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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.198.79514" ID-Pensoft-Pub="1314-2003-198-1" ID-Pensoft-UUID="486F1F1B4F5854D2831AAA341B9A322C" ModsDocID="1314-2003-198-1" checkinTime="1654168732655" checkinUser="pensoft" docAuthor="Aubriot, Xavier &amp; Knapp, Sandra" docDate="2022" docId="E185CA406919B28C42E92C27E13F6C6C" docLanguage="en" docName="PhytoKeys 198: 1-270" docOrigin="PhytoKeys 198" docPubDate="2022-06-01" docSource="http://dx.doi.org/10.3897/phytokeys.198.79514" docTitle="Solanum deflexicarpum C. Y. Wu &amp; S. C. Huang, Acta Phytotax. Sin. 16 (2): 73. 1978." docType="treatment" docVersion="1" id="486F1F1B4F5854D2831AAA341B9A322C" lastPageNumber="1" masterDocId="486F1F1B4F5854D2831AAA341B9A322C" masterDocTitle="A revision of the &quot; spiny solanums &quot; of Tropical Asia (Solanum, the Leptostemonum Clade, Solanaceae)" masterLastPageNumber="270" masterPageNumber="1" pageNumber="1" updateTime="1654168732655" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>A revision of the &quot; spiny solanums &quot; of Tropical Asia (Solanum, the Leptostemonum Clade, Solanaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aubriot, Xavier</mods:namePart>
<mods:affiliation>Universite Paris-Saclay, CNRS, AgroParisTech, Ecologie Systematique et Evolution, 91190, Gif-sur-Yvette, France &amp; The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7698-3945</mods:nameIdentifier>
<mods:affiliation>The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>198</mods:number>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.198.79514</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.198.79514</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-198-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">486F1F1B4F5854D2831AAA341B9A322C</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:E185CA406919B28C42E92C27E13F6C6C" httpUri="http://treatment.plazi.org/id/E185CA406919B28C42E92C27E13F6C6C" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
11.
<taxonomicName authority="C. Y. Wu &amp; S. C. Huang, Acta Phytotax. Sin. 16 (2): 73. 1978." authorityName="C. Y. Wu &amp; S. C. Huang, Acta Phytotax. Sin. 16 (2): 73. 1978." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexicarpum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexicarpum">Solanum deflexicarpum C.Y.Wu &amp; S.C.Huang, Acta Phytotax. Sin. 16(2): 73. 1978.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Solanum deflexicarpum C. Y. Wu &amp; S. C. Huang A herbarium specimen collected in China in 2007 (Knapp et al. 10130, BM 001019378) B detail of a fertile branch (Knapp et al. 10130, China) C detail view of a flower (Knapp et al. 10130, China) D detail view of an infructescence (Knapp et al. 10130, China). Photograph credits: A CC-BY, © copyright The Trustees of the Natural History Museum, London B-D S. Knapp." figureDoi="10.3897/phytokeys.198.79514.figure18" httpUri="https://binary.pensoft.net/fig/689711" pageId="0" pageNumber="1">Fig. 18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum indicum subsp. var. var. recurvatum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="indicum" subSpecies="var." variety="recurvatum">Solanum indicum L. var. recurvatum</taxonomicName>
C.Y.Wu &amp; S.C.Huang, Acta Phytotax. Sin. 16(2): 73. 1978. Type. China. Yunnan: Dali, &quot;Bohai city [transl. from the label]&quot;, 21 Feb 1957,
<emphasis italics="true" pageId="0" pageNumber="1">Yunnan expedition 5005</emphasis>
(holotype: KUN [KUN184304]; isotypes: IBK [IBK00241973], IBSC [IBSC0004706], PE [PE00031389, PE02079736], SZ [SZ00254985, SZ00254997]).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1947-08-29" collectorName="G. Feng" country="China" location="Xishou" municipality="Xiangping Mountain" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<collectingCountry name="China">China</collectingCountry>
.
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
: &quot;
<collectingMunicipality>Xiangping Mountain</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E185CA406919B28C42E92C27E13F6C6C:A084CAF03A8AB98125804D67135730D2" country="China" municipality="Xiangping Mountain" name="Xishou" stateProvince="Yunnan">Xishou</location>
[transl. from protologue]&quot;,
<collectingDate value="1947-08-29">29 Aug 1947</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>G. Feng</collectorName>
11429
</emphasis>
(
<typeStatus>holotype</typeStatus>
: KUN [KUN484279]; isotypes: A [A00310057], PE [PE00730648], WUK [WUK0194818])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shrubs to 2 m tall, unarmed or rarely sparsely armed. Stems erect, terete, moderately stellate-pubescent; prickles, if present, to 5 mm long, to 3 mm wide at the base, broad-based, curved, pale yellowish tan, sparsely stellate-pubescent near the base; pubescence of sessile to very short-stalked porrect-stellate trichomes, the rays 5-7, 0.1-0.4 mm long, the midpoints absent or to 0.5 mm long; new growth densely stellate-pubescent, the trichomes white and tangled, soon deciduous and the stems glabrate; bark of older stems greyish brown to grey. Sympodial units difoliate, the leaves geminate, or not geminate, the leaves of a pair equal in size and shape. Leaves simple, shallowly lobed, the blades 5-11 cm long, 2.5-7.5 cm wide, ca. 2 times longer than wide, elliptic, chartaceous, discolorous, unarmed or sparsely armed along the midrib and major veins with small curved prickles; adaxial surface evenly and sparsely to moderately stellate-pubescent with white-cream sessile porrect trichomes, the rays 4-6, 0.1-0.2 mm long, the midpoints 0.5-1.3 mm long; abaxial surface with similar sessile porrect-stellate trichomes, but with some short-stalked stellate trichomes with stalks to 0.5 mm and longer rays to 0.5 mm long; major veins 3-4 pairs, densely pubescent especially abaxially; base abruptly truncate, somewhat oblique; margins shallowly lobed, the lobes 3-4 on each side, 0.3-1 cm long, broadly deltate, apically rounded, the sinuses less than halfway to the midrib; apex acute; petioles 1-2 cm long, 1/4-1/2 as long as the leaf blades, sparsely prickly and moderately pubescent, the prickles 0-3, like those of the stems, the pubescence of sessile stellate-trichomes like those of the stems. Inflorescences 1-3 cm long, internodal and lateral, unbranched, with 5-10 flowers, only 1 or 2 flowers open at any one time, pubescent with sessile stellate-porrect trichomes like those of the stems, with 4-6 rays ca. 0.2 mm long and midpoints to 0.5 mm long, unarmed or with a few small prickles to 3 mm long; peduncle 0.1-0.8 cm long, unarmed; pedicels 0.35-0.5 cm long, 0.5-1 mm in diameter at the base, ca. 1 mm in diameter at the apex, spreading and slightly nodding at anthesis, unarmed or with a few small prickles to 1 mm long, densely stellate-pubescent like the inflorescence axes, articulated at the base; pedicel scars irregularly spaced 3-4 mm apart. Buds ellipsoid and somewhat tapering, strongly exserted from the calyx before anthesis. Flowers 5-merous, apparently all perfect, but some distal flowers may be short-styled. Calyx with the tube 2-2.5 mm long, conical, the lobes 1-2 mm long, ca. 1 mm wide, deltate, unarmed or with a few small prickles to 3 mm long and densely stellate-pubescent abaxially with sessile porrect-stellate trichomes like those of the pedicels. Corolla 0.8-1 cm in diameter, white, deeply stellate, lobed 3/4 of the way to the base, interpetalar tissue present, the lobes 4-4.5 mm long, 2-2.5 mm wide, deltate, spreading at anthesis, mostly glabrous adaxially but with a few stellate trichomes at the tips, densely stellate-pubescent abaxially with densely tangled sessile trichomes where exposed in bud, these densest at the tips. Stamens equal; anthers 3-4 mm long, ca. 2 mm wide, slightly tapering, yellow, glabrous, poricidal at the tips, the pores directed distally, not elongating to slits with drying; filament tube minute, glabrous; free portion of the filaments ca. 0.5 mm long, glabrous. Ovary conical, glabrous; style 5-6 mm long, glabrous; stigma capitate, the surfaces minutely papillose, bright green in live plants. Fruit a globose berry, several to many per infructescence, 1-1.2 cm in diameter, orange-red when ripe, the pericarp thin and shiny, glabrous; fruiting pedicels 0.8-1 cm long, 1-1.2 mm in diameter at the base, ca. 2 mm in diameter at the apex, somewhat woody, sharply deflexed and nodding, unarmed or with small prickles; fruiting calyx not accrescent, the lobes often breaking off. Seeds 50-60 per berry, 2-3 mm long, 2-3 mm wide, flattened reniform, pale tan or yellowish brown, the surfaces minutely pitted, the testal cells with sinuate margins. Chromosome number: not known.</paragraph>
<caption doi="10.3897/phytokeys.198.79514.figure18" httpUri="https://binary.pensoft.net/fig/689711" pageId="0" pageNumber="1" start="Figure 18" startId="F18">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexicarpum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum deflexicarpum</emphasis>
</taxonomicName>
C.Y.Wu &amp; S.C.Huang
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
herbarium specimen collected in China in 2007 (
<emphasis italics="true" pageId="0" pageNumber="1">Knapp et al. 10130</emphasis>
, BM001019378)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
detail of a fertile branch (
<emphasis italics="true" pageId="0" pageNumber="1">Knapp et al. 10130</emphasis>
, China)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
detail view of a flower (
<emphasis italics="true" pageId="0" pageNumber="1">Knapp et al. 10130</emphasis>
, China)
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
detail view of an infructescence (
<emphasis italics="true" pageId="0" pageNumber="1">Knapp et al. 10130</emphasis>
, China). Photograph credits:
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
CC-BY, © copyright The Trustees of the Natural History Museum, London
<emphasis bold="true" pageId="0" pageNumber="1">B-D</emphasis>
S. Knapp.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of S. deflexicarpum." figureDoi="10.3897/phytokeys.198.79514.figure19" httpUri="https://binary.pensoft.net/fig/689712" pageId="0" pageNumber="1">19</figureCitation>
).
</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexicarpum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum deflexicarpum</emphasis>
</taxonomicName>
is endemic to China (Yunnan province) and only known from few collections; potentially also present in northeastern Myanmar (
<emphasis italics="true" pageId="0" pageNumber="1">Andrew s.n.</emphasis>
collected in
<normalizedToken originalValue="Poneshee">'Poneshee'</normalizedToken>
close to the boarder with Myanmar).
</paragraph>
<caption doi="10.3897/phytokeys.198.79514.figure19" httpUri="https://binary.pensoft.net/fig/689712" pageId="0" pageNumber="1" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 19.</emphasis>
Distribution of
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. deflexicarpum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">S. deflexicarpum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexicarpum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum deflexicarpum</emphasis>
</taxonomicName>
grows in open vegetation and roadsides in semideciduous tropical forests, from 1,255 to 1,500 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names and uses">
<paragraph pageId="0" pageNumber="1">Common names and uses.</paragraph>
<paragraph pageId="0" pageNumber="1">
China. ku ci (
<bibRefCitation author="Zhang, Z-Y" journalOrPublisher="National Taiwan University" pageId="0" pageNumber="1" refId="B280" refString="Zhang, Z-Y, Lu, A, D'Arcy, WG, 1994. Solanaceae. In: Wu ZY, Raven PH (Eds) Flora of China. Vol. 17 (Verbenaceae through Solanaceae). Science Press &amp; Missouri Botanical Garden Press, Beijing &amp; St. Louis." title="Solanaceae. In: Wu ZY, Raven PH (Eds) Flora of China. Vol. 17 (Verbenaceae through Solanaceae). Science Press &amp; Missouri Botanical Garden Press, Beijing &amp; St. Louis." year="1994">Zhang et al. 1994</bibRefCitation>
); Yunnan: ku ci, wan bing ci tian qie (pedicel curved and spiny, high altitude) [Mandarin] (
<emphasis italics="true" pageId="0" pageNumber="1">Feng 11429</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(
<bibRefCitation author="Hutchings, A" journalOrPublisher="AGRIVITA Journal of Agricultural Science" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B105" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
).
</emphasis>
Near Threatened (NT). EOO (65,755 km2); AOO (16 km2).
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexicarpum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum deflexicarpum</emphasis>
</taxonomicName>
occurs over a relatively wide range, but there are very few collections; the paucity of collections suggest that the species is of some conservation concern, meriting further study.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexicarpum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum deflexicarpum</emphasis>
</taxonomicName>
is morphologically similar to the sympatric and very widespread
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. violaceum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="violaceum">
<emphasis italics="true" pageId="0" pageNumber="1">S. violaceum</emphasis>
</taxonomicName>
and to the Indian endemics
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. hovei" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hovei">
<emphasis italics="true" pageId="0" pageNumber="1">S. hovei</emphasis>
</taxonomicName>
and
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. multiflorum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="multiflorum">
<emphasis italics="true" pageId="0" pageNumber="1">S. multiflorum</emphasis>
</taxonomicName>
; in the analyses of
<bibRefCitation DOI="https://doi.org/10.1111/boj.12412" author="Aubriot, X" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="199 - 223" refId="B8" refString="Aubriot, X, Singh, P, Knapp, S, 2016a. Tropical Asian species show the Old World clade of &quot;spiny solanums&quot; (subgenus Leptostemonum pro parte: Solanaceae) is not monophyletic. Botanical Journal of the Linnean Society 181 (2): 199 - 223, DOI: https://doi.org/10.1111/boj.12412" title="Tropical Asian species show the Old World clade of &quot; spiny solanums &quot; (subgenus Leptostemonum pro parte: Solanaceae) is not monophyletic." url="https://doi.org/10.1111/boj.12412" volume="181" year="2016 a">Aubriot et al. (2016a)</bibRefCitation>
it is part of a monophyletic group with those species. It differs from
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. violaceum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="violaceum">
<emphasis italics="true" pageId="0" pageNumber="1">S. violaceum</emphasis>
</taxonomicName>
in its condensed inflorescences with white rather than violet flowers and strongly down-curved fruiting pedicels; the fruiting pedicels of
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. violaceum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="violaceum">
<emphasis italics="true" pageId="0" pageNumber="1">S. violaceum</emphasis>
</taxonomicName>
are more spaced, straight and are strongly spreading from the axis.
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum deflexiflorum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="deflexiflorum">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum deflexiflorum</emphasis>
</taxonomicName>
differs from
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. hovei" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hovei">
<emphasis italics="true" pageId="0" pageNumber="1">S. hovei</emphasis>
</taxonomicName>
in its recurved fruiting pedicels (straight in
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. hovei" order="Solanales" pageId="0" pageNumber="1" rank="species" species="hovei">
<emphasis italics="true" pageId="0" pageNumber="1">S. hovei</emphasis>
</taxonomicName>
) and truncate, oblique (versus attenuate) leaf bases. The inflorescences of
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. multiflorum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="multiflorum">
<emphasis italics="true" pageId="0" pageNumber="1">S. multiflorum</emphasis>
</taxonomicName>
are similarly condensed, but
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. deflexicarpum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="deflexicarpum">
<emphasis italics="true" pageId="0" pageNumber="1">S. deflexicarpum</emphasis>
</taxonomicName>
differs from that species in its more compact trichomes, fewer flowers per inflorescence, and its distribution in southern China.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">See Suppl. materials 1-3.</paragraph>
</subSubSection>
</treatment>
</document>