140 lines
15 KiB
XML
140 lines
15 KiB
XML
<document id="4737949BE2929100069AE1B48E8F4A80" ID-DOI="10.5281/zenodo.213326" ID-GBIF-Dataset="0d399975-ca69-4a60-ad1f-e967e4b4f3ea" ID-ISSN="1175-5326" ID-Zenodo-Dep="213326" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460551797102" checkinUser="plazi" docAuthor="Martino, Emanuela Di & Taylor, Paul D" docDate="2012" docId="1373087CFFD8FFA4C6A396A72545F79C" docLanguage="en" docName="zt03534p020.pdf" docOrigin="Zootaxa 3534" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pyrisinellidae" docType="treatment" docVersion="8" lastPageNumber="3" masterDocId="EF4A7004FFDAFFA6C634977F2000F27C" masterDocTitle="Pyrisinellidae, a new family of anascan cheilostome bryozoans" masterLastPageNumber="20" masterPageNumber="1" pageNumber="3" updateTime="1698318438355" updateUser="plazi">
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<mods:mods id="D48208A9652D09F9F08270076EE72799" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CD87ED85F39B7FD1F996601BA0FDE49C">
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<mods:title id="BC2E4B6916843F16F8F024168B5EAEC5">Pyrisinellidae, a new family of anascan cheilostome bryozoans</mods:title>
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<mods:name id="79D5C468A1C01DA1A87150027604F4EC" type="personal">
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<mods:roleTerm id="EDD549FBA74842FBB729DFEB70112C26">Author</mods:roleTerm>
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<mods:namePart id="F06BA6A08F326DDB86CDA829EE0FA92B">Martino, Emanuela Di</mods:namePart>
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<mods:name id="0B644F6337C50EFDE7E906D406B4C4D9" type="personal">
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<mods:roleTerm id="06AA9F5C8E0FE503C01FC110805721EF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8AD9BB91CFF47F0225D1BB6D2EF0311A">Taylor, Paul D</mods:namePart>
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<mods:typeOfResource id="7904299D19225020E7B17FA9C62A4D4F">text</mods:typeOfResource>
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<mods:title id="1E2C6CB2AE2F3295445342980AC54051">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C1FD31195C69BEE86A5E9B9D1FDF5A4F">
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<mods:date id="D5C0B12A46042A5F559AA306F88A070C">2012</mods:date>
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<mods:detail id="7F86011BE0C205798B39C44CFA0A5C6E" type="volume">
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<mods:number id="69B2AB2FBA700773263345A0874C2424">3534</mods:number>
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<mods:start id="D1297EEE0703A71E72A88FEBC1F2B051">1</mods:start>
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<mods:identifier id="E6251D91E8CBB6DE7FEFE61CE49E1B6B" type="DOI">10.5281/zenodo.213326</mods:identifier>
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<mods:identifier id="4B33E97B6256944BCDC3F11115758ADA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="041F6EFECD053F093B743A744FF22DE2" type="Zenodo-Dep">213326</mods:identifier>
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<treatment id="1373087CFFD8FFA4C6A396A72545F79C" ID-DOI="http://doi.org/10.5281/zenodo.5623583" ID-GBIF-Taxon="119530381" ID-Zenodo-Dep="5623583" LSID="urn:lsid:plazi:treatment:1373087CFFD8FFA4C6A396A72545F79C" httpUri="http://treatment.plazi.org/id/1373087CFFD8FFA4C6A396A72545F79C" lastPageNumber="3" pageId="2" pageNumber="3">
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<subSubSection id="D3C0EAE1FFD8FFA4C6A396A7221BF38E" box="[151,539,472,498]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="9B65B96AFFD8FFA4C6A396A7221BF38E" blockId="2.[151,539,472,498]" box="[151,539,472,498]" pageId="2" pageNumber="3">
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<heading id="C02D0E06FFD8FFA4C6A396A7221BF38E" bold="true" box="[151,539,472,498]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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<emphasis id="A9AE6578FFD8FFA4C6A396A7221BF38E" bold="true" box="[151,539,472,498]" pageId="2" pageNumber="3">
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Family
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<taxonomicName id="5CDAC2E9FFD8FFA4C6C296A721A1F38E" ID-CoL="858BN" box="[246,417,472,498]" class="Gymnolaemata" family="Pyrisinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="family">Pyrisinellidae</taxonomicName>
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fam. nov.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3C0EAE1FFD8FFA4C6A3956024A0F12B" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="9B65B96AFFD8FFA4C6A3956024A0F12B" blockId="2.[151,1437,543,1504]" pageId="2" pageNumber="3">
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<emphasis id="A9AE6578FFD8FFA4C6A395602111F044" bold="true" box="[151,273,543,568]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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Colony encrusting and multiserial. Autozooids distinct, oval, separated by deep furrows. Gymnocyst narrow, visible around entire circumference of zooid, often enlarged proximally. Cryptocyst extensively developed, depressed, flat and granular. Frontal surface of autozooids marked by pear-shaped ridge formed proximally and laterally by mural rim and distally by distal rim of opesia. Opesiules absent or present but few in number and small. Orifice trifoliate or semielliptical. Oral spines present, closely spaced, forming an arch distally around the orifice. Cryptocystal closure plates often developed. Ovicell hyperstomial, resting on distal zooid and indenting its mural rim. Growing edge stepped, revealing windows of pore chambers, that of the distal pore chamber the largest, ovoidal and facing frontally. Avicularia absent or present and variously adventitious, interzooidal or vicarious; rostrum acuminate or rounded; pivotal bar entirely calcified or represented by two condyles.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3C0EAE1FFD8FFA4C6F3941B2246F100" box="[199,582,868,892]" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="9B65B96AFFD8FFA4C6F3941B2246F100" blockId="2.[151,1437,543,1504]" box="[199,582,868,892]" pageId="2" pageNumber="3">
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<emphasis id="A9AE6578FFD8FFA4C6F3941B2154F100" bold="true" box="[199,340,868,892]" pageId="2" pageNumber="3">
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<typeStatus id="446107C8FFD8FFA4C6F3941B2102F100" box="[199,258,868,892]" pageId="2" pageNumber="3">Type</typeStatus>
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genus.
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</emphasis>
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<taxonomicName id="5CDAC2E9FFD8FFA4C76F941B21D6F100" box="[347,470,868,892]" class="Gymnolaemata" family="Pyrisinellidae" genus="Pyrisinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="genus" status="gen. nov.">
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<emphasis id="A9AE6578FFD8FFA4C76F941B21D6F100" box="[347,470,868,892]" italics="true" pageId="2" pageNumber="3">Pyrisinella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B29DD803FFD8FFA4C7E9941B2246F100" box="[477,582,868,892]" pageId="2" pageNumber="3" rank="genus">
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<emphasis id="A9AE6578FFD8FFA4C7E9941B2246F100" bold="true" box="[477,582,868,892]" pageId="2" pageNumber="3">gen. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3C0EAE1FFD8FFA4C6F394F72170F750" pageId="2" pageNumber="3" type="discussion">
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<paragraph id="9B65B96AFFD8FFA4C6F394F72170F750" blockId="2.[151,1437,543,1504]" pageId="2" pageNumber="3">
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<emphasis id="A9AE6578FFD8FFA4C6F394F7213BF1DC" bold="true" box="[199,315,904,928]" pageId="2" pageNumber="3">Remarks.</emphasis>
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This new family is erected to accommodate two new genera (
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<taxonomicName id="5CDAC2E9FFD8FFA4C23594F7247CF1DC" box="[1025,1148,904,928]" class="Gymnolaemata" family="Pyrisinellidae" genus="Pyrisinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="4" phylum="Bryozoa" rank="genus" status="gen. nov.">
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<emphasis id="A9AE6578FFD8FFA4C23594F7247CF1DC" box="[1025,1148,904,928]" italics="true" pageId="2" pageNumber="3">Pyrisinella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="5CDAC2E9FFD8FFA4C28094F72538F1DC" box="[1204,1336,904,928]" class="Gymnolaemata" family="Pyrisinellidae" genus="Spinisinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="6" phylum="Bryozoa" rank="genus" status="gen. nov.">
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<emphasis id="A9AE6578FFD8FFA4C28094F72538F1DC" box="[1204,1336,904,928]" italics="true" pageId="2" pageNumber="3">Spinisinella</emphasis>
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</taxonomicName>
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) plus an existing genus (
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<emphasis id="A9AE6578FFD8FFA4C77F94D321C6F1B8" box="[331,454,940,964]" italics="true" pageId="2" pageNumber="3">Setosinella</emphasis>
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) which share a sufficient numbers of skeletal morphological characters to suggest that they represent a clade of anascan-grade neocheilostomes distinguishable from other ‘calloporids’. In all three genera, the small autozooids have a salient mural rim extending to the proximolateral corners of the opesia and joining with the opesial rim to form a pear-shaped ridge around the cryptocyst and opesia, an arch of oral spines over the orifice, and a prominent hyperstomial ovicell. A fourth genus,
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<taxonomicName id="5CDAC2E9FFD8FFA4C5C29342246BF628" box="[1014,1131,1085,1108]" class="Gymnolaemata" family="Calloporidae" genus="Megapora" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="genus">
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<emphasis id="A9AE6578FFD8FFA4C5C29342246BF628" box="[1014,1131,1085,1108]" italics="true" pageId="2" pageNumber="3">Megapora</emphasis>
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</taxonomicName>
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, is tentatively referred to
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<taxonomicName id="5CDAC2E9FFD8FFA4C6A393202134F60B" box="[151,308,1119,1143]" class="Gymnolaemata" family="Pyrisinellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="family">Pyrisinellidae</taxonomicName>
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, although it differs from the other three genera in that the proximal part of the pear-shaped ridge is located within the cryptocyst rather than representing a true mural rim marking the boundary between the cryptocystal and gymnocystal components of the frontal wall. Although a few genera of
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<taxonomicName id="5CDAC2E9FFD8FFA4C24B93D8259CF6C3" authority="Gray, 1848" authorityName="Gray" authorityYear="1848" box="[1151,1436,1191,1215]" class="Gymnolaemata" family="Microporidae" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="family">
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Microporidae
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<bibRefCitation id="FF4BC49BFFD8FFA4C32B93D8259CF6C3" author="Gray" box="[1311,1436,1191,1215]" pageId="2" pageNumber="3" refString="Gray, J. E. (1848) List of the specimens of British animals in the collections of the British Museum. Part 1. Centrionae or radiated animals. Trustees of the British Museum (Natural History), London, 173 pp. [Polyzoa pp. 91 - 151]." type="book chapter" year="1848">Gray, 1848</bibRefCitation>
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</taxonomicName>
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possess a similar pear-shaped ridge, they differ from pyrisinellids in one or more of the following characters: immersed ovicells, lack of gymnocyst, and presence of connecting tubes between the zooids (as in
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<taxonomicName id="5CDAC2E9FFD8FFA4C365938F2163F750" authority="Lamouroux, 1816" authorityName="Lamouroux" authorityYear="1816" class="Gymnolaemata" family="Microporidae" genus="Mollia" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="genus">
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<emphasis id="A9AE6578FFD8FFA4C365938F259BF774" box="[1361,1435,1264,1288]" italics="true" pageId="2" pageNumber="3">Mollia</emphasis>
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Lamouroux, 1816
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D3C0EAE1FFD8FFA4C6F392472545F79C" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="9B65B96AFFD8FFA4C6F392472264F7C0" blockId="2.[151,1437,543,1504]" pageId="2" pageNumber="3">
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<taxonomicName id="5CDAC2E9FFD8FFA4C6F392472142F72C" ID-CoL="85HSD" box="[199,322,1336,1360]" class="Gymnolaemata" family="Pyrisinellidae" genus="Pyrisinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="4" phylum="Bryozoa" rank="genus" status="gen. nov.">
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<emphasis id="A9AE6578FFD8FFA4C6F392472142F72C" box="[199,322,1336,1360]" italics="true" pageId="2" pageNumber="3">Pyrisinella</emphasis>
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</taxonomicName>
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is chosen as the
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<typeStatus id="446107C8FFD8FFA4C4289246224CF72D" box="[540,588,1337,1361]" pageId="2" pageNumber="3">type</typeStatus>
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genus of the new family for the following reasons: (1)
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<taxonomicName id="5CDAC2E9FFD8FFA4C2CC9247257DF72C" ID-CoL="85JB6" box="[1272,1405,1336,1360]" class="Gymnolaemata" family="Pyrisinellidae" genus="Spinisinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="6" phylum="Bryozoa" rank="genus" status="gen. nov.">
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<emphasis id="A9AE6578FFD8FFA4C2CC9247257DF72C" box="[1272,1405,1336,1360]" italics="true" pageId="2" pageNumber="3">Spinisinella</emphasis>
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</taxonomicName>
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is represented by only a single small colony whereas
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<taxonomicName id="5CDAC2E9FFD8FFA4C4E392232352F708" ID-CoL="85HSD" box="[727,850,1372,1396]" class="Gymnolaemata" family="Pyrisinellidae" genus="Pyrisinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="4" phylum="Bryozoa" rank="genus" status="gen. nov.">
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<emphasis id="A9AE6578FFD8FFA4C4E392232352F708" box="[727,850,1372,1396]" italics="true" pageId="2" pageNumber="3">Pyrisinella</emphasis>
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</taxonomicName>
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is abundant; (2) using
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<emphasis id="A9AE6578FFD8FFA4C263922324D2F708" box="[1111,1234,1372,1396]" italics="true" pageId="2" pageNumber="3">Setosinella</emphasis>
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as the
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<typeStatus id="446107C8FFD8FFA4C31792222553F709" box="[1315,1363,1373,1397]" pageId="2" pageNumber="3">type</typeStatus>
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genus would result in a family name confusingly similar to
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<taxonomicName id="5CDAC2E9FFD8FFA4C52092FF245BF7E4" ID-CoL="84KRX" authority="Levinsen, 1909" authorityName="Levinsen" authorityYear="1909" box="[788,1115,1408,1432]" class="Gymnolaemata" family="Setosellidae" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="family">
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Setosellidae
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<bibRefCitation id="FF4BC49BFFD8FFA4C59392FF245BF7E4" author="Levinsen" box="[935,1115,1408,1432]" pageId="2" pageNumber="3" refString="Levinsen, G. M. R. (1909) Morphological and systematic studies on the cheilostomatous Bryozoa. Nationale Forfatterers Forlag, Copenhagen, 431 pp." type="book" year="1909">Levinsen, 1909</bibRefCitation>
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</taxonomicName>
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; and (3)
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<taxonomicName id="5CDAC2E9FFD8FFA4C2F892FE2541F7E4" ID-CoL="84Q4Q" box="[1228,1345,1409,1432]" class="Gymnolaemata" family="Calloporidae" genus="Megapora" kingdom="Animalia" order="Cheilostomatida" pageId="2" pageNumber="3" phylum="Bryozoa" rank="genus">
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<emphasis id="A9AE6578FFD8FFA4C2F892FE2541F7E4" box="[1228,1345,1409,1432]" italics="true" pageId="2" pageNumber="3">Megapora</emphasis>
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</taxonomicName>
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is only provisionally assigned to the new family.
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</paragraph>
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<paragraph id="9B65B96AFFD8FFA4C6F392B82545F79C" blockId="2.[151,1437,543,1504]" box="[199,1349,1479,1504]" pageId="2" pageNumber="3">
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<emphasis id="A9AE6578FFD8FFA4C6F392B82217F79C" bold="true" box="[199,535,1479,1504]" pageId="2" pageNumber="3">Stratigraphical distribution.</emphasis>
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Cretaceous (Cenomanian or Turonian) to Miocene (Langhian),?Recent.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |