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<document id="3784BD47D7E94E792B3520EB70BDCFA0" ID-DOI="10.11646/zootaxa.5316.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8154005" ID-ZooBank="BC922E16-2614-4F3D-AD82-87A845DE7E2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1689581758737" checkinUser="plazi" docAuthor="Zwick, Peter &amp; Zwick, Andreas" docDate="2023" docId="E12C876C4A11FFF8FF4FF98EFB2D0BB8" docLanguage="en" docName="zootaxa.5316.1.1.pdf" docOrigin="Zootaxa 5316 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5316.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neoperla proxima Zwick &amp; Zwick 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="125" masterDocId="1D15FF144A6AFF84FFD8FF9CFF910F7D" masterDocTitle="Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data" masterLastPageNumber="194" masterPageNumber="1" pageNumber="124" updateTime="1699540694688" updateUser="plazi">
<mods:mods id="BBEE63A97BC2F9CFEAEE74BB5B91DF09" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A3AA7ABDCB3BADF115653A555AD449BE">
<mods:title id="6C037B6049A20A7733F0322690BBC24F">Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data</mods:title>
</mods:titleInfo>
<mods:name id="E625E28C9E533D04A4ED53B9847E2C24" type="personal">
<mods:role id="9EACE414C65EC26D83AF9D53D0868012">
<mods:roleTerm id="0D0B1F9DDD54F51A6F82FF2728228D7B">Author</mods:roleTerm>
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<mods:namePart id="53D6B9BE02F30EF848EBC78D8596FAE9">Zwick, Peter</mods:namePart>
<mods:nameIdentifier id="B92CDC620BD2E8B9FE7D6AF9BDD18086" type="ORCID">0000-0002-6730-0011</mods:nameIdentifier>
<mods:nameIdentifier id="E4C52AB59B558B438CE95FB86B2D95EC" type="email">pleco-p.zwick@t-online.de</mods:nameIdentifier>
</mods:name>
<mods:name id="5F5C2654FD879B7668310A931CF0B41B" type="personal">
<mods:role id="C58DDC7897A3C3B59E1466F6FD7EDBA5">
<mods:roleTerm id="9E52734E0E5681E224202A31AD71464F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CBAC58CE512A2BADEE069EE5E6060ABA">Zwick, Andreas</mods:namePart>
<mods:nameIdentifier id="1A39E5AD92848E669B155ACF918FFFDC" type="ORCID">0000-0002-7532-1752</mods:nameIdentifier>
<mods:nameIdentifier id="8ABE403AD4284ABDC3FA3E6B6CE55D4B" type="email">andreas.zwick@csiro.au</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8ED82C541842F0143429E325F1056CB6">text</mods:typeOfResource>
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<mods:titleInfo id="09679B7A5372686CDF9DA2B1CED686D3">
<mods:title id="C84CFFFBA2112CDF9029AD3AE2A9EDC7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7B065B4FCCB8DD1660A813D91291E82F">
<mods:date id="B14A9F9AFAB02758783A302B3177A55D">2023</mods:date>
<mods:detail id="14D5C975ACC40A90D7225A26AE16B8F6" type="pubDate">
<mods:number id="0CB153157510F6548D8B29B0F442ED6B">2023-07-17</mods:number>
</mods:detail>
<mods:detail id="1EB36A4C3CEE7CCE4A0C898708EACEF2" type="volume">
<mods:number id="C0C149C7FB4D9F78477B678FD64C5C6C">5316</mods:number>
</mods:detail>
<mods:detail id="399F0715F60932CF340D93B31A05B321" type="issue">
<mods:number id="A739005EA915706B45C88BD9AAF59941">1</mods:number>
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<mods:start id="610D8494D2C73C3C6DB81BC7FE320979">1</mods:start>
<mods:end id="2C0FC868F8C18F8039F816E8F0B6DF28">194</mods:end>
</mods:extent>
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<mods:location id="C84866455158F833E3A83D2FF208E24F">
<mods:url id="CFA15E9122CA31026BD679A6D915664B">http://dx.doi.org/10.11646/zootaxa.5316.1.1</mods:url>
</mods:location>
<mods:classification id="D10652D4CBF47D23C1FDEF81B210B157">journal article</mods:classification>
<mods:identifier id="229F99F3482BCFC3F6D03FDEC68B60DB" type="DOI">10.11646/zootaxa.5316.1.1</mods:identifier>
<mods:identifier id="EA9B97E8F8707224AE78CD6105F4B1E4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="84D7EF14F723BD19DA9D9B6612A5A305" type="Zenodo-Dep">8154005</mods:identifier>
<mods:identifier id="31DC0A790B71A5C9AEEEA3D4CBFA8424" type="ZooBank">BC922E16-2614-4F3D-AD82-87A845DE7E2B</mods:identifier>
</mods:mods>
<treatment id="E12C876C4A11FFF8FF4FF98EFB2D0BB8" LSID="urn:lsid:plazi:treatment:E12C876C4A11FFF8FF4FF98EFB2D0BB8" httpUri="http://treatment.plazi.org/id/E12C876C4A11FFF8FF4FF98EFB2D0BB8" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
<subSubSection id="219F65F14A11FFFFFF4FF98EFE790950" box="[151,488,1554,1581]" pageId="123" pageNumber="124" type="nomenclature">
<paragraph id="693A367A4A11FFFFFF4FF98EFE790950" blockId="123.[151,488,1554,1617]" box="[151,488,1554,1581]" pageId="123" pageNumber="124">
<emphasis id="5BF1EA684A11FFFFFF4FF98EFE790950" bold="true" box="[151,488,1554,1581]" pageId="123" pageNumber="124">
56.
<taxonomicName id="AE854DF94A11FFFFFF1BF98EFE0F0950" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[195,414,1554,1581]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="proxima" status="sp. nov.">
<emphasis id="5BF1EA684A11FFFFFF1BF98EFE0F0950" bold="true" box="[195,414,1554,1581]" italics="true" pageId="123" pageNumber="124">Neoperla proxima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C257134A11FFFFFE7EF98FFE790950" box="[422,488,1555,1581]" pageId="123" pageNumber="124" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A11FFFFFF4FF9ABFE2B092C" box="[151,442,1591,1617]" pageId="123" pageNumber="124" type="description">
<paragraph id="693A367A4A11FFFFFF4FF9ABFE2B092C" blockId="123.[151,488,1554,1617]" box="[151,442,1591,1617]" pageId="123" pageNumber="124">
(
<figureCitation id="F1BE2AFF4A11FFFFFF47F9ABFED5092C" box="[159,324,1591,1617]" captionStart="FIGURES 313–322" captionStartId="122.[152,270,1787,1811]" captionTargetBox="[251,1336,187,1757]" captionTargetId="figure-32@122.[246,1341,181,1762]" captionTargetPageId="122" captionText="FIGURES 313–322. Males in the Neoperla dubia-complex: N. dubia Klapálek (from Makak, Cameroon): 313, terminal segments, dorsal view; 314, 315, enlarged centre of tergites 7 and 8, dorsal and lateral; 316, contracted, 317, everted penis with detail of distal armature (not to scale, from Ikom, Nigeria). Neoperla proxima n. sp.: 318, abdominal tip in dorsal view; 317, tip of penis with everted endophallus (D. R. Congo, Odzala N. P.). Neoperla vicina n. sp.: 320, 321, abdominal tip, dorsal and lateral views; 322, penis tip with everted endophallus in lateral view, and a detail of external spines in oblique lateral view (D. R. Congo, Bukavu). Scales in 317 and 322 are 100µm, scale in 320 is 0.5mm. The tergites, hemitergite 10 and the cercus are labeled." figureDoi="http://doi.org/10.5281/zenodo.8154118" httpUri="https://zenodo.org/record/8154118/files/figure.png" pageId="123" pageNumber="124">Figs. 318–319</figureCitation>
,
<figureCitation id="F1BE2AFF4A11FFFFFE97F9ABFE22092C" box="[335,435,1591,1617]" captionStart="FIGURES 323–331" captionStartId="125.[152,270,1838,1862]" captionTargetBox="[235,1343,189,1806]" captionTargetId="figure-17@125.[229,1357,181,1814]" captionTargetPageId="125" captionText="FIGURES 323–331. Neoperla dubia-complex, femalesand eggs.N.dubia Klapálek:323, genital situs; 324, eggwith moderately raised costae (Ghana, Agumatsa waterfalls); 325–327, egg with flanged costae (D. R. Congo, Odzala N. P.). Neoperla proxima n. sp.: 328, genital situs (D. R.Congo, N of Matadi), 329, egg (Cameroon, Sangmelina). Neoperla vicina n. sp. (D.R. Congo): 330, female situs, 331,egg, arrow points at end of raised part of costa (paratype, Walikale, Z2101). Scales are 100µm, similar structures shown to the same scale. Top view of egg (326) based on combined halves of a photograph. Sa, spermatheca; SSt, spermathecal stalk; V, vagina." figureDoi="http://doi.org/10.5281/zenodo.8154124" httpUri="https://zenodo.org/record/8154124/files/figure.png" pageId="123" pageNumber="124">328–329</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A11FFF8FF4FF9E3FD800C44" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="693A367A4A11FFFFFF4FF9E3FEE7080C" blockId="123.[151,1437,1662,2013]" pageId="123" pageNumber="124">
<emphasis id="5BF1EA684A11FFFFFF4FF9E3FED109E5" bold="true" box="[151,320,1662,1689]" pageId="123" pageNumber="124">Type material</emphasis>
:
<materialsCitation id="D9ED3C274A11FFFFFE89F9E3FEE7080C" collectingDate="1997-03-03" collectingDateMax="2013-10-24" collectingDateMin="1997-03-03" collectionCode="ZMHUB, SMNS, SMTD" collectorName="V. Sinaev &amp; M. Nuss &amp; Cameroon &amp; Sangmelima &amp; Mintom &amp; A. Zwick &amp; D. Bartsch" country="Democratic Republic of the Congo" county="Kongo" elevation="560" latitude="2.7886221" location="Dja River" longLatPrecision="1" longitude="11.97896" municipality="Odzala Nat. Park" pageId="123" pageNumber="124" specimenCode="NEOP257, NEOP258, PZ17.46, NEOP259, NEOP260, NEOP261, NEOP262, NEOP263, NEOP264, NEOP265, NEOP266" specimenCount="11" specimenCount-female="9" stateProvince="Uige" typeStatus="holotype">
<emphasis id="5BF1EA684A11FFFFFE89F9E3FC0A09E5" bold="true" box="[337,923,1662,1689]" pageId="123" pageNumber="124">
<typeStatus id="B63E88D84A11FFFFFE89F9E3FE2F09E4" box="[337,446,1663,1689]" pageId="123" pageNumber="124" type="holotype">Holotype</typeStatus>
<emphasis id="5BF1EA684A11FFFFFE1FF9E3FE7209E4" bold="true" box="[455,483,1663,1689]" italics="true" pageId="123" pageNumber="124">J,</emphasis>
<collectingCountry id="119276EA4A11FFFFFE35F9E2FC0709E5" box="[493,918,1662,1689]" name="Democratic Republic of the Congo" pageId="123" pageNumber="124">Democratic Republic of the Congo</collectingCountry>
:
</emphasis>
<collectingCounty id="805B4EF64A11FFFFFC7DF9E3FC6509E4" box="[933,1012,1663,1689]" pageId="123" pageNumber="124">Kongo</collectingCounty>
,
<collectingMunicipality id="895EAC004A11FFFFFBD9F9E3FB5409E4" box="[1025,1221,1663,1689]" pageId="123" pageNumber="124">Odzala Nat. Park</collectingMunicipality>
[
<geoCoordinate id="0CB150BD4A11FFFFFB0EF9E3FABA09E4" box="[1238,1323,1663,1689]" degrees="1.19" direction="north" orientation="latitude" pageId="123" pageNumber="124" precision="555" value="1.19">1.19°N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A11FFFFFAE1F9E2FA0909E4" box="[1337,1432,1662,1689]" degrees="14.85" direction="east" orientation="longitude" pageId="123" pageNumber="124" precision="555" value="14.85">14.85°E</geoCoordinate>
;
<quantity id="AE7D9B9F4A11FFFFFF4FF93EFF4D09C0" box="[151,220,1698,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="123" pageNumber="124" unit="m" value="560.0">
<elevation id="E2A8D1494A11FFFFFF4FF93EFF4D09C0" box="[151,220,1698,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.6" pageId="123" pageNumber="124" unit="m" value="560.0">560m</elevation>
</quantity>
], 29.9.–
<date id="1D3B10BA4A11FFFFFEEBF93FFE0609C0" box="[307,407,1699,1725]" pageId="123" pageNumber="124" value="1997-03-03">
<collectingDate id="0D7FE9524A11FFFFFEEBF93FFE0609C0" box="[307,407,1699,1725]" pageId="123" pageNumber="124" value="1997-03-03">3.3.1997</collectingDate>
</date>
leg.
<collectorName id="C47053AC4A11FFFFFE09F93FFDA409C0" box="[465,565,1699,1725]" pageId="123" pageNumber="124">V.Sinaev</collectorName>
(
<specimenCode id="39239E014A11FFFFFD9DF93EFD2C09C0" box="[581,701,1698,1725]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP257</specimenCode>
;
<collectionCode id="0F94AEBF4A11FFFFFD11F93FFCA209C0" box="[713,819,1699,1725]" pageId="123" pageNumber="124">ZMHUB</collectionCode>
).
<specimenCount id="7F83FDF34A11FFFFFC9FF93FFC5909C0" box="[839,968,1699,1725]" pageId="123" pageNumber="124" type="generic" typeStatus="paratype">
2
<emphasis id="5BF1EA684A11FFFFFC85F93FFC5909C0" bold="true" box="[861,968,1699,1725]" pageId="123" pageNumber="124">paratype</emphasis>
</specimenCount>
<emphasis id="5BF1EA684A11FFFFFC09F93FFC7C09C1" box="[977,1005,1699,1724]" italics="true" pageId="123" pageNumber="124">J,</emphasis>
the same data (
<collectionCode id="0F94AEBF4A11FFFFFB78F93FFA9609C0" box="[1184,1287,1699,1725]" pageId="123" pageNumber="124">ZMHUB</collectionCode>
and
<collectionCode id="0F94AEBF4A11FFFFFA98F93FFA0009C0" box="[1344,1425,1699,1725]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="123" pageNumber="124">SMNS</collectionCode>
).
<emphasis id="5BF1EA684A11FFFFFF4FF95AFDA0099C" bold="true" box="[151,561,1734,1764]" pageId="123" pageNumber="124">
Republic of Angola.
<specimenCount id="7F83FDF34A11FFFFFE4AF954FE240999" box="[402,437,1736,1764]" pageId="123" pageNumber="124" type="female">1♀</specimenCount>
<typeStatus id="B63E88D84A11FFFFFE67F95BFDBD099C" box="[447,556,1735,1761]" pageId="123" pageNumber="124" type="paratype">paratype</typeStatus>
,
</emphasis>
Angola, prov.
<collectingRegion id="AB41F8984A11FFFFFD07F95BFC89099C" box="[735,792,1735,1761]" country="Angola" name="Uige" pageId="123" pageNumber="124">Uíge</collectingRegion>
, Finca
<geoCoordinate id="0CB150BD4A11FFFFFCB6F95BFC6E099C" box="[878,1023,1735,1761]" degrees="7" direction="south" minutes="38" orientation="latitude" pageId="123" pageNumber="124" precision="1" seconds="08.6" value="-7.635722">7°38’08.6’’S</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A11FFFFFBD6F95AFB3C099C" box="[1038,1197,1734,1761]" degrees="14" direction="east" minutes="56" orientation="longitude" pageId="123" pageNumber="124" precision="1" seconds="25.8" value="14.9405">14°56’25.8’’E</geoCoordinate>
<quantity id="AE7D9B9F4A11FFFFFB6FF95AFB68099C" box="[1207,1273,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="123" pageNumber="124" unit="m" value="645.0">
<elevation id="E2A8D1494A11FFFFFB6FF95AFB68099C" box="[1207,1273,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="123" pageNumber="124" unit="m" value="645.0">645m</elevation>
</quantity>
,
<date id="1D3B10BA4A11FFFFFADFF95BFAEB099C" box="[1287,1402,1735,1761]" pageId="123" pageNumber="124" value="2013-10-24">
<collectingDate id="0D7FE9524A11FFFFFADFF95BFAEB099C" box="[1287,1402,1735,1761]" pageId="123" pageNumber="124" value="2013-10-24">24.x.2013</collectingDate>
</date>
, at light, leg.
<collectorName id="C47053AC4A11FFFFFED7F977FEF70878" box="[271,358,1771,1797]" pageId="123" pageNumber="124">M.Nuss</collectorName>
(
<specimenCode id="39239E014A11FFFFFEA0F976FE610878" box="[376,496,1770,1797]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP258</specimenCode>
,
<collectionCode id="0F94AEBF4A11FFFFFE26F977FDDE0878" box="[510,591,1771,1797]" pageId="123" pageNumber="124">SMTD</collectionCode>
; slide
<specimenCode id="39239E014A11FFFFFD44F977FC900878" box="[668,769,1771,1797]" collectionCode="PZ" pageId="123" pageNumber="124">PZ17.46</specimenCode>
).
<emphasis id="5BF1EA684A11FFFFFCCEF976FBA50879" bold="true" box="[790,1076,1770,1796]" pageId="123" pageNumber="124">
<collectingCountry id="119276EA4A11FFFFFCCEF976FBBE0879" box="[790,1071,1770,1796]" name="Cameroon" pageId="123" pageNumber="124">Republic of Cameroon</collectingCountry>
:
</emphasis>
<specimenCount id="7F83FDF34A11FFFFFBE6F977FBF00878" box="[1086,1121,1771,1797]" pageId="123" pageNumber="124" type="female">8♀</specimenCount>
<emphasis id="5BF1EA684A11FFFFFBB3F977FB700878" bold="true" box="[1131,1249,1771,1797]" pageId="123" pageNumber="124">
<typeStatus id="B63E88D84A11FFFFFBB3F977FB700878" box="[1131,1249,1771,1797]" pageId="123" pageNumber="124" type="paratype">paratypes</typeStatus>
</emphasis>
:
<geoCoordinate id="0CB150BD4A11FFFFFB2BF977FA060878" box="[1267,1431,1771,1797]" degrees="2" direction="north" minutes="47" orientation="latitude" pageId="123" pageNumber="124" precision="1" seconds="19.04" value="2.7886221">2°47’19.04’’N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A11FFFFFF4FF893FE8A0854" box="[151,283,1807,1833]" degrees="11.97896" direction="east" orientation="longitude" pageId="123" pageNumber="124" precision="1" value="11.97896">11.97896 E</geoCoordinate>
,
<collectorName id="C47053AC4A11FFFFFEF1F893FE300854" box="[297,417,1807,1833]" pageId="123" pageNumber="124">Cameroon</collectorName>
, S-Reg.,
<collectorName id="C47053AC4A11FFFFFDD3F893FD060854" box="[523,663,1807,1833]" pageId="123" pageNumber="124">Sangmélima</collectorName>
<quantity id="AE7D9B9F4A11FFFFFD78F893FD4E0854" box="[672,735,1807,1833]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="123" pageNumber="124" unit="km" value="17.0">17km</quantity>
on road to
<location id="6C5A60A14A11FFFFFCBFF893FC470854" LSID="urn:lsid:plazi:treatment:E12C876C4A11FFF8FF4FF98EFB2D0BB8:6C5A60A14A11FFFFFCBFF893FC470854" box="[871,982,1807,1833]" country="Democratic Republic of the Congo" county="Kongo" latitude="2.7886221" longLatPrecision="1" longitude="11.97896" municipality="Odzala Nat. Park" name="Dja River" pageId="123" pageNumber="124" stateProvince="Uige">Dja River</location>
NEbN
<collectorName id="C47053AC4A11FFFFFBE8F893FB1C0854" box="[1072,1165,1807,1833]" pageId="123" pageNumber="124">Mintom</collectorName>
,
<elevation id="E2A8D1494A11FFFFFB42F892FA940854" box="[1178,1285,1806,1833]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="123" pageNumber="124" unit="m" value="570.0">
<quantity id="AE7D9B9F4A11FFFFFB42F892FB4B0854" box="[1178,1242,1806,1833]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="123" pageNumber="124" unit="m" value="570.0">570m</quantity>
asl
</elevation>
,
<date id="1D3B10BA4A11FFFFFACAF893FA090854" box="[1298,1432,1807,1833]" pageId="123" pageNumber="124" value="2011-11-04">
<collectingDate id="0D7FE9524A11FFFFFACAF893FA090854" box="[1298,1432,1807,1833]" pageId="123" pageNumber="124" value="2011-11-04">4 Nov.2011</collectingDate>
</date>
, MV-lamp,
<collectorName id="C47053AC4A11FFFFFEC8F8AFFEE20830" box="[272,371,1843,1869]" pageId="123" pageNumber="124">A.Zwick</collectorName>
&amp;
<collectorName id="C47053AC4A11FFFFFE4FF8AFFD990830" box="[407,520,1843,1869]" pageId="123" pageNumber="124">D.Bartsch</collectorName>
(
<specimenCode id="39239E014A11FFFFFDCFF8AEFD1E0830" box="[535,655,1842,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP259</specimenCode>
,
<specimenCode id="39239E014A11FFFFFD42F8AFFC810830" box="[666,784,1843,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP260</specimenCode>
,
<specimenCode id="39239E014A11FFFFFCC4F8AFFC030830" box="[796,914,1843,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP261</specimenCode>
,
<specimenCode id="39239E014A11FFFFFC45F8AFFB820830" box="[925,1043,1843,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP262</specimenCode>
,
<specimenCode id="39239E014A11FFFFFBC7F8AFFB040830" box="[1055,1173,1843,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP263</specimenCode>
,
<specimenCode id="39239E014A11FFFFFB79F8AFFA870830" box="[1185,1302,1843,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP264</specimenCode>
,
<specimenCode id="39239E014A11FFFFFAFAF8AEFA080830" box="[1314,1433,1842,1869]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP265</specimenCode>
,
<specimenCode id="39239E014A11FFFFFF4FF8CBFE9F080C" box="[151,270,1879,1905]" collectionCode="NEOP" pageId="123" pageNumber="124">NEOP266</specimenCode>
;
<collectionCode id="0F94AEBF4A11FFFFFEC1F8CBFEFB080C" box="[281,362,1879,1905]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="123" pageNumber="124">SMNS</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="693A367A4A11FFFFFF1FF8E6FA0D08A0" blockId="123.[151,1437,1662,2013]" pageId="123" pageNumber="124">
<emphasis id="5BF1EA684A11FFFFFF1FF8E6FD8808E9" bold="true" box="[199,537,1914,1940]" pageId="123" pageNumber="124">Additional material studied:</emphasis>
<materialsCitation id="D9ED3C274A11FFFFFDC7F8E6FB8A08C4" collectingDate="1939-05" collectionCode="SMNS, MRAC" collectorName="Coll. Mus. Congo W. Kivu Lubongola &amp; Dr. Hautman" country="Democratic Republic of the Congo" latitude="-2.98693" location="Shabunda" longLatPrecision="1" longitude="27.55371" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1">
<emphasis id="5BF1EA684A11FFFFFDC7F8E6FC2808E9" bold="true" box="[543,953,1914,1941]" pageId="123" pageNumber="124">
<collectingCountry id="119276EA4A11FFFFFDC7F8E6FC2808E9" box="[543,953,1914,1941]" name="Democratic Republic of the Congo" pageId="123" pageNumber="124">Democratic Republic of the Congo</collectingCountry>
</emphasis>
:
<specimenCount id="7F83FDF34A11FFFFFC1FF8E7FC7D08E8" box="[967,1004,1915,1941]" pageId="123" pageNumber="124" type="female">1♀</specimenCount>
:
<collectorName id="C47053AC4A11FFFFFC20F8E7FA0D08E8" box="[1016,1436,1915,1941]" pageId="123" pageNumber="124">Coll. Mus. Congo W.Kivu Lubongola</collectorName>
pr.
<location id="6C5A60A14A11FFFFFF67F803FEBC08C4" LSID="urn:lsid:plazi:treatment:E12C876C4A11FFF8FF4FF98EFB2D0BB8:6C5A60A14A11FFFFFF67F803FEBC08C4" box="[191,301,1951,1977]" country="Democratic Republic of the Congo" latitude="-2.98693" longLatPrecision="1" longitude="27.55371" name="Shabunda" pageId="123" pageNumber="124">Shabunda</location>
[
<geoCoordinate id="0CB150BD4A11FFFFFE98F803FE3808C4" box="[320,425,1951,1977]" degrees="2.98693" direction="south" orientation="latitude" pageId="123" pageNumber="124" precision="1" value="-2.98693">-2.98693</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A11FFFFFE61F802FDB408C4" box="[441,549,1950,1977]" degrees="27.55371" direction="east" orientation="longitude" pageId="123" pageNumber="124" precision="1" value="27.55371">27.55371</geoCoordinate>
]
<date id="1D3B10BA4A11FFFFFDEEF803FD1808C4" box="[566,649,1951,1977]" pageId="123" pageNumber="124" value="1939-05">
<collectingDate id="0D7FE9524A11FFFFFDEEF803FD1808C4" box="[566,649,1951,1977]" pageId="123" pageNumber="124" value="1939-05">V-1939</collectingDate>
</date>
<collectorName id="C47053AC4A11FFFFFD4CF803FCBB08C4" box="[660,810,1951,1977]" pageId="123" pageNumber="124">Dr. Hautman</collectorName>
(
<collectionCode id="0F94AEBF4A11FFFFFCE4F803FC1B08C4" box="[828,906,1951,1977]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="123" pageNumber="124">SMNS</collectionCode>
ex
<collectionCode id="0F94AEBF4A11FFFFFC63F803FB8208C4" box="[955,1043,1951,1977]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="123" pageNumber="124" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="D9ED3C274A11FFFFFBF1F803FBDB08A0" collectingDate="1992-11" collectionCode="SMNS" collectorName="Rep. Congo P. N. d'Odzala &amp; Mbdanza &amp; G. Carpaneto &amp; R. Fochetti" country="Democratic Republic of the Congo" pageId="123" pageNumber="124" specimenCount="3" specimenCount-female="3">
<specimenCount id="7F83FDF34A11FFFFFBF1F803FBDE08C4" box="[1065,1103,1951,1977]" pageId="123" pageNumber="124" type="female">3♀</specimenCount>
:
<collectorName id="C47053AC4A11FFFFFB87F803FA0908C4" box="[1119,1432,1951,1977]" pageId="123" pageNumber="124">Rep. Congo P.N. d’Odzala</collectorName>
,
<collectorName id="C47053AC4A11FFFFFF4FF85FFF6E08A0" box="[151,255,1987,2013]" pageId="123" pageNumber="124">Mbdanza</collectorName>
<date id="1D3B10BA4A11FFFFFED2F85FFEF608A0" box="[266,359,1987,2013]" pageId="123" pageNumber="124" value="1992-11">
<collectingDate id="0D7FE9524A11FFFFFED2F85FFEF608A0" box="[266,359,1987,2013]" pageId="123" pageNumber="124" value="1992-11">XI.1992</collectingDate>
</date>
<collectorName id="C47053AC4A11FFFFFEA9F85FFD9B08A0" box="[369,522,1987,2013]" pageId="123" pageNumber="124">G. Carpaneto</collectorName>
(
<collectionCode id="0F94AEBF4A11FFFFFDC4F85FFDFD08A0" box="[540,620,1987,2013]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="123" pageNumber="124">SMNS</collectionCode>
, gift
<collectorName id="C47053AC4A11FFFFFD75F85FFCA108A0" box="[685,816,1987,2013]" pageId="123" pageNumber="124">R. Fochetti</collectorName>
; slides Z16/19&amp;20). 1
<emphasis id="5BF1EA684A11FFFFFBEDF85FFBDB08A0" bold="true" box="[1077,1098,1987,2013]" italics="true" pageId="123" pageNumber="124">J</emphasis>
</materialsCitation>
<emphasis id="5BF1EA684A11FFFFFB91F85FFBC108A0" bold="true" box="[1097,1104,1987,2013]" italics="true" pageId="123" pageNumber="124">,</emphasis>
<specimenCount id="7F83FDF34A11FFFFFB83F85FFB1108A0" box="[1115,1152,1987,2013]" pageId="123" pageNumber="124" type="female">1♀</specimenCount>
: B. Congo, 12 ml. N of
</paragraph>
<paragraph id="693A367A4A16FFF8FF4FFF0BFD8D0E60" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<materialsCitation id="D9ED3C274A16FFF8FF4FFF0BFD890E60" collectingDate="1957-07-28" collectingDateMax="1993-11-19" collectingDateMin="1957-07-28" collectionCode="SMNS" collectorName="Matadi &amp; E. S. Ross &amp; R. E. Leech &amp; Ref. No." country="Ghana" county="Ghana" latitude="9.17" location="Niandan" longLatPrecision="780" longitude="-10.07" municipality="Agumatsa" pageId="124" pageNumber="125" specimenCode="CASENT 8413072" specimenCount="5" specimenCount-female="5" stateProvince="Volta">
<collectorName id="C47053AC4A16FFF8FF4FFF0BFF770FCC" box="[151,230,151,177]" pageId="124" pageNumber="125">Matadi</collectorName>
[
<geoCoordinate id="0CB150BD4A16FFF8FF21FF0AFED20FCC" box="[249,323,150,177]" degrees="5.85" direction="north" orientation="latitude" pageId="124" pageNumber="125" precision="555" value="5.85">5.85N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A16FFF8FE8BFF0BFE370FCC" box="[339,422,151,177]" degrees="13.46" direction="east" orientation="longitude" pageId="124" pageNumber="125" precision="555" value="13.46">13.46E</geoCoordinate>
]
<date id="1D3B10BA4A16FFF8FE6FFF0AFDD40FCC" box="[439,581,150,177]" pageId="124" pageNumber="125" value="1957-07-28">
<collectingDate id="0D7FE9524A16FFF8FE6FFF0AFDD40FCC" box="[439,581,150,177]" pageId="124" pageNumber="125" value="1957-07-28">VII-28-1957</collectingDate>
</date>
<collectorName id="C47053AC4A16FFF8FD88FF0BFD240FCC" box="[592,693,151,177]" pageId="124" pageNumber="125">E.S.Ross</collectorName>
&amp;
<collectorName id="C47053AC4A16FFF8FD3AFF0BFCC90FCC" box="[738,856,151,177]" pageId="124" pageNumber="125">R.E.Leech</collectorName>
(
<specimenCode id="39239E014A16FFF8FCB3FF0BFBD70FCC" box="[875,1094,151,177]" collectionCode="CASENT" pageId="124" pageNumber="125">CASENT 8413072</specimenCode>
and CASENT 8413079;slide Z18.37).
<emphasis id="5BF1EA684A16FFF8FED8FF26FE7C0FA9" bold="true" box="[256,493,186,212]" pageId="124" pageNumber="125">
<collectingCountry id="119276EA4A16FFF8FED8FF26FE790FA9" box="[256,488,186,212]" name="Ghana" pageId="124" pageNumber="125">Republic of Ghana</collectingCountry>
:
</emphasis>
<specimenCount id="7F83FDF34A16FFF8FE2DFF27FD8D0FA8" box="[501,540,187,213]" pageId="124" pageNumber="125" type="female">2♀</specimenCount>
:
<collectingCounty id="805B4EF64A16FFF8FDF0FF27FDE50FA8" box="[552,628,187,213]" pageId="124" pageNumber="125">Ghana</collectingCounty>
,
<collectingRegion id="AB41F8984A16FFF8FD58FF27FC860FA8" box="[640,791,187,213]" country="Ghana" name="Volta" pageId="124" pageNumber="125">Volta Region</collectingRegion>
,
<collectingMunicipality id="895EAC004A16FFF8FCF9FF27FC020FA8" box="[801,915,187,213]" pageId="124" pageNumber="125">Agumatsa</collectingMunicipality>
waterfalls, St. 10,
<date id="1D3B10BA4A16FFF8FBB2FF27FB7B0FA8" box="[1130,1258,187,213]" pageId="124" pageNumber="125" value="1993-11-19">
<collectingDate id="0D7FE9524A16FFF8FBB2FF27FB7B0FA8" box="[1130,1258,187,213]" pageId="124" pageNumber="125" value="1993-11-19">19.11.1993</collectingDate>
</date>
, J.
<collectingMethod id="B0C44E6D4A16FFF8FAC8FF27FE900F84" pageId="124" pageNumber="125">Kjaerendsen light trap</collectingMethod>
;
<collectorName id="C47053AC4A16FFF8FED4FF43FEFD0F84" box="[268,364,223,249]" pageId="124" pageNumber="125">Ref. No.</collectorName>
2 (slide GH10/6).
<emphasis id="5BF1EA684A16FFF8FDE6FF42FCBE0F85" bold="true" box="[574,815,222,248]" pageId="124" pageNumber="125">
<collectingCountry id="119276EA4A16FFF8FDE6FF42FCBB0F85" box="[574,810,222,248]" name="Guinea-Bissau" pageId="124" pageNumber="125">Republic of Guinea</collectingCountry>
:
</emphasis>
<specimenCount id="7F83FDF34A16FFF8FCEEFF43FCCD0F84" box="[822,860,223,249]" pageId="124" pageNumber="125" type="female">3♀</specimenCount>
:
<location id="6C5A60A14A16FFF8FCB0FF43FC560F84" LSID="urn:lsid:plazi:treatment:E12C876C4A11FFF8FF4FF98EFB2D0BB8:6C5A60A14A16FFF8FCB0FF43FC560F84" box="[872,967,223,249]" country="Ghana" county="Ghana" latitude="9.17" longLatPrecision="780" longitude="-10.07" municipality="Agumatsa" name="Niandan" pageId="124" pageNumber="125" stateProvince="Volta">Niandan</location>
[
<geoCoordinate id="0CB150BD4A16FFF8FC0DFF43FBBB0F84" box="[981,1066,223,249]" degrees="9.17" direction="north" orientation="latitude" pageId="124" pageNumber="125" precision="555" value="9.17">9.17°N</geoCoordinate>
,
<geoCoordinate id="0CB150BD4A16FFF8FBEEFF43FB050F84" box="[1078,1172,223,249]" degrees="10.07" direction="west" orientation="longitude" pageId="124" pageNumber="125" precision="555" value="-10.07">10.07W</geoCoordinate>
],
<location id="6C5A60A14A16FFF8FB7FFF43FA850F84" LSID="urn:lsid:plazi:treatment:E12C876C4A11FFF8FF4FF98EFB2D0BB8:6C5A60A14A16FFF8FB7FFF43FA850F84" box="[1191,1300,223,249]" country="Ghana" county="Ghana" latitude="9.17" longLatPrecision="780" longitude="-10.07" municipality="Agumatsa" name="Bambaya" pageId="124" pageNumber="125" stateProvince="Volta">Bambaya</location>
,
<date id="1D3B10BA4A16FFF8FAC6FF42FA0D0F84" box="[1310,1436,222,249]" pageId="124" pageNumber="125" value="1984-10-15">
<collectingDate id="0D7FE9524A16FFF8FAC6FF42FA0D0F84" box="[1310,1436,222,249]" pageId="124" pageNumber="125" value="1984-10-15">15.10.1984</collectingDate>
</date>
(gift Élouard, slide 86/14,
<collectionCode id="0F94AEBF4A16FFF8FE67FE9FFD810E60" box="[447,528,259,285]" country="Germany" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="124" pageNumber="125">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFEBAFE480E18" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFEBAFEB70E3D" bold="true" box="[199,294,294,320]" pageId="124" pageNumber="125">Habitus</emphasis>
. WL 9.7–10.0 mm. Ochre to light brown, including wings and appendages. Ocellar rings narrow, only a brown spot between ocelli.
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFEF3FBAA0ED0" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFEF3FE820EF5" bold="true" box="[199,275,366,393]" pageId="124" pageNumber="125">Male (</emphasis>
<figureCitation id="F1BE2AFF4A16FFF8FECCFEF3FE290EF4" box="[276,440,367,393]" captionStart="FIGURES 313–322" captionStartId="122.[152,270,1787,1811]" captionTargetBox="[251,1336,187,1757]" captionTargetId="figure-32@122.[246,1341,181,1762]" captionTargetPageId="122" captionText="FIGURES 313–322. Males in the Neoperla dubia-complex: N. dubia Klapálek (from Makak, Cameroon): 313, terminal segments, dorsal view; 314, 315, enlarged centre of tergites 7 and 8, dorsal and lateral; 316, contracted, 317, everted penis with detail of distal armature (not to scale, from Ikom, Nigeria). Neoperla proxima n. sp.: 318, abdominal tip in dorsal view; 317, tip of penis with everted endophallus (D. R. Congo, Odzala N. P.). Neoperla vicina n. sp.: 320, 321, abdominal tip, dorsal and lateral views; 322, penis tip with everted endophallus in lateral view, and a detail of external spines in oblique lateral view (D. R. Congo, Bukavu). Scales in 317 and 322 are 100µm, scale in 320 is 0.5mm. The tergites, hemitergite 10 and the cercus are labeled." figureDoi="http://doi.org/10.5281/zenodo.8154118" httpUri="https://zenodo.org/record/8154118/files/figure.png" pageId="124" pageNumber="125">Figs. 318–319</figureCitation>
). As for complex. Mediocaudal process of T7 a small button-like structure with some SB. HT10 long, slender, in dorsal view straight, a distinct small button at tip (
<figureCitation id="F1BE2AFF4A16FFF8FC14FE0FFBBF0ED0" box="[972,1070,403,429]" captionStart="FIGURES 313–322" captionStartId="122.[152,270,1787,1811]" captionTargetBox="[251,1336,187,1757]" captionTargetId="figure-32@122.[246,1341,181,1762]" captionTargetPageId="122" captionText="FIGURES 313–322. Males in the Neoperla dubia-complex: N. dubia Klapálek (from Makak, Cameroon): 313, terminal segments, dorsal view; 314, 315, enlarged centre of tergites 7 and 8, dorsal and lateral; 316, contracted, 317, everted penis with detail of distal armature (not to scale, from Ikom, Nigeria). Neoperla proxima n. sp.: 318, abdominal tip in dorsal view; 317, tip of penis with everted endophallus (D. R. Congo, Odzala N. P.). Neoperla vicina n. sp.: 320, 321, abdominal tip, dorsal and lateral views; 322, penis tip with everted endophallus in lateral view, and a detail of external spines in oblique lateral view (D. R. Congo, Bukavu). Scales in 317 and 322 are 100µm, scale in 320 is 0.5mm. The tergites, hemitergite 10 and the cercus are labeled." figureDoi="http://doi.org/10.5281/zenodo.8154118" httpUri="https://zenodo.org/record/8154118/files/figure.png" pageId="124" pageNumber="125">Fig. 318</figureCitation>
).
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFE2BFEE40D1C" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
Penis (
<figureCitation id="F1BE2AFF4A16FFF8FECCFE2BFEE50EAC" box="[276,372,439,465]" captionStart="FIGURES 313–322" captionStartId="122.[152,270,1787,1811]" captionTargetBox="[251,1336,187,1757]" captionTargetId="figure-32@122.[246,1341,181,1762]" captionTargetPageId="122" captionText="FIGURES 313–322. Males in the Neoperla dubia-complex: N. dubia Klapálek (from Makak, Cameroon): 313, terminal segments, dorsal view; 314, 315, enlarged centre of tergites 7 and 8, dorsal and lateral; 316, contracted, 317, everted penis with detail of distal armature (not to scale, from Ikom, Nigeria). Neoperla proxima n. sp.: 318, abdominal tip in dorsal view; 317, tip of penis with everted endophallus (D. R. Congo, Odzala N. P.). Neoperla vicina n. sp.: 320, 321, abdominal tip, dorsal and lateral views; 322, penis tip with everted endophallus in lateral view, and a detail of external spines in oblique lateral view (D. R. Congo, Bukavu). Scales in 317 and 322 are 100µm, scale in 320 is 0.5mm. The tergites, hemitergite 10 and the cercus are labeled." figureDoi="http://doi.org/10.5281/zenodo.8154118" httpUri="https://zenodo.org/record/8154118/files/figure.png" pageId="124" pageNumber="125">Fig. 319</figureCitation>
) stout, straight in dorsal, sinuous in side view, middle bulging ventrad. The large dorsal notch at the tube apex with many strong conical teeth, tips pointing caudad. Endophallus with long bare base, strongly curved, tip crossing over tube. A band of strong teeth covers most of convex face to near the end. Sides are bare, distal half of ventral face with about seven rows of stout teeth. Size of teeth and number of rows decrease caudally, the short tip is bare.
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFDF7FDFD0DD4" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFDF7FE8F0DF8" bold="true" box="[199,286,619,645]" pageId="124" pageNumber="125">Female</emphasis>
(
<figureCitation id="F1BE2AFF4A16FFF8FEE8FDF7FE050DF8" box="[304,404,619,645]" captionStart="FIGURES 323–331" captionStartId="125.[152,270,1838,1862]" captionTargetBox="[235,1343,189,1806]" captionTargetId="figure-17@125.[229,1357,181,1814]" captionTargetPageId="125" captionText="FIGURES 323–331. Neoperla dubia-complex, femalesand eggs.N.dubia Klapálek:323, genital situs; 324, eggwith moderately raised costae (Ghana, Agumatsa waterfalls); 325–327, egg with flanged costae (D. R. Congo, Odzala N. P.). Neoperla proxima n. sp.: 328, genital situs (D. R.Congo, N of Matadi), 329, egg (Cameroon, Sangmelina). Neoperla vicina n. sp. (D.R. Congo): 330, female situs, 331,egg, arrow points at end of raised part of costa (paratype, Walikale, Z2101). Scales are 100µm, similar structures shown to the same scale. Top view of egg (326) based on combined halves of a photograph. Sa, spermatheca; SSt, spermathecal stalk; V, vagina." figureDoi="http://doi.org/10.5281/zenodo.8154124" httpUri="https://zenodo.org/record/8154124/files/figure.png" pageId="124" pageNumber="125">Fig. 328</figureCitation>
). As for the complex. SSt longer than in other members of the complex forming almost 2 rings, narrow end-section is a slender tail.
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFD2FFD800C44" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFD2FFF670DB0" bold="true" box="[199,246,691,717]" pageId="124" pageNumber="125">Egg</emphasis>
(
<figureCitation id="F1BE2AFF4A16FFF8FEDCFD2FFEF40DB0" box="[260,357,691,717]" captionStart="FIGURES 323–331" captionStartId="125.[152,270,1838,1862]" captionTargetBox="[235,1343,189,1806]" captionTargetId="figure-17@125.[229,1357,181,1814]" captionTargetPageId="125" captionText="FIGURES 323–331. Neoperla dubia-complex, femalesand eggs.N.dubia Klapálek:323, genital situs; 324, eggwith moderately raised costae (Ghana, Agumatsa waterfalls); 325–327, egg with flanged costae (D. R. Congo, Odzala N. P.). Neoperla proxima n. sp.: 328, genital situs (D. R.Congo, N of Matadi), 329, egg (Cameroon, Sangmelina). Neoperla vicina n. sp. (D.R. Congo): 330, female situs, 331,egg, arrow points at end of raised part of costa (paratype, Walikale, Z2101). Scales are 100µm, similar structures shown to the same scale. Top view of egg (326) based on combined halves of a photograph. Sa, spermatheca; SSt, spermathecal stalk; V, vagina." figureDoi="http://doi.org/10.5281/zenodo.8154124" httpUri="https://zenodo.org/record/8154124/files/figure.png" pageId="124" pageNumber="125">Fig. 329</figureCitation>
). Oval, 1.8 times longer than wide, on average 378µm long in specimens from the
<collectingCountry id="119276EA4A16FFF8FB2DFD2FFAD30DB0" box="[1269,1346,691,717]" name="Democratic Republic of the Congo" pageId="124" pageNumber="125">Congo</collectingCountry>
, 424µm in the Guinean female. Anchor pole wider than operculum, walls of cells low. About 20 straight striae, sulci wide, surface irregularly punctate, rough. Costae are distinct ridges but not flanged, their fine ends form a delicate network on the narrow conical operculum.
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A16FFF8FF1FFCDEFCDD0BDC" pageId="124" pageNumber="125" type="description">
<paragraph id="693A367A4A16FFF8FF1FFCDEFDC40CD8" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFCDEFEA90C21" bold="true" box="[199,312,834,860]" pageId="124" pageNumber="125">Variation</emphasis>
. Females from
<collectingCountry id="119276EA4A16FFF8FE2FFCDFFDFD0C20" box="[503,620,835,861]" name="Cameroon" pageId="124" pageNumber="125">Cameroon</collectingCountry>
(identified by DNA) are apparently juvenile, they have no or vague marks on S8. Both marks are distinct in specimens from the
<collectingCountry id="119276EA4A16FFF8FC9AFCFBFC1C0CFC" box="[834,909,871,897]" name="Democratic Republic of the Congo" pageId="124" pageNumber="125">Congo</collectingCountry>
and
<collectingCountry id="119276EA4A16FFF8FC1CFCFBFB880CFC" box="[964,1049,871,897]" name="Guinea-Bissau" pageId="124" pageNumber="125">Guinea</collectingCountry>
. Egg size and collar width may be characteristic of individual populations.
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFC32FD8B0B00" blockId="124.[151,1437,150,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFC32FE950CB5" bold="true" box="[199,260,942,968]" pageId="124" pageNumber="125">DNA</emphasis>
(
<figureCitation id="F1BE2AFF4A16FFF8FECBFC33FEEF0CB4" box="[275,382,943,969]" captionStart="FIGURE 492" captionStartId="178.[152,255,1943,1967]" captionTargetBox="[361,1187,182,1910]" captionTargetPageId="178" captionText="FIGURE 492. Simplified overview of codon-model maximum likelihood phylogram based on 356 mitochondrial genome and COX1 DNA barcode sequences (all-NT). Terminal labels include information on the species only. Clades of particular interest highlighted with colours that correspond to the clades in other trees. The full tree is shown in Figs 493-498." figureDoi="http://doi.org/10.5281/zenodo.8154202" httpUri="https://zenodo.org/record/8154202/files/figure.png" pageId="124" pageNumber="125">Figs. 492</figureCitation>
, 497). A total of
<specimenCount id="7F83FDF34A16FFF8FDE0FC33FD440CB4" box="[568,725,943,969]" pageId="124" pageNumber="125" type="generic">ten specimens</specimenCount>
from
<collectingCountry id="119276EA4A16FFF8FCC0FC33FC010CB4" box="[792,912,943,969]" name="Cameroon" pageId="124" pageNumber="125">Cameroon</collectingCountry>
, the D. R. Congo, and
<collectingCountry id="119276EA4A16FFF8FB53FC33FB4C0CB4" box="[1163,1245,943,969]" name="Angola" pageId="124" pageNumber="125">Angola</collectingCountry>
were sequenced, including the male
<typeStatus id="B63E88D84A16FFF8FEB7FC4FFE400C90" box="[367,465,979,1005]" pageId="124" pageNumber="125" type="holotype">holotype</typeStatus>
from the D. R. Congo with only a partial COX1 DNA barcode fragment, which might be responsible for the basal-most placement of this specimen. This sampling represents the largest part of the geographic spread of this species and provides very strong support (94.4/100/100) for monophyly. The species is very strongly supported (69.1/100/100) as sister to
<taxonomicName id="AE854DF94A16FFF8FD27FBA3FB860B24" authority="Klapalek. Both" authorityName="Klapalek. Both" authorityYear="1909" box="[767,1047,1087,1113]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="5BF1EA684A16FFF8FD27FBA3FCF30B24" box="[767,866,1087,1113]" italics="true" pageId="124" pageNumber="125">N. dubia</emphasis>
Klapálek. Both
</taxonomicName>
sexes are represented, but female samples are vastly more abundant.
</paragraph>
<paragraph id="693A367A4A16FFF8FF1FFB1BFCDD0BDC" blockId="124.[151,1437,150,1221]" box="[199,844,1159,1185]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFB1BFE9B0BDC" bold="true" box="[199,266,1159,1185]" pageId="124" pageNumber="125">Notes</emphasis>
. Ranges of
<taxonomicName id="AE854DF94A16FFF8FE48FB1BFD9D0BDD" authority="Zwick &amp; Zwick, 2023" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[400,524,1159,1184]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="proxima" status="sp. nov.">
<emphasis id="5BF1EA684A16FFF8FE48FB1BFD9D0BDD" box="[400,524,1159,1184]" italics="true" pageId="124" pageNumber="125">N. proxima</emphasis>
</taxonomicName>
<emphasis id="5BF1EA684A16FFF8FDCCFB1BFDC30BDC" bold="true" box="[532,594,1159,1185]" pageId="124" pageNumber="125">
<taxonomicNameLabel id="40C257134A16FFF8FDCCFB1BFDC30BDC" box="[532,594,1159,1185]" pageId="124" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="AE854DF94A16FFF8FD51FB1BFD7B0BDC" authorityName="Klapalek. Both" authorityYear="1909" box="[649,746,1159,1185]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="5BF1EA684A16FFF8FD51FB1BFD7B0BDC" box="[649,746,1159,1185]" italics="true" pageId="124" pageNumber="125">N. dubia</emphasis>
</taxonomicName>
overlap.
</paragraph>
</subSubSection>
<subSubSection id="219F65F14A16FFF8FF1FFB37FB2D0BB8" box="[199,1212,1195,1221]" pageId="124" pageNumber="125" type="etymology">
<paragraph id="693A367A4A16FFF8FF1FFB37FB2D0BB8" blockId="124.[151,1437,150,1221]" box="[199,1212,1195,1221]" pageId="124" pageNumber="125">
<emphasis id="5BF1EA684A16FFF8FF1FFB37FED90BB8" bold="true" box="[199,328,1195,1221]" pageId="124" pageNumber="125">Etymology</emphasis>
. The closest (Lat.
<taxonomicName id="AE854DF94A16FFF8FDC0FB37FDE50BB9" authorityName="Zwick &amp; Zwick" authorityYear="2023" box="[536,628,1195,1220]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="proxima">
<emphasis id="5BF1EA684A16FFF8FDC0FB37FDE50BB9" box="[536,628,1195,1220]" italics="true" pageId="124" pageNumber="125">proxima</emphasis>
</taxonomicName>
) relative of
<emphasis id="5BF1EA684A16FFF8FD26FB37FCF40BB8" box="[766,869,1195,1221]" italics="true" pageId="124" pageNumber="125">
<taxonomicName id="AE854DF94A16FFF8FD26FB37FCF30BB8" authorityName="Klapalek. Both" authorityYear="1909" box="[766,866,1195,1221]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="dubia">N. dubia</taxonomicName>
,
</emphasis>
an adjective in female gender.
</paragraph>
</subSubSection>
</treatment>
</document>
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