treatments-xml/data/1F/53/FD/1F53FD42D67CFF8CFF1975C6FC86FC9C.xml
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<document id="0BA843AD89422AA2EA3DEC1925FB672F" ID-DOI="10.5281/zenodo.194870" ID-GBIF-Dataset="3e0d4d97-b5d0-472e-bc4d-87134c7565b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="194870" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460218486540" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2010" docId="1F53FD42D67CFF8CFF1975C6FC86FC9C" docLanguage="en" docName="zt02438p036.pdf" docOrigin="Zootaxa 2438" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptobasis candelaria Alayo 1968" docType="treatment" docVersion="13" lastPageNumber="7" masterDocId="E36A853AD679FF8AFF8E703CFF92FFC8" masterDocTitle="Redefinition of Leptobasis Selys with the synonymy of Chrysobasis Rácenis and description of L. mauffrayi sp. nov. from Peru (Odonata: Coenagrionidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="6" updateTime="1698237345539" updateUser="plazi">
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<mods:title id="67CD4529925092BA7B666E185FAC4065">Redefinition of Leptobasis Selys with the synonymy of Chrysobasis Rácenis and description of L. mauffrayi sp. nov. from Peru (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="47D8E316523A0C7C4E302DA51F544C2A">Garrison, Rosser W.</mods:namePart>
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<mods:namePart id="9D97E4A05A5EC4D4ECC0D6E44F72FA42">Ellenrieder, Natalia Von</mods:namePart>
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<mods:date id="A03D1BDC4C6FA0F963A69800C4820FBE">2010</mods:date>
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<treatment id="1F53FD42D67CFF8CFF1975C6FC86FC9C" ID-DOI="http://doi.org/10.5281/zenodo.6203230" ID-GBIF-Taxon="119384944" ID-Zenodo-Dep="6203230" LSID="urn:lsid:plazi:treatment:1F53FD42D67CFF8CFF1975C6FC86FC9C" httpUri="http://treatment.plazi.org/id/1F53FD42D67CFF8CFF1975C6FC86FC9C" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="DFE01FDFD67CFF8FFF1975C6FD35F9FF" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="97454C54D67CFF8FFF1975C6FDA0F9DC" blockId="5.[151,679,1530,1591]" box="[151,562,1530,1556]" pageId="5" pageNumber="6">
<heading id="CC0DFB38D67CFF8FFF1975C6FDA0F9DC" bold="true" box="[151,562,1530,1556]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="50FA37D7D67CFF8FFF1975C6FDA0F9DC" ID-CoL="3TJZH" authority="Alayo 1968" authorityName="Alayo" authorityYear="1968" box="[151,562,1530,1556]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67CFF8FFF1975C6FDA0F9DC" bold="true" box="[151,562,1530,1556]" pageId="5" pageNumber="6">
<emphasis id="A58E9046D67CFF8FFF1975C6FE32F9DC" bold="true" box="[151,416,1530,1556]" italics="true" pageId="5" pageNumber="6">Leptobasis candelaria</emphasis>
Alayo 1968
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="97454C54D67CFF8FFF197621FD35F9FF" blockId="5.[151,679,1530,1591]" box="[151,679,1565,1591]" pageId="5" pageNumber="6">
<figureCitation id="0FC150D1D67CFF8FFF197621FF76F9FF" box="[151,228,1565,1591]" captionStart="FIGURE 1" captionStartId="17.[151,255,1606,1630]" captionTargetBox="[229,1344,194,1567]" captionTargetId="figure@17.[227,1360,175,1582]" captionTargetPageId="17" captionText="FIGURE 1. Head, dorsal view of right half. (a) Leptobasis candelaria, male, Mexico, Veracruz, E of Córdoba; (b) L. guanacaste, male paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. mauffrayi, male holotype, Peru, Manu, Pakitza; (d) L. vacillans, male, Dominican Republic, La Vega, S of La Vega." httpUri="https://zenodo.org/record/194871/files/figure.png" pageId="5" pageNumber="6">Figs. 1</figureCitation>
a, 2b, 3b, 5a, 6b, 10, 19a, 20b, 21b, 27
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67CFF8FFF19765CFF4FF82F" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="97454C54D67CFF8FFF19765CFB8AF8FF" blockId="5.[151,1436,1631,1847]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="B7EA6B7AD67CFF8FFF19765CFB8AF8FF" pageId="5" pageNumber="6">
<taxonomicName id="50FA37D7D67CFF8FFF19765CFDC9F9BF" ID-CoL="3TJZH" authority="Alayo 1968a: 83" authorityName="Alayo" authorityPageNumber="83" authorityYear="1968" box="[151,603,1632,1655]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67CFF8FFF19765CFE1EF9BF" box="[151,396,1632,1655]" italics="true" pageId="5" pageNumber="6">Leptobasis candelaria</emphasis>
<treatmentCitation id="165B6A45D67CFF8FFE14765CFDC9F9BF" author="Alayo" box="[410,603,1632,1655]" page="83" pageId="5" pageNumber="6" year="1968">
<bibRefCitation id="F36B31A5D67CFF8FFE14765CFDBDF9BF" author="Alayo" box="[410,559,1632,1655]" pageId="5" pageNumber="13" refString="Alayo, D. P. (1968 a) Las libelulas de Cuba (Insecta-Odonata). Torreia (Nueva Serie), 2 (Parte I: Texto), 1 - 102." type="journal article" year="1968" yearSuffix="a">Alayo 1968a</bibRefCitation>
: 83
</treatmentCitation>
</taxonomicName>
(description:
<typeStatus id="4841F2F6D67CFF8FFD70765CFCC9F9BF" box="[766,859,1632,1655]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
3,
<taxonomicName id="50FA37D7D67CFF8FFC1C765CFB9BF9BF" ID-CoL="3TJZH" box="[914,1033,1632,1655]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="35" phylum="Arthropoda" rank="species" species="candelaria">Candelaria</taxonomicName>
, Pinar del Río,
<collectingCountry id="EFED0CC4D67CFF8FFB45765CFA94F9BF" box="[1227,1286,1632,1655]" name="Cuba" pageId="5" pageNumber="6">Cuba</collectingCountry>
, in IES [not examined]); 1968b: 45, 51 (illustrations of
<typeStatus id="4841F2F6D67CFF8FFD0176BCFD7EF95F" box="[655,748,1664,1687]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
3);
<emphasis id="A58E9046D67CFF8FFC9376BCFCA7F95F" box="[797,821,1664,1687]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFCB576BCFB91F95F" author="Paulson" box="[827,1027,1664,1687]" page="254" pageId="5" pageNumber="6" year="1982">
<bibRefCitation id="F36B31A5D67CFF8FFCB576BCFC5CF95F" author="Paulson" box="[827,974,1664,1687]" pageId="5" pageNumber="16" refString="Paulson, D. R. (1982) Odonata. In: Hurlbert, S. H. &amp; Villalobos-Figueroa, A. (Eds), Aquatic biota of Mexico, Central America and the West Indies. San Diego State University, San Diego, California, pp. 249 - 277." type="book chapter" year="1982">Paulson 1982</bibRefCitation>
: 254
</treatmentCitation>
, 260 (
<collectingCountry id="EFED0CC4D67CFF8FFBCC76BCFB0AF95F" box="[1090,1176,1664,1687]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
, West Indies);
<emphasis id="A58E9046D67CFF8FFABA76BCFADEF95F" box="[1332,1356,1664,1687]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFADC76BCFE01F97F" author="Davies" page="77" pageId="5" pageNumber="6" year="1984">
<bibRefCitation id="F36B31A5D67CFF8FFADC76BCFEFBF97F" author="Davies" pageId="5" pageNumber="14" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The dragonflies of the world: a systematic list of the extant species of Odonata. Volume 1. Zygoptera, Anisozygoptera. Societas Internationalis Odonatologica Rapid Communications (Supplement), 3, 1 - 127." type="journal article" year="1984">Davies &amp; Tobin 1984</bibRefCitation>
: 77
</treatmentCitation>
(catalog);
<emphasis id="A58E9046D67CFF8FFD81769CFDB5F97F" box="[527,551,1696,1719]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFDBC769CFCE2F97F" author="Gonzalez-Soriano" box="[562,880,1696,1719]" page="296" pageId="5" pageNumber="6" year="1993">
<bibRefCitation id="F36B31A5D67CFF8FFDBC769CFCA5F97F" author="Gonzalez-Soriano" box="[562,823,1696,1719]" pageId="5" pageNumber="15" refString="Gonzalez-Soriano, E. (1993) Odonata de Mexico: Situacion actual y perspectivas de estudio. Revista de la Sociedad Mexicana de Historia Natural, Volumen Especial, 44, 291 - 302." type="journal article" year="1993">González-Soriano 1993</bibRefCitation>
: 296
</treatmentCitation>
;
<emphasis id="A58E9046D67CFF8FFC0E769CFC0AF97F" box="[896,920,1696,1719]" italics="true" pageId="5" pageNumber="6"></emphasis>
<bibRefCitation id="F36B31A5D67CFF8FFC2D769CFBA8F97F" author="Bridges" box="[931,1082,1696,1719]" pageId="5" pageNumber="13" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group, and species-group names of the Odonata of the world (third edition). Bridges, Urbana, Illinois, 905 pp." type="book" year="1994">Bridges 1994</bibRefCitation>
:
<date id="E3446A94D67CFF8FFBC6769CFB1CF97F" box="[1096,1166,1696,1719]" pageId="5" pageNumber="6" value="1943-07">VII.43</date>
(catalog);
<emphasis id="A58E9046D67CFF8FFA82769CFAB6F97F" box="[1292,1316,1696,1719]" italics="true" pageId="5" pageNumber="6"></emphasis>
González-
<treatmentCitation id="165B6A45D67CFF8FFF4976FCFDFCF91F" author="Soriano" box="[199,622,1728,1751]" page="164" pageId="5" pageNumber="6" year="1996">
<bibRefCitation id="F36B31A5D67CFF8FFF4976FCFDA9F91F" author="Soriano" box="[199,571,1728,1751]" pageId="5" pageNumber="15" refString="Gonzalez-Soriano, E. &amp; Novelo-Gutierrez, R. (1996) Odonata. In: Llorente Bousquets, J., Garcia Aldrete, A. N. &amp; Gonzalez Soriano, E. (Eds.), Biodiversidad, taxonomia y biogeografia de artropodos de Mexico: Hacia una sintesis de su conocimiento. Universidad Autonoma de Mexico, Mexico City, pp. 147 - 167." type="book chapter" year="1996">Soriano &amp; Novelo-Gutiérrez 1996</bibRefCitation>
: 164
</treatmentCitation>
(
<collectingCountry id="EFED0CC4D67CFF8FFDF076FCFD47F91F" box="[638,725,1728,1751]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
);
<emphasis id="A58E9046D67CFF8FFD6676FCFC92F91F" box="[744,768,1728,1751]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFC8676FCFC33F91F" author="Flint" box="[776,929,1728,1751]" page="19" pageId="5" pageNumber="6" year="1996">
<bibRefCitation id="F36B31A5D67CFF8FFC8676FCFCE9F91F" author="Flint" box="[776,891,1728,1751]" pageId="5" pageNumber="14" refString="Flint, O. S., Jr. (1996) The Odonata of Cuba, with a report on a recent collection and checklist of the Cuban species. Cocuyo, 5, 17 - 20." type="journal article" year="1996">Flint 1996</bibRefCitation>
: 19
</treatmentCitation>
(checklist);
<emphasis id="A58E9046D67CFF8FFBA776FCFBD3F91F" box="[1065,1089,1728,1751]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFBC676FCFABCF91F" author="Steinmann" box="[1096,1326,1728,1751]" page="287" pageId="5" pageNumber="6" year="1997">
<bibRefCitation id="F36B31A5D67CFF8FFBC676FCFB69F91F" author="Steinmann" box="[1096,1275,1728,1751]" pageId="5" pageNumber="16" refString="Steinmann, H. (1997). World catalogue of Odonata. Volume I. Zygoptera. In: Wermuth, H. &amp; Fischer, M. (Eds.), Das Tierreich. The Animal Kingdom. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen, Volume 110. Walter de Gruyter, Berlin, 500 pp." type="book" year="1997">Steinmann 1997</bibRefCitation>
: 287
</treatmentCitation>
(catalog);
<emphasis id="A58E9046D67CFF8FFF4976DCFF4DF93F" box="[199,223,1760,1783]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFF6876DCFE1EF93F" author="Tsuda" box="[230,396,1760,1783]" page="38" pageId="5" pageNumber="6" year="2000">
<bibRefCitation id="F36B31A5D67CFF8FFF6876DCFEF7F93F" author="Tsuda" box="[230,357,1760,1783]" pageId="5" pageNumber="16" refString="Tsuda, S. (2000) A distributional list of world Odonata 2000. Privately published, Osaka, 430 pp." type="book" year="2000">Tsuda 2000</bibRefCitation>
: 38
</treatmentCitation>
(catalog);
<emphasis id="A58E9046D67CFF8FFD8E76DCFD8AF93F" box="[512,536,1760,1783]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFDAE76DCFD40F93F" author="Forster" box="[544,722,1760,1783]" page="62" pageId="5" pageNumber="6" year="2001">
<bibRefCitation id="F36B31A5D67CFF8FFDAE76DCFD3EF93F" author="Forster" box="[544,684,1760,1783]" pageId="5" pageNumber="14" refString="Forster, S. (2001) The dragonflies of Central America exclusive of Mexico and the West Indies. A guide to their identification [Second edition]. Odonatological monographs 2, Gunnar Rehfeldt, Wolfenbuttel, Germany, 141 pp." type="book" year="2001">Förster 2001</bibRefCitation>
: 62
</treatmentCitation>
(key to species);
<emphasis id="A58E9046D67CFF8FFC0776DCFC33F93F" box="[905,929,1760,1783]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFC2676DCFA1CF93F" author="Trapero" box="[936,1422,1760,1783]" page="27" pageId="5" pageNumber="6" year="2003">
<bibRefCitation id="F36B31A5D67CFF8FFC2676DCFAFBF93F" author="Trapero" box="[936,1385,1760,1783]" pageId="5" pageNumber="16" refString="Trapero Quintana, A. D. &amp; Naranjo Lopez, C. (2003) Revision of the order Odonata in Cuba. Bulletin of American Odonatology, 7 (2), 23 - 40." type="journal article" year="2003">Trapero Quintana &amp; Naranjo López 2003</bibRefCitation>
: 27
</treatmentCitation>
28 (
<collectingCountry id="EFED0CC4D67CFF8FFF7D773CFEA3F8DF" box="[243,305,1792,1815]" name="Cuba" pageId="5" pageNumber="6">Cuba</collectingCountry>
);
<emphasis id="A58E9046D67CFF8FFEC8773CFECCF8DF" box="[326,350,1792,1815]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFEE9773CFCF4F8DF" author="Trapero" box="[359,870,1792,1815]" page="179" pageId="5" pageNumber="6" year="2004">
<bibRefCitation id="F36B31A5D67CFF8FFEE9773CFCA3F8DF" author="Trapero" box="[359,817,1792,1815]" pageId="5" pageNumber="16" refString="Trapero Quintana, A. D. &amp; Naranjo Lopez, C. (2004) Clave de identificacion para los adultos de las especies del Orden Odonata presentes en Cuba. Boletin de la Sociedad Entomologica Aragonesa, 35, 171 - 180." type="journal article" year="2004">Trapero Quintana &amp; Naranjo López 2004</bibRefCitation>
: 179
</treatmentCitation>
(in key);
<emphasis id="A58E9046D67CFF8FFC59773CFC7DF8DF" box="[983,1007,1792,1815]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFC76773CFEA0F8FF" author="Gonzalez-Soriano" page="129" pageId="5" pageNumber="6" year="2007">
<bibRefCitation id="F36B31A5D67CFF8FFC76773CFE92F8FF" author="Gonzalez-Soriano" pageId="5" pageNumber="15" refString="Gonzalez-Soriano, E. &amp; Novelo-Gutierrez, R. (2007) Odonata of Mexico revisited, In: Tyagi, B. K. (Ed.), Odonata: biology of dragonflies. Pawan Kumar, Scientific Publishers, Jodhpur, India, pp. 105 - 136." type="book chapter" year="2007">González-Soriano &amp; Novelo-Gutiérrez 2007</bibRefCitation>
: 129
</treatmentCitation>
(
<collectingCountry id="EFED0CC4D67CFF8FFECE771CFE05F8FF" box="[320,407,1824,1847]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
);
<emphasis id="A58E9046D67CFF8FFE24771CFE50F8FF" box="[426,450,1824,1847]" italics="true" pageId="5" pageNumber="6"></emphasis>
<treatmentCitation id="165B6A45D67CFF8FFE46771CFC26F8FF" author="Novelo-Gutierrez" box="[456,948,1824,1847]" page="686" pageId="5" pageNumber="6" year="2009">
<bibRefCitation id="F36B31A5D67CFF8FFE46771CFC10F8FF" author="Novelo-Gutierrez" box="[456,898,1824,1847]" pageId="5" pageNumber="15" refString="Novelo-Gutierrez, R. &amp; Gomez-Anaya, J. A. (2009) A comparative study of Odonata (Insecta) assemblages along an altitudinal gradient in the Sierra de Coalcoman Mountains, Michoacan, Mexico. Biodiversity Conservation, 18, 679 - 698." type="journal article" year="2009">Novelo-Gutiérrez &amp; Gómez-Anaya 2009</bibRefCitation>
: 686
</treatmentCitation>
(record).
</treatmentCitationGroup>
</paragraph>
<paragraph id="97454C54D67CFF8FFF197761FF4FF82F" blockId="5.[151,1436,1884,2023]" pageId="5" pageNumber="6">
<emphasis id="A58E9046D67CFF8FFF197761FE3AF8BF" bold="true" box="[151,424,1885,1911]" pageId="5" pageNumber="6">Specimens examined.</emphasis>
Total 2 3, 2 Ƥ: 1 3, 1 Ƥ,
<collectingCountry id="EFED0CC4D67CFF8FFD637761FCD9F8BF" box="[749,843,1885,1911]" name="Mexico" pageId="5" pageNumber="6">Mexico</collectingCountry>
, Veracruz State, stream
<quantity id="5002E1B1D67CFF8FFBE17761FB51F8BF" box="[1135,1219,1885,1911]" metricMagnitude="3" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="6" unit="km" value="6.9">6.9 km</quantity>
E of Córdoba, by Mex. Hwy. 150,
<geoCoordinate id="F2CE2A93D67CFF8FFED077BEFE79F854" box="[350,491,1922,1948]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="15" value="18.89">18°53'24&quot;N</geoCoordinate>
,
<geoCoordinate id="F2CE2A93D67CFF8FFE7677BEFD1AF854" box="[504,648,1922,1948]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="15" value="-96.92">96°55'12&quot;W</geoCoordinate>
, 12 xiii 1976, leg. R.W. &amp; J.A. Garrison; 1 3, 1 Ƥ,
<collectingCountry id="EFED0CC4D67CFF8FFB7E77BEFAACF854" box="[1264,1342,1922,1948]" name="Belize" pageId="5" pageNumber="6">Belize</collectingCountry>
, Toledo District, Monkey River,
<geoCoordinate id="F2CE2A93D67CFF8FFE357794FDB2F80A" box="[443,544,1960,1986]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="16.35">16°21'N</geoCoordinate>
,
<geoCoordinate id="F2CE2A93D67CFF8FFDA37794FD07F80A" box="[557,661,1960,1986]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="-88.48333">88°29'W</geoCoordinate>
,
<date id="E3446A94D67CFF8FFD2D7794FCA4F80A" box="[675,822,1960,1986]" pageId="5" pageNumber="6" value="1995-08-27">27 viii 1995</date>
, leg. Tineke Boomsma, All specimens deposited in RWG.
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67FFF8CFF4870ABFD10FCE7" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="97454C54D67FFF8CFF4870ABFCF3FD2C" blockId="6.[151,1437,151,852]" pageId="6" pageNumber="7">
<emphasis id="A58E9046D67FFF8CFF4870ABFEC3FF7C" bold="true" box="[198,337,151,180]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
The combination of metafemoral armature (
<figureCitation id="0FC150D1D67FFF8CFC1170A4FC7CFF7C" box="[927,1006,152,180]" captionStart="FIGURE 5" captionStartId="19.[151,256,1573,1597]" captionTargetBox="[163,1418,187,1542]" captionTargetId="figure@19.[151,1436,172,1552]" captionTargetPageId="19" captionText="FIGURE 5. Male metafemur. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. vacillans, Dominican Republic, Jarabacoa." httpUri="https://zenodo.org/record/194873/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
a) and convoluted, foliate cercus (
<figureCitation id="0FC150D1D67FFF8CFF2E70FCFE99FF14" box="[160,267,192,220]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="6" pageNumber="7">Figs. 20</figureCitation>
b, 21b) will distinguish males from all others except for
<taxonomicName id="50FA37D7D67FFF8CFC6F70FCFB07FF14" box="[993,1173,192,220]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67FFF8CFC6F70FCFB07FF14" box="[993,1173,192,220]" italics="true" pageId="6" pageNumber="7">L. guanacaste</emphasis>
</taxonomicName>
. The proximal large metafemoral spur of
<taxonomicName id="50FA37D7D67FFF8CFE2A70D4FDC0FECC" box="[420,594,232,260]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67FFF8CFE2A70D4FDC0FECC" box="[420,594,232,260]" italics="true" pageId="6" pageNumber="7">L. candelaria</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67FFF8CFDEA70D4FD23FECC" box="[612,689,232,260]" captionStart="FIGURE 5" captionStartId="19.[151,256,1573,1597]" captionTargetBox="[163,1418,187,1542]" captionTargetId="figure@19.[151,1436,172,1552]" captionTargetPageId="19" captionText="FIGURE 5. Male metafemur. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. vacillans, Dominican Republic, Jarabacoa." httpUri="https://zenodo.org/record/194873/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
a) is not as long as that of
<taxonomicName id="50FA37D7D67FFF8CFB8970D4FB2EFECC" box="[1031,1212,232,260]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67FFF8CFB8970D4FB2EFECC" box="[1031,1212,232,260]" italics="true" pageId="6" pageNumber="7">L. guanacaste</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67FFF8CFB4370D4FA8BFECC" box="[1229,1305,232,260]" captionStart="FIGURE 5" captionStartId="19.[151,256,1573,1597]" captionTargetBox="[163,1418,187,1542]" captionTargetId="figure@19.[151,1436,172,1552]" captionTargetPageId="19" captionText="FIGURE 5. Male metafemur. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. vacillans, Dominican Republic, Jarabacoa." httpUri="https://zenodo.org/record/194873/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
b) and the shape of the cerci differ between the two species. The ventro-basal process in
<taxonomicName id="50FA37D7D67FFF8CFB4B712CFAEEFEE4" box="[1221,1404,272,300]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67FFF8CFB4B712CFAEEFEE4" box="[1221,1404,272,300]" italics="true" pageId="6" pageNumber="7">L. candelaria</emphasis>
</taxonomicName>
is projected ventro-posteriorly and is visible in medio-dorsal view when the cerci are spread (
<figureCitation id="0FC150D1D67FFF8CFA937104FAEEFE9C" box="[1309,1404,312,340]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="6" pageNumber="7">Fig. 20</figureCitation>
b); this structure is ventrally curved and hidden from view when the cercus is viewed medio-dorsally in
<taxonomicName id="50FA37D7D67FFF8CFF1971B4FED8FE6C" box="[151,330,392,420]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67FFF8CFF1971B4FED8FE6C" box="[151,330,392,420]" italics="true" pageId="6" pageNumber="7">L. guanacaste</emphasis>
</taxonomicName>
(
<figureCitation id="0FC150D1D67FFF8CFED471B4FE2BFE6C" box="[346,441,392,420]" captionStart="FIGURE 20" captionStartId="27.[151,256,1327,1351]" captionTargetBox="[154,1425,183,1295]" captionTargetId="figure@27.[151,1436,180,1306]" captionTargetPageId="27" captionText="FIGURE 20. Male S 10, medio-dorsal view. (a) Leptobasis buchholzi, Colombia, Puerto Colombia; (b) L. candelaria, Belize, Toledo, Monkey River; (c) L. guanacaste, holotype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Costa Rica, Heredia, Finca La Selva; (e) L. mauffrayi, holotype, Peru, Madre de Dios, Pakitza; (f) L. melinogaster, USA, Texas, Hidalgo, Santa Ana Natural Wildlife Reserve; (g) L. raineyi, Trinidad, St. Andrew, 1 mi W of Cumuto; (h) L. vacillans, Puerto Rico, Mayagüez, Mayagüez." httpUri="https://zenodo.org/record/194881/files/figure.png" pageId="6" pageNumber="7">Fig. 20</figureCitation>
c). Females of
<taxonomicName id="50FA37D7D67FFF8CFDFF71B4FC8FFE6C" box="[625,797,392,420]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67FFF8CFDFF71B4FC8FFE6C" box="[625,797,392,420]" italics="true" pageId="6" pageNumber="7">L. candelaria</emphasis>
</taxonomicName>
and
<taxonomicName id="50FA37D7D67FFF8CFCD471B4FB9FFE6C" box="[858,1037,392,420]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67FFF8CFCD471B4FB9FFE6C" box="[858,1037,392,420]" italics="true" pageId="6" pageNumber="7">L. guanacaste</emphasis>
</taxonomicName>
are unique in having the dorsal margin of ovipositor concave in lateral view (
<figureCitation id="0FC150D1D67FFF8CFD55718CFCD1FE04" box="[731,835,432,460]" captionStart="FIGURE 19" captionStartId="26.[151,255,1251,1275]" captionTargetBox="[153,1435,199,1216]" captionTargetId="figure@26.[151,1436,194,1230]" captionTargetPageId="26" captionText="FIGURE 19. Female S 7 10, lateral view. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, allotype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. lucifer, Mexico, Campeche, Laguna Zoh; (d) L. mauffrayi, allotype, Peru, Madre de Dios, Pakitza; (e) L. raineyi, Trinidad, St. George, Indian Walk Ride; (f) L. vacillans, Mexico, Morelos, La Fuente." httpUri="https://zenodo.org/record/194880/files/figure.png" pageId="6" pageNumber="7">Figs. 19</figureCitation>
a, b); this structure is linear in all other species (
<figureCitation id="0FC150D1D67FFF8CFF2E71E4FE98FE3C" box="[160,266,472,500]" captionStart="FIGURE 19" captionStartId="26.[151,255,1251,1275]" captionTargetBox="[153,1435,199,1216]" captionTargetId="figure@26.[151,1436,194,1230]" captionTargetPageId="26" captionText="FIGURE 19. Female S 7 10, lateral view. (a) Leptobasis candelaria, Belize, Toledo, Monkey River; (b) L. guanacaste, allotype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. lucifer, Mexico, Campeche, Laguna Zoh; (d) L. mauffrayi, allotype, Peru, Madre de Dios, Pakitza; (e) L. raineyi, Trinidad, St. George, Indian Walk Ride; (f) L. vacillans, Mexico, Morelos, La Fuente." httpUri="https://zenodo.org/record/194880/files/figure.png" pageId="6" pageNumber="7">Figs. 19</figureCitation>
cf). The female of
<taxonomicName id="50FA37D7D67FFF8CFD8271E4FD2BFE3C" box="[524,697,472,500]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67FFF8CFD8271E4FD2BFE3C" box="[524,697,472,500]" italics="true" pageId="6" pageNumber="7">L. candelaria</emphasis>
</taxonomicName>
differs by the unique morphology of its prothoracic hind lobe; its middle lobe is expanded with lateral margins angulate and overlying a smaller inferior margin (
<figureCitation id="0FC150D1D67FFF8CFE8B7214FEC6FD8C" box="[261,340,552,580]" captionStart="FIGURE 2" captionStartId="17.[151,255,1702,1726]" captionText="FIGURE 2. Female pronotum, dorsal view. (a) Leptobasis buchholzi, paratype, Venezuela, Guárico, Pericoco [redrawn from a draft by J. De Marmels]; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Mexico, Campeche, Laguna Zoh; (e) L. mauffrayi, allotype, Peru, Manu, Pakitza; (f) L. melinogaster, USA, Texas, Kleberg, King Ranch; (g) L. raineyi, Trinidad, St. George, Indian Walk Ride; (h) L. vacillans, Cuba, Cienfuegos, E of Pepito Rey." pageId="6" pageNumber="7">Fig. 2</figureCitation>
b). In
<taxonomicName id="50FA37D7D67FFF8CFE2D7214FDC8FD8C" box="[419,602,552,580]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67FFF8CFE2D7214FDC8FD8C" box="[419,602,552,580]" italics="true" pageId="6" pageNumber="7">L. guanacaste</emphasis>
</taxonomicName>
, the hind lobe is upright, of even width, and its lateral arms are deflexed ventrally diverging from the carinate propleural extension (
<figureCitation id="0FC150D1D67FFF8CFB83726CFBCEFDA4" box="[1037,1116,592,620]" captionStart="FIGURE 2" captionStartId="17.[151,255,1702,1726]" captionText="FIGURE 2. Female pronotum, dorsal view. (a) Leptobasis buchholzi, paratype, Venezuela, Guárico, Pericoco [redrawn from a draft by J. De Marmels]; (b) L. candelaria, female, Belize, Toledo, Monkey River; (c) L. guanacaste, paratype, Costa Rica, Guanacaste, Hacienda Taboga; (d) L. lucifer, Mexico, Campeche, Laguna Zoh; (e) L. mauffrayi, allotype, Peru, Manu, Pakitza; (f) L. melinogaster, USA, Texas, Kleberg, King Ranch; (g) L. raineyi, Trinidad, St. George, Indian Walk Ride; (h) L. vacillans, Cuba, Cienfuegos, E of Pepito Rey." pageId="6" pageNumber="7">Fig. 2</figureCitation>
c). Mature individuals of
<taxonomicName id="50FA37D7D67FFF8CFF197244FEDEFD5C" box="[151,332,632,660]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="A58E9046D67FFF8CFF197244FEDEFD5C" box="[151,332,632,660]" italics="true" pageId="6" pageNumber="7">L. candelaria</emphasis>
</taxonomicName>
have broad mid-dorsal and humeral dark stripes and the pale postocular spots are round (
<figureCitation id="0FC150D1D67FFF8CFF7B729CFEADFD74" box="[245,319,672,700]" captionStart="FIGURE 1" captionStartId="17.[151,255,1606,1630]" captionTargetBox="[229,1344,194,1567]" captionTargetId="figure@17.[227,1360,175,1582]" captionTargetPageId="17" captionText="FIGURE 1. Head, dorsal view of right half. (a) Leptobasis candelaria, male, Mexico, Veracruz, E of Córdoba; (b) L. guanacaste, male paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. mauffrayi, male holotype, Peru, Manu, Pakitza; (d) L. vacillans, male, Dominican Republic, La Vega, S of La Vega." httpUri="https://zenodo.org/record/194871/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
a); in mature
<taxonomicName id="50FA37D7D67FFF8CFE7C729CFD3AFD74" box="[498,680,672,700]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="A58E9046D67FFF8CFE7C729CFD3AFD74" box="[498,680,672,700]" italics="true" pageId="6" pageNumber="7">L. guanacaste</emphasis>
</taxonomicName>
, the dark antehumeral stripe is lacking and the postocular spots are represented by narrow pale triangles (
<figureCitation id="0FC150D1D67FFF8CFD7D72F4FCAEFD2C" box="[755,828,712,740]" captionStart="FIGURE 1" captionStartId="17.[151,255,1606,1630]" captionTargetBox="[229,1344,194,1567]" captionTargetId="figure@17.[227,1360,175,1582]" captionTargetPageId="17" captionText="FIGURE 1. Head, dorsal view of right half. (a) Leptobasis candelaria, male, Mexico, Veracruz, E of Córdoba; (b) L. guanacaste, male paratype, Costa Rica, Guanacaste, Hacienda Taboga; (c) L. mauffrayi, male holotype, Peru, Manu, Pakitza; (d) L. vacillans, male, Dominican Republic, La Vega, S of La Vega." httpUri="https://zenodo.org/record/194871/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
b).
</paragraph>
<paragraph id="97454C54D67FFF8CFF4872D3FD10FCE7" blockId="6.[151,1437,151,852]" pageId="6" pageNumber="7">
<emphasis id="A58E9046D67FFF8CFF4872D3FECCFCC1" bold="true" box="[198,350,751,777]" pageId="6" pageNumber="7">Dimensions.</emphasis>
Males (
<emphasis id="A58E9046D67FFF8CFE4A72CCFE41FCC1" box="[452,467,752,777]" italics="true" pageId="6" pageNumber="7">n</emphasis>
= 2): Hw 15.015.6; abdomen 28; total length 3334. Females (
<emphasis id="A58E9046D67FFF8CFB6D72CCFB60FCC1" box="[1251,1266,752,777]" italics="true" pageId="6" pageNumber="7">n</emphasis>
2): Hw 17.2 18.1; abdomen 2628; total length 3233.
</paragraph>
</subSubSection>
<subSubSection id="DFE01FDFD67FFF8CFF487306FC86FC9C" box="[198,788,826,852]" pageId="6" pageNumber="7" type="distribution">
<paragraph id="97454C54D67FFF8CFF487306FC86FC9C" blockId="6.[151,1437,151,852]" box="[198,788,826,852]" pageId="6" pageNumber="7">
<emphasis id="A58E9046D67FFF8CFF487306FEF5FC9C" bold="true" box="[198,359,826,852]" pageId="6" pageNumber="7">Distribution.</emphasis>
<collectingCountry id="EFED0CC4D67FFF8CFEE07306FE3DFC9C" box="[366,431,826,852]" name="Cuba" pageId="6" pageNumber="7">Cuba</collectingCountry>
,
<collectingCountry id="EFED0CC4D67FFF8CFE347306FD85FC9C" box="[442,535,826,852]" name="Mexico" pageId="6" pageNumber="7">Mexico</collectingCountry>
, and
<collectingCountry id="EFED0CC4D67FFF8CFDDA7306FD0DFC9C" box="[596,671,826,852]" name="Belize" pageId="6" pageNumber="7">Belize</collectingCountry>
(
<figureCitation id="0FC150D1D67FFF8CFD217306FC95FC9C" box="[687,775,826,852]" captionStart="FIGURE 27" captionStartId="34.[151,254,1739,1763]" captionTargetBox="[160,1423,522,1711]" captionTargetId="figure@34.[151,1436,514,1718]" captionTargetPageId="34" captionText="FIGURE 27. Distribution of Leptobasis buchholzi, L. candelaria, L. guanacaste, L. lucifer, and L. melinogaster." httpUri="https://zenodo.org/record/194886/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>