with information of coloration would help to distinguish the
<taxonomicName id="4C295EE2FFACBF30AFA2E8124E8C9D4D" box="[964,1167,1632,1657]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AFA2E8124E8C9D4D" box="[964,1167,1632,1657]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
species complex.
</paragraph>
<paragraph id="8B962561FFACBF30ACA1E8FA48F49C5D" blockId="34.[151,1437,152,2018]" pageId="34" pageNumber="35">
<bibRefCitation id="EFB85890FFACBF30ACA1E8FA4B5A9D96" author="Yang" box="[199,345,1672,1698]" pageId="34" pageNumber="49" refString="Yang, S. (1996) New species and new records of porcellanids crabs (Crustacea: Decapoda: Anomura) from Nansha Islands, China. Studies on marine fauna and flora and biogeography of the Nansha Islands and neighboring waters, II, 258 - 269. [In Chinese with English summary]" type="book chapter" year="1996">Yang (1996)</bibRefCitation>
described a new species,
<taxonomicName id="4C295EE2FFACBF30AEEBE8FA49939D95" box="[653,912,1672,1697]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="parabiunguiculatus">
<emphasis id="B95DF973FFACBF30AEEBE8FA49939D95" box="[653,912,1672,1697]" italics="true" pageId="34" pageNumber="35">P. parabiunguiculatus</emphasis>
</taxonomicName>
, from the Nansha Islands, South
<collectingCountry id="F33E65F1FFACBF30A97AE8FA4F619D96" box="[1308,1378,1672,1698]" name="China" pageId="34" pageNumber="35">China</collectingCountry>
Sea. This species was reported to differ from “
<taxonomicName id="4C295EE2FFACBF30AE18E8C2494A9DFD" box="[638,841,1712,1737]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AE18E8C2494A9DFD" box="[638,841,1712,1737]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
” in the median lobe of the rostrum being narrower and more prominent, merus of the cheliped with a well developed, flexor distal lobe, and presence of pleopods in males. In addition to the smooth dorsal surface of the larger chela, these characters agree with those of the male specimens examined including the
<typeStatus id="54929BC3FFACBF30AE1AE95A48DA9C76" box="[636,729,1832,1858]" pageId="34" pageNumber="35" type="neotype">neotype</typeStatus>
of
<taxonomicName id="4C295EE2FFACBF30AF62E95A49D29C75" box="[772,977,1832,1857]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AF62E95A49D29C75" box="[772,977,1832,1857]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
. Therefore,
<taxonomicName id="4C295EE2FFACBF30A800E95A4F6A9C75" box="[1126,1385,1832,1857]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="parabiunguiculatus">
<emphasis id="B95DF973FFACBF30A800E95A4F6A9C75" box="[1126,1385,1832,1857]" italics="true" pageId="34" pageNumber="35">P. parabiunguiculatus</emphasis>
</taxonomicName>
can be treated as a junior synonym of
<emphasis id="B95DF973FFACBF30AE42E92248F49C5D" box="[548,759,1872,1897]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="4C295EE2FFACBF30AE42E92248F09C5D" box="[548,755,1872,1897]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">P. biunguiculatus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B962561FFACBF31ACA1E90A49BC9A36" blockId="34.[151,1437,152,2018]" lastBlockId="35.[151,1436,152,418]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
The specimens examined of
<taxonomicName id="4C295EE2FFACBF30AE7EE90A48E09CA5" box="[536,739,1912,1937]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30AE7EE90A48E09CA5" box="[536,739,1912,1937]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
have a rounded, longitudinal ridge on the proximal ventral surface of the palm of the cheliped (
<figureCitation id="131239E4FFACBF30AE5DE9D248939C8E" box="[571,656,1952,1978]" captionStart="FIGURE 10" captionStartId="33.[151,255,1730,1754]" captionTargetBox="[171,1412,184,1678]" captionTargetId="figure@33.[151,1436,166,1706]" captionTargetPageId="33" captionText="FIGURE 10. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A E); female, CL 2.5 mm, New Caledonia, Koumac (F H). A, F, right larger cheliped, dorsal view; B, G, same, chela, dorsoextensor view; C, same, ventral view; D, left smaller cheliped, dorsal view; E, H, same, chela, dorso-extensor view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178050/files/figure.png" pageId="34" pageNumber="35">Fig. 10</figureCitation>
C), which is a similar structure observed in the material of
<taxonomicName id="4C295EE2FFACBF30A95BE9D24BCC9CD6" authority="Zehntner, 1894" authorityName="Zehntner" authorityYear="1894" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="triunguiculatus">
<emphasis id="B95DF973FFACBF30A95BE9D24B129CD5" italics="true" pageId="34" pageNumber="35">P. triunguiculatus</emphasis>
<bibRefCitation id="EFB85890FFACBF30AD7FE9BA4BCC9CD6" author="Zehntner" box="[281,463,1992,2018]" pageId="34" pageNumber="49" refString="Zehntner, L. (1894) Crustaces de l'Archipel malais. Voyage de MM. M. Bedot et C. Pictet dans l'Archipel malais. Revue Suisse de Zoologie et Annales du Musee d'Histoire Naturelle de Geneve, 2, 135 - 214, pls. 7 - 9." type="journal article" year="1894">Zehntner, 1894</bibRefCitation>
</taxonomicName>
. However,
<taxonomicName id="4C295EE2FFACBF30AE33E9BA49259CD5" box="[597,806,1992,2017]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="triunguiculatus">
<emphasis id="B95DF973FFACBF30AE33E9BA49259CD5" box="[597,806,1992,2017]" italics="true" pageId="34" pageNumber="35">P. triunguiculatus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C295EE2FFACBF30A856E9BA4EFF9CD5" box="[1072,1276,1992,2017]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFACBF30A856E9BA4EFF9CD5" box="[1072,1276,1992,2017]" italics="true" pageId="34" pageNumber="35">P. biunguiculatus</emphasis>
</taxonomicName>
by the longitudinal ridge being better developed and more strongly crested as well as the median lobe of the rostrum being narrower and more strongly produced and carpi of the chelipeds being rather transverse or convex on the dorsoflexor margin (the margin is weakly concave in
<taxonomicName id="4C295EE2FFADBF31AE85EE9A49AD9A35" box="[739,942,232,257]" class="Malacostraca" family="Porcellanidae" genus="Polyonyx" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="biunguiculatus">
<emphasis id="B95DF973FFADBF31AE85EE9A49AD9A35" box="[739,942,232,257]" italics="true" pageId="35" pageNumber="36">P. biunguiculatus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B962561FFADBF31ACA1EF624E109A66" blockId="35.[151,1436,152,418]" pageId="35" pageNumber="36">
In the female specimen examined, the dactylus of the right second ambulatory leg is largely deformed and does not show a bifurcate form as seen in other ambulatory dactyli (
<figureCitation id="131239E4FFADBF31AFD5EF4A49F99A66" box="[947,1018,312,338]" captionStart="FIGURE 9" captionStartId="32.[151,255,1730,1754]" captionTargetBox="[164,1404,209,1692]" captionTargetId="figure@32.[151,1436,194,1706]" captionTargetPageId="32" captionText="FIGURE 9. Polyonyx biunguiculatus (Dana, 1852), neotype, male, CL 2.8 mm, Loyalty Islands, St. 1462 (A H); female, CL 2.5 mm, New Caledonia, Koumac (I). A, carapace, and ocular and antennal peduncles, dorsal view; B, rostrum, anterior view; C, larger cheliped, dorsal view; C, third and forth thoracic sternites, ventral view (setae omitted from right side); D, left anterior part of pterygostomial flap, ocular and antennal peduncles, and basal article of antennular peduncle, ventrofrontal view; E, left third maxilliped, ventral view (setae omitted); F, left first ambulatory leg, lateral view; G, same, dactylus and distal part of propodus, lateral view; H, left second ambulatory leg, lateral view; dactylus and distal part of propodus, lateral view; I, right second ambulatory leg, lateral view. Scales equal 1.0 mm." httpUri="https://zenodo.org/record/178049/files/figure.png" pageId="35" pageNumber="36">
Fig.
<date id="FF9703A1FFADBF31AF8CEF4A49F99A66" box="[1002,1018,312,338]" pageId="35" pageNumber="36">9</date>
</figureCitation>
I).
</paragraph>
</subSubSection>
<subSubSection id="C33376EAFFADBF31ACA1EF2D49839A96" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8B962561FFADBF31ACA1EF2D49839A96" blockId="35.[151,1436,152,418]" pageId="35" pageNumber="36">
<emphasis id="B95DF973FFADBF31ACA1EF2D4B649A4D" bold="true" box="[199,359,351,377]" pageId="35" pageNumber="36">Distribution.</emphasis>
Presently known with certainty only from
<collectingCountry id="F33E65F1FFADBF31AF0EEF124E1C9A4E" box="[872,1055,352,378]" name="New Caledonia" pageId="35" pageNumber="36">New Caledonia</collectingCountry>
(
<typeStatus id="54929BC3FFADBF31A856EF124E679A4E" box="[1072,1124,352,378]" pageId="35" pageNumber="36">type</typeStatus>
locality), Loyalty Islands, and Nansha Islands in South
<collectingCountry id="F33E65F1FFADBF31AD8AEFFA48319A96" box="[492,562,392,418]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
Sea; at depths of
<quantity id="4CD18884FFADBF31AF64EFFA497F9A96" box="[770,892,392,418]" metricMagnitude="1" metricUnit="m" metricValue="6.6" metricValueMax="12.0" metricValueMin="1.2" pageId="35" pageNumber="36" unit="m" value="66.0" valueMax="120.0" valueMin="12.0">
12
<specimenCount id="9D2FEEE8FFADBF31AF48EFFA497F9A96" box="[814,892,392,418]" pageId="35" pageNumber="36" type="generic">120 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>