171 lines
21 KiB
XML
171 lines
21 KiB
XML
<document id="F97C487458FAD07A1DC597B123AA156F" ID-DOI="10.11646/zootaxa.5316.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8154005" ID-ZooBank="BC922E16-2614-4F3D-AD82-87A845DE7E2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689581758737" checkinUser="plazi" docAuthor="Zwick, Peter & Zwick, Andreas" docDate="2023" docId="E12C876C4A25FFD4FF4FF99DFC360EF4" docLanguage="en" docName="zootaxa.5316.1.1.pdf" docOrigin="Zootaxa 5316 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5316.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neoperla aethiopica Zwick & Zwick 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="81" masterDocId="1D15FF144A6AFF84FFD8FF9CFF910F7D" masterDocTitle="Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data" masterLastPageNumber="194" masterPageNumber="1" pageNumber="80" updateTime="1699540694688" updateUser="plazi">
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<mods:titleInfo id="D230098A93921A6A2915140A771FC404">
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<mods:title id="42659CD83E02588D19825EA4F72C4E4A">Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data</mods:title>
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<mods:name id="2617FEDD4574E224090A03C1C18259D3" type="personal">
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<mods:namePart id="4E36F91BEDCBF64EDBEC780D87EEEFC0">Zwick, Peter</mods:namePart>
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<mods:nameIdentifier id="504DFDDCD9C75E7B6279AEFAF59E765A" type="ORCID">0000-0002-6730-0011</mods:nameIdentifier>
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<mods:nameIdentifier id="4B38BBB90F65BFC4B1CBDF62BF0EECF7" type="email">pleco-p.zwick@t-online.de</mods:nameIdentifier>
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<mods:namePart id="84F3F3F5FEB2CCE46CF2C37FE49214A2">Zwick, Andreas</mods:namePart>
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<mods:nameIdentifier id="87BA07E587EE80CECBE505B276AE2CF9" type="ORCID">0000-0002-7532-1752</mods:nameIdentifier>
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<mods:nameIdentifier id="3038B9E6B5983D8DB3589F8BE94B0212" type="email">andreas.zwick@csiro.au</mods:nameIdentifier>
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<mods:typeOfResource id="6F8BA29EE4FB95AD63E73A8D014B6705">text</mods:typeOfResource>
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<mods:title id="3AC5A7143874D72FE783B41BEAFC46CA">Zootaxa</mods:title>
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<mods:part id="A4BDB3F977F0890AFB42314A271855F8">
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<mods:date id="A15CA439432A89F36673B3F84392AE71">2023</mods:date>
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<mods:number id="7C73E1E68531698773997EC92BC28D7F">2023-07-17</mods:number>
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<mods:number id="7B8EF1531B2C3CC3026D54456CF000D9">5316</mods:number>
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<mods:detail id="1D9D63D1AFF21A6A9ED7A2A524DFE940" type="issue">
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<mods:number id="C3BD7A7F10FC5F31C8219EE64C155244">1</mods:number>
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<mods:url id="F60E2A0A62D0221D7A7947F74EBDB9C9">http://dx.doi.org/10.11646/zootaxa.5316.1.1</mods:url>
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<mods:classification id="E1BBD86971655CBA4968DCD47804EFE7">journal article</mods:classification>
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<mods:identifier id="01C96BF2D62F7156EFDA0C8FEEFC726A" type="DOI">10.11646/zootaxa.5316.1.1</mods:identifier>
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<treatment id="E12C876C4A25FFD4FF4FF99DFC360EF4" ID-DOI="http://doi.org/10.5281/zenodo.8154218" ID-Zenodo-Dep="8154218" LSID="urn:lsid:plazi:treatment:E12C876C4A25FFD4FF4FF99DFC360EF4" httpUri="http://treatment.plazi.org/id/E12C876C4A25FFD4FF4FF99DFC360EF4" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
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<subSubSection id="219F65F14A25FFCBFF4FF99DFD900966" box="[151,513,1537,1564]" pageId="79" pageNumber="80" type="nomenclature">
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<paragraph id="693A367A4A25FFCBFF4FF99DFD900966" blockId="79.[151,513,1537,1599]" box="[151,513,1537,1564]" pageId="79" pageNumber="80">
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<emphasis id="5BF1EA684A25FFCBFF4FF99DFD900966" bold="true" box="[151,513,1537,1564]" pageId="79" pageNumber="80">
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32.
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<taxonomicName id="AE854DF94A25FFCBFF1BF99DFE270961" authority="Zwick & Zwick, 2023" authorityName="Zwick & Zwick" authorityYear="2023" box="[195,438,1537,1564]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
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<emphasis id="5BF1EA684A25FFCBFF1BF99DFE270961" bold="true" box="[195,438,1537,1564]" italics="true" pageId="79" pageNumber="80">Neoperla aethiopica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="40C257134A25FFCBFE66F99DFD900966" box="[446,513,1537,1563]" pageId="79" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="219F65F14A25FFCBFF4FF9B9FEDB0942" box="[151,330,1573,1599]" pageId="79" pageNumber="80" type="description">
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<paragraph id="693A367A4A25FFCBFF4FF9B9FEDB0942" blockId="79.[151,513,1537,1599]" box="[151,330,1573,1599]" pageId="79" pageNumber="80">
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||
(
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<figureCitation id="F1BE2AFF4A25FFCBFF47F9B9FED30942" box="[159,322,1573,1599]" captionStart="FIGURES 177–180" captionStartId="79.[152,270,1323,1347]" captionTargetBox="[160,1416,187,1292]" captionTargetId="figure-17@79.[154,1433,181,1299]" captionTargetPageId="79" captionText="FIGURES 177–180. Neoperla aethiopica n. sp. and N. occulta n. sp.. N. aethiopica: 177, penis of holotype (NEOP074); 178, enlarged distal section of the endophallus of the paratype. N. occulta: 179, distal half of penis with everted endophallus (Bayanga LP1); 180, distal part of contracted penis (Bayanga PJ, NEOP058). The curved line connects corresponding structures. Scales are 100µm. bbs, bare basal stretch; bl, basolateral, dd, dorsodistal, dl, dorsolateral spine groups." figureDoi="http://doi.org/10.5281/zenodo.8154068" httpUri="https://zenodo.org/record/8154068/files/figure.png" pageId="79" pageNumber="80">Figs. 177–178</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="219F65F14A25FFD4FF4FF9F1FDEC0E18" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="materials_examined">
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<paragraph id="693A367A4A25FFCBFF4FF9F1FAD009D6" blockId="79.[151,1437,1645,2032]" pageId="79" pageNumber="80">
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<emphasis id="5BF1EA684A25FFCBFF4FF9F1FE3209FA" bold="true" box="[151,419,1645,1671]" pageId="79" pageNumber="80">Type material studied</emphasis>
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:
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<materialsCitation id="D9ED3C274A25FFCBFE6EF9F3FAD009D6" collectingDate="1973-07" collectionCode="SMNS" collectorName="G. de Rougemont" country="Ethiopia" elevation="1600" location="Illubabor prov." pageId="79" pageNumber="80" specimenCode="NEOP074" specimenCount="2" typeStatus="holotype">
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♁
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<emphasis id="5BF1EA684A25FFCBFE0EF9F1FDD209FA" bold="true" box="[470,579,1645,1671]" pageId="79" pageNumber="80">
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<typeStatus id="B63E88D84A25FFCBFE0EF9F1FDD209FA" box="[470,579,1645,1671]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
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</emphasis>
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(
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<specimenCode id="39239E014A25FFCBFD8DF9F1FD5E09FA" box="[597,719,1645,1671]" collectionCode="NEOP" pageId="79" pageNumber="80">NEOP074</specimenCode>
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), 1 ♁
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<emphasis id="5BF1EA684A25FFCBFCC5F9F1FC1909FA" bold="true" box="[797,904,1645,1671]" pageId="79" pageNumber="80">
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||
<typeStatus id="B63E88D84A25FFCBFCC5F9F1FC1909FA" box="[797,904,1645,1671]" pageId="79" pageNumber="80" type="paratype">paratype</typeStatus>
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||
</emphasis>
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||
:
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<emphasis id="5BF1EA684A25FFCBFC43F9F1FA0409FA" bold="true" box="[923,1429,1645,1671]" pageId="79" pageNumber="80">
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<collectingCountry id="119276EA4A25FFCBFC43F9F1FA0409FA" box="[923,1429,1645,1671]" name="Ethiopia" pageId="79" pageNumber="80">Federal Democratic Republic of Ethiopia</collectingCountry>
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</emphasis>
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,
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<locationDeviation id="78882F584A25FFCBFF4FF90DFEE909D6" box="[151,376,1681,1708]" location="Chora" pageId="79" pageNumber="80">
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<quantity id="AE7D9B9F4A25FFCBFF4FF90DFF4609D1" box="[151,215,1681,1708]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="79" pageNumber="80" unit="km" value="15.0">15km</quantity>
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NW of Chora
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</locationDeviation>
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,
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<location id="6C5A60A14A25FFCBFE5CF90DFDBC09D1" LSID="urn:lsid:plazi:treatment:E12C876C4A25FFD4FF4FF99DFC360EF4:6C5A60A14A25FFCBFE5CF90DFDBC09D1" box="[388,557,1681,1708]" country="Ethiopia" name="Illubabor prov." pageId="79" pageNumber="80">Illubabor prov.</location>
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,
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<quantity id="AE7D9B9F4A25FFCBFDE0F90DFD1909D6" box="[568,648,1681,1707]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="79" pageNumber="80" unit="m" value="1600.0">
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||
<elevation id="E2A8D1494A25FFCBFDE0F90DFD1909D6" box="[568,648,1681,1707]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="79" pageNumber="80" unit="m" value="1600.0">1600m</elevation>
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</quantity>
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,
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<date id="1D3B10BA4A25FFCBFD4CF90DFD6209D6" box="[660,755,1681,1707]" pageId="79" pageNumber="80" value="1973-07">
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||
<collectingDate id="0D7FE9524A25FFCBFD4CF90DFD6209D6" box="[660,755,1681,1707]" pageId="79" pageNumber="80" value="1973-07">vii.1973</collectingDate>
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</date>
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,
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<collectorName id="C47053AC4A25FFCBFD26F90DFC2F09D1" box="[766,958,1681,1708]" pageId="79" pageNumber="80">G.de Rougemont</collectorName>
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||
(slides Z16.166 & 166A;
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<collectionCode id="0F94AEBF4A25FFCBFB3CF90DFAA409D6" box="[1252,1333,1681,1707]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="79" pageNumber="80">SMNS</collectionCode>
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||
).
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</materialsCitation>
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</paragraph>
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<paragraph id="693A367A4A25FFCBFF1FF929FEB9098E" blockId="79.[151,1437,1645,2032]" pageId="79" pageNumber="80">
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<emphasis id="5BF1EA684A25FFCBFF1FF929FEB709B2" bold="true" box="[199,294,1717,1743]" pageId="79" pageNumber="80">Habitus</emphasis>
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. WL 12.0–
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<quantity id="AE7D9B9F4A25FFCBFE7EF929FD9409B2" box="[422,517,1717,1743]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="79" pageNumber="80" unit="mm" value="12.3">12.3mm</quantity>
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. Ochre colour, legs a little darker, brownish, wings infuscate. A black ring around each ocellus.
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</paragraph>
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<paragraph id="693A367A4A25FFCBFF1FF961FDAC08FE" blockId="79.[151,1437,1645,2032]" pageId="79" pageNumber="80">
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<emphasis id="5BF1EA684A25FFCBFF1FF961FE95086A" bold="true" box="[199,260,1789,1815]" pageId="79" pageNumber="80">Male</emphasis>
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||
(
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<figureCitation id="F1BE2AFF4A25FFCBFECFF961FE50086A" box="[279,449,1789,1815]" captionStart="FIGURES 177–180" captionStartId="79.[152,270,1323,1347]" captionTargetBox="[160,1416,187,1292]" captionTargetId="figure-17@79.[154,1433,181,1299]" captionTargetPageId="79" captionText="FIGURES 177–180. Neoperla aethiopica n. sp. and N. occulta n. sp.. N. aethiopica: 177, penis of holotype (NEOP074); 178, enlarged distal section of the endophallus of the paratype. N. occulta: 179, distal half of penis with everted endophallus (Bayanga LP1); 180, distal part of contracted penis (Bayanga PJ, NEOP058). The curved line connects corresponding structures. Scales are 100µm. bbs, bare basal stretch; bl, basolateral, dd, dorsodistal, dl, dorsolateral spine groups." figureDoi="http://doi.org/10.5281/zenodo.8154068" httpUri="https://zenodo.org/record/8154068/files/figure.png" pageId="79" pageNumber="80">Figs. 177–178</figureCitation>
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). Hind tibiae unmodified. T7 with rather long triangular projection. T8 flat, front of median sclerite band as wide as the T7 process above it, narrowing caudally, with some SB. T9 unmodified, the paramedian pilose humps not much raised. Callus of HT10 inconspicuous, tongue-shaped. Tip of HT10 sinuous, short, the sharp apex points cephalad.
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</paragraph>
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<paragraph id="693A367A4A25FFD4FF1FF811FC810FA8" blockId="79.[151,1437,1645,2032]" lastBlockId="80.[151,1436,151,393]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
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Penis tubular, slender, smooth except one or two single spinelets on sides, about one third from tip (bl in
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<figureCitation id="F1BE2AFF4A25FFCBFAA8F811FF5408B6" captionStart="FIGURES 177–180" captionStartId="79.[152,270,1323,1347]" captionTargetBox="[160,1416,187,1292]" captionTargetId="figure-17@79.[154,1433,181,1299]" captionTargetPageId="79" captionText="FIGURES 177–180. Neoperla aethiopica n. sp. and N. occulta n. sp.. N. aethiopica: 177, penis of holotype (NEOP074); 178, enlarged distal section of the endophallus of the paratype. N. occulta: 179, distal half of penis with everted endophallus (Bayanga LP1); 180, distal part of contracted penis (Bayanga PJ, NEOP058). The curved line connects corresponding structures. Scales are 100µm. bbs, bare basal stretch; bl, basolateral, dd, dorsodistal, dl, dorsolateral spine groups." figureDoi="http://doi.org/10.5281/zenodo.8154068" httpUri="https://zenodo.org/record/8154068/files/figure.png" pageId="79" pageNumber="80">Fig. 177</figureCitation>
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). Distally, is a rough dorsal patch of spines, plus a smaller spine patch on each side (dd and dl, respectively in
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<figureCitation id="F1BE2AFF4A25FFCBFF6EF849FE8B0892" box="[182,282,2005,2031]" captionStart="FIGURES 177–180" captionStartId="79.[152,270,1323,1347]" captionTargetBox="[160,1416,187,1292]" captionTargetId="figure-17@79.[154,1433,181,1299]" captionTargetPageId="79" captionText="FIGURES 177–180. Neoperla aethiopica n. sp. and N. occulta n. sp.. N. aethiopica: 177, penis of holotype (NEOP074); 178, enlarged distal section of the endophallus of the paratype. N. occulta: 179, distal half of penis with everted endophallus (Bayanga LP1); 180, distal part of contracted penis (Bayanga PJ, NEOP058). The curved line connects corresponding structures. Scales are 100µm. bbs, bare basal stretch; bl, basolateral, dd, dorsodistal, dl, dorsolateral spine groups." figureDoi="http://doi.org/10.5281/zenodo.8154068" httpUri="https://zenodo.org/record/8154068/files/figure.png" pageId="79" pageNumber="80">Fig. 177</figureCitation>
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). Endophallus almost straight, long base bare, distal half narrowing. Sides bare, the armature begins dorsolaterally as band of large hooks, distal hooks smaller, more numerous. Conical apex with many dorsal spines and a short ventral row of single spines (
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<figureCitation id="F1BE2AFF4A3AFFD4FD86FF27FC950FA8" box="[606,772,187,213]" captionStart="FIGURES 177–180" captionStartId="79.[152,270,1323,1347]" captionTargetBox="[160,1416,187,1292]" captionTargetId="figure-17@79.[154,1433,181,1299]" captionTargetPageId="79" captionText="FIGURES 177–180. Neoperla aethiopica n. sp. and N. occulta n. sp.. N. aethiopica: 177, penis of holotype (NEOP074); 178, enlarged distal section of the endophallus of the paratype. N. occulta: 179, distal half of penis with everted endophallus (Bayanga LP1); 180, distal part of contracted penis (Bayanga PJ, NEOP058). The curved line connects corresponding structures. Scales are 100µm. bbs, bare basal stretch; bl, basolateral, dd, dorsodistal, dl, dorsolateral spine groups." figureDoi="http://doi.org/10.5281/zenodo.8154068" httpUri="https://zenodo.org/record/8154068/files/figure.png" pageId="80" pageNumber="81">Figs. 177–178</figureCitation>
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).
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</paragraph>
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<paragraph id="693A367A4A3AFFD4FF1FFF43FE330F84" blockId="80.[151,1436,151,393]" box="[199,418,223,249]" pageId="80" pageNumber="81">
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||
<emphasis id="5BF1EA684A3AFFD4FF1FFF43FE8F0F84" bold="true" box="[199,286,223,249]" pageId="80" pageNumber="81">Female</emphasis>
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. Unknown.
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</paragraph>
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<paragraph id="693A367A4A3AFFD4FF1FFE9EFDEC0E18" blockId="80.[151,1436,151,393]" pageId="80" pageNumber="81">
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||
<emphasis id="5BF1EA684A3AFFD4FF1FFE9EFE950E61" bold="true" box="[199,260,258,284]" pageId="80" pageNumber="81">DNA</emphasis>
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(
|
||
<figureCitation id="F1BE2AFF4A3AFFD4FECCFE9FFE290E60" box="[276,440,259,285]" captionStart-0="FIGURE 491" captionStart-1="FIGURE 492" captionStartId-0="177.[152,255,1835,1859]" captionStartId-1="178.[152,255,1943,1967]" captionTargetBox-0="[176,1411,312,1806]" captionTargetBox-1="[361,1187,182,1910]" captionTargetId-0="graphics-120@177.[176,904,321,1800]" captionTargetPageId-0="177" captionTargetPageId-1="178" captionText-0="FIGURE 491. Codon-model maximum likelihood phylogram based on 83 largely complete mitochondrial genome sequences only (mt-NT). Statistical support values (SH-aLRT, UFBoot) from this and other analyses summarised by squares as explained in the figure. Terminal labels include information on the species, type status, gender, originating country, unique sequence identifier (NEOP###), sequencing approach and total basepairs of 13 mitochondrial, protein-coding genes. Clades of particular interest highlighted with colours that correspond to the clades in other trees. HT = holotype; PT = paratype; PCR = Polymerase Chaine Reaction; WGS = Whole Genome Shotgun." captionText-1="FIGURE 492. Simplified overview of codon-model maximum likelihood phylogram based on 356 mitochondrial genome and COX1 DNA barcode sequences (all-NT). Terminal labels include information on the species only. Clades of particular interest highlighted with colours that correspond to the clades in other trees. The full tree is shown in Figs 493-498." figureDoi-0="http://doi.org/10.5281/zenodo.8154198" figureDoi-1="http://doi.org/10.5281/zenodo.8154202" httpUri-0="https://zenodo.org/record/8154198/files/figure.png" httpUri-1="https://zenodo.org/record/8154202/files/figure.png" pageId="80" pageNumber="81">Figs. 491–492</figureCitation>
|
||
,
|
||
<figureCitation id="F1BE2AFF4A3AFFD4FE1CFE9FFE620E60" box="[452,499,259,285]" captionStart="FIGURE 494" captionStartId="180.[152,255,1112,1136]" captionText="FIGURE 494. Partial (clade F) codon-model maximum likelihood phylogram based on 356 mitochondrial genome and COX1 DNA barcode sequences (all-NT). See Fig. 493 for common details." pageId="80" pageNumber="81">494</figureCitation>
|
||
).
|
||
<materialsCitation id="D9ED3C274A3AFFD4FDDEFE9FFDE10E18" country="Ethiopia" location="The" pageId="80" pageNumber="81" specimenCount="1" typeStatus="holotype">
|
||
Only the male
|
||
<typeStatus id="B63E88D84A3AFFD4FD75FE9FFC9E0E60" box="[685,783,259,285]" pageId="80" pageNumber="81" type="holotype">holotype</typeStatus>
|
||
from
|
||
<collectingCountry id="119276EA4A3AFFD4FC8CFE9FFC220E60" box="[852,947,259,285]" name="Ethiopia" pageId="80" pageNumber="81">Ethiopia</collectingCountry>
|
||
was sequenced with the genome skimming approach, resulting in 10,983bp of mitochondrial, protein-coding genes.
|
||
<location id="6C5A60A14A3AFFD4FC06FEBBFB9B0E3C" LSID="urn:lsid:plazi:treatment:E12C876C4A25FFD4FF4FF99DFC360EF4:6C5A60A14A3AFFD4FC06FEBBFB9B0E3C" box="[990,1034,295,321]" country="Ethiopia" name="The" pageId="80" pageNumber="81">The</location>
|
||
species is very strongly supported (98.2/99/99) as sister to
|
||
<taxonomicName id="AE854DF94A3AFFD4FE7FFED7FDBE0E18" authority="Zwick & Zwick, 2023" authorityName="Zwick & Zwick" authorityYear="2023" box="[423,559,331,357]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pilulifera" status="sp. nov.">
|
||
<emphasis id="5BF1EA684A3AFFD4FE7FFED7FDBE0E18" box="[423,559,331,357]" italics="true" pageId="80" pageNumber="81">N. pilulifera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="40C257134A3AFFD4FDEEFED7FDE10E18" box="[566,624,331,357]" pageId="80" pageNumber="81" rank="species">
|
||
<emphasis id="5BF1EA684A3AFFD4FDEEFED7FDE10E18" bold="true" box="[566,624,331,357]" pageId="80" pageNumber="81">n. sp</emphasis>
|
||
</taxonomicNameLabel>
|
||
</materialsCitation>
|
||
<emphasis id="5BF1EA684A3AFFD4FDA8FED7FDE40E18" bold="true" box="[624,629,331,357]" pageId="80" pageNumber="81">.</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="219F65F14A3AFFD4FF1FFEF3FC360EF4" box="[199,935,367,393]" pageId="80" pageNumber="81" type="etymology">
|
||
<paragraph id="693A367A4A3AFFD4FF1FFEF3FC360EF4" blockId="80.[151,1436,151,393]" box="[199,935,367,393]" pageId="80" pageNumber="81">
|
||
<emphasis id="5BF1EA684A3AFFD4FF1FFEF3FED90EF4" bold="true" box="[199,328,367,393]" pageId="80" pageNumber="81">Etymology</emphasis>
|
||
. The name was derived from the patria of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |