195 lines
27 KiB
XML
195 lines
27 KiB
XML
<document id="E285699A4673BE20170EAD840F72933B" ID-DOI="10.5281/zenodo.175414" ID-GBIF-Dataset="f485f2e0-6053-49c6-9863-4c71023258b3" ID-ISSN="11755326" ID-Zenodo-Dep="175414" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459808583420" checkinUser="plazi" docAuthor="Thomas, Richard & Hedges, Blair" docDate="2007" docId="004FE949FFBAFFC2FF3FFF52FCC92736" docLanguage="en" docName="zt01400p026.pdf" docOrigin="Zootaxa 1400" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Typhlops biminiensis" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FC769131FFB4FFCCFFA8FFC5FF942112" masterDocTitle="Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba" masterLastPageNumber="26" masterPageNumber="1" pageNumber="15" updateTime="1698220755238" updateUser="plazi">
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<mods:title id="E94CEB84EAB27E7B152BA3F89A1EB1F0">Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba</mods:title>
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<mods:name id="6E1A678BB7CB6ED9332F3DBD4DEDF643" type="personal">
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<mods:namePart id="345B0DCD4E20306A4753913C2F643D0D">Thomas, Richard</mods:namePart>
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<mods:namePart id="19F776561731D24B07CCCCD231E944B4">Hedges, Blair</mods:namePart>
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<mods:title id="40E2D21304535791EDADBEA1E0D39A26">Zootaxa</mods:title>
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<mods:date id="D9EF8FB739346A671FD2A5F4DC1E136A">2007</mods:date>
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<mods:identifier id="4D0B8A09B1F6CE00448C7F18700D612E" type="DOI">10.5281/zenodo.175414</mods:identifier>
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<treatment id="004FE949FFBAFFC2FF3FFF52FCC92736" ID-DOI="http://doi.org/10.5281/zenodo.5677327" ID-GBIF-Taxon="119353297" ID-Zenodo-Dep="5677327" LSID="urn:lsid:plazi:treatment:004FE949FFBAFFC2FF3FFF52FCC92736" httpUri="http://treatment.plazi.org/id/004FE949FFBAFFC2FF3FFF52FCC92736" lastPageNumber="15" pageId="14" pageNumber="15">
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<subSubSection id="C0FC0BD4FFBAFFC2FF3FFF52FD9821A3" box="[151,524,151,177]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8859585FFFBAFFC2FF3FFF52FD9821A3" blockId="14.[151,524,151,177]" box="[151,524,151,177]" pageId="14" pageNumber="15">
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<heading id="D311EF33FFBAFFC2FF3FFF52FD9821A3" bold="true" box="[151,524,151,177]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="BA92844DFFBAFFC2FF3FFF52FD9821A3" bold="true" box="[151,524,151,177]" pageId="14" pageNumber="15">
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<taxonomicName id="4FE623DCFFBAFFC2FF3FFF52FE0221A3" box="[151,406,151,177]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFBAFFC2FF3FFF52FE0221A3" bold="true" box="[151,406,151,177]" italics="true" pageId="14" pageNumber="15">Typhlops biminiensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="10DD44DAFFBAFFC2FE0EFF52FE7921A3" box="[422,493,151,177]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
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A)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C0FC0BD4FFBAFFC2FF3FFF27FC36226E" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8859585FFFBAFFC2FF3FFF27FC36226E" blockId="14.[151,1439,226,1572]" pageId="14" pageNumber="15">
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<emphasis id="BA92844DFFBAFFC2FF3FFF27FD2621E9" box="[151,690,226,251]" italics="true" pageId="14" pageNumber="15">
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Diagnosis:
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<taxonomicName id="4FE623DCFFBAFFC2FEB7FF27FDCB21E9" box="[287,607,226,251]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="biminiensis" subSpecies="sensu">
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Typhlops biminiensis
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<taxonomicNameLabel id="A1A13936FFBAFFC2FDB6FF27FDCB21E9" box="[542,607,226,251]" pageId="14" pageNumber="15" sensu="stricto">sensu</taxonomicNameLabel>
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</taxonomicName>
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stricto
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</emphasis>
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is one of the largest West Indian species of
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<taxonomicName id="4FE623DCFFBAFFC2FB12FF27FAB521E9" box="[1210,1313,226,251]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
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<emphasis id="BA92844DFFBAFFC2FB12FF27FAB521E9" box="[1210,1313,226,251]" italics="true" pageId="14" pageNumber="15">Typhlops</emphasis>
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</taxonomicName>
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, TL (total length) to
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<quantity id="4F1EF5BAFFBAFFC2FEA7FECFFEE02036" box="[271,372,266,292]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="14" pageNumber="15" unit="mm" value="370.0">370 mm</quantity>
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. The preocular contacts supralabials 2 and 3, which is a characteristic of the
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<taxonomicName id="4FE623DCFFBAFFC2FB51FECFFA0F2031" box="[1273,1435,266,291]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFBAFFC2FB51FECFFA0F2031" box="[1273,1435,266,291]" italics="true" pageId="14" pageNumber="15">T. biminiensis</emphasis>
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</taxonomicName>
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Group (
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<bibRefCitation id="EC7725AEFFBAFFC2FF59FEF7FE2D205E" author="Richmond" box="[241,441,306,332]" pageId="14" pageNumber="26" refString="Richmond, N. D. (1955) The blind snakes (Typhlops) of Bimini, Bahama Islands, British West Indies, with description of a new species. American Museum Novitates, 1 - 7." type="book chapter" year="1955">Richmond, 1955</bibRefCitation>
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;
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<bibRefCitation id="EC7725AEFFBAFFC2FE6CFEF7FDF9205E" author="Thomas" box="[452,621,306,332]" pageId="14" pageNumber="26" refString="Thomas, R. (1968) The Typhlops biminiensis group of Antillean blindsnakes. Copeia, 1968, 713 - 722." type="journal article" year="1968">Thomas, 1968</bibRefCitation>
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) of West Indian species. This feature also occurs in all mainland Central and South American species (
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<bibRefCitation id="EC7725AEFFBAFFC2FD8CFE9FFCCD2066" author="Dixon" box="[548,857,346,372]" pageId="14" pageNumber="25" refString="Dixon, J. R. & Hendricks, F. S. (1979) The wormsnakes (Family Typhlopidae) of the Neotropics, exclusive of the Antilles. Zoologische Verhandelingen, 1 - 39." type="book chapter" year="1979">Dixon & Hendricks, 1979</bibRefCitation>
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) but not most West Indian species, which belong to another West Indian radiation of
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<taxonomicName id="4FE623DCFFBAFFC2FDEFFE47FD3A2089" box="[583,686,386,411]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
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<emphasis id="BA92844DFFBAFFC2FDEFFE47FD3A2089" box="[583,686,386,411]" italics="true" pageId="14" pageNumber="15">Typhlops</emphasis>
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</taxonomicName>
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, the major Antillean radiation (MAR) (
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<bibRefCitation id="EC7725AEFFBAFFC2FB38FE47FAD5208E" author="Thomas" box="[1168,1345,386,412]" pageId="14" pageNumber="26" refString="Thomas, R. (1976) Systematics of Antillean blindsnakes of the genus Typhlops (Serpentes: Typhlopidae). Ph. D. dissertation. Louisiana State University, Baton Rouge, 288 pp." type="book" year="1976">Thomas, 1976</bibRefCitation>
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). Aside from the preocularcontact character,
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<taxonomicName id="4FE623DCFFBAFFC2FDE6FE6FFD7B20D1" box="[590,751,426,451]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFBAFFC2FDE6FE6FFD7B20D1" box="[590,751,426,451]" italics="true" pageId="14" pageNumber="15">T. biminiensis</emphasis>
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</taxonomicName>
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is distinguished from members of the MAR by its broadly angled preocular (93–125°) and its broad rostral scale (RW1/RL1 0.81–1.08). The only currently recognized member of the Major Antillean Radiation occurring within the range of
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<taxonomicName id="4FE623DCFFBAFFC2FBADFE3FFB3F2301" box="[1029,1195,506,531]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFBAFFC2FBADFE3FFB3F2301" box="[1029,1195,506,531]" italics="true" pageId="14" pageNumber="15">T. biminiensis</emphasis>
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</taxonomicName>
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is
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<taxonomicName id="4FE623DCFFBAFFC2FB7CFE3FFAED2301" box="[1236,1401,506,532]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="lumbricalis">
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<emphasis id="BA92844DFFBAFFC2FB7CFE3FFB762301" box="[1236,1250,506,531]" italics="true" pageId="14" pageNumber="15">T</emphasis>
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.
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<emphasis id="BA92844DFFBAFFC2FB5CFE3FFAED2301" box="[1268,1401,506,531]" italics="true" pageId="14" pageNumber="15">lumbricalis</emphasis>
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</taxonomicName>
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of
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<collectingCountry id="F0F118CFFFBAFFC2FF3FFDE7FF41232E" box="[151,213,546,572]" name="Cuba" pageId="14" pageNumber="15">Cuba</collectingCountry>
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and the Great and Little
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<collectingCountry id="F0F118CFFFBAFFC2FDAFFDE7FDF3232E" box="[519,615,546,572]" name="Bahamas" pageId="14" pageNumber="15">Bahama</collectingCountry>
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Bank islands. From described members of the
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<taxonomicName id="4FE623DCFFBAFFC2FB0CFDE7FA0F2329" box="[1188,1435,546,571]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
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<emphasis id="BA92844DFFBAFFC2FB0CFDE7FA0F2329" box="[1188,1435,546,571]" italics="true" pageId="14" pageNumber="15">Typhlops biminiensis</emphasis>
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</taxonomicName>
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group (
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<taxonomicName id="4FE623DCFFBAFFC2FF44FD8FFE0A2371" box="[236,414,586,611]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="caymanensis">
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<emphasis id="BA92844DFFBAFFC2FF44FD8FFE0A2371" box="[236,414,586,611]" italics="true" pageId="14" pageNumber="15">T. caymanensis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4FE623DCFFBAFFC2FE04FD8FFDBF2371" box="[428,555,586,611]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="epactius">
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<emphasis id="BA92844DFFBAFFC2FE04FD8FFDBF2371" box="[428,555,586,611]" italics="true" pageId="14" pageNumber="15">T. epactius</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="4FE623DCFFBAFFC2FDC3FD8FFC972371" box="[619,771,586,611]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="paradoxus">
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<emphasis id="BA92844DFFBAFFC2FDC3FD8FFC972371" box="[619,771,586,611]" italics="true" pageId="14" pageNumber="15">T. paradoxus</emphasis>
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</taxonomicName>
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),
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<emphasis id="BA92844DFFBAFFC2FCB3FD8FFBCC2371" box="[795,1112,586,611]" italics="true" pageId="14" pageNumber="15">
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<taxonomicName id="4FE623DCFFBAFFC2FCB3FD8FFB912371" box="[795,1029,586,611]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="biminiensis" subSpecies="sensu">
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T. biminiensis
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<taxonomicNameLabel id="A1A13936FFBAFFC2FC6CFD8FFB912371" box="[964,1029,586,611]" pageId="14" pageNumber="15" sensu="stricto">sensu</taxonomicNameLabel>
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</taxonomicName>
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stricto
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</emphasis>
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differs in its larger size and broad, nearly round, rostral scale (
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<figureCitation id="10DD44DAFFBAFFC2FD84FDB7FDE4239E" box="[556,624,626,652]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
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A) with RW1/RL1 values of over 0.81. It further differs from
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<taxonomicName id="4FE623DCFFBAFFC2FAE4FDB7FE9023A1" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="caymanensis">
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<emphasis id="BA92844DFFBAFFC2FAE4FDB7FE9023A1" italics="true" pageId="14" pageNumber="15">T. caymanensis</emphasis>
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</taxonomicName>
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by its greater number of scale rows (24 or 22 initially versus 20 rows for
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<taxonomicName id="4FE623DCFFBAFFC2FB0EFD5FFAC923A1" box="[1190,1373,666,691]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="caymanensis">
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<emphasis id="BA92844DFFBAFFC2FB0EFD5FFAC923A1" box="[1190,1373,666,691]" italics="true" pageId="14" pageNumber="15">T. caymanensis</emphasis>
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</taxonomicName>
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) and greater number of middorsal scales (454–537 versus 351–408). From the Cuban species (
|
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<figureCitation id="10DD44DAFFBAFFC2FB17FD07FA9223CE" box="[1215,1286,706,732]" captionStart="FIGURE 8" captionStartId="13.[151,255,1722,1746]" captionTargetBox="[273,1319,349,1708]" captionTargetId="figure@13.[273,1320,349,1709]" captionTargetPageId="13" captionText="FIGURE 8. Head scalation in the Typhlops biminiensis Group, including seven new species from Cuba. (A) T. biminiensis (ASFS V 11655), from Elbow Cay, Cay Sal Bank, Bahamas. (B) T. paradoxus (ASFS V 12067), from Great Inagua, Bahamas. (C) T. epactius (ASFS V 11673), from Cayman Brac, Cayman Islands. (D) T. caymanensis (ASFS V 17112), from Grand Cayman, Cayman Islands. (E) T. perimychus (USNM 564800), holotype, from Guantanamo Bay Naval Station, Cuba. (F) T. arator (MNHNCu 191), holotype, from El Narigon, La Habana, Province, Cuba). (G) T. anousius (MNHNCu 4550), holotype, from 3.5 km E Tortuguilla, Guantánamo Province, Cuba. (H) T. notorachius (MNHNCu 4551), holotype, from 9.4 km W Imias, Guantánamo Province, Cuba. (I) T. contorhinus (MNHNCu 4552), holotype, from 4.9 km S La Tinta, Guantánamo Province, Cuba. (J) T. anchaurus (MNHNCu 4553), holotype, from within 1 km of Cueva de Agua (2.5 km airline NW Maisí), Guantánamo Province, Cuba. (K) T. satelles (SMF 61303), holotype, from Rancho Luna, about 12 km airline S Cienfuegos, Cienfuegos Province, Cuba." httpUri="https://zenodo.org/record/175421/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
) it differs in having a larger rostral size (RW1+RL3;
|
||
<figureCitation id="10DD44DAFFBAFFC2FDDAFD2FFD522216" box="[626,710,746,772]" captionStart="FIGURE 10" captionStartId="17.[151,255,1319,1343]" captionTargetBox="[293,1229,214,1276]" captionTargetId="figure@17.[279,1308,194,1295]" captionTargetPageId="17" captionText="FIGURE 10. Variation in measurements of Typhlops biminiensis and Cuban species of the Typhlops biminiensis Group. For all graphs (A – F), the following species are indicated by symbols: T. anchaurus (open diamond), T. anousius (closed square), T. arator (closed diamond), T. biminiensis (open circle), T. contorhinus (open triangle), T. notorachius (closed triangle), T. paradoxus (open square), T. perimychus (closed circle), T. satelles (“ X ”). Trendlines are indicated on some graphs for T. biminiensis and T. perimychus. See text for definition of abbreviations." httpUri="https://zenodo.org/record/175423/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
|
||
A), longer snout (NE;
|
||
<figureCitation id="10DD44DAFFBAFFC2FC67FD2FFBB02216" box="[975,1060,746,772]" captionStart="FIGURE 10" captionStartId="17.[151,255,1319,1343]" captionTargetBox="[293,1229,214,1276]" captionTargetId="figure@17.[279,1308,194,1295]" captionTargetPageId="17" captionText="FIGURE 10. Variation in measurements of Typhlops biminiensis and Cuban species of the Typhlops biminiensis Group. For all graphs (A – F), the following species are indicated by symbols: T. anchaurus (open diamond), T. anousius (closed square), T. arator (closed diamond), T. biminiensis (open circle), T. contorhinus (open triangle), T. notorachius (closed triangle), T. paradoxus (open square), T. perimychus (closed circle), T. satelles (“ X ”). Trendlines are indicated on some graphs for T. biminiensis and T. perimychus. See text for definition of abbreviations." httpUri="https://zenodo.org/record/175423/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
|
||
B), and smaller preocular apical diameter (PD;
|
||
<figureCitation id="10DD44DAFFBAFFC2FEE0FCD7FE34223E" box="[328,416,786,812]" captionStart="FIGURE 10" captionStartId="17.[151,255,1319,1343]" captionTargetBox="[293,1229,214,1276]" captionTargetId="figure@17.[279,1308,194,1295]" captionTargetPageId="17" captionText="FIGURE 10. Variation in measurements of Typhlops biminiensis and Cuban species of the Typhlops biminiensis Group. For all graphs (A – F), the following species are indicated by symbols: T. anchaurus (open diamond), T. anousius (closed square), T. arator (closed diamond), T. biminiensis (open circle), T. contorhinus (open triangle), T. notorachius (closed triangle), T. paradoxus (open square), T. perimychus (closed circle), T. satelles (“ X ”). Trendlines are indicated on some graphs for T. biminiensis and T. perimychus. See text for definition of abbreviations." httpUri="https://zenodo.org/record/175423/files/figure.png" pageId="14" pageNumber="15">Fig. 10</figureCitation>
|
||
C). Because the new Cuban species can thus be distinguished from
|
||
<taxonomicName id="4FE623DCFFBAFFC2FB78FCD7FAE12239" box="[1232,1397,786,811]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
|
||
<emphasis id="BA92844DFFBAFFC2FB78FCD7FAE12239" box="[1232,1397,786,811]" italics="true" pageId="14" pageNumber="15">T. biminiensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4FE623DCFFBAFFC2FA2EFCD7FEB92241" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="caymanensis">
|
||
<emphasis id="BA92844DFFBAFFC2FA2EFCD7FEB92241" italics="true" pageId="14" pageNumber="15">T. caymanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4FE623DCFFBAFFC2FEE8FCFFFE512241" box="[320,453,826,851]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="epactius">
|
||
<emphasis id="BA92844DFFBAFFC2FEE8FCFFFE512241" box="[320,453,826,851]" italics="true" pageId="14" pageNumber="15">T. epactius</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4FE623DCFFBAFFC2FDA7FCFFFD382241" box="[527,684,826,851]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="paradoxus">
|
||
<emphasis id="BA92844DFFBAFFC2FDA7FCFFFD382241" box="[527,684,826,851]" italics="true" pageId="14" pageNumber="15">T. paradoxus</emphasis>
|
||
</taxonomicName>
|
||
, our discussion below will focus mostly on diagnostic traits among the Cuban species previously confused with
|
||
<taxonomicName id="4FE623DCFFBAFFC2FD50FCA7FC0E2269" box="[760,922,866,891]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
|
||
<emphasis id="BA92844DFFBAFFC2FD50FCA7FC0E2269" box="[760,922,866,891]" italics="true" pageId="14" pageNumber="15">T. biminiensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0FC0BD4FFBAFFC2FF6EFC4FFB352496" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8859585FFFBAFFC2FF6EFC4FFB352496" blockId="14.[151,1439,226,1572]" pageId="14" pageNumber="15">
|
||
<emphasis id="BA92844DFFBAFFC2FF6EFC4FFECF22B1" box="[198,347,906,931]" italics="true" pageId="14" pageNumber="15">Description:</emphasis>
|
||
Snout rounded, slightly lobed. Rostral broad and rounded in dorsal aspect, nearly as broad as long in most specimens (RW1/RL1 0.81–1.08), broadly oval, almost circular; not flared on apex; minimal umbo; no labial flare. PA 93–125°, apex rounded; lower portion of preocular contacting upper labials two and three. Anterior nasal width as a proportion of RW1 0.30–0.37. OL 1/2 height, OS 0.09–0.28. Rostronasal pattern strongly divergent. Postocular 3–4 times higher than long, rarely two postoculars as a result of a second scale encroaching minimally on the postocular space. First parietal standard, spanning two scale rows or slightly less. Second parietal present and similar in size to the first, or absent. TL to
|
||
<quantity id="4F1EF5BAFFBAFFC2FB29FBBFFB732586" box="[1153,1255,1146,1172]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="14" pageNumber="15" unit="mm" value="380.0">380 mm</quantity>
|
||
. TL/TA 42–60. TL/MBD 39–51. Middorsal scales 454–537. Scale rows 22 with no posterior reduction or 24 reducing to 22 at about midbody or not reducing at all. Coloration bicolor with dorsal pigmentation (pale brown) ending abruptly along a midlateral or dorsolateral line by dropping out of pigmentation on individual scales, resulting in a jagged line of separation between pigmented and nonpigmented scales; or color fading gradually over 2– 3 scale rows, without dropping out of pigment; a combination of these effects may occur with a gradual transition anteriorly and dropping out of pigmented scales posteriorly. Hemipenes attenuate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C0FC0BD4FFBAFFC2FF6EFA56FCC92736" pageId="14" pageNumber="15" type="distribution">
|
||
<paragraph id="8859585FFFBAFFC2FF6EFA56FEC624C6" blockId="14.[151,1439,226,1572]" pageId="14" pageNumber="15">
|
||
<emphasis id="BA92844DFFBAFFC2FF6EFA56FEF424BE" box="[198,352,1427,1452]" italics="true" pageId="14" pageNumber="15">Distribution:</emphasis>
|
||
<collectingCountry id="F0F118CFFFBAFFC2FECFFA57FE4324BE" box="[359,471,1426,1452]" name="Bahamas" pageId="14" pageNumber="15">Bahamas</collectingCountry>
|
||
: Bimini Islands, Andros Island, New Providence, Elbow Cay (Cay Sal Bank), and Ragged Islands.
|
||
</paragraph>
|
||
<paragraph id="8859585FFFBAFFC2FF6EFA27FCC92736" blockId="14.[151,1439,226,1572]" pageId="14" pageNumber="15">
|
||
The first of the new species having affinities with
|
||
<taxonomicName id="4FE623DCFFBAFFC2FC88FA26FC5724EE" box="[800,963,1507,1532]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="biminiensis">
|
||
<emphasis id="BA92844DFFBAFFC2FC88FA26FC5724EE" box="[800,963,1507,1532]" italics="true" pageId="14" pageNumber="15">T. biminiensis</emphasis>
|
||
</taxonomicName>
|
||
occurs on the southern coast of eastern
|
||
<collectingCountry id="F0F118CFFFBAFFC2FF3FF9CFFF4C2736" box="[151,216,1546,1572]" name="Cuba" pageId="14" pageNumber="15">Cuba</collectingCountry>
|
||
, in the region of Guantánamo Bay. It may be known as
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |