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<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.5.63435" ID-GBIF-Dataset="bfba5087-74d4-44e8-b783-94db25e3442a" ID-GBIF-Taxon="184941086" ID-Pensoft-Pub="2535-0730-2-193" ID-Pensoft-UUID="21773559522D5D9DA4B1A7DA51E4F2A6" ID-ZooBank="984E15D880E04B7DA84F92BB0AD4EA73" ModsDocID="2535-0730-5-2-193" checkinTime="1629846914602" checkinUser="pensoft" docAuthor="Zahiri, Reza, Nazari, Vazrick, Rajaei, Hossein, Wiemers, Martin, Fatahi, Maryam, Seidel, Matthias, Dalsgaard, Thure &amp; Husemann, Martin" docDate="2021" docId="0F1FEDD9DFC455BBB6E093ADBDACD9E8" docLanguage="en" docName="EvolutSyst 5(2): 193-261" docOrigin="Evolutionary Systematics 5 (2)" docPubDate="2021-08-20" docSource="http://dx.doi.org/10.3897/evolsyst.5.63435" docTitle="Melitaea tenuicula Verity 1919" docType="treatment" docVersion="3" id="21773559522D5D9DA4B1A7DA51E4F2A6" lastPageNumber="193" masterDocId="21773559522D5D9DA4B1A7DA51E4F2A6" masterDocTitle="? An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea" masterLastPageNumber="261" masterPageNumber="193" pageId="0" pageNumber="193" updateTime="1643429667741" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>? An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zahiri, Reza</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6274-6973</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany &amp; Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., K 1 A 0 C 6, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nazari, Vazrick</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9064-8959</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rajaei, Hossein</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3940-3734</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiemers, Martin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5272-3903</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fatahi, Maryam</mods:namePart>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Seidel, Matthias</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4913-8778</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dalsgaard, Thure</mods:namePart>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Husemann, Martin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5536-6681</mods:nameIdentifier>
<mods:affiliation>Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-20</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>5</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>193</mods:start>
<mods:end>261</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/evolsyst.5.63435</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.5.63435</mods:identifier>
<mods:identifier type="Pensoft-Pub">2535-0730-2-193</mods:identifier>
<mods:identifier type="ZooBank">984E15D880E04B7DA84F92BB0AD4EA73</mods:identifier>
<mods:identifier type="Pensoft-UUID">21773559522D5D9DA4B1A7DA51E4F2A6</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="184941086" LSID="urn:lsid:plazi:treatment:0F1FEDD9DFC455BBB6E093ADBDACD9E8" httpUri="http://treatment.plazi.org/id/0F1FEDD9DFC455BBB6E093ADBDACD9E8" lastPageNumber="193" pageId="0" pageNumber="193">
<subSubSection pageId="0" pageNumber="193" type="nomenclature">
<paragraph pageId="0" pageNumber="193">
?110.
<taxonomicName LSID="0F1FEDD9-DFC4-55BB-B6E0-93ADBDACD9E8" authority="Verity, 1919" authorityName="Verity" authorityYear="1919" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea tenuicula" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="tenuicula">Melitaea tenuicula Verity, 1919</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="original combination">
<paragraph pageId="0" pageNumber="193">Original combination.</paragraph>
<paragraph pageId="0" pageNumber="193">
&quot;
<taxonomicName baseAuthorityName="Rottemburg" baseAuthorityYear="1775" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea athalia" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="athalia">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea athalia</emphasis>
</taxonomicName>
race tenuicula mihi&quot; Verity, 1919 Ent. Rec. 31: 193-194.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="current combination">
<paragraph pageId="0" pageNumber="193">Current combination.</paragraph>
<paragraph pageId="0" pageNumber="193">
<emphasis bold="true" pageId="0" pageNumber="193">
<taxonomicName class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea athalia subsp. tenuicula" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="athalia" subSpecies="tenuicula">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea athalia tenuicula</emphasis>
</taxonomicName>
Verity, 1919
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="current status">
<paragraph pageId="0" pageNumber="193">Current status.</paragraph>
<paragraph pageId="0" pageNumber="193">Valid subspecies.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="materials_examined">
<paragraph pageId="0" pageNumber="193">Original material.</paragraph>
<paragraph pageId="0" pageNumber="193">
<materialsCitation collectingDate="1937-06-15" collectingDateMax="1937-06-28" collectingDateMin="1937-06-15" specimenCount="1" typeStatus="Type">
Labelled as
<normalizedToken originalValue="“Type”">
&quot;
<typeStatus>Type</typeStatus>
&quot;
</normalizedToken>
6?? (ZMH 833455-833460) (Fig.
<figureCitation captionStart="Plate 16" captionStartId="F16" captionText="Plate 16. 110. Melitaea athalia tenuicula Verity, 1919; 111. Melitaea varia varissima Verity, 1914; 112. Melitaea cinxia riloensis Zuellich, 1936; 113. Melitaea deione ab. mediofasciata Bubacek, 1926; 114. Melitaea diamina ab. johni Stauder, 1922; 115. Melitaea trivia balcanica Zuellich, 1936; 116. Melitaea trivia catapelioides Stauder, 1918; 117. Melitaea trivia ignasiti de Sagarra, 1926. a. Dorsal view, b. ventral view." httpUri="https://binary.pensoft.net/fig/579725" pageId="0" pageNumber="193">110</figureCitation>
). &quot;
<emphasis italics="true" pageId="0" pageNumber="193">tenuicula</emphasis>
Vrty&quot; // helvetica
<normalizedToken originalValue="Rühl”">Ruehl&quot;</normalizedToken>
// &quot;Marche / Bolognola /
<collectingDate value="1937-06-28">28 VI 1937</collectingDate>
/ Querci.&quot; // [blank label] // &quot;ZMH 833455&quot;; &quot;
<emphasis italics="true" pageId="0" pageNumber="193">tenuicula</emphasis>
Vrty&quot; // helvetica
<normalizedToken originalValue="Rühl”">Ruehl&quot;</normalizedToken>
// &quot;Marche / Bolognola /
<collectingDate value="1937-06-15">15 VI 1937</collectingDate>
/ Querci.&quot; // [blank label] // &quot;ZMH 833456&quot;; &quot;
<emphasis italics="true" pageId="0" pageNumber="193">tenuicula</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate value="1937-06-17">17 VI 1937</collectingDate>
/ Querci.&quot; // [blank label] // &quot;ZMH 833457&quot;; &quot;
<emphasis italics="true" pageId="0" pageNumber="193">tenuicula</emphasis>
Vrty&quot; // helvetica
<normalizedToken originalValue="Rühl”">Ruehl&quot;</normalizedToken>
// &quot;Marche / Bolognola /
<collectingDate value="1937-06-18">18 VI 1937</collectingDate>
/ Querci.&quot; // [blank label] // &quot;ZMH 833458&quot;; &quot;
<emphasis italics="true" pageId="0" pageNumber="193">tenuicula</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate value="1937-06-28">28 VI 1937</collectingDate>
/ Querci.&quot; // [blank label] // &quot;ZMH 833459&quot;; &quot;
<emphasis italics="true" pageId="0" pageNumber="193">tenuicula</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate value="1937-06-27">27 VI 1937</collectingDate>
/ Querci.&quot; // [blank label] // &quot;ZMH 833460
</materialsCitation>
&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="original locality">
<paragraph pageId="0" pageNumber="193">Original locality.</paragraph>
<paragraph pageId="0" pageNumber="193">Italy: &quot;Bolognola in the Sibillini Mountains, 1200 m&quot;.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="193" type="remarks">
<paragraph pageId="0" pageNumber="193">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="193">
<bibRefCitation author="Verity, R" journalOrPublisher="Entomologist's Record" pageId="0" pageNumber="193" pagination="193 - 201" refId="B97" refString="Verity, R, 1919. Seasonal Polymorphism and Races of Some European Grypocera and Rhopalocera. Entomologist's Record 31: 193 - 201" title="Seasonal Polymorphism and Races of Some European Grypocera and Rhopalocera." volume="31" year="1919">Verity (1919)</bibRefCitation>
proposed this name as a
<normalizedToken originalValue="“race”">&quot;race&quot;</normalizedToken>
of
<taxonomicName baseAuthorityName="Rottemburg" baseAuthorityYear="1775" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea athalia" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="athalia">
<emphasis italics="true" pageId="0" pageNumber="193">Melitaea athalia</emphasis>
</taxonomicName>
(Rottemburg, 1775). According to
<bibRefCitation author="Kudrna, O" journalOrPublisher="Journal of Research on the Lepidoptera" pageId="0" pageNumber="193" pagination="1 - 105" refId="B56" refString="Kudrna, O, 1983. An annotated catalogue of the butterflies named by Roger Verity. Journal of Research on the Lepidoptera 21 (1): 1 - 105" title="An annotated catalogue of the butterflies named by Roger Verity." volume="21" year="1983">Kudrna (1983</bibRefCitation>
: 83-84), this name is available as subspecific. The year on the specimen labels (1937) indicate specimens were collected 18 years after description (1919) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as
<normalizedToken originalValue="“types”">&quot;types&quot;</normalizedToken>
is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as &quot;Not a Type&quot; and are treated here as
<normalizedToken originalValue="“Non-type”">&quot;Non-type&quot;</normalizedToken>
specimens
</paragraph>
</subSubSection>
</treatment>
</document>
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