252 lines
18 KiB
XML
252 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.158.50848" ID-GBIF-Dataset="4d1dd045-c029-47d4-b441-a7fd49443918" ID-PMC="PMC7483337" ID-Pensoft-Pub="1314-2003-158-1" ID-Pensoft-UUID="8480BB61E8B2553ABA8D418072EC52B6" ID-PubMed="32973385" ModsDocID="1314-2003-158-1" checkinTime="1599212532520" checkinUser="pensoft" docAuthor="Calvo, Joel & Moreira-Munoz, Andres" docDate="2020" docId="97F64DB226675409B52ADAFA143C0081" docLanguage="en" docName="PhytoKeys 158: 1-106" docOrigin="PhytoKeys 158" docSource="http://dx.doi.org/10.3897/phytokeys.158.50848" docTitle="Xenophyllum esquilachense V. A. Funk, Novon 7 (3): 239. 1997" docType="treatment" docVersion="4" id="8480BB61E8B2553ABA8D418072EC52B6" lastPageNumber="1" masterDocId="8480BB61E8B2553ABA8D418072EC52B6" masterDocTitle="Taxonomic revision of the Andean genus Xenophyllum (Compositae, Senecioneae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="1" updateTime="1668139998449" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Andean genus Xenophyllum (Compositae, Senecioneae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Calvo, Joel</mods:namePart>
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<mods:affiliation>Instituto de Geografia, Facultad de Ciencias del Mar y Geografia, Pontificia Universidad Catolica de Valparaiso, Avenida Brasil 2241, 2362807 Valparaiso, Chile</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-2340-7666</mods:nameIdentifier>
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<mods:nameIdentifier type="email">calvocasas@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Moreira-Munoz, Andres</mods:namePart>
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<mods:affiliation>Instituto de Geografia, Facultad de Ciencias del Mar y Geografia, Pontificia Universidad Catolica de Valparaiso, Avenida Brasil 2241, 2362807 Valparaiso, Chile</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-9136-1391</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>158</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>106</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.158.50848</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.158.50848</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-158-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8480BB61E8B2553ABA8D418072EC52B6</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="167491855" LSID="urn:lsid:plazi:treatment:97F64DB226675409B52ADAFA143C0081" httpUri="http://treatment.plazi.org/id/97F64DB226675409B52ADAFA143C0081" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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22.
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<taxonomicName LSID="97F64DB2-2667-5409-B52A-DAFA143C0081" authority="(Cuatrec.) V. A. Funk, Novon 7 (3): 239. 1997." class="Magnoliopsida" family="Asteraceae" genus="Xenophyllum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Xenophyllum esquilachense" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="esquilachense">Xenophyllum esquilachense (Cuatrec.) V.A.Funk, Novon 7(3): 239. 1997.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Werneria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Werneria esquilachensis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="esquilachensis">Werneria esquilachensis</taxonomicName>
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Cuatrec., Brittonia 8: 192. 1956. Type. Peru. Puno: San Antonio de Esquilache, 4725 m, 12 May 1937,
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<emphasis italics="true" pageId="0" pageNumber="1">D. Stafford 716</emphasis>
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(holotype: K s.n.!).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Ricardi & Marticor" authorityYear="1964" class="Magnoliopsida" family="Asteraceae" genus="Senecio" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Senecio pfisteri" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pfisteri">Senecio pfisteri</taxonomicName>
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Ricardi & Martic., Gayana, Bot. 11: 25. 1964. Type. Chile. Arica-Parinacota: Arica, Portezuelo de
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<normalizedToken originalValue="Chapiquiña">Chapiquina</normalizedToken>
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, faldeos al lado norte del campamento, 4400 m, 10 Feb 1964,
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<emphasis italics="true" pageId="0" pageNumber="1">C. Marticorena, O. Matthei & M. Quezada 116</emphasis>
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(holotype: CONC-29863!; isotype: CONC s.n.!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Suffruticose plant, forming clumps of erect or decumbent stems.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Rhizomes</emphasis>
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4-8
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<normalizedToken originalValue="×">x</normalizedToken>
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0.1-0.2 cm, horizontal to oblique, glabrous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Stems</emphasis>
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5-10 cm tall, branched, glabrescent, usually with leaves restricted to the upper part.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Leaves</emphasis>
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somewhat distantly arranged, abruptly narrowed at the base; leaf laminas 11.2-15.5
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<normalizedToken originalValue="×">x</normalizedToken>
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3.3-4.6 mm, broadly spatulate, 3-forked at the apex, entire and scabrous-ciliate (cilia ca. 0.17 mm long), flat in cross section, glabrous, 1-nerved above (barely visible), 1-nerved beneath, somewhat fleshy, matte, papillose; leaf lobes 2.4-3.9
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<normalizedToken originalValue="×">x</normalizedToken>
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0.7-1 mm, with 2 to 3 rough mucronate teeth up to 1.4 mm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Capitula</emphasis>
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disciform, erect, sessile to subsessile.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Involucres</emphasis>
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10.4-15.1
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<normalizedToken originalValue="×">x</normalizedToken>
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6.5-7.4 mm, cupuliform; involucral bracts 8 to 12, 5.4-7.2
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<normalizedToken originalValue="×">x</normalizedToken>
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1.3-2.5 mm, acute to obtuse at the apex, greenish, sometimes purple-edged.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Peripheral florets</emphasis>
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ca. 10, pistillate; corollas reduced to a vestigial tube ca. 2.1 mm long or without corolla.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Disc florets</emphasis>
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ca. 15; corollas 6-10.6 mm long, purplish; style branches truncate with a crown of sweeping trichomes, purplish.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Achenes</emphasis>
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cylindrical, glabrous (immature); pappus ca. 11.3 mm long, barbellate, whitish. Chromosome number unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="iconography">
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<paragraph pageId="0" pageNumber="1">Iconography.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.382.3.5" author="Calvo, J" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" pagination="288 - 292" refId="B9" refString="Calvo, J, Munoz-Schick, M, Moreira-Munoz, A, 2018. Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species. Phytotaxa 382 (3): 288 - 292, DOI: https://doi.org/10.11646/phytotaxa.382.3.5" title="Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species." url="https://doi.org/10.11646/phytotaxa.382.3.5" volume="382" year="2018">Calvo et al. (2018</bibRefCitation>
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: 289, fig. 1, as photo).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Southern Peru to central-western Bolivia. Bolivia (Oruro), Chile (Arica-Parinacota), Peru (Arequipa, Moquegua, Puno, Tacna). It grows in rocky outcrops, steep volcanic slopes, and open shrublands of the subhumid and dry puna ecoregions, between elevations of 3700-4950 m (Fig.
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<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Distribution map of Xenophyllum staffordiae (left hand) and X. esquilachense (right hand)." figureDoi="10.3897/phytokeys.158.50848.figure27" httpUri="https://binary.pensoft.net/fig/448907" pageId="0" pageNumber="1">27</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="phenology">
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<paragraph pageId="0" pageNumber="1">Phenology.</paragraph>
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<paragraph pageId="0" pageNumber="1">Flowering from February to June.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">It is named after the Peruvian village of San Antonio de Esquilache (western Puno), where the type material was collected in 1937 by Dora Stafford.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="notes">
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<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="1">This is the unique species within the genus displaying disciform capitula. They have ca. 10, peripheral, pistillate florets with a corolla reduced to a vestigial tube ca. 2.1 mm long or without corolla, and therefore, the style is exsert in its whole length or almost so. Another distinctive feature is the morphology of the leaves; they are 3-forked, abruptly narrowed at the base, and marginally scabrous-ciliate. Each lobe usually has 2 to 3 rough mucronate teeth. Any confusion with the other members of the genus is unlikely.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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See
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<bibRefCitation DOI="https://doi.org/10.11646/phytotaxa.382.3.5" author="Calvo, J" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="1" pagination="288 - 292" refId="B9" refString="Calvo, J, Munoz-Schick, M, Moreira-Munoz, A, 2018. Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species. Phytotaxa 382 (3): 288 - 292, DOI: https://doi.org/10.11646/phytotaxa.382.3.5" title="Towards a better understanding of Xenophyllum esquilachense (Senecioneae, Compositae), a poorly known Andean species." url="https://doi.org/10.11646/phytotaxa.382.3.5" volume="382" year="2018">Calvo et al. (2018)</bibRefCitation>
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for further details on the type material of the name
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<taxonomicName authorityName="Ricardi & Marticor" authorityYear="1964" class="Magnoliopsida" family="Asteraceae" genus="Senecio" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Senecio pfisteri" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pfisteri">
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||
<emphasis italics="true" pageId="0" pageNumber="1">Senecio pfisteri</emphasis>
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</taxonomicName>
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Ricardi & Martic.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Additional specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Bolivia. Oruro</emphasis>
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: Challapata, Livichuco, Tarpata, estribaciones del cerro Toro en el collado antes de las lagunas,
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<geoCoordinate degrees="18" direction="south" minutes="57" orientation="latitude" precision="925" value="-18.95">18°57'S</geoCoordinate>
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,
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<geoCoordinate degrees="66" direction="west" minutes="24" orientation="longitude" precision="925" value="-66.4">66°24'W</geoCoordinate>
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, 15 Feb 2019,
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<emphasis italics="true" pageId="0" pageNumber="1">
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J. Calvo & M.
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<normalizedToken originalValue="Zárate">Zarate</normalizedToken>
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7877
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</emphasis>
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(BOLV); prov. Eduardo Abaroa, Challapata, comunidad Churacani,
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<geoCoordinate degrees="18" direction="south" minutes="55" orientation="latitude" precision="925" value="-18.916666">18°55'S</geoCoordinate>
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,
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<geoCoordinate degrees="66" direction="west" minutes="40" orientation="longitude" precision="925" value="-66.666664">66°40'W</geoCoordinate>
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, 1 Apr 2018,
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<emphasis italics="true" pageId="0" pageNumber="1">
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M.
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<normalizedToken originalValue="Guzmán">Guzman</normalizedToken>
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124
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</emphasis>
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(LPB); Abaroa, Challapata, cerro Toro Ichurata, en
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<normalizedToken originalValue="línea">linea</normalizedToken>
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recta a 34.38 km al E,
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<geoCoordinate degrees="18" direction="south" minutes="57" orientation="latitude" precision="925" value="-18.95">18°57'S</geoCoordinate>
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,
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<geoCoordinate degrees="66" direction="west" minutes="27" orientation="longitude" precision="925" value="-66.45">66°27'W</geoCoordinate>
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, 27 Feb 2016,
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<emphasis italics="true" pageId="0" pageNumber="1">F. Zenteno, L. Moya & D. Villalba 16745</emphasis>
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(LPB).
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<emphasis bold="true" pageId="0" pageNumber="1">Chile. Arica-Parinacota</emphasis>
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: quebrada Cataguanchuta,
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<geoCoordinate degrees="18" direction="south" minutes="14" orientation="latitude" precision="925" value="-18.233334">18°14'S</geoCoordinate>
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,
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<geoCoordinate degrees="69" direction="west" minutes="31" orientation="longitude" precision="925" value="-69.51667">69°31'W</geoCoordinate>
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, 17 Dec 1988,
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<emphasis italics="true" pageId="0" pageNumber="1">E. Belmonte 88644</emphasis>
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(CONC); pasado azufrera Tacora,
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<geoCoordinate degrees="17" direction="south" minutes="42" orientation="latitude" precision="925" value="-17.7">17°42'S</geoCoordinate>
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,
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<geoCoordinate degrees="69" direction="west" minutes="49" orientation="longitude" precision="925" value="-69.816666">69°49'W</geoCoordinate>
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, 17 Mar 2015,
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<emphasis italics="true" pageId="0" pageNumber="1">
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A.
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<normalizedToken originalValue="Moreira-Muñoz">Moreira-Munoz</normalizedToken>
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& F. Luebert 2407
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</emphasis>
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(SGO); Portezuelo de
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<normalizedToken originalValue="Chapiquiña">Chapiquina</normalizedToken>
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,
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<geoCoordinate degrees="18" direction="south" minutes="19" orientation="latitude" precision="925" value="-18.316668">18°19'S</geoCoordinate>
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,
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<geoCoordinate degrees="69" direction="west" minutes="30" orientation="longitude" precision="925" value="-69.5">69°30'W</geoCoordinate>
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, 26 Mar 1961,
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<emphasis italics="true" pageId="0" pageNumber="1">M. Ricardi, C. Marticorena & O. Matthei 207</emphasis>
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(CONC).
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<emphasis bold="true" pageId="0" pageNumber="1">Peru. Arequipa</emphasis>
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: Chiguata, faldas del nevado Pichu Pichu,
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<geoCoordinate degrees="16" direction="south" minutes="27" orientation="latitude" precision="925" value="-16.45">16°27'S</geoCoordinate>
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,
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<geoCoordinate degrees="71" direction="west" minutes="27" orientation="longitude" precision="925" value="-71.45">71°27'W</geoCoordinate>
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, 20 Jun 2004,
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<emphasis italics="true" pageId="0" pageNumber="1">V. Quipuscoa et al. 2949</emphasis>
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(HSP); Chiguata, ca. 2 km arriba del
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<normalizedToken originalValue="túnel">tunel</normalizedToken>
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del Simbral, carretera a Salinas,
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<geoCoordinate degrees="16" direction="south" minutes="22" orientation="latitude" precision="925" value="-16.366667">16°22'S</geoCoordinate>
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,
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<geoCoordinate degrees="71" direction="west" minutes="17" orientation="longitude" precision="925" value="-71.28333">71°17'W</geoCoordinate>
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, 15 Apr 2018,
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<emphasis italics="true" pageId="0" pageNumber="1">V. Quipuscoa & M. Balvin 7541</emphasis>
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(HSP);
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<emphasis bold="true" pageId="0" pageNumber="1">Moquegua</emphasis>
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: Mariscal Nieto, Carumas, cerca a un acueducto,
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<geoCoordinate degrees="16" direction="south" minutes="50" orientation="latitude" precision="925" value="-16.833334">16°50'S</geoCoordinate>
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,
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<geoCoordinate degrees="70" direction="west" minutes="34" orientation="longitude" precision="925" value="-70.566666">70°34'W</geoCoordinate>
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, 15 Jun 2013,
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<emphasis italics="true" pageId="0" pageNumber="1">
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H.
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<normalizedToken originalValue="Beltrán">Beltran</normalizedToken>
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7734
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</emphasis>
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(USM); General
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<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
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Cerro, Yunga, Sura-Perusa,
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<geoCoordinate degrees="16" direction="south" minutes="11" orientation="latitude" precision="925" value="-16.183332">16°11'S</geoCoordinate>
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,
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<geoCoordinate degrees="70" direction="west" minutes="38" orientation="longitude" precision="925" value="-70.63333">70°38'W</geoCoordinate>
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, 13 Apr 2012,
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<emphasis italics="true" pageId="0" pageNumber="1">D. Montesinos 3804</emphasis>
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(HSP); General
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<normalizedToken originalValue="Sánchez">Sanchez</normalizedToken>
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Cerro, Yunga, Perusa,
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<geoCoordinate degrees="16" direction="south" minutes="11" orientation="latitude" precision="925" value="-16.183332">16°11'S</geoCoordinate>
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,
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<geoCoordinate degrees="70" direction="west" minutes="37" orientation="longitude" precision="925" value="-70.61667">70°37'W</geoCoordinate>
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, 3 Mar 2018,
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<emphasis italics="true" pageId="0" pageNumber="1">D. Montesinos & J. Calvo 5958</emphasis>
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(HSP);
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<emphasis bold="true" pageId="0" pageNumber="1">Tacna</emphasis>
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: Tarata, Poma, carretera Tarata-Puno,
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<geoCoordinate degrees="17" direction="south" minutes="25" orientation="latitude" precision="925" value="-17.416666">17°25'S</geoCoordinate>
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,
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<geoCoordinate degrees="69" direction="west" minutes="56" orientation="longitude" precision="925" value="-69.933334">69°56'W</geoCoordinate>
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, 25 Mar 1998,
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<emphasis italics="true" pageId="0" pageNumber="1">A. Cano 8139</emphasis>
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(USM).
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |