131 lines
12 KiB
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131 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZBK="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2018" docId="139897831E9A021F68721075000F3FAC" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina simbaiensis Shaverdo & Balke, sp. n." docType="treatment" docUuid="11CCE13F-F7DC-40A0-967D-C7F70EB423C2" docUuidSource="ZooBank" docVersion="7" lastPageNumber="38" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="37" updateTime="1668166558239" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>803</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>7</mods:start>
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<mods:end>70</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
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<mods:identifier type="ZBK">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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<mods:identifier type="ZooBank">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154125609" LSID="urn:lsid:zoobank.org:act:11CCE13F-F7DC-40A0-967D-C7F70EB423C2" httpUri="http://treatment.plazi.org/id/139897831E9A021F68721075000F3FAC" lastPageId="31" lastPageNumber="38" pageId="30" pageNumber="37">
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<subSubSection pageId="30" pageNumber="37" type="nomenclature">
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<paragraph pageId="30" pageNumber="37">
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21.
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<taxonomicName LSID="http://zoobank.org/11CCE13F-F7DC-40A0-967D-C7F70EB423C2" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina simbaiensis" order="Coleoptera" pageId="30" pageNumber="37" phylum="Arthropoda" rank="species" species="simbaiensis">Exocelina simbaiensis Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="30" pageNumber="37">sp. n.</taxonomicNameLabel>
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Figs 19, 43
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="37" type="reference_group">
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<paragraph pageId="30" pageNumber="37">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="30" pageNumber="37" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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undescribed sp. MB3315:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
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: supplementary figs 1-4, table 2;
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</paragraph>
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<paragraph pageId="30" pageNumber="37">
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary figs S1-S2, table S3.
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</paragraph>
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<paragraph pageId="30" pageNumber="37">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina simbaiensis" order="Coleoptera" pageId="30" pageNumber="37" phylum="Arthropoda" rank="species" species="simbaiensis">Exocelina simbaiensis</taxonomicName>
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_New_Guinea_MB3315:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary information S5-S6.
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</paragraph>
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<paragraph pageId="30" pageNumber="37">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina inengensis" order="Coleoptera" pageId="30" pageNumber="37" phylum="Arthropoda" rank="species" species="inengensis">Exocelina inengensis</taxonomicName>
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MB3309:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary figs S1-S2, table S3, and information S5-S6.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="37" type="type locality">
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<paragraph pageId="30" pageNumber="37">Type locality.</paragraph>
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<paragraph pageId="30" pageNumber="37">
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Papua New Guinea: Western Highlands Province, Simbai, Ineng River,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-5.249">05°14.94'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="144.547">144°32.82'E</geoCoordinate>
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, 2,000 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="37" type="type material">
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<paragraph pageId="30" pageNumber="37">Type material.</paragraph>
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<paragraph pageId="30" pageNumber="37">Holotype: male "Papua New Guinea: Western Highlands, Simbai, Ineng River, 2000m, 27.ii.2007, 05.14.943S 144.32.818E, Kinibel (PNG 135)" (ZSM). Paratypes: 4 males, 9 females with the same label as the holotype, one male with an additional green label "M.Balke 3309" (NHMW, ZSM). 1 male, 1 female "Papua New Guinea: Western Highlands, Simbai area, 2200m, 6.iii.2007, 05.18.752S 144.31.849E, Kinibel (PNG 149)", male additionally with "M.Balke 3316" [green] (ZSM). 2 females "Papua New Guinea: Western Highlands, Simbai area, 2500m, 8.iii.2007, 05.14.202S 144.33.651E, Kinibel (PNG 150)" (NHMW, ZSM). 1 female "M.Balke 3315" [green], "Papua New Guinea: Western Highlands, Jimi Valley, above Sendiap Station, 2000m, 6.iii.2007, 05.19.314S 144.31.266E, Kinibel (PNG 148)" (ZSM).</paragraph>
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</subSubSection>
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<subSubSection pageId="30" pageNumber="37" type="description">
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<paragraph pageId="30" pageNumber="37">Description.</paragraph>
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<paragraph pageId="30" pageNumber="37">Body size and form: Beetle small to medium-sized: TL-H 3.3-4.15 mm, TL 3.65-4.5 mm, MW 1.8-2.15 mm (holotype: TL-H 3.9 mm, TL 4.25 mm, MW 2.0 mm), with rather oblong habitus.</paragraph>
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<paragraph pageId="30" pageNumber="37">Coloration: Dark brown to piceous, sometimes with reddish pronotum. Head dark brown to piceous, paler anteriorly. Pronotum dark brown to piceous, with reddish brown lateral sides and sometimes anteriorly and narrowly posteriorly. Elytra dark brown to piceous, sometimes with narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish (Fig. 19).</paragraph>
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<paragraph pageId="30" pageNumber="37">Surface sculpture: Matt dorsally. Head with dense punctation (no spaces between punctures or spaces 2 times size of punctures), finer and sparser anteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum and elytra with slightly sparser and finer punctation than on head. Head, pronotum and elytra with strongly impressed microreticulation. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine, sparse but distinct punctation.</paragraph>
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<paragraph pageId="30" pageNumber="37">Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded.</paragraph>
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<paragraph pageId="30" pageNumber="37">
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Male: Antennomere simple, slightly stout (Fig. 19). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, slightly concave ventrally, with anterior band of ca 40 and irregular posterior row of 17 relatively long setae (Fig. 43D). Median lobe in lateral view slightly curved, its apex thickened, bent downwards; in ventral view, slightly broadened medially, with broadly pointed apex. Paramere very slightly concave on dorsal side and with dense setae on subdistal part; proximal setae inconspicuous (Fig. 43
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Abdominal ventrite 6 with 7-12 lateral striae on each side.
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</paragraph>
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<paragraph pageId="30" pageNumber="37">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
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<paragraph pageId="30" pageNumber="37">Variability. The specimens from Ineng River are larger: TL-H 3.5-4.15 mm; remaining specimens: TL-H 3.3-3.7 mm. In addition, the population from Ineng River shows variability in how strongly apex of the median lobe bent downwards in lateral view.</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="38" type="affinities">
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<paragraph pageId="31" pageNumber="38">
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<pageBreakToken pageId="31" pageNumber="38" start="start">Affinities</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="31" pageNumber="38">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina simbaiensis" order="Coleoptera" pageId="31" pageNumber="38" phylum="Arthropoda" rank="species" species="simbaiensis">Exocelina simbaiensis</taxonomicName>
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sp. n. is similar to
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<taxonomicName lsidName="E. yoginofi" pageId="31" pageNumber="38" rank="species" species="yoginofi">E. yoginofi</taxonomicName>
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sp. n. in body form, size, and coloration, but differs from it in distinctly stronger punctation and microreticulation and in thickened apex of the median lobe.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="38" type="distribution">
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<paragraph pageId="31" pageNumber="38">Distribution.</paragraph>
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<paragraph pageId="31" pageNumber="38">Papua New Guinea: Western Highlands Province, near Simbai (Fig. 50).</paragraph>
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</subSubSection>
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<subSubSection pageId="31" pageNumber="38" type="etymology">
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<paragraph pageId="31" pageNumber="38">Etymology.</paragraph>
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<paragraph pageId="31" pageNumber="38">The species is named after Simbai area. The name is an adjective in the nominative singular.</paragraph>
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</subSubSection>
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</treatment>
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</document> |