342 lines
24 KiB
XML
342 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.222.87779" ID-Pensoft-Pub="1314-2003-222-75" ID-Pensoft-UUID="348E6053A6D55787993694B23682FDE9" ModsDocID="1314-2003-222-75" checkinTime="1679617121532" checkinUser="pensoft" docAuthor="Maas, Paul J. M., Maas-van de Kamer, Hiltje, Andre, Thiago, Skinner, David, Valderrama, Eugenio & Specht, Chelsea D." docDate="2023" docId="191887D029815EFAA87A8804961FE2FE" docLanguage="en" docName="PhytoKeys 222: 75-127" docOrigin="PhytoKeys 222" docPubDate="2023-03-22" docSource="http://dx.doi.org/10.3897/phytokeys.222.87779" docTitle="Costus rubineus D. Skinner & Maas 2023, sp. nov." docType="treatment" docVersion="1" id="348E6053A6D55787993694B23682FDE9" lastPageNumber="75" masterDocId="348E6053A6D55787993694B23682FDE9" masterDocTitle="Eighteen new species of Neotropical Costaceae (Zingiberales)" masterLastPageNumber="127" masterPageNumber="75" pageNumber="75" updateTime="1679617121532" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eighteen new species of Neotropical Costaceae (Zingiberales)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maas, Paul J. M.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maas-van de Kamer, Hiltje</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Andre, Thiago</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4148-3662</mods:nameIdentifier>
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<mods:affiliation>Universidade de Brasilia, Departamento de Botanica, Campus Universitario Darcy Ribeiro, Asa Norte, Brasilia (DF), Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skinner, David</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4585-925X</mods:nameIdentifier>
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<mods:affiliation>Le Jardin Ombrage, Tallahassee, (Private botanical garden, Botanic Gardens Conservation International - BGCI - registration ID 50148), Florida, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Valderrama, Eugenio</mods:namePart>
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<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Specht, Chelsea D.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7746-512X</mods:nameIdentifier>
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<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
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<mods:nameIdentifier type="email">cdspecht@cornell.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>222</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>127</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.222.87779</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.222.87779</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-222-75</mods:identifier>
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<mods:identifier type="Pensoft-UUID">348E6053A6D55787993694B23682FDE9</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:191887D029815EFAA87A8804961FE2FE" httpUri="http://treatment.plazi.org/id/191887D029815EFAA87A8804961FE2FE" lastPageNumber="75" pageId="0" pageNumber="75">
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<subSubSection pageId="0" pageNumber="75" type="nomenclature">
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName LSID="191887D0-2981-5EFA-A87A-8804961FE2FE" authority="D. Skinner & Maas" authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus" status="sp. nov.">Costus rubineus D.Skinner & Maas</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="75">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="diagnosis">
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<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus rubineus</emphasis>
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</taxonomicName>
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sp. nov. (Fig.
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<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costus rubineus D. Skinner & Maas A plant in habitat near Villa Rica, Pasco, Peru B inflorescence with flowers showing the labellum with characteristic lobulate apex (a) C adaxial leaf midrib showing absence of line of hairs D bract and flower detail from plant in habitat at Alto Churumazu, Pasco, Peru showing the full flower with subtending bract (flower), bract (br), bracteole subtending the capsule (ovary plus calyx) (brl / ca), stigma and style as attached to the young capsule (gyn), petals, single fertile stamen (st) and labellum (lab). Seeds from this plant were collected and cultivated as D. Skinner R 3421 E plant in habitat at 1980 meters near Chinchao, Huanuco, Peru, the original collection site of " C. ruber " as indicated in Ruiz' journal F inflorescence with lower bracts removed and single flower photographed in the wild near Chinchao, Peru. Photos A-F by Dave Skinner." figureDoi="10.3897/phytokeys.222.87779.figure19" httpUri="https://binary.pensoft.net/fig/828358" pageId="0" pageNumber="75">19</figureCitation>
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) looks somewhat similar to
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. scaber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="scaber">
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<emphasis italics="true" pageId="0" pageNumber="75">C. scaber</emphasis>
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</taxonomicName>
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Ruiz & Pav. but differs from that species by a larger calyx (9-15 vs. 3-7 mm), larger bracteole (20-28 vs. 9-12 mm), larger corolla (50-60 vs. 35-40 mm), and a labellum which is distinctly lobulate at the apex vs. entire in
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. scaber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="scaber">
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<emphasis italics="true" pageId="0" pageNumber="75">C. scaber</emphasis>
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</taxonomicName>
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; furthermore it is restricted to high elevations whereas
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. scaber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="scaber">
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<emphasis italics="true" pageId="0" pageNumber="75">C. scaber</emphasis>
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</taxonomicName>
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is mostly a lowland species;
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
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<emphasis italics="true" pageId="0" pageNumber="75">C. rubineus</emphasis>
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</taxonomicName>
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also lacks the row of hairs along the adaxial midvein of the lamina, which is so typical for
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. scaber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="scaber">
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<emphasis italics="true" pageId="0" pageNumber="75">C. scaber</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<caption doi="10.3897/phytokeys.222.87779.figure19" httpUri="https://binary.pensoft.net/fig/828358" pageId="0" pageNumber="75" start="Figure 19" startId="F19">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 19.</emphasis>
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<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus rubineus</emphasis>
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</taxonomicName>
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D.Skinner & Maas
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<emphasis bold="true" pageId="0" pageNumber="75">A</emphasis>
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plant in habitat near Villa Rica, Pasco, Peru
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<emphasis bold="true" pageId="0" pageNumber="75">B</emphasis>
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inflorescence with flowers showing the labellum with characteristic lobulate apex (a)
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<emphasis bold="true" pageId="0" pageNumber="75">C</emphasis>
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adaxial leaf midrib showing absence of line of hairs
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<emphasis bold="true" pageId="0" pageNumber="75">D</emphasis>
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bract and flower detail from plant in habitat at Alto Churumazu, Pasco,
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<normalizedToken originalValue="Perú">Peru</normalizedToken>
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showing the full flower with subtending bract (flower), bract (br), bracteole subtending the capsule (ovary plus calyx) (brl/ca), stigma and style as attached to the young capsule (gyn), petals, single fertile stamen (st) and labellum (lab). Seeds from this plant were collected and cultivated as
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<emphasis italics="true" pageId="0" pageNumber="75">D.Skinner R3421</emphasis>
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<emphasis bold="true" pageId="0" pageNumber="75">E</emphasis>
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plant in habitat at 1980 meters near Chinchao,
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<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
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,
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<normalizedToken originalValue="Perú">Peru</normalizedToken>
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, the original collection site of "
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. ruber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="ruber">
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<emphasis italics="true" pageId="0" pageNumber="75">C. ruber</emphasis>
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</taxonomicName>
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" as indicated in
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<normalizedToken originalValue="Ruiz’">Ruiz'</normalizedToken>
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journal
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<emphasis bold="true" pageId="0" pageNumber="75">F</emphasis>
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inflorescence with lower bracts removed and single flower photographed in the wild near Chinchao,
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<normalizedToken originalValue="Perú">Peru</normalizedToken>
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. Photos
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<emphasis bold="true" pageId="0" pageNumber="75">A-F</emphasis>
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by Dave Skinner.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="materials_examined">
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<paragraph pageId="0" pageNumber="75">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<materialsCitation collectingDate="2005-03-17" collectorName="Rojas G., Vasquez Chavez, Francis" country="Peru" elevation="1961" location="Gramazu" specimenCount="1" stateProvince="Pasco" typeStatus="holotype">
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<collectingCountry name="Peru">Peru</collectingCountry>
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,
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<collectingRegion country="Peru" name="Pasco">Pasco</collectingRegion>
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: Prov. Oxapampa,
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<normalizedToken originalValue="Gramazú">
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<location LSID="urn:lsid:plazi:treatment:191887D029815EFAA87A8804961FE2FE:443DEFD8B9B467736573AC509ED6039E" country="Peru" name="Gramazu" stateProvince="Pasco">Gramazu</location>
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</normalizedToken>
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,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.9609999999999999" unit="m" value="1961.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.9609999999999999" unit="m" value="1961.0">1961 m</elevation>
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</quantity>
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,
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<collectingDate value="2005-03-17">17 Mar 2005</collectingDate>
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,
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<emphasis italics="true" pageId="0" pageNumber="75">
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<collectorName>Rojas G.</collectorName>
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</emphasis>
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,
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<emphasis italics="true" pageId="0" pageNumber="75">
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<collectorName>
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<normalizedToken originalValue="Vásquez">Vasquez</normalizedToken>
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<normalizedToken originalValue="Chávez">Chavez</normalizedToken>
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</collectorName>
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&
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<collectorName>Francis</collectorName>
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3574
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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L4196133; isotypes HUT, MO-2152325, USM)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="description">
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<paragraph pageId="0" pageNumber="75">Description.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Herb</emphasis>
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0.5-4 m tall.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Leaves</emphasis>
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sheaths 15-30 mm diam; ligule truncate to slightly lobed, 2-7 mm long; petiole 3-8 mm long; sheaths, ligule and petiole densely sericeous to glabrous; lamina narrowly elliptic, 20-37
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<normalizedToken originalValue="×">x</normalizedToken>
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5-13 cm, adaxially glabrous, abaxially rather densely villose to almost glabrous, base obtuse to slightly cordate, apex acuminate (acumen 10-15 mm long) to acute.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Inflorescence</emphasis>
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ovoid, 7-11
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<normalizedToken originalValue="×">x</normalizedToken>
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4-6 cm, terminating a leafy shoot; bracts, bracteole, and calyx rather densely puberulous to glabrous, ovary and capsule densely sericeous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Flowers</emphasis>
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abaxially oriented; bracts dark ruby red to red, coriaceous, broadly ovate, 3-4
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<normalizedToken originalValue="×">x</normalizedToken>
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2.5-4 cm, apex obtuse, callus 6-10 mm long; bracteole boat-shaped, 20-28 mm long; calyx red to pink, 9-15 mm long, lobes very shallowly triangular, 2-3 mm long; corolla pink to orange, 50-60 mm long, glabrous, lobes narrowly elliptic, 40-50 mm long; labellum yellow, distal part red, lateral lobes rolled inwards and forming a curved tube ca. 10 mm diam, oblong-obovate when spread out, 30-50
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<normalizedToken originalValue="×">x</normalizedToken>
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18-20 mm, 5-lobulate, lobules narrowly ovate-triangular, ca. 4 mm long; stamen yellow, 30-45
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<normalizedToken originalValue="×">x</normalizedToken>
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7-10 mm, not or slightly exceeding the labellum, apex red, irregularly dentate, anther 8-9 mm long
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Capsule</emphasis>
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obovoid, ca. 13 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="distribution">
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<paragraph pageId="0" pageNumber="75">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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Peru (
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<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
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, Pasco) (Fig.
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<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution maps for new species of Costaceae. Costus gibbosus - Costus whiskeycola m A Costus gibbosus D. Skinner & Maas B Costus mollissimus Maas & H. Maas C Costus obscurus D. Skinner & Maas D Costus oreophilus Maas & D. Skinner E Costus pitalito C. D. Specht & H. Maas F Costus prancei Maas & H. Maas G Costus pseudospiralis Maas & H. Maas H Costus rubineus D. Skinner & Maas I Costus whiskeycola Maas & H. Maas." figureDoi="10.3897/phytokeys.222.87779.figure22" httpUri="https://binary.pensoft.net/fig/828361" pageId="0" pageNumber="75">22H</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="habitat">
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<paragraph pageId="0" pageNumber="75">Habitat and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="75">In primary or secondary, premontane or montane forests, at elevations of (300-)1400-2200 m. Flowering year-round.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="etymology">
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<paragraph pageId="0" pageNumber="75">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus rubineus</emphasis>
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</taxonomicName>
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sp. nov. is named in reference to the Spanish explorer Hipolito Ruiz who visited in August 1780 the place called Chinchao in the Peruvian department of
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<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
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. In his journal, he mentioned the collection of a plant he referred to as "
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<taxonomicName class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus ruber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="ruber">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus ruber</emphasis>
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</taxonomicName>
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", but his collection was lost in a shipwreck off the coast of Portugal, and this
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<normalizedToken originalValue="species’">species'</normalizedToken>
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name was never again mentioned in the journal. Since there is already a
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. ruber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="ruber">
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<emphasis italics="true" pageId="0" pageNumber="75">C. ruber</emphasis>
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</taxonomicName>
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C.Wright ex Griseb. (a synonym of
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. pulverulentus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="pulverulentus">
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<emphasis italics="true" pageId="0" pageNumber="75">C. pulverulentus</emphasis>
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</taxonomicName>
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C.Presl), we cannot use that name but are using the very apt name of
|
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
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<emphasis italics="true" pageId="0" pageNumber="75">C. rubineus</emphasis>
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</taxonomicName>
|
||
to reflect the beautiful ruby-red colour of the bracts. The first author of this species visited the Chinchao region in 2016 and found at that locality specimen which he believed to be the same species as Ruiz mentioned in his journal (
|
||
<bibRefCitation author="Skinner, D" journalOrPublisher="Heliconia Society Bulletin" pageId="0" pageNumber="75" pagination="7 - 14" refId="B27" refString="Skinner, D, 2016. Following Ruiz. Heliconia Society Bulletin 22 (4): 7 - 14" title="Following Ruiz." volume="22" year="2016">Skinner 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="75" type="paratypes">
|
||
<paragraph pageId="0" pageNumber="75">Paratypes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="75">
|
||
<emphasis bold="true" pageId="0" pageNumber="75">
|
||
Peru.
|
||
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
|
||
</emphasis>
|
||
: Cordillera Azul, ca. 42.7 km E of Tingo Maria on road to Pucallpa, 5650 ft, 21 Nov 1979,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Davidson & Jones 9395</emphasis>
|
||
(U).
|
||
<emphasis bold="true" pageId="0" pageNumber="75">Pasco</emphasis>
|
||
: Prov. Oxapampa, Distr. Huancabamba, Zona de Amortiguamiento del Parque Nacional
|
||
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
|
||
, Sector alto tunqui, 1781 m, 10 Dec 2006,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Castillo et al. 642</emphasis>
|
||
(AMAZ, HUT, L, MO, USM); Prov. Oxapampa, Pichis Valley, new road between Santa Rose de Chivis and Puerto
|
||
<normalizedToken originalValue="Bermúdez">Bermudez</normalizedToken>
|
||
, 300-400 m, 30 Sept. 1982,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Foster 8950A</emphasis>
|
||
(MO); Prov. and Distr. Oxapampa, Carretera Oxapampa-Tsachopem, 1850 m, 15 Nov 2004,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Monteagudo Mendoza et al. 7658</emphasis>
|
||
(HUT, MO, U, USM); Prov. Oxapampa, Distr. Huancabamba, Carretera hacia Pozuzo-Zona de Amortiguamiento, Parque Nacional
|
||
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
|
||
, 1390-1630 m, 22 Mar 2005,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Ortiz V. et al. 521</emphasis>
|
||
(L, MO, USM); Prov. Oxapampa, 5 km SE of Oxapampa, Oswald
|
||
<normalizedToken originalValue="Müller">Mueller</normalizedToken>
|
||
property, 1850 m, 13-16 Dec 1982,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Smith 2946</emphasis>
|
||
(MO, U), idem, 24 Dec 1983,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Smith 5360</emphasis>
|
||
(HUT, MO, U, USM); Distr. Oxapampa, La Suiza, 10 Dec. 2002, 2200 m,
|
||
<emphasis italics="true" pageId="0" pageNumber="75">
|
||
<normalizedToken originalValue="Vásquez">Vasquez</normalizedToken>
|
||
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
|
||
& Rojas 27782
|
||
</emphasis>
|
||
(F, HUT, MO, U, USM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="75" type="notes">
|
||
<paragraph pageId="0" pageNumber="75">Notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="75">
|
||
<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Costus rubineus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is a species mostly restricted to relatively high elevations (except for
|
||
<emphasis italics="true" pageId="0" pageNumber="75">Foster 8950A</emphasis>
|
||
, which was found at 300 m) in the departments of Pasco, San
|
||
<normalizedToken originalValue="Martín">Martin</normalizedToken>
|
||
, and
|
||
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
|
||
, Peru. It looks, at first glance, somewhat similar to
|
||
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. spiralis" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="spiralis">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">C. spiralis</emphasis>
|
||
</taxonomicName>
|
||
(Jacq.) Roscoe, but it differs from that species by its abaxially oriented flowers and a very short ligule. The living plants look similar to
|
||
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. scaber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="scaber">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">C. scaber</emphasis>
|
||
</taxonomicName>
|
||
Ruiz & Pav., but that species differs by a much shorter calyx (3-7 mm vs. 9-15 mm long in
|
||
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">C. rubineus</emphasis>
|
||
</taxonomicName>
|
||
) and the presence of a row of erect hairs on the adaxially of the lamina (lacking in
|
||
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. rubineus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="rubineus">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">C. rubineus</emphasis>
|
||
</taxonomicName>
|
||
). There are often several flowers at the same time at anthesis, whereas in
|
||
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. scaber" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="scaber">
|
||
<emphasis italics="true" pageId="0" pageNumber="75">C. scaber</emphasis>
|
||
</taxonomicName>
|
||
, there is usually just one flower at anthesis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |