treatments-xml/data/E1/2C/87/E12C876C4A19FFF1FF4FF9AAFB5C0887.xml
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<document id="91141EDD18D90A5C2294AB4DD256FDEE" ID-DOI="10.11646/zootaxa.5316.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8154005" ID-ZooBank="BC922E16-2614-4F3D-AD82-87A845DE7E2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689581758737" checkinUser="plazi" docAuthor="Zwick, Peter &amp; Zwick, Andreas" docDate="2023" docId="E12C876C4A19FFF1FF4FF9AAFB5C0887" docLanguage="en" docName="zootaxa.5316.1.1.pdf" docOrigin="Zootaxa 5316 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5316.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neoperla orthonema Zwick &amp; Zwick 2023, n. sp." docType="treatment" docVersion="4" lastPageNumber="118" masterDocId="1D15FF144A6AFF84FFD8FF9CFF910F7D" masterDocTitle="Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data" masterLastPageNumber="194" masterPageNumber="1" pageNumber="116" updateTime="1701827539936" updateUser="ExternalLinkService">
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<mods:title id="71D854BA02646AB05116EE2DA3C101CE">Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data</mods:title>
</mods:titleInfo>
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<mods:namePart id="D029823808B20270BB9BC6BD86659267">Zwick, Peter</mods:namePart>
<mods:nameIdentifier id="D0D57AF8C686E6098C45F73B8EAD0DAE" type="ORCID">0000-0002-6730-0011</mods:nameIdentifier>
<mods:nameIdentifier id="353F40BB9E192030DA653EFD8F2EDA0E" type="email">pleco-p.zwick@t-online.de</mods:nameIdentifier>
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<mods:namePart id="A1D4F394A6307803C90923C2A55ABDC7">Zwick, Andreas</mods:namePart>
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<mods:nameIdentifier id="F671BFFC74BB683A1935351319FB740E" type="email">andreas.zwick@csiro.au</mods:nameIdentifier>
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<mods:title id="C7CB2D4AEF152A1E8D917C185FD1353B">Zootaxa</mods:title>
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<mods:date id="7CE2FF5BD9EF024AC3292580043D2717">2023</mods:date>
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<mods:number id="B6BAD075E1A82B4E867B34726EAF1110">2023-07-17</mods:number>
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<treatment id="E12C876C4A19FFF1FF4FF9AAFB5C0887" ID-DOI="http://doi.org/10.5281/zenodo.8154255" ID-Zenodo-Dep="8154255" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887" httpUri="http://treatment.plazi.org/id/E12C876C4A19FFF1FF4FF9AAFB5C0887" lastPageId="117" lastPageNumber="118" pageId="115" pageNumber="116">
<subSubSection id="219F65F14A19FFF7FF4FF9AAFD96092C" box="[151,519,1590,1617]" pageId="115" pageNumber="116" type="nomenclature">
<paragraph id="693A367A4A19FFF7FF4FF9AAFD96092C" blockId="115.[151,519,1590,1653]" box="[151,519,1590,1617]" pageId="115" pageNumber="116">
<emphasis id="5BF1EA684A19FFF7FF4FF9AAFD96092C" bold="true" box="[151,519,1590,1617]" pageId="115" pageNumber="116">
52.
<taxonomicName id="AE854DF94A19FFF7FF1BF9AAFE2C092C" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[195,445,1590,1617]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="orthonema" status="sp. nov.">
<emphasis id="5BF1EA684A19FFF7FF1BF9AAFE2C092C" bold="true" box="[195,445,1590,1617]" italics="true" pageId="115" pageNumber="116">Neoperla orthonema</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C257134A19FFF7FE1DF9ABFD96092C" box="[453,519,1591,1617]" pageId="115" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A19FFF7FF4FF9C7FEDA0908" box="[151,331,1627,1653]" pageId="115" pageNumber="116" type="description">
<paragraph id="693A367A4A19FFF7FF4FF9C7FEDA0908" blockId="115.[151,519,1590,1653]" box="[151,331,1627,1653]" pageId="115" pageNumber="116">
(
<figureCitation id="F1BE2AFF4A19FFF7FF47F9C7FED30908" box="[159,322,1627,1653]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="115" pageNumber="116">Figs. 294304</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A19FFF1FF4FF93FFAE008CF" lastPageId="117" lastPageNumber="118" pageId="115" pageNumber="116" type="materials_examined">
<paragraph id="693A367A4A19FFF0FF4FF93FFD090EB2" blockId="115.[151,1437,1698,2015]" lastBlockId="116.[151,1437,151,2037]" lastPageId="116" lastPageNumber="117" pageId="115" pageNumber="116">
<emphasis id="5BF1EA684A19FFF7FF4FF93FFC8E09C0" bold="true" box="[151,799,1698,1725]" pageId="115" pageNumber="116">
Type material studied:
<collectingCountry id="119276EA4A19FFF7FE71F93EFD1D09C1" box="[425,652,1698,1724]" name="Ghana" pageId="115" pageNumber="116">Republic of Ghana</collectingCountry>
: J
<typeStatus id="B63E88D84A19FFF7FD6AF93FFC8E09C0" box="[690,799,1699,1725]" pageId="115" pageNumber="116" type="holotype">Holotype</typeStatus>
</emphasis>
(
<collectionCode id="0F94AEBF4A19FFF7FCF4F93FFCED09C0" box="[812,892,1699,1725]" pageId="115" pageNumber="116">NEOP</collectionCode>
171, slide Z17_037_sp_Ax \GH.10), Ghana
<collectingRegion id="AB41F8984A19FFF7FABAF93FFF70099C" country="Ghana" name="Volta" pageId="115" pageNumber="116">Volta region</collectingRegion>
, Agumatsa waterfalls, Wli, St. 10
<date id="1D3B10BA4A19FFF7FDBEF95BFD73099C" box="[614,738,1735,1761]" pageId="115" pageNumber="116" value="1993-11-19">19.11.1993</date>
H. Olsvik light trap; NUFU project (
<collectionCode id="0F94AEBF4A19FFF7FB51F95BFB71099C" box="[1161,1248,1735,1761]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="115" pageNumber="116">ZMBN</collectionCode>
).
<specimenCount id="7F83FDF34A19FFF7FB2CF95BFA86099C" box="[1268,1303,1735,1761]" pageId="115" pageNumber="116" type="female">3♀</specimenCount>
<emphasis id="5BF1EA684A19FFF7FAC7F95BFA04099C" bold="true" box="[1311,1429,1735,1761]" pageId="115" pageNumber="116">
<typeStatus id="B63E88D84A19FFF7FAC7F95BFA04099C" box="[1311,1429,1735,1761]" pageId="115" pageNumber="116" type="paratype">paratypes</typeStatus>
</emphasis>
, Ghana, same data as
<typeStatus id="B63E88D84A19FFF7FE54F977FE7F0878" box="[396,494,1771,1797]" pageId="115" pageNumber="116" type="holotype">holotype</typeStatus>
(slides GH.10\2, GH.11\2,
<collectionCode id="0F94AEBF4A19FFF7FCF4F977FC130878" box="[812,898,1771,1797]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="115" pageNumber="116">ZMBN</collectionCode>
).
<specimenCount id="7F83FDF34A19FFF7FC4EF977FC280878" box="[918,953,1771,1797]" pageId="115" pageNumber="116" type="female">1♀</specimenCount>
<typeStatus id="B63E88D84A19FFF7FC1BF977FBB90878" box="[963,1064,1771,1797]" pageId="115" pageNumber="116" type="paratype">Paratype</typeStatus>
, Ghana
<collectingRegion id="AB41F8984A19FFF7FB5FF977FA8E0878" box="[1159,1311,1771,1797]" country="Ghana" name="Volta" pageId="115" pageNumber="116">Volta Region</collectingRegion>
, Wli Falls, Afegame [
<geoCoordinate id="0CB150BD4A19FFF7FEC8F893FEF40854" box="[272,357,1807,1833]" degrees="7.12" direction="north" orientation="latitude" pageId="115" pageNumber="116" precision="555" value="7.12">7.12°N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A19FFF7FEADF892FE590854" box="[373,456,1806,1833]" degrees="0.59" direction="east" orientation="longitude" pageId="115" pageNumber="116" precision="555" value="0.59">0.59°E</geoCoordinate>
], St. 12
<date id="1D3B10BA4A19FFF7FDF7F893FD3C0854" box="[559,685,1807,1833]" pageId="115" pageNumber="116" value="1993-07-03">07.03.1993</date>
J.S.Amakye &amp; J. Kjaerendsen light trap; NUFU-project
<collectionCode id="0F94AEBF4A19FFF7FA93F893FA0D0854" box="[1355,1436,1807,1833]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="115" pageNumber="116">ZMBN</collectionCode>
Sample no. 1 (
<collectionCode id="0F94AEBF4A19FFF7FE9FF8AFFE040830" box="[327,405,1843,1869]" pageId="115" pageNumber="116">NEOP</collectionCode>
169, slide GH.09\1,
<collectionCode id="0F94AEBF4A19FFF7FD5BF8AFFD470830" box="[643,726,1843,1869]" pageId="115" pageNumber="116">ZMNS</collectionCode>
);
<specimenCount id="7F83FDF34A19FFF7FD36F8AFFC800830" box="[750,785,1843,1869]" pageId="115" pageNumber="116" type="female">4♀</specimenCount>
<emphasis id="5BF1EA684A19FFF7FCC5F8AFFC020830" bold="true" box="[797,915,1843,1869]" pageId="115" pageNumber="116">
<typeStatus id="B63E88D84A19FFF7FCC5F8AFFC020830" box="[797,915,1843,1869]" pageId="115" pageNumber="116" type="paratype">paratypes</typeStatus>
</emphasis>
, Ghana Western region, Ankasa game prod. Reserve [
<geoCoordinate id="0CB150BD4A19FFF7FF27F8CAFEDB080C" box="[255,330,1878,1905]" degrees="5.44" direction="north" orientation="latitude" pageId="115" pageNumber="116" precision="555" value="5.44">5.44N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A19FFF7FE8EF8CBFE30080C" box="[342,417,1879,1905]" degrees="2.07" direction="west" orientation="longitude" pageId="115" pageNumber="116" precision="555" value="-2.07">2.07W</geoCoordinate>
,
<quantity id="AE7D9B9F4A19FFF7FE75F8CBFE75080C" box="[429,484,1879,1905]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="115" pageNumber="116" unit="m" value="47.0">47m</quantity>
], 1016.12.93 St. 12 J. Kjaerendsen &amp; T. Andersen light trap; NUFU-project
<collectionCode id="0F94AEBF4A19FFF7FA93F8CBFA0D080C" box="[1355,1436,1879,1905]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="115" pageNumber="116">ZMBN</collectionCode>
Ref. No. 3 (GH.01/17,
<collectionCode id="0F94AEBF4A19FFF7FE7FF8E7FE6408E8" box="[423,501,1915,1941]" pageId="115" pageNumber="116">NEOP</collectionCode>
164;
<emphasis id="5BF1EA684A19FFF7FDF4F8E6FDEB08E9" bold="true" box="[556,634,1914,1940]" pageId="115" pageNumber="116">
<collectionCode id="0F94AEBF4A19FFF7FDF4F8E6FDEB08E9" box="[556,634,1914,1940]" pageId="115" pageNumber="116">NEOP</collectionCode>
</emphasis>
168, slides GH.01/6, /9, /16, /17;
<collectionCode id="0F94AEBF4A19FFF7FBDDF8E7FBC708E8" box="[1029,1110,1915,1941]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="115" pageNumber="116">SMNS</collectionCode>
); 1♁
<emphasis id="5BF1EA684A19FFF7FB42F8E7FA9408E8" bold="true" box="[1178,1285,1915,1941]" pageId="115" pageNumber="116">
<typeStatus id="B63E88D84A19FFF7FB42F8E7FA9408E8" box="[1178,1285,1915,1941]" pageId="115" pageNumber="116" type="paratype">paratype</typeStatus>
</emphasis>
(
<collectionCode id="0F94AEBF4A19FFF7FACFF8E7FAF708E8" box="[1303,1382,1915,1941]" pageId="115" pageNumber="116">NEOP</collectionCode>
170),
<specimenCount id="7F83FDF34A19FFF7FF4FF803FF2B08C4" box="[151,186,1951,1977]" pageId="115" pageNumber="116" type="female">1♀</specimenCount>
<emphasis id="5BF1EA684A19FFF7FF1DF803FEA108C4" bold="true" box="[197,304,1951,1977]" pageId="115" pageNumber="116">
<typeStatus id="B63E88D84A19FFF7FF1DF803FEA108C4" box="[197,304,1951,1977]" pageId="115" pageNumber="116" type="paratype">paratype</typeStatus>
</emphasis>
(
<collectionCode id="0F94AEBF4A19FFF7FE9AF802FE0008C4" box="[322,401,1950,1977]" pageId="115" pageNumber="116">NEOP</collectionCode>
165, slide GH12\1),
<specimenCount id="7F83FDF34A19FFF7FDA5F803FD3108C4" box="[637,672,1951,1977]" pageId="115" pageNumber="116" type="female">1♀</specimenCount>
(Slide Z17.073) Ghana
<collectingRegion id="AB41F8984A19FFF7FC65F803FBC708C4" box="[957,1110,1951,1977]" country="Ghana" name="Volta" pageId="115" pageNumber="116">Volta Region</collectingRegion>
, Wli Falls, Afegame, St. 10
<date id="1D3B10BA4A19FFF7FF4FF85FFE8408A0" box="[151,277,1987,2013]" pageId="115" pageNumber="116" value="1993-08-03">08.03.1993</date>
J.S.Amakye &amp; J. Kjaerendsen light trap; NUFU-project
<collectionCode id="0F94AEBF4A19FFF7FC47F85FFC6108A0" box="[927,1008,1987,2013]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="115" pageNumber="116">ZMBN</collectionCode>
Sample no. 12; 1♁
<emphasis id="5BF1EA684A19FFF7FB00F85FFAD208A0" bold="true" box="[1240,1347,1987,2013]" pageId="115" pageNumber="116">
<typeStatus id="B63E88D84A19FFF7FB00F85FFAD208A0" box="[1240,1347,1987,2013]" pageId="115" pageNumber="116" type="paratype">paratype</typeStatus>
</emphasis>
, Ghana
<collectingRegion id="AB41F8984A1EFFF0FF4FFF0BFEA00FCC" box="[151,305,151,177]" country="Ghana" name="Volta" pageId="116" pageNumber="117">Volta Region</collectingRegion>
, Wli Falls, Afegame, St. 12
<date id="1D3B10BA4A1EFFF0FD50FF0BFC970FCC" box="[648,774,151,177]" pageId="116" pageNumber="117" value="1993-07-03">07.03.1993</date>
J.S.Amakye &amp; J. Kjaerendsen light trap; NUFU-project
<collectionCode id="0F94AEBF4A1EFFF0FF4FFF27FF790FA8" box="[151,232,187,213]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="116" pageNumber="117">ZMBN</collectionCode>
Sample no. 1); 1♁
<emphasis id="5BF1EA684A1EFFF0FE0DFF27FDD10FA8" bold="true" box="[469,576,187,213]" pageId="116" pageNumber="117">
<typeStatus id="B63E88D84A1EFFF0FE0DFF27FDD10FA8" box="[469,576,187,213]" pageId="116" pageNumber="117" type="paratype">paratype</typeStatus>
</emphasis>
, Ghana
<collectingRegion id="AB41F8984A1EFFF0FD7CFF27FCAC0FA8" box="[676,829,187,213]" country="Ghana" name="Volta" pageId="116" pageNumber="117">Volta Region</collectingRegion>
, Wli Falls, Afegame, St. 12
<date id="1D3B10BA4A1EFFF0FB55FF27FA9A0FA8" box="[1165,1291,187,213]" pageId="116" pageNumber="117" value="1993-07-03">07.03.1993</date>
J.S.Amakye &amp; J. Kjaerendsen light trap; NUFU-project
<collectionCode id="0F94AEBF4A1EFFF0FD4DFF43FD770F84" box="[661,742,223,249]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="116" pageNumber="117">ZMBN</collectionCode>
Sample no. 1 (
<collectionCode id="0F94AEBF4A1EFFF0FC46FF43FC7D0F84" box="[926,1004,223,249]" pageId="116" pageNumber="117">NEOP</collectionCode>
166, slide 98\9,
<collectionCode id="0F94AEBF4A1EFFF0FB7FFF43FB690F84" box="[1191,1272,223,249]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">SMNS</collectionCode>
).
<emphasis id="5BF1EA684A1EFFF0FAD7FF42FE8A0E61" bold="true" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FAD7FF42FE870E61" name="Cameroon" pageId="116" pageNumber="117">Republic of Cameroon</collectingCountry>
,
</emphasis>
4♁ p
<emphasis id="5BF1EA684A1EFFF0FEBAFE9FFE580E60" bold="true" box="[354,457,259,285]" pageId="116" pageNumber="117">aratypes</emphasis>
: Muyuka [
<geoCoordinate id="0CB150BD4A1EFFF0FD91FE9FFD3E0E60" box="[585,687,259,285]" degrees="4.7217" direction="north" orientation="latitude" pageId="116" pageNumber="117" precision="5" value="4.7217">4.7217N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A1EFFF0FD66FE9FFCB00E60" box="[702,801,259,285]" degrees="9.6408" direction="east" orientation="longitude" pageId="116" pageNumber="117" precision="5" value="9.6408">9.6408E</geoCoordinate>
;
<quantity id="AE7D9B9F4A1EFFF0FCF7FE9EFCE20E60" box="[815,883,258,285]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="116" pageNumber="117" unit="m" value="305.0">305m</quantity>
], Victoria Div., BRIT. CAMEROONS,VI-25-49 B.Malkin at Light (Slides Z17.021-023;
<collectionCode id="0F94AEBF4A1EFFF0FDBBFEBBFD310E3C" box="[611,672,295,321]" country="Germany" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="116" pageNumber="117" type="Museum">ZMB</collectionCode>
&amp;
<collectionCode id="0F94AEBF4A1EFFF0FD1DFEBBFC870E3C" box="[709,790,295,321]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">SMNS</collectionCode>
); 1♁
<emphasis id="5BF1EA684A1EFFF0FC8EFEBBFC590E3C" bold="true" box="[854,968,295,321]" pageId="116" pageNumber="117">
<typeStatus id="B63E88D84A1EFFF0FC8EFEBBFC520E3C" box="[854,963,295,321]" pageId="116" pageNumber="117" type="paratype">paratype</typeStatus>
,
</emphasis>
<quantity id="AE7D9B9F4A1EFFF0FC09FEBBFB940E3C" box="[977,1029,295,321]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="116" pageNumber="117" unit="mi" value="7.0">7 mi</quantity>
S Ebolowa [
<geoCoordinate id="0CB150BD4A1EFFF0FB4FFEBBFB730E3C" box="[1175,1250,295,321]" degrees="2.92" direction="north" orientation="latitude" pageId="116" pageNumber="117" precision="555" value="2.92">2.92N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A1EFFF0FB36FEBBFAD30E3C" box="[1262,1346,295,321]" degrees="11.16" direction="east" orientation="longitude" pageId="116" pageNumber="117" precision="555" value="11.16">11.16E</geoCoordinate>
],
<quantity id="AE7D9B9F4A1EFFF0FA8DFEBAFA090E3C" box="[1365,1432,294,321]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="116" pageNumber="117" unit="m" value="580.0">580m</quantity>
,
<date id="1D3B10BA4A1EFFF0FF4FFED6FED20E18" box="[151,323,330,357]" pageId="116" pageNumber="117" value="1966-10-15" valueMax="1966-10-17" valueMin="1966-10-15">15.17-X-1966</date>
Ross &amp; Leech CASENT8413020 (penis slide).
<emphasis id="5BF1EA684A1EFFF0FCB4FED5FB5E0E19" bold="true" box="[876,1231,329,356]" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FCB4FED5FB5B0E19" box="[876,1226,329,356]" name="Ivory Coast" pageId="116" pageNumber="117">République de Côte dIvoire</collectingCountry>
:
</emphasis>
1
<emphasis id="5BF1EA684A1EFFF0FB3FFED6FAE60E19" bold="true" box="[1255,1399,330,356]" pageId="116" pageNumber="117">
J
<typeStatus id="B63E88D84A1EFFF0FADDFED6FAE30E19" box="[1285,1394,330,356]" pageId="116" pageNumber="117" type="paratype">paratype</typeStatus>
,
</emphasis>
<quantity id="AE7D9B9F4A1EFFF0FA58FED6FF2A0EF5" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="116" pageNumber="117" unit="km" value="15.0">15 km</quantity>
N Lakota [
<geoCoordinate id="0CB150BD4A1EFFF0FEE6FEF2FE180EF4" box="[318,393,366,393]" degrees="5.86" direction="north" orientation="latitude" pageId="116" pageNumber="117" precision="555" value="5.86">5.86N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A1EFFF0FE4EFEF2FE750EF4" box="[406,484,366,393]" degrees="5.68" direction="west" orientation="longitude" pageId="116" pageNumber="117" precision="555" value="-5.68">5.68W</geoCoordinate>
]
<date id="1D3B10BA4A1EFFF0FE2CFEF2FD160EF5" box="[500,647,366,392]" pageId="116" pageNumber="117" value="1971-10-23">23.Oct. 1971</date>
white light trap J.A.Gruwell (
<collectionCode id="0F94AEBF4A1EFFF0FC39FEF2FBA50EF5" box="[993,1076,366,392]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="116" pageNumber="117" type="Museum">USNM</collectionCode>
152) (
<figureCitation id="F1BE2AFF4A1EFFF0FB59FEF2FAB20EF5" box="[1153,1315,366,392]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Figs 294299</figureCitation>
).
<emphasis id="5BF1EA684A1EFFF0FAEFFEF2FE980ED1" bold="true" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FAEFFEF2FE950ED1" name="Togo" pageId="116" pageNumber="117">Togolese Republic</collectingCountry>
,
</emphasis>
1♁
<emphasis id="5BF1EA684A1EFFF0FEE2FE0EFE340ED1" bold="true" box="[314,421,402,428]" pageId="116" pageNumber="117">
<typeStatus id="B63E88D84A1EFFF0FEE2FE0EFE340ED1" box="[314,421,402,428]" pageId="116" pageNumber="117" type="paratype">paratype</typeStatus>
</emphasis>
, penis in microvial,
<specimenCount id="7F83FDF34A1EFFF0FD53FE0EFD3E0ED1" box="[651,687,402,428]" pageId="116" pageNumber="117" type="female">2♀</specimenCount>
,
<emphasis id="5BF1EA684A1EFFF0FD64FE0EFCA30ED1" bold="true" box="[700,818,402,428]" pageId="116" pageNumber="117">
<typeStatus id="B63E88D84A1EFFF0FD64FE0EFCA30ED1" box="[700,818,402,428]" pageId="116" pageNumber="117" type="paratype">paratypes</typeStatus>
</emphasis>
(Z16/131,
<collectionCode id="0F94AEBF4A1EFFF0FC77FE0EFC6A0ED1" box="[943,1019,402,428]" pageId="116" pageNumber="117">NEOP</collectionCode>
167 and Z16/132,
<collectionCode id="0F94AEBF4A1EFFF0FB1DFE0EFA850ED1" box="[1221,1300,402,428]" pageId="116" pageNumber="117">NEOP</collectionCode>
161;
<collectionCode id="0F94AEBF4A1EFFF0FA9FFE0EFA040ED1" box="[1351,1429,402,428]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">SMNS</collectionCode>
) Amoutchou,
<date id="1D3B10BA4A1EFFF0FEF1FE29FE380EAD" box="[297,425,437,464]" pageId="116" pageNumber="117" value="1985-11-28">28.11.1985</date>
, V. Landa (Z16.132).
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FFE45FC0C0CDD" blockId="116.[151,1437,151,2037]" pageId="116" pageNumber="117">
<materialsCitation id="D9ED3C274A1EFFF0FF1FFE45FEA80D46" collectingDate="1993-11-03" collectionCode="SMNS" collectorName="J. S. Amakye &amp; Afr. Z &amp; Afr. &amp; Ghana" county="St." location="Ghana" municipality="Afegame" pageId="116" pageNumber="117" specimenCount="8" specimenCount-female="8">
<emphasis id="5BF1EA684A1EFFF0FF1FFE45FD8F0E8E" bold="true" box="[199,542,473,499]" pageId="116" pageNumber="117">Additional material studied:</emphasis>
<specimenCount id="7F83FDF34A1EFFF0FDFEFE45FDDB0E8E" box="[550,586,473,499]" pageId="116" pageNumber="117" type="female">3♀</specimenCount>
,
<location id="6C5A60A14A1EFFF0FD81FE45FD340E8E" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FD81FE45FD340E8E" box="[601,677,473,499]" county="St." municipality="Afegame" name="Ghana" pageId="116" pageNumber="117">Ghana</location>
, same data as holotype,
<specimenCount id="7F83FDF34A1EFFF0FC65FE45FC710E8E" box="[957,992,473,499]" pageId="116" pageNumber="117" type="female">4♀</specimenCount>
slides,
<location id="6C5A60A14A1EFFF0FBE0FE45FAB10E8E" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FBE0FE45FAB10E8E" box="[1080,1312,473,499]" county="St." municipality="Afegame" name="Ghana Volta Region" pageId="116" pageNumber="117">Ghana Volta Region</location>
,
<location id="6C5A60A14A1EFFF0FAF4FE45FA090E8E" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FAF4FE45FA090E8E" box="[1324,1432,473,499]" county="St." municipality="Afegame" name="Wli Falls" pageId="116" pageNumber="117">Wli Falls</location>
,
<collectingMunicipality id="895EAC004A1EFFF0FF4FFE61FE900D6A" box="[151,257,509,535]" pageId="116" pageNumber="117">Afegame</collectingMunicipality>
,
<collectingCounty id="805B4EF64A1EFFF0FED3FE61FEB80D6A" box="[267,297,509,535]" pageId="116" pageNumber="117">St.</collectingCounty>
6
<date id="1D3B10BA4A1EFFF0FE9BFE61FE510D6A" box="[323,448,509,535]" pageId="116" pageNumber="117" value="1993-11-03">
<collectingDate id="0D7FE9524A1EFFF0FE9BFE61FE510D6A" box="[323,448,509,535]" pageId="116" pageNumber="117" value="1993-11-03">11.03.1993</collectingDate>
</date>
<collectorName id="C47053AC4A1EFFF0FE1EFE61FDDC0D6A" box="[454,589,509,535]" pageId="116" pageNumber="117">J.S.Amakye</collectorName>
&amp; J.
<collectingMethod id="B0C44E6D4A1EFFF0FD5FFE61FCEF0D65" box="[647,894,509,536]" pageId="116" pageNumber="117">Kjaerendsen light trap</collectingMethod>
(slides
<collectorName id="C47053AC4A1EFFF0FC09FE61FB800D6A" box="[977,1041,509,535]" pageId="116" pageNumber="117">Afr.Z</collectorName>
98\09,
<collectorName id="C47053AC4A1EFFF0FB82FE61FB1B0D6A" box="[1114,1162,509,535]" pageId="116" pageNumber="117">Afr.</collectorName>
Z98\14; GH.03/3,
<collectorName id="C47053AC4A1EFFF0FA8BFE61FA0D0D6A" box="[1363,1436,509,535]" pageId="116" pageNumber="117">Ghana</collectorName>
<specimenCount id="7F83FDF34A1EFFF0FF4FFDBDFF3D0D46" box="[151,172,545,571]" pageId="116" pageNumber="117" type="female"></specimenCount>
14;
<collectionCode id="0F94AEBF4A1EFFF0FF38FDBDFEA00D46" box="[224,305,545,571]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">SMNS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A1EFFF0FE9CFDBDFBC30DDB" collectingDate="1993-11-03" collectionCode="ZMBN" collectorName="J. S. Amakye &amp; Slide Z &amp; St. &amp; J. Kjaerendsen &amp; Ref. No." county="Reserve" elevation="47" latitude="5.44" location="Ghana Volta Region" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" pageId="116" pageNumber="117" specimenCode="NEOP163" specimenCount="11" specimenCount-female="11">
<location id="6C5A60A14A1EFFF0FE9CFDBDFDBD0D46" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FE9CFDBDFDBD0D46" box="[324,556,545,571]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Ghana Volta Region" pageId="116" pageNumber="117">Ghana Volta Region</location>
,
<location id="6C5A60A14A1EFFF0FDEFFDBDFD330D46" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FDEFFDBDFD330D46" box="[567,674,545,571]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Wli Falls" pageId="116" pageNumber="117">Wli Falls</location>
,
<location id="6C5A60A14A1EFFF0FD75FDBDFC860D46" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FD75FDBDFC860D46" box="[685,791,545,571]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Afegame" pageId="116" pageNumber="117">Afegame</location>
,
<location id="6C5A60A14A1EFFF0FCFAFDBDFCD00D46" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FCFAFDBDFCD00D46" box="[802,833,545,571]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="St." pageId="116" pageNumber="117">St.</location>
10
<date id="1D3B10BA4A1EFFF0FCB4FDBDFC7B0D46" box="[876,1002,545,571]" pageId="116" pageNumber="117" value="1993-11-03">
<collectingDate id="0D7FE9524A1EFFF0FCB4FDBDFC7B0D46" box="[876,1002,545,571]" pageId="116" pageNumber="117" value="1993-11-03">11.03.1993</collectingDate>
</date>
<collectorName id="C47053AC4A1EFFF0FC29FDBDFBEA0D46" box="[1009,1147,545,571]" pageId="116" pageNumber="117">J.S.Amakye</collectorName>
, J.
<collectingMethod id="B0C44E6D4A1EFFF0FB7AFDBDFA0C0D46" box="[1186,1437,545,571]" pageId="116" pageNumber="117">Kjaerendsen light trap</collectingMethod>
(
<specimenCount id="7F83FDF34A1EFFF0FF7AFDD9FF240D22" box="[162,181,581,607]" pageId="116" pageNumber="117" type="female"></specimenCount>
slides:
<location id="6C5A60A14A1EFFF0FED2FDD8FEC20D23" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FED2FDD8FEC20D23" box="[266,339,580,606]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Ghana" pageId="116" pageNumber="117">Ghana</location>
<specimenCount id="7F83FDF34A1EFFF0FE82FDD9FEFD0D22" box="[346,364,581,607]" pageId="116" pageNumber="117" type="female"></specimenCount>
2,
<location id="6C5A60A14A1EFFF0FE53FDD8FE450D23" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FE53FDD8FE450D23" box="[395,468,580,606]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Ghana" pageId="116" pageNumber="117">Ghana</location>
<specimenCount id="7F83FDF34A1EFFF0FE02FDD9FE7D0D22" box="[474,492,581,607]" pageId="116" pageNumber="117" type="female"></specimenCount>
4, Z17/73;
<location id="6C5A60A14A1EFFF0FDB2FDD8FD220D23" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FDB2FDD8FD220D23" box="[618,691,580,606]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Ghana" pageId="116" pageNumber="117">Ghana</location>
<specimenCount id="7F83FDF34A1EFFF0FD62FDD9FD5A0D22" box="[698,715,581,607]" pageId="116" pageNumber="117" type="female"></specimenCount>
10;
<specimenCount id="7F83FDF34A1EFFF0FD21FDD9FC8D0D22" box="[761,796,581,607]" pageId="116" pageNumber="117" type="female">1♀</specimenCount>
(
<collectorName id="C47053AC4A1EFFF0FCF3FDD8FCEE0D22" box="[811,895,580,607]" pageId="116" pageNumber="117">Slide Z</collectorName>
17.073); 3♁,
<specimenCount id="7F83FDF34A1EFFF0FBCBFDD9FBA90D22" box="[1043,1080,581,607]" pageId="116" pageNumber="117" type="female">4♀</specimenCount>
,
<location id="6C5A60A14A1EFFF0FB9CFDD8FAAE0D23" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FB9CFDD8FAAE0D23" box="[1092,1343,580,607]" county="Reserve" latitude="5.44" longLatPrecision="783" longitude="-2.07" municipality="Ankasa" name="Ghana Western region" pageId="116" pageNumber="117">Ghana Western region</location>
,
<collectingMunicipality id="895EAC004A1EFFF0FA90FDD8FA0D0D23" box="[1352,1436,580,606]" pageId="116" pageNumber="117">Ankasa</collectingMunicipality>
game prod.
<collectingCounty id="805B4EF64A1EFFF0FEF9FDF5FEEA0DFE" box="[289,379,617,643]" pageId="116" pageNumber="117">Reserve</collectingCounty>
[
<geoCoordinate id="0CB150BD4A1EFFF0FE53FDF4FE470DFE" box="[395,470,616,643]" degrees="5.44" direction="north" orientation="latitude" pageId="116" pageNumber="117" precision="555" value="5.44">5.44N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A1EFFF0FE3CFDF4FDBE0DFF" box="[484,559,616,642]" degrees="2.07" direction="west" orientation="longitude" pageId="116" pageNumber="117" precision="555" value="-2.07">2.07W</geoCoordinate>
,
<quantity id="AE7D9B9F4A1EFFF0FDE5FDF4FDE50DFF" box="[573,628,616,642]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="116" pageNumber="117" unit="m" value="47.0">
<elevation id="E2A8D1494A1EFFF0FDE5FDF4FDE50DFF" box="[573,628,616,642]" metricMagnitude="1" metricUnit="m" metricValue="4.7" pageId="116" pageNumber="117" unit="m" value="47.0">47m</elevation>
</quantity>
], 1016.12.93
<collectorName id="C47053AC4A1EFFF0FCC6FDF4FCAD0DFF" box="[798,828,616,642]" pageId="116" pageNumber="117">St.</collectorName>
12
<collectorName id="C47053AC4A1EFFF0FCB3FDF5FB830DFF" box="[875,1042,616,643]" pageId="116" pageNumber="117">J. Kjaerendsen</collectorName>
&amp; T.
<collectingMethod id="B0C44E6D4A1EFFF0FB80FDF4FAA80DFE" box="[1112,1337,616,643]" pageId="116" pageNumber="117">Andersen light trap</collectingMethod>
; NUFUproject
<collectionCode id="0F94AEBF4A1EFFF0FF37FD11FED10DDA" box="[239,320,653,679]" country="Norway" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="116" pageNumber="117">ZMBN</collectionCode>
<collectorName id="C47053AC4A1EFFF0FE92FD10FE3C0DDA" box="[330,429,652,679]" pageId="116" pageNumber="117">Ref. No.</collectorName>
3 (slides GH1\1820); 1♁ (penis in mv),
<specimenCount id="7F83FDF34A1EFFF0FC43FD10FC2F0DDB" box="[923,958,652,678]" pageId="116" pageNumber="117" type="female">2♀</specimenCount>
(
<specimenCode id="39239E014A1EFFF0FC08FD10FBDB0DDB" box="[976,1098,652,678]" collectionCode="NEOP" pageId="116" pageNumber="117">NEOP163</specimenCode>
)
</materialsCitation>
:
<materialsCitation id="D9ED3C274A1EFFF0FBB8FD10FA970D90" collectingDate="1993-05-11" collectingDateMax="2005-06-08" collectingDateMin="1993-05-11" collectionCode="BYU" collectorName="Ghana Subri R. &amp; Uni Bergen. &amp; K. B. Miller" county="Nkwanta Camp" latitude="5.2761" location="Western Reg." longLatPrecision="5" longitude="-1.7481" municipality="Ankasa Resource Res." pageId="116" pageNumber="117" specimenCount="1" specimenCount-female="1">
<collectorName id="C47053AC4A1EFFF0FBB8FD10FA850DDA" box="[1120,1300,652,679]" pageId="116" pageNumber="117">Ghana Subri R.</collectorName>
[
<geoCoordinate id="0CB150BD4A1EFFF0FAFCFD10FA150DDA" box="[1316,1412,652,679]" degrees="5.22851" direction="north" orientation="latitude" pageId="116" pageNumber="117" precision="1" value="5.22851">5.22851</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A1EFFF0FA4BFD11FF780DB7" degrees="1.7481" direction="west" orientation="longitude" pageId="116" pageNumber="117" precision="5" value="-1.7481">- 1.7481</geoCoordinate>
],
<date id="1D3B10BA4A1EFFF0FF27FD33FEED0DB7" box="[255,380,687,714]" pageId="116" pageNumber="117" value="1993-05-11">
<collectingDate id="0D7FE9524A1EFFF0FF27FD33FEED0DB7" box="[255,380,687,714]" pageId="116" pageNumber="117" value="1993-05-11">05.11.1993</collectingDate>
</date>
<collectingMethod id="B0C44E6D4A1EFFF0FE5EFD2CFE620DB7" box="[390,499,688,714]" pageId="116" pageNumber="117">light trap</collectingMethod>
; NUFU project,
<collectorName id="C47053AC4A1EFFF0FD6FFD2CFCD50DB7" box="[695,836,688,714]" pageId="116" pageNumber="117">Uni Bergen.</collectorName>
4♁,
<specimenCount id="7F83FDF34A1EFFF0FC5AFD2CFC340DB7" box="[898,933,688,714]" pageId="116" pageNumber="117" type="female">1♀</specimenCount>
, GHANA,
<location id="6C5A60A14A1EFFF0FBF4FD2CFB580DB7" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FBF4FD2CFB580DB7" box="[1068,1225,688,714]" county="Nkwanta Camp" latitude="5.2761" longLatPrecision="5" longitude="-1.7481" municipality="Ankasa Resource Res." name="Western Reg." pageId="116" pageNumber="117">Western Reg.</location>
,
<collectingMunicipality id="895EAC004A1EFFF0FB0EFD2CFF5A0D93" pageId="116" pageNumber="117">Ankasa Resource Res.</collectingMunicipality>
,
<collectingCounty id="805B4EF64A1EFFF0FF0FFD4FFE190D93" box="[215,392,723,750]" pageId="116" pageNumber="117">Nkwanta Camp</collectingCounty>
<geoCoordinate id="0CB150BD4A1EFFF0FE48FD4FFDB30D93" box="[400,546,723,750]" degrees="05" direction="north" minutes="16.566" orientation="latitude" pageId="116" pageNumber="117" precision="1" value="5.2761">05°16.566N</geoCoordinate>
002°18342
<date id="1D3B10BA4A1EFFF0FD6CFD48FCDF0D93" box="[692,846,723,750]" pageId="116" pageNumber="117" value="2005-06-07" valueMax="2005-06-08" valueMin="2005-06-07">
<collectingDate id="0D7FE9524A1EFFF0FD6CFD48FCDF0D93" box="[692,846,723,750]" pageId="116" pageNumber="117" value="2005-06-07" valueMax="2005-06-08" valueMin="2005-06-07">78 Jun 2005</collectingDate>
</date>
<collectorName id="C47053AC4A1EFFF0FC8EFD4FFC430D90" box="[854,978,723,749]" pageId="116" pageNumber="117">K.B.Miller</collectorName>
, colr. (
<collectionCode id="0F94AEBF4A1EFFF0FBF8FD48FBCE0D93" box="[1056,1119,724,750]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="116" pageNumber="117" type="University or college">BYU</collectionCode>
, slide Z18.76)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A1EFFF0FAC9FD4FFB800C6F" collectionCode="SMNS" collectorName="J. M. Elouard" country="Guinea-Bissau" location="Kousankoro" pageId="116" pageNumber="117" specimenCount="1">
<emphasis id="5BF1EA684A1EFFF0FAC9FD4FFF690C6C" bold="true" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FAC9FD4FFF630C6C" name="Guinea-Bissau" pageId="116" pageNumber="117">Republic of Guinea</collectingCountry>
:
</emphasis>
1♁,
<location id="6C5A60A14A1EFFF0FEF6FD6BFE260C6C" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FEF6FD6BFE260C6C" box="[302,439,759,785]" country="Guinea-Bissau" name="Kousankoro" pageId="116" pageNumber="117">Kousankoro</location>
no.2, 21.10.84 (
<collectionCode id="0F94AEBF4A1EFFF0FDB3FD6BFD2A0C6C" box="[619,699,759,785]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">SMNS</collectionCode>
, gift
<collectorName id="C47053AC4A1EFFF0FD2AFD65FC110C69" box="[754,896,761,788]" pageId="116" pageNumber="117">J.M.Élouard</collectorName>
; slide 16/95)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A1EFFF0FBC3FD65FD510C4A" collectorName="J. M. Elouard" country="Guinea-Bissau" location="Guinea" pageId="116" pageNumber="117" specimenCount="1">
4♁,
<location id="6C5A60A14A1EFFF0FB93FD6BFB0E0C6C" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FB93FD6BFB0E0C6C" box="[1099,1183,759,785]" country="Guinea-Bissau" name="Guinea" pageId="116" pageNumber="117">Guinea</location>
, piège lumineuse, sans étiquette (slides Z18.66 &amp; 67, gift
<collectorName id="C47053AC4A1EFFF0FDC3FC87FD290C4A" box="[539,696,795,823]" pageId="116" pageNumber="117">J. M. Élouard</collectorName>
)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A1EFFF0FD12FC87FD260C24" collectionCode="USNM" country="Liberia" county="Blickenstaff" elevation="349" latitude="7.11" location="Suakoko" longLatPrecision="782" longitude="-9.59" municipality="Jan" pageId="116" pageNumber="117" specimenCode="USNM 8, 11, 12" specimenCount="2" specimenCount-female="2">
<emphasis id="5BF1EA684A1EFFF0FD12FC87FC2C0C48" bold="true" box="[714,957,795,821]" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FD12FC87FC290C48" box="[714,952,795,821]" name="Liberia" pageId="116" pageNumber="117">Republic of Liberia</collectingCountry>
:
</emphasis>
2♁,
<specimenCount id="7F83FDF34A1EFFF0FC2DFC87FB8B0C48" box="[1013,1050,795,821]" pageId="116" pageNumber="117" type="female">2♀</specimenCount>
,
<location id="6C5A60A14A1EFFF0FBFEFC87FB190C48" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FBFEFC87FB190C48" box="[1062,1160,795,821]" country="Liberia" county="Blickenstaff" latitude="7.11" longLatPrecision="782" longitude="-9.59" municipality="Jan" name="Suakoko" pageId="116" pageNumber="117">Suakoko</location>
[
<geoCoordinate id="0CB150BD4A1EFFF0FB4EFC87FB7A0C48" box="[1174,1259,795,821]" degrees="7.11" direction="north" orientation="latitude" pageId="116" pageNumber="117" precision="555" value="7.11">7.11°N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A1EFFF0FB2EFC87FADD0C4B" box="[1270,1356,795,822]" degrees="9.59" direction="west" orientation="longitude" pageId="116" pageNumber="117" precision="555" value="-9.59">9.59°W</geoCoordinate>
<quantity id="AE7D9B9F4A1EFFF0FA8BFC87FA070C48" box="[1363,1430,795,821]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="116" pageNumber="117" unit="m" value="349.0">
<elevation id="E2A8D1494A1EFFF0FA8BFC87FA070C48" box="[1363,1430,795,821]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="116" pageNumber="117" unit="m" value="349.0">349m</elevation>
</quantity>
] 31
<collectingMunicipality id="895EAC004A1EFFF0FF63FCA3FF710C24" box="[187,224,831,857]" pageId="116" pageNumber="117">Jan</collectingMunicipality>
52
<collectingCounty id="805B4EF64A1EFFF0FED5FCA3FE040C24" box="[269,405,831,857]" pageId="116" pageNumber="117">Blickenstaff</collectingCounty>
(slides
<specimenCode id="39239E014A1EFFF0FE37FCA3FDC60C24" box="[495,599,831,857]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="116" pageNumber="117" type="Museum">USNM 8</specimenCode>
,
<specimenCode id="39239E014A1EFFF0FDBDFCA3FD130C24" box="[613,642,831,857]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="116" pageNumber="117" type="Museum">11</specimenCode>
,
<specimenCode id="39239E014A1EFFF0FD57FCA3FD3F0C24" box="[655,686,831,857]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="116" pageNumber="117" type="Museum">12</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="D9ED3C274A1EFFF0FD1CFCDCFA940C24" box="[708,1285,830,859]" collectingDate="1952-05-03" collectingDateMax="1952-05-09" collectingDateMin="1952-05-03" collectionCode="USNM" country="Liberia" pageId="116" pageNumber="117" specimenCode="USNM 41" specimenCount="1">
1♁, same, except
<date id="1D3B10BA4A1EFFF0FC48FCA3FBA70C24" box="[912,1078,830,857]" pageId="116" pageNumber="117" value="1952-05-03" valueMax="1952-05-09" valueMin="1952-05-03">
<collectingDate id="0D7FE9524A1EFFF0FC48FCA3FBA70C24" box="[912,1078,830,857]" pageId="116" pageNumber="117" value="1952-05-03" valueMax="1952-05-09" valueMin="1952-05-03">39 May 1952</collectingDate>
</date>
(slide
<specimenCode id="39239E014A1EFFF0FB5CFCA3FB6D0C24" box="[1156,1276,831,857]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="116" pageNumber="117" type="Museum">USNM 41</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A1EFFF0FAC9FCA2FABC0C01" collectionCode="USNM" collectorName="Van Zwaluvenburg &amp; McGough" country="Sierra Leone" location="Njala Sierra Leone Afr." pageId="116" pageNumber="117" specimenCode="USNM 13" specimenCount="1">
<emphasis id="5BF1EA684A1EFFF0FAC9FCA2FEAB0C01" bold="true" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FAC9FCA2FEA40C01" name="Sierra Leone" pageId="116" pageNumber="117">Republic of Sierra Leone</collectingCountry>
:
</emphasis>
1♁,
<location id="6C5A60A14A1EFFF0FEA8FCFEFDE50C01" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FEA8FCFEFDE50C01" box="[368,628,866,893]" country="Sierra Leone" name="Njala Sierra Leone Afr." pageId="116" pageNumber="117">Njala Sierra Leone Afr.</location>
XI-16XII-4 35 \
<collectorName id="C47053AC4A1EFFF0FCEFFCFFFB9A0C01" box="[823,1035,866,893]" pageId="116" pageNumber="117">Van Zwaluvenburg</collectorName>
&amp;
<collectorName id="C47053AC4A1EFFF0FBF5FCFEFB0E0C01" box="[1069,1183,866,892]" pageId="116" pageNumber="117">McGough</collectorName>
(
<specimenCode id="39239E014A1EFFF0FB74FCFEFAB20C01" box="[1196,1315,866,892]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="116" pageNumber="117" type="Museum">USNM 13</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A1EFFF0FAEFFCFEFC080CDD" collectingDate="1985-11-28" collectionCode="SMNS" collectorName="Landa" country="Togo" location="Amoutchou" pageId="116" pageNumber="117" specimenCode="Z16.132" specimenCount="1" specimenCount-female="1">
<emphasis id="5BF1EA684A1EFFF0FAEFFCFEFE9D0CDD" bold="true" pageId="116" pageNumber="117">
<collectingCountry id="119276EA4A1EFFF0FAEFFCFEFE960CDD" name="Togo" pageId="116" pageNumber="117">Togolese Republic</collectingCountry>
:
</emphasis>
<specimenCount id="7F83FDF34A1EFFF0FECBFC1AFEA70CDD" box="[275,310,902,928]" pageId="116" pageNumber="117" type="female">1♀</specimenCount>
,
<location id="6C5A60A14A1EFFF0FE9AFC1AFE580CDD" LSID="urn:lsid:plazi:treatment:E12C876C4A19FFF1FF4FF9AAFB5C0887:6C5A60A14A1EFFF0FE9AFC1AFE580CDD" box="[322,457,902,928]" country="Togo" name="Amoutchou" pageId="116" pageNumber="117">Amoutchou</location>
,
<date id="1D3B10BA4A1EFFF0FE0DFC1AFDC40CDC" box="[469,597,902,929]" pageId="116" pageNumber="117" value="1985-11-28">
<collectingDate id="0D7FE9524A1EFFF0FE0DFC1AFDC40CDC" box="[469,597,902,929]" pageId="116" pageNumber="117" value="1985-11-28">28.11.1985</collectingDate>
</date>
, V.
<collectorName id="C47053AC4A1EFFF0FDA6FC1AFD550CDD" box="[638,708,902,928]" pageId="116" pageNumber="117">Landa</collectorName>
(
<collectionCode id="0F94AEBF4A1EFFF0FD0BFC1AFCB20CDD" box="[723,803,902,928]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">SMNS</collectionCode>
,
<specimenCode id="39239E014A1EFFF0FCF6FC1AFC000CDD" box="[814,913,902,928]" collectionCode="SMNS" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="116" pageNumber="117">Z16.132</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FFC36FCCC0C9A" blockId="116.[151,1437,151,2037]" pageId="116" pageNumber="117">
<emphasis id="5BF1EA684A1EFFF0FF1FFC36FEBC0CB9" bold="true" box="[199,301,938,964]" pageId="116" pageNumber="117">Habitus.</emphasis>
WL of males 8.211.0mm, of females 11.713.0mm. Uniformly yellowish ochre, the brown ocellar mark is small. Tip of flagellum faintly infuscate, cercus yellow.
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FFC6DFCF80BE7" blockId="116.[151,1437,151,2037]" pageId="116" pageNumber="117">
<emphasis id="5BF1EA684A1EFFF0FF1FFC6DFE950B76" bold="true" box="[199,260,1009,1035]" pageId="116" pageNumber="117">Male</emphasis>
(
<figureCitation id="F1BE2AFF4A1EFFF0FECAFC6DFE290B76" box="[274,440,1009,1035]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Figs. 294300</figureCitation>
). Sternites unmodified. A triangular sclerite on T7 has a narrow caudal process with single SB at tip and a smooth keel along rear face (
<figureCitation id="F1BE2AFF4A1EFFF0FD45FB89FC930B52" box="[669,770,1045,1071]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 294</figureCitation>
); some SB in the intersegmental fold where the sclerite is articulated with T8. Sclerite on T8 T-shaped, with a distinct hump with numerous SB in front (
<figureCitation id="F1BE2AFF4A1EFFF0FB14FBA5FABE0B29" box="[1228,1327,1081,1108]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 295</figureCitation>
). T9 with a median depression between pilose lateral swellings with many SB. HT10 short, regularly curved in dorsal and lateral views, sharply pointed, mediobasal callus tongue-shaped.
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FFB38FCF90AE9" blockId="116.[151,1437,151,2037]" pageId="116" pageNumber="117">
Penis (
<figureCitation id="F1BE2AFF4A1EFFF0FECBFB38FE270BC3" box="[275,438,1188,1214]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Figs. 296299</figureCitation>
) a straight slender tube, distal third membranous except for long lateral sclerites between paired dorsal spine patches and a pair of more distal ventral spine patches (
<figureCitation id="F1BE2AFF4A1EFFF0FC08FB54FBA00B9F" box="[976,1073,1224,1250]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 300</figureCitation>
). Tips of spines point in opposite directions, spines in the ventrodistal groups point towards the penis base. Base of endophallus dorsomedially (
<figureCitation id="F1BE2AFF4A1EFFF0FAB7FB77FF540A54" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 296</figureCitation>
: eb) with patch of erect slender sharp spines. There are scattered slender spines on the wide endophallus base. Distally, the endophallus is a narrow tube with three spine rows: two regular rows of conical spines dorsally plus a single regular row of more slender and locally also longer spines ventrally (
<figureCitation id="F1BE2AFF4A1EFFF0FC24FACBFBCF0A0C" box="[1020,1118,1367,1393]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 296</figureCitation>
, inset). Differences between rows diminish distally, all continue to the tip of the endophallus.
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FFA02FDF00AC5" blockId="116.[151,1437,151,2037]" box="[199,609,1438,1465]" pageId="116" pageNumber="117">
<emphasis id="5BF1EA684A1EFFF0FF1FFA02FE8F0AC5" bold="true" box="[199,286,1438,1464]" pageId="116" pageNumber="117">Female</emphasis>
. As for the group (
<figureCitation id="F1BE2AFF4A1EFFF0FE2AFA02FDC40AC5" box="[498,597,1438,1464]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 301</figureCitation>
).
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FFA5EFE8309CF" blockId="116.[151,1437,151,2037]" pageId="116" pageNumber="117">
<emphasis id="5BF1EA684A1EFFF0FF1FFA5EFF670AA1" bold="true" box="[199,246,1474,1500]" pageId="116" pageNumber="117">Egg</emphasis>
(
<figureCitation id="F1BE2AFF4A1EFFF0FEDCFA5EFE380AA1" box="[260,425,1474,1500]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Figs. 302304</figureCitation>
). Size 345380µm long, on average 1.85 times longer than wide. Numerous (approximately 30) straight striae, each consisting of a wide rib associated with a slightly narrower rib (
<figureCitation id="F1BE2AFF4A1EFFF0FB4EFA79FB6A097D" box="[1174,1275,1509,1536]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 303</figureCitation>
). The ribs are separated by a single row of micropunctures. All ribs are impunctate, micropyles are located on top of the narrow ribs (similar to
<figureCitation id="F1BE2AFF4A1EFFF0FE90F9B1FE3B093A" box="[328,426,1581,1607]" captionStart="FIGURES 309312" captionStartId="120.[152,270,1800,1824]" captionTargetBox="[199,1388,191,1770]" captionTargetId="figure-32@120.[193,1394,181,1776]" captionTargetPageId="120" captionText="FIGURES 309312. Neoperla spironema n. sp.: 309, 310, holotype; 311, a paratype; both from Ankasa Game. Res; 312, SEMdetail of chorion with the 2 kinds of costae and a micropyle (Ivory Coast, USNM; from Moore, 1991). Scales are 100µm, or as indicated. mi, micropyle; sa, spermatheca; SSt, spermathecal stalk; V, vagina." figureDoi="http://doi.org/10.5281/zenodo.8154112" httpUri="https://zenodo.org/record/8154112/files/figure.png" pageId="116" pageNumber="117">Fig. 312</figureCitation>
, after
<bibRefCitation id="0D144B8B4A1EFFF0FE2AF9B2FD13093A" author="Moore, T." box="[498,642,1581,1608]" pageId="116" pageNumber="117" refId="ref155424" refString="Moore, T. (1991) Eggs of African Neoperla (Plecoptera: Perlidae). Honors Research Paper at Mississippi College, Clinton. Unpublished manuscript, III + 40 pp." type="book" year="1991">Moore 1991</bibRefCitation>
). No collar, the wide ribs end in oval depressions on a thickened ring around the anchor pole, the narrow ribs end between them, slightly more distant from the edge (
<figureCitation id="F1BE2AFF4A1EFFF0FB0BF9CDFAA70916" box="[1235,1334,1617,1643]" captionStart="FIGURES 294304" captionStartId="117.[152,270,1599,1623]" captionTargetBox="[153,1423,184,1555]" captionTargetId="figure-17@117.[152,1435,178,1576]" captionTargetPageId="117" captionText="FIGURES 294304. Neoperla orthonema n. sp.: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines." figureDoi="http://doi.org/10.5281/zenodo.8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="116" pageNumber="117">Fig. 304</figureCitation>
). Anchor cavity shallow. Operculum bluntly rounded, with faint reticulation, low impunctate strips separate finely punctate polygones.
</paragraph>
<paragraph id="693A367A4A1EFFF0FF1FF920FB140889" blockId="116.[151,1437,151,2037]" pageId="116" pageNumber="117">
<emphasis id="5BF1EA684A1EFFF0FF1FF920FE9509AB" bold="true" box="[199,260,1724,1750]" pageId="116" pageNumber="117">DNA</emphasis>
(
<figureCitation id="F1BE2AFF4A1EFFF0FECBF920FE2709AB" box="[275,438,1724,1750]" captionStart-0="FIGURE 491" captionStart-1="FIGURE 492" captionStartId-0="177.[152,255,1835,1859]" captionStartId-1="178.[152,255,1943,1967]" captionTargetBox-0="[176,1411,312,1806]" captionTargetBox-1="[361,1187,182,1910]" captionTargetId-0="graphics-120@177.[176,904,321,1800]" captionTargetPageId-0="177" captionTargetPageId-1="178" captionText-0="FIGURE 491. Codon-model maximum likelihood phylogram based on 83 largely complete mitochondrial genome sequences only (mt-NT). Statistical support values (SH-aLRT, UFBoot) from this and other analyses summarised by squares as explained in the figure. Terminal labels include information on the species, type status, gender, originating country, unique sequence identifier (NEOP###), sequencing approach and total basepairs of 13 mitochondrial, protein-coding genes. Clades of particular interest highlighted with colours that correspond to the clades in other trees. HT = holotype; PT = paratype; PCR = Polymerase Chaine Reaction; WGS = Whole Genome Shotgun." captionText-1="FIGURE 492. Simplified overview of codon-model maximum likelihood phylogram based on 356 mitochondrial genome and COX1 DNA barcode sequences (all-NT). Terminal labels include information on the species only. Clades of particular interest highlighted with colours that correspond to the clades in other trees. The full tree is shown in Figs 493-498." figureDoi-0="http://doi.org/10.5281/zenodo.8154198" figureDoi-1="http://doi.org/10.5281/zenodo.8154202" httpUri-0="https://zenodo.org/record/8154198/files/figure.png" httpUri-1="https://zenodo.org/record/8154202/files/figure.png" pageId="116" pageNumber="117">Figs. 491492</figureCitation>
,
<figureCitation id="F1BE2AFF4A1EFFF0FE19F920FE6109AB" box="[449,496,1724,1750]" captionStart="FIGURE 496" captionStartId="182.[152,255,1882,1906]" captionTargetBox="[481,485,328,1836]" captionTargetPageId="182" captionText="FIGURE 496. Partial (clades G, H) codon-model maximum likelihood phylogram based on 356 mitochondrial genome and COX1 DNA barcode sequences (all-NT). See Fig. 493 for common details." figureDoi="http://doi.org/10.5281/zenodo.10274184" httpUri="https://zenodo.org/record/10274184/files/figure.png" pageId="116" pageNumber="117">496</figureCitation>
). A total of
<specimenCount id="7F83FDF34A1EFFF0FDA8F920FC9909AB" box="[624,776,1724,1750]" pageId="116" pageNumber="117" type="generic">10 specimens</specimenCount>
from
<collectingCountry id="119276EA4A1EFFF0FC93F921FC1509AA" box="[843,900,1725,1751]" name="Togo" pageId="116" pageNumber="117">Togo</collectingCountry>
and
<collectingCountry id="119276EA4A1EFFF0FC61F920FB9309AB" box="[953,1026,1724,1750]" name="Ghana" pageId="116" pageNumber="117">Ghana</collectingCountry>
were sequenced for the COX1 DNA barcode fragment and with the genome-skimming approach, providing limited representation of the geographical distribution of the species. This also includes the male
<typeStatus id="B63E88D84A1EFFF0FD22F89FFCCD0860" box="[762,860,1795,1821]" pageId="116" pageNumber="117" type="holotype">holotype</typeStatus>
from
<collectingCountry id="119276EA4A1EFFF0FC78F89FFC780860" box="[928,1001,1795,1821]" name="Ghana" pageId="116" pageNumber="117">Ghana</collectingCountry>
for the COX1 DNA barcode fragment. The monophyly of the species is very strongly supported (99.7/99/9), but its placement as sister to
<taxonomicName id="AE854DF94A1EFFF0FB05F8B4FA11083C" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[1245,1408,1831,1857]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="filamentosa" status="sp. nov.">
<emphasis id="5BF1EA684A1EFFF0FB05F8B4FA11083C" box="[1245,1408,1831,1857]" italics="true" pageId="116" pageNumber="117">N. filamentosa</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1EFFF0FA5EF8BBFF290818" bold="true" pageId="116" pageNumber="117">
<taxonomicNameLabel id="40C257134A1EFFF0FA5EF8BBFF290818" pageId="116" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is only poorly supported (43.9/90/0). The poor support for this placement is linked to the inclusion of
<collectionCode id="0F94AEBF4A1EFFF0FAFAF8D7FAE10818" box="[1314,1392,1867,1893]" pageId="116" pageNumber="117">NEOP</collectionCode>
167, which includes particularly badly fragmented mitochondrial genome assembly data and has the potential for the inadvertent inclusion of cross contaminant fragments, potentially resulting in instability of the placement of this specimen relative to
<taxonomicName id="AE854DF94A1EFFF0FE59F82BFDB408AD" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[385,549,1974,2000]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="filamentosa" status="sp. nov.">
<emphasis id="5BF1EA684A1EFFF0FE59F82BFDB408AD" box="[385,549,1974,2000]" italics="true" pageId="116" pageNumber="117">N. filamentosa</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1EFFF0FDF4F82AFDFA08AD" bold="true" box="[556,619,1974,2000]" pageId="116" pageNumber="117">
<taxonomicNameLabel id="40C257134A1EFFF0FDF4F82AFDFA08AD" box="[556,619,1974,2000]" pageId="116" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
.. Together,
<taxonomicName id="AE854DF94A1EFFF0FD29F82BFC1A08AD" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[753,907,1974,2000]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="orthonema" status="sp. nov.">
<emphasis id="5BF1EA684A1EFFF0FD29F82BFC1A08AD" box="[753,907,1974,2000]" italics="true" pageId="116" pageNumber="117">N. orthonema</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1EFFF0FC4AF82AFC4008AD" bold="true" box="[914,977,1974,2000]" pageId="116" pageNumber="117">
<taxonomicNameLabel id="40C257134A1EFFF0FC4AF82AFC4008AD" box="[914,977,1974,2000]" pageId="116" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="AE854DF94A1EFFF0FBD1F82BFB3C08AD" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[1033,1197,1974,2000]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="filamentosa" status="sp. nov.">
<emphasis id="5BF1EA684A1EFFF0FBD1F82BFB3C08AD" box="[1033,1197,1974,2000]" italics="true" pageId="116" pageNumber="117">N. filamentosa</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1EFFF0FB6CF82AFB6208AD" bold="true" box="[1204,1267,1974,2000]" pageId="116" pageNumber="117">
<taxonomicNameLabel id="40C257134A1EFFF0FB6CF82AFB6208AD" box="[1204,1267,1974,2000]" pageId="116" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are maximally supported (100/100/100) as sister to
<taxonomicName id="AE854DF94A1EFFF0FDE9F847FD560889" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[561,711,2011,2036]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="spironema" status="sp. nov.">
<emphasis id="5BF1EA684A1EFFF0FDE9F847FD560889" box="[561,711,2011,2036]" italics="true" pageId="116" pageNumber="117">N. spironema</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1EFFF0FD16F846FC9C0889" bold="true" box="[718,781,2010,2036]" pageId="116" pageNumber="117">
<taxonomicNameLabel id="40C257134A1EFFF0FD16F846FC9C0889" box="[718,781,2010,2036]" pageId="116" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. Both sexes are well represented.
</paragraph>
<caption id="3DFA66F24A1FFFF1FF40F9A3FD220852" ID-DOI="http://doi.org/10.5281/zenodo.8154106" ID-Zenodo-Dep="8154106" httpUri="https://zenodo.org/record/8154106/files/figure.png" pageId="117" pageNumber="118" startId="117.[152,270,1599,1623]" targetBox="[153,1423,184,1555]" targetPageId="117" targetType="figure">
<paragraph id="693A367A4A1FFFF1FF40F9A3FD220852" blockId="117.[151,1436,1599,1840]" pageId="117" pageNumber="118">
<emphasis id="5BF1EA684A1FFFF1FF40F9A3FEE7092A" bold="true" box="[152,374,1599,1623]" pageId="117" pageNumber="118">FIGURES 294304.</emphasis>
<taxonomicName id="AE854DF94A1FFFF1FE58F9A3FDC7092A" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[384,598,1599,1623]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="orthonema" status="sp. nov.">
<emphasis id="5BF1EA684A1FFFF1FE58F9A3FDC7092A" box="[384,598,1599,1623]" italics="true" pageId="117" pageNumber="118">Neoperla orthonema</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1FFFF1FD87F9DCFD0A0925" bold="true" box="[607,667,1600,1624]" pageId="117" pageNumber="118">
<taxonomicNameLabel id="40C257134A1FFFF1FD87F9DCFD0A0925" box="[607,667,1600,1624]" pageId="117" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: Male paratype from Lakota, Ivory Coast: 294, 295, abdominal tip; 296, everted penis, with detail from middle of endophallus; 297299, apex of contracted penis, in dorsal, ventral and lateral views (from left); 300, lateral view of penis apex, beginning eversion (♁ from Kousankoro); 301, female genital situs (vagina flipped back to show the anchor pattern on S8) and 302, egg, Ankasa game prod. res., Ghana; 303, view onto edge of chorion fragment showing variable size of costae (Ivory Coast, Lakota); 304, detail of collar with end of striae (Wli Falls, Ghana). Scales are 100µm. Tergites, sternites and hemitergite 10 are labeled. d, dorsal spines; eb, spines at base of endophallus; sa, spermatheca; SSt, spermathecal stalk; V, vagina; vs, ventral spines.
</paragraph>
</caption>
<paragraph id="693A367A4A1FFFF1FF1FF8E9FAE008CF" blockId="117.[151,1437,1908,2043]" pageId="117" pageNumber="118">
<emphasis id="5BF1EA684A1FFFF1FF1FF8E9FE9B08F2" bold="true" box="[199,266,1909,1935]" pageId="117" pageNumber="118">Notes</emphasis>
. In all species of
<taxonomicName id="AE854DF94A1FFFF1FE09F8E8FDA808F3" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[465,569,1908,1934]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="5BF1EA684A1FFFF1FE09F8E8FDA808F3" box="[465,569,1908,1934]" italics="true" pageId="117" pageNumber="118">Neoperla</emphasis>
</taxonomicName>
, micropyles are at the bottom of sulci but in both
<taxonomicName id="AE854DF94A1FFFF1FBB3F8E9FA9408F3" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[1131,1285,1908,1934]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="orthonema" status="sp. nov.">
<emphasis id="5BF1EA684A1FFFF1FBB3F8E9FA9408F3" box="[1131,1285,1908,1934]" italics="true" pageId="117" pageNumber="118">N. orthonema</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1FFFF1FAD4F8E9FADA08F2" bold="true" box="[1292,1355,1909,1935]" pageId="117" pageNumber="118">
<taxonomicNameLabel id="40C257134A1FFFF1FAD4F8E9FADA08F2" box="[1292,1355,1909,1935]" pageId="117" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="AE854DF94A1FFFF1FA5BF8E9FE9D08CF" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="spironema" status="sp. nov.">
<emphasis id="5BF1EA684A1FFFF1FA5BF8E9FE9D08CF" italics="true" pageId="117" pageNumber="118">N. spironema</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A1FFFF1FECBF805FEC308CE" bold="true" box="[275,338,1945,1971]" pageId="117" pageNumber="118">
<taxonomicNameLabel id="40C257134A1FFFF1FECBF805FEC308CE" box="[275,338,1945,1971]" pageId="117" pageNumber="118" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
they are on top of narrow low ribs (
<figureCitation id="F1BE2AFF4A1FFFF1FD3FF804FCD808CE" box="[743,841,1944,1971]" captionStart="FIGURES 309312" captionStartId="120.[152,270,1800,1824]" captionTargetBox="[199,1388,191,1770]" captionTargetId="figure-32@120.[193,1394,181,1776]" captionTargetPageId="120" captionText="FIGURES 309312. Neoperla spironema n. sp.: 309, 310, holotype; 311, a paratype; both from Ankasa Game. Res; 312, SEMdetail of chorion with the 2 kinds of costae and a micropyle (Ivory Coast, USNM; from Moore, 1991). Scales are 100µm, or as indicated. mi, micropyle; sa, spermatheca; SSt, spermathecal stalk; V, vagina." figureDoi="http://doi.org/10.5281/zenodo.8154112" httpUri="https://zenodo.org/record/8154112/files/figure.png" pageId="117" pageNumber="118">Fig. 312</figureCitation>
), which apparently are secondarily uplifted sulci.
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A1FFFF1FF1FF821FB5C0887" pageId="117" pageNumber="118" type="etymology">
<paragraph id="693A367A4A1FFFF1FF1FF821FB5C0887" blockId="117.[151,1437,1908,2043]" pageId="117" pageNumber="118">
<emphasis id="5BF1EA684A1FFFF1FF1FF821FED908AA" bold="true" box="[199,328,1981,2007]" pageId="117" pageNumber="118">Etymology</emphasis>
. The name is a composite noun in apposition, formed from the Greek stems
<emphasis id="5BF1EA684A1FFFF1FB1BF820FA9108AB" box="[1219,1280,1980,2006]" italics="true" pageId="117" pageNumber="118">ortho</emphasis>
, straight, and
<emphasis id="5BF1EA684A1FFFF1FF4FF87DFF450887" box="[151,212,2017,2042]" italics="true" pageId="117" pageNumber="118">nema</emphasis>
, thread, alluding to the straight egg striae and the threads supporting the cap of the anchor.
</paragraph>
</subSubSection>
</treatment>
</document>