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<document id="CBA46C27665A76170654E946DD2E5127" ID-CLB-Dataset="6063" ID-DOI="10.11646/zootaxa.4933.4.6" ID-GBIF-Dataset="dba46bab-d899-4ded-9310-ca8c53317f6d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4556718" ID-ZooBank="D714E5D4-BE5A-49D3-AED5-9A0B38413959" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614068583615" checkinUser="plazi" docAuthor="Zhu, Wenhui &amp; Li, Shuqiang" docDate="2021" docId="E12C0973F540FFED2C89FA27FCD3FBA9" docLanguage="en" docName="zootaxa.4933.4.6.pdf" docOrigin="Zootaxa 4933 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana bubeng Zhu &amp; Li 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="561" masterDocId="1D15710BF541FFE92C1EFFCEFF91FFBB" masterDocTitle="Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China" masterLastPageNumber="566" masterPageNumber="557" pageNumber="558" updateTime="1698930772463" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C19AD72D5E5E1757C6070B6CCE380741">
<mods:title id="0FBFF201FDBEFE06920917897553F70E">Two new spiders of the genus Belisana (Araneae: Pholcidae) from Xishuangbanna Rainforest, China</mods:title>
</mods:titleInfo>
<mods:name id="FBFEC24F8C20B33DFFAF692DFD05B0D7" type="personal">
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<mods:namePart id="1A85BB9642172B41822CC3EDF4EB22A8">Zhu, Wenhui</mods:namePart>
<mods:nameIdentifier id="E98ACE134B4AE4740CD7EEB93B7932A8" type="ORCID">0000-0001-5650-1670</mods:nameIdentifier>
<mods:affiliation id="324BEC35CE21CBB099B1ACA84055808B">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
<mods:nameIdentifier id="CDF0CCEFF651CDA0800E50B9A89B6B36" type="email">zhuwh1016@outlook.com</mods:nameIdentifier>
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<mods:name id="69C7D18B7516761B0F3A666061133605" type="personal">
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<mods:namePart id="0040FEDC77B1C3CA42A3D87D97FB9327">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="2F3864B17DB3CAA4D0A8CC5FE93D45D0" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="EC250895B6406747234E89B1BF9691F6">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
<mods:nameIdentifier id="751CE996AF6048F15B07CEF979B92751" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource id="CFDDB2D30F9F25BF2216C61DE8312FF1">text</mods:typeOfResource>
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<mods:title id="F56F3D94E29289148B01E9DEFB62698E">Zootaxa</mods:title>
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<mods:part id="01B301B079FAF82F2791037CA8C78A61">
<mods:date id="BB120F4A5BEE49D73E1DD40496475F69">2021</mods:date>
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<mods:number id="8F9F1334C6B4566CFF43953E6AFBD9CA">2021-02-23</mods:number>
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<mods:number id="4166D857DAD93E703D75F498B2E306EE">4933</mods:number>
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<treatment id="E12C0973F540FFED2C89FA27FCD3FBA9" ID-DOI="http://doi.org/10.5281/zenodo.4556720" ID-GBIF-Taxon="182753405" ID-Zenodo-Dep="4556720" LSID="urn:lsid:plazi:treatment:E12C0973F540FFED2C89FA27FCD3FBA9" httpUri="http://treatment.plazi.org/id/E12C0973F540FFED2C89FA27FCD3FBA9" lastPageId="4" lastPageNumber="561" pageId="1" pageNumber="558">
<subSubSection id="219FEBEEF540FFE82C89FA27FE28F9B8" box="[151,441,1513,1540]" pageId="1" pageNumber="558" type="nomenclature">
<paragraph id="693AB865F540FFE82C89FA27FE28F9B8" blockId="1.[151,441,1513,1576]" box="[151,441,1513,1540]" pageId="1" pageNumber="558">
<heading id="32720F09F540FFE82C89FA27FE28F9B8" bold="true" box="[151,441,1513,1540]" fontSize="11" level="1" pageId="1" pageNumber="558" reason="1">
<emphasis id="5BF16477F540FFE82C89FA27FE28F9B8" bold="true" box="[151,441,1513,1540]" pageId="1" pageNumber="558">
<taxonomicName id="AE85C3E6F540FFE82C89FA27FEC8F9BF" ID-CoL="98YWF" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[151,345,1513,1540]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="558" phylum="Arthropoda" rank="species" species="bubeng" status="sp. nov.">
<emphasis id="5BF16477F540FFE82C89FA27FEC8F9BF" bold="true" box="[151,345,1513,1540]" italics="true" pageId="1" pageNumber="558">Belisana bubeng</emphasis>
</taxonomicName>
<taxonomicNameLabel id="40C2D90CF540FFE82D7EFA27FE28F9B8" box="[352,441,1513,1539]" pageId="1" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="219FEBEEF540FFE82C89F9C3FEDAF993" box="[151,331,1549,1576]" pageId="1" pageNumber="558" type="description">
<paragraph id="693AB865F540FFE82C89F9C3FEDAF993" blockId="1.[151,441,1513,1576]" box="[151,331,1549,1576]" pageId="1" pageNumber="558">
<figureCitation id="F1BEA4E0F540FFE82C89F9C3FF6BF99C" box="[151,250,1549,1575]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1824,1848]" captionStartId-1="3.[151,250,1874,1898]" captionTargetBox-0="[243,1344,179,1798]" captionTargetBox-1="[226,1361,181,1851]" captionTargetId-0="figure-18@2.[243,1344,179,1798]" captionTargetId-1="figure-17@3.[226,1361,181,1851]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." captionText-1="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.4556722" figureDoi-1="http://doi.org/10.5281/zenodo.4556724" httpUri-0="https://zenodo.org/record/4556722/files/figure.png" httpUri-1="https://zenodo.org/record/4556724/files/figure.png" pageId="1" pageNumber="558">Figs 1–2</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82D18F9C3FE83F993" box="[262,274,1549,1576]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5</figureCitation>
A–D
</paragraph>
</subSubSection>
<subSubSection id="219FEBEEF540FFE82C89F99EFB53F909" pageId="1" pageNumber="558" type="materials_examined">
<paragraph id="693AB865F540FFE82C89F99EFB53F909" blockId="1.[151,1437,1616,2039]" pageId="1" pageNumber="558">
<emphasis id="5BF16477F540FFE82C89F99EFED6F9D1" bold="true" box="[151,327,1616,1642]" pageId="1" pageNumber="558">Type material.</emphasis>
<materialsCitation id="D9EDB238F540FFE82D4EF99EFE05F909" ID-GBIF-Occurrence="3302850305" collectingDate="2012-07-11" collectionCode="IZCAS" collectorName="Zhigang Chen &amp; Qingyuan Zhao" country="China" county="Mengla County" elevation="30" latitude="21.611223" location="Bubeng Village" longLatPrecision="1" longitude="101.57378" municipality="Xishuangbanna" pageId="1" pageNumber="558" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="5BF16477F540FFE82D4EF99EFE57F9D1" bold="true" box="[336,454,1616,1642]" pageId="1" pageNumber="558">
<typeStatus id="B63E06C7F540FFE82D4EF99EFE50F9D1" box="[336,449,1616,1642]" pageId="1" pageNumber="558" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="0F9420A0F540FFE82E07F99EFDFDF9D1" box="[537,620,1616,1642]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="558">IZCAS</collectionCode>
Ar41821), seasonal rainforest (
<geoCoordinate id="0CB1DEA2F540FFE82FCAF99EFB12F9D1" box="[980,1155,1616,1642]" degrees="21" direction="north" minutes="36" orientation="latitude" pageId="1" pageNumber="558" precision="1" seconds="40.40" value="21.611223">21°36’40.40”N</geoCoordinate>
,
<geoCoordinate id="0CB1DEA2F540FFE8288FF99EFADAF9D0" box="[1169,1355,1616,1643]" degrees="101" direction="east" minutes="34" orientation="longitude" pageId="1" pageNumber="558" precision="1" seconds="25.60" value="101.57378">101°34’25.60”E</geoCoordinate>
, elevation 855 ±
<quantity id="AE7D1580F540FFE82D0CF9BAFEDEF934" box="[274,335,1652,1679]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="558" unit="m" value="30.0">
<elevation id="E2A85F56F540FFE82D0CF9BAFEDEF934" box="[274,335,1652,1679]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="558" unit="m" value="30.0">30 m</elevation>
</quantity>
),
<location id="6C5AEEBEF540FFE82D7DF9BAFD87F935" LSID="urn:lsid:plazi:treatment:E12C0973F540FFED2C89FA27FCD3FBA9:6C5AEEBEF540FFE82D7DF9BAFD87F935" box="[355,534,1652,1678]" country="China" county="Mengla County" latitude="21.611223" longLatPrecision="1" longitude="101.57378" municipality="Xishuangbanna" name="Bubeng Village" pageId="1" pageNumber="558" stateProvince="Yunnan">Bubeng Village</location>
,
<collectingCounty id="805BC0E9F540FFE82E3FF9BAFD43F935" box="[545,722,1652,1678]" pageId="1" pageNumber="558">Mengla County</collectingCounty>
,
<collectingMunicipality id="895E221FF540FFE82EC3F9BAFC1FF935" box="[733,910,1652,1678]" pageId="1" pageNumber="558">Xishuangbanna</collectingMunicipality>
,
<emphasis id="5BF16477F540FFE82F87F9BAFC66F935" bold="true" box="[921,1015,1652,1678]" pageId="1" pageNumber="558">
<collectingRegion id="AB417687F540FFE82F87F9BAFC66F935" box="[921,1015,1652,1678]" country="China" name="Yunnan" pageId="1" pageNumber="558">Yunnan</collectingRegion>
</emphasis>
,
<emphasis id="5BF16477F540FFE82818F9BAFBDEF935" bold="true" box="[1030,1103,1652,1678]" pageId="1" pageNumber="558">
<collectingCountry id="1192F8F5F540FFE82818F9BAFBDEF935" box="[1030,1103,1652,1678]" name="China" pageId="1" pageNumber="558">China</collectingCountry>
</emphasis>
,
<date id="1D3B9EA5F540FFE82843F9BAFB60F935" box="[1117,1265,1652,1678]" pageId="1" pageNumber="558" value="2012-07-11">
<collectingDate id="0D7F674DF540FFE82843F9BAFB60F935" box="[1117,1265,1652,1678]" pageId="1" pageNumber="558" value="2012-07-11">11 July 2012</collectingDate>
</date>
,
<collectorName id="C470DDB3F540FFE828E2F9BAFA0DF935" box="[1276,1436,1652,1678]" pageId="1" pageNumber="558">Zhigang Chen</collectorName>
&amp;
<collectorName id="C470DDB3F540FFE82CAAF956FEF2F909" box="[180,355,1688,1714]" pageId="1" pageNumber="558">Qingyuan Zhao</collectorName>
leg.
</materialsCitation>
<materialsCitation id="D9EDB238F540FFE82D84F956FB2FF909" ID-GBIF-Occurrence="3302850304" box="[410,1214,1688,1715]" collectingDate="2012-07-11" collectionCode="IZCAS" collectorName="Zhigang Chen &amp; Qingyuan Zhao" country="China" county="Mengla County" elevation="30" latitude="21.611223" location="Bubeng Village" longLatPrecision="1" longitude="101.57378" municipality="Xishuangbanna" pageId="1" pageNumber="558" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="5BF16477F540FFE82D84F956FD8AF909" bold="true" box="[410,539,1688,1714]" pageId="1" pageNumber="558">
<typeStatus id="B63E06C7F540FFE82D84F956FD87F909" box="[410,534,1688,1714]" pageId="1" pageNumber="558" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="7F8373ECF540FFE82E3CF956FD1FF909" box="[546,654,1688,1714]" pageId="1" pageNumber="558" type="female">2 females</specimenCount>
(
<collectionCode id="0F9420A0F540FFE82E83F956FD61F909" box="[669,752,1688,1714]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="558">IZCAS</collectionCode>
Ar41822–41823), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="219FEBEEF540FFE82CD9F972FBEDF96D" box="[199,1148,1724,1751]" pageId="1" pageNumber="558" type="etymology">
<paragraph id="693AB865F540FFE82CD9F972FBEDF96D" blockId="1.[151,1437,1616,2039]" box="[199,1148,1724,1751]" pageId="1" pageNumber="558">
<emphasis id="5BF16477F540FFE82CD9F972FEDFF96D" bold="true" box="[199,334,1724,1750]" pageId="1" pageNumber="558">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="B63E06C7F540FFE82EABF973FD74F96C" box="[693,741,1725,1751]" pageId="1" pageNumber="558">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="219FEBEEF540FFE82CD9F92EFC60F84D" pageId="1" pageNumber="558" type="diagnosis">
<paragraph id="693AB865F540FFE82CD9F92EFC60F84D" blockId="1.[151,1437,1616,2039]" pageId="1" pageNumber="558">
<emphasis id="5BF16477F540FFE82CD9F92EFED3F941" bold="true" box="[199,322,1760,1786]" pageId="1" pageNumber="558">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="AE85C3E6F540FFE82E52F92FFCD0F941" authority="Yao &amp; Li, 2018" authorityName="Yao &amp; Li" authorityYear="2018" box="[588,833,1760,1787]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="558" phylum="Arthropoda" rank="species" species="cas">
<emphasis id="5BF16477F540FFE82E52F92FFD1EF941" box="[588,655,1761,1786]" italics="true" pageId="1" pageNumber="558">B. cas</emphasis>
<bibRefCitation id="0D14C594F540FFE82E8BF92FFCD0F941" author="Yao, Z. &amp; Zhu, K. &amp; Du, Z. &amp; Li, S." box="[661,833,1760,1787]" pageId="1" pageNumber="558" pagination="243 - 262" refId="ref4193" refString="Yao, Z., Zhu, K., Du, Z. &amp; Li, S. (2018) The Belisana spiders (Araneae: Pholcidae) from Xishuangbanna Tropical Botanical Garden, Yunnan, China. Zootaxa, 4425 (2), 243 - 262. https: // doi. org / 10.11646 / zootaxa. 4425.2.3" type="journal article" year="2018">Yao &amp; Li, 2018</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="0D14C594F540FFE82F67F92FFBCAF940" author="Yao, Z. &amp; Zhu, K. &amp; Du, Z. &amp; Li, S." box="[889,1115,1760,1787]" pageId="1" pageNumber="558" pagination="243 - 262" refId="ref4193" refString="Yao, Z., Zhu, K., Du, Z. &amp; Li, S. (2018) The Belisana spiders (Araneae: Pholcidae) from Xishuangbanna Tropical Botanical Garden, Yunnan, China. Zootaxa, 4425 (2), 243 - 262. https: // doi. org / 10.11646 / zootaxa. 4425.2.3" type="journal article" year="2018">
Yao
<emphasis id="5BF16477F540FFE82FB5F92FFC4AF941" box="[939,987,1760,1786]" italics="true" pageId="1" pageNumber="558">et al</emphasis>
. 2018: 250
</bibRefCitation>
, figs 7–8, 11D, 12D) by having similar external female genitalia (
<figureCitation id="F1BEA4E0F540FFE82E28F8CAFD03F8A5" box="[566,658,1796,1822]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="1" pageNumber="558">Figs 2A</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82E83F8CAFD51F8A4" box="[669,704,1796,1823]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5A</figureCitation>
) but females can be distinguished by the presence of a short scape (arrow s in
<figureCitation id="F1BEA4E0F540FFE82D07F8E6FEE5F8F9" box="[281,372,1832,1858]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="1" pageNumber="558">Figs 2B</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82D9EF8E6FE30F8F8" box="[384,417,1832,1859]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5B</figureCitation>
) and narrow pore plates (arrow pp in
<figureCitation id="F1BEA4E0F540FFE82F52F8E6FC36F8F9" box="[844,935,1832,1858]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="1" pageNumber="558">Figs 2B</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82FADF8E6FC44F8F8" box="[947,981,1832,1859]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5B</figureCitation>
) and by the absence of serrated sclerites on vulva. Males can be distinguished by the presence of a bifid subdisto-prolateral sclerite (arrow
<quantity id="AE7D1580F540FFE828C9F882FA90F8DD" box="[1239,1281,1868,1894]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="558" unit="in" value="1.0">1 in</quantity>
<figureCitation id="F1BEA4E0F540FFE82919F882FACEF8DD" box="[1287,1375,1868,1894]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="1" pageNumber="558">Figs 1C</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82975F882FA1FF8DC" box="[1387,1422,1868,1895]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5C</figureCitation>
), nearly trapezoidal subdisto-ventral sclerite (arrow
<quantity id="AE7D1580F540FFE82ED2F8BEFD66F831" box="[716,759,1904,1930]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="558" unit="in" value="2.0">2 in</quantity>
<figureCitation id="F1BEA4E0F540FFE82EE0F8BEFCC6F831" box="[766,855,1904,1930]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="1" pageNumber="558">Figs 1C</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82F7AF8BEFC16F830" box="[868,903,1904,1931]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5C</figureCitation>
), curved subdisto-prolateral sclerite (arrow
<quantity id="AE7D1580F540FFE8296FF8BEFA0DF831" box="[1393,1436,1904,1930]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="1" pageNumber="558" unit="in" value="3.0">3 in</quantity>
<figureCitation id="F1BEA4E0F540FFE82C89F85AFF60F815" box="[151,241,1940,1966]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="1" pageNumber="558">Figs 1C</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82CE1F85AFEB2F814" box="[255,291,1940,1967]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5C</figureCitation>
), fan-shaped disto-retrolateral membranous lamella (arrow
<quantity id="AE7D1580F540FFE82FDAF85AFC61F815" box="[964,1008,1940,1966]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="558" unit="in" value="1.0">1 in</quantity>
<figureCitation id="F1BEA4E0F540FFE82FE6F85AFBC5F815" box="[1016,1108,1940,1966]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="1" pageNumber="558">Figs 1D</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE8287CF85AFB16F814" box="[1122,1159,1940,1967]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5D</figureCitation>
), small disto-retrolateral sclerite (arrow
<quantity id="AE7D1580F540FFE82D5DF876FEFEF869" box="[323,367,1976,2002]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="558" unit="in" value="2.0">2 in</quantity>
<figureCitation id="F1BEA4E0F540FFE82D69F876FE42F869" box="[375,467,1976,2002]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="1" pageNumber="558">Figs 1D</figureCitation>
,
<figureCitation id="F1BEA4E0F540FFE82DFFF876FD95F868" box="[481,516,1976,2003]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="1" pageNumber="558">5D</figureCitation>
) on the procursus, longer male cheliceral distal apophyses (arrow da in
<figureCitation id="F1BEA4E0F540FFE82929F876FA1EF869" box="[1335,1423,1976,2002]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="1" pageNumber="558">Fig. 2B</figureCitation>
), and by the presence of a long distal spine on the embolus (arrow in
<figureCitation id="F1BEA4E0F540FFE82F93F812FC75F84D" box="[909,996,2012,2038]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="1" pageNumber="558">Fig. 2C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="3DFAE8EDF543FFEB2C89F8EEFDE7F856" ID-DOI="http://doi.org/10.5281/zenodo.4556722" ID-Zenodo-Dep="4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="2" pageNumber="559" startId="2.[151,250,1824,1848]" targetBox="[243,1344,179,1798]" targetPageId="2">
<paragraph id="693AB865F543FFEB2C89F8EEFDE7F856" blockId="2.[151,1437,1824,2029]" pageId="2" pageNumber="559">
<emphasis id="5BF16477F543FFEB2C89F8EEFE82F883" bold="true" box="[151,275,1824,1848]" pageId="2" pageNumber="559">FIGURE 1.</emphasis>
<taxonomicName id="AE85C3E6F543FFEB2D06F8EEFE53F883" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[280,450,1824,1848]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="559" phylum="Arthropoda" rank="species" species="bubeng" status="sp. nov.">
<emphasis id="5BF16477F543FFEB2D06F8EEFE53F883" box="[280,450,1824,1848]" italics="true" pageId="2" pageNumber="559">Belisana bubeng</emphasis>
</taxonomicName>
<emphasis id="5BF16477F543FFEB2DD9F8EFFD86F882" bold="true" box="[455,535,1825,1849]" pageId="2" pageNumber="559">
<taxonomicNameLabel id="40C2D90CF543FFEB2DD9F8EFFD86F882" box="[455,535,1825,1849]" pageId="2" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D).
</paragraph>
</caption>
<caption id="3DFAE8EDF542FFEA2C89F89CFE03F840" ID-DOI="http://doi.org/10.5281/zenodo.4556724" ID-Zenodo-Dep="4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="3" pageNumber="560" startId="3.[151,250,1874,1898]" targetBox="[226,1361,181,1851]" targetPageId="3">
<paragraph id="693AB865F542FFEA2C89F89CFE03F840" blockId="3.[151,1437,1874,2043]" pageId="3" pageNumber="560">
<emphasis id="5BF16477F542FFEA2C89F89CFE85F8D0" bold="true" box="[151,276,1874,1899]" pageId="3" pageNumber="560">FIGURE 2.</emphasis>
<taxonomicName id="AE85C3E6F542FFEA2D02F89DFE59F8D0" authority="Zhu &amp; Li, 2021" authorityName="Zhu &amp; Li" authorityYear="2021" box="[284,456,1875,1899]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="560" phylum="Arthropoda" rank="species" species="bubeng" status="sp. nov.">
<emphasis id="5BF16477F542FFEA2D02F89DFE59F8D0" box="[284,456,1875,1899]" italics="true" pageId="3" pageNumber="560">Belisana bubeng</emphasis>
</taxonomicName>
<emphasis id="5BF16477F542FFEA2DD1F89DFDB3F8D0" bold="true" box="[463,546,1875,1899]" pageId="3" pageNumber="560">
<taxonomicNameLabel id="40C2D90CF542FFEA2DD1F89DFDB3F8D0" box="[463,546,1875,1899]" pageId="3" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H).
</paragraph>
</caption>
<subSubSection id="219FEBEEF545FFED2CD9FF58FC15FC56" pageId="4" pageNumber="561" type="description">
<paragraph id="693AB865F545FFED2CD9FF58FD59FD76" blockId="4.[151,1437,150,1042]" pageId="4" pageNumber="561">
<emphasis id="5BF16477F545FFED2CD9FF58FE0EFF0A" bold="true" box="[199,415,150,177]" pageId="4" pageNumber="561">Description. Male</emphasis>
(
<emphasis id="5BF16477F545FFED2DACFF59FD88FF0A" bold="true" box="[434,537,151,177]" pageId="4" pageNumber="561">
<typeStatus id="B63E06C7F545FFED2DACFF59FD88FF0A" box="[434,537,151,177]" pageId="4" pageNumber="561" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar41821): Total length 1.62 (1.70 with clypeus), carapace 0.79 long, 0.66 wide, opisthosoma 0.91 long, 0.69 wide. Leg I: – (1.78 + 0.28 + – + – + –), leg II: – (1.64 + 0.25 + 1.60 + 2.59 + –), leg III: 4.50 (1.34 + 0.16 + 1.09 + 1.44 + 0.47), femur IV: 1.68, other segments missing. Distance PME-PME 0.23, diameter PME 0.06, distance PME-ALE 0.07, AME absent. Sternum approximately as wide as long (0.50). Habitus as in
<figureCitation id="F1BEA4E0F545FFED2D2EFEE9FEE5FEFA" box="[304,372,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2</figureCitation>
E–F. Carapace pale, with brown lateral margins and small, brown median marks; sternum pale, without marks. Legs pale, without darker rings. Opisthosoma pale, without spots. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Chelicerae (
<figureCitation id="F1BEA4E0F545FFED2EA5FEA1FC83FE32" box="[699,786,367,393]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Fig. 2D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses, each with a sclerotized and pointed tip (distance between tips of distal apophyses: 0.23). Pedipalps as in
<figureCitation id="F1BEA4E0F545FFED2D8EFE79FE43FE6A" box="[400,466,439,465]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1</figureCitation>
A–B; coxa unmodified; trochanter with short ventral apophysis with small retrolateral apophysis (arrow
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<figureCitation id="F1BEA4E0F545FFED2D88FE15FE7DFE4E" box="[406,492,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Fig. 1B</figureCitation>
); femur with small proximo-dorsal apophysis (arrow
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<figureCitation id="F1BEA4E0F545FFED289EFE15FB47FE4E" box="[1152,1238,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Fig. 1B</figureCitation>
); procursus (
<figureCitation id="F1BEA4E0F545FFED2975FE15FF31FDA2" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1</figureCitation>
A–D, 5C–D) simple proximally but complex distally, with bifid subdisto-prolateral sclerite (arrow
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<figureCitation id="F1BEA4E0F545FFED2922FE31FA07FDA2" box="[1340,1430,511,537]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1C</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2C89FDECFF2BFD86" box="[151,186,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5C</figureCitation>
), nearly trapezoidal subdisto-ventral sclerite (arrow
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<figureCitation id="F1BEA4E0F545FFED2F2CFDEDFC1AFD86" box="[818,907,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1C</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2F89FDECFC2AFD86" box="[919,955,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5C</figureCitation>
), curved subdisto-prolateral sclerite (arrow
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<figureCitation id="F1BEA4E0F545FFED2CD0FD89FEB8FDDA" box="[206,297,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1C</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2D27FD88FECDFDDA" box="[313,348,582,609]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5C</figureCitation>
), fan-shaped disto-retrolateral membranous lamella (arrow
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<figureCitation id="F1BEA4E0F545FFED2822FD89FB08FDDA" box="[1084,1177,583,609]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1D</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED28B6FD88FB5CFDDA" box="[1192,1229,582,609]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5D</figureCitation>
), and small distoretrolateral sclerite (arrow
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<figureCitation id="F1BEA4E0F545FFED2DE8FDA5FDC0FD3E" box="[502,593,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1824,1848]" captionTargetBox="[243,1344,179,1798]" captionTargetId="figure-18@2.[243,1344,179,1798]" captionTargetPageId="2" captionText="FIGURE 1. Belisana bubeng sp. nov., holotype male.A–B. Pedipalp (Bulb removed) (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis on pedipalpal trochanter, arrow 2 points at small proximo-dorsal apophysis on pedipalpal femur); C–D. Distal part of procursus (C. Prolateral view, arrow 1 points at bifid subdisto-prolateral sclerite, arrow 2 points at nearly trapezoidal subdisto-ventral sclerite, arrow 3 points at curved subdisto-prolateral sclerite; D. Retrolateral view, arrow 1 points at fan-shaped disto-retrolateral membranous lamella, arrow 2 points at small disto-retrolateral sclerite). pr = procursus. Scale bars: 0.05 (A–B), 0.02 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.4556722" httpUri="https://zenodo.org/record/4556722/files/figure.png" pageId="4" pageNumber="561">Figs 1D</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2E40FDA4FD12FD3E" box="[606,643,618,645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5D</figureCitation>
); bulb (
<figureCitation id="F1BEA4E0F545FFED2EC4FDA5FCBEFD3E" box="[730,815,619,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Fig. 2C</figureCitation>
) with cone-shaped elongation, with distally sclerotized apophysis (arrow ba in
<figureCitation id="F1BEA4E0F545FFED2DBEFD41FE69FD12" box="[416,504,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Fig. 2C</figureCitation>
), and embolus with a long distal spine (arrow in
<figureCitation id="F1BEA4E0F545FFED2801FD41FBE6FD12" box="[1055,1143,655,681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Fig. 2C</figureCitation>
). Legs with short, vertical setae on metatarsi, without spines or curved setae.
</paragraph>
<paragraph id="693AB865F545FFED2CD9FD19FB61FC1E" blockId="4.[151,1437,150,1042]" pageId="4" pageNumber="561">
<emphasis id="5BF16477F545FFED2CD9FD19FE8FFD4A" bold="true" box="[199,286,727,753]" pageId="4" pageNumber="561">Female</emphasis>
(IZCAS Ar41822): Similar to male, habitus as in
<figureCitation id="F1BEA4E0F545FFED2F7EFD19FC36FD4A" box="[864,935,727,753]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2</figureCitation>
G–H. Total length 1.66 (1.74 with clypeus), carapace 0.78 long, 0.68 wide, opisthosoma 0.96 long, 0.73 wide. Tibia I: 2.44; tibia I L/d: 28. Distance PME-PME 0.24, diameter PME 0.06, distance PME-ALE 0.07, AME absent. Sternum approximately as wide as long (0.53). External female genitalia (
<figureCitation id="F1BEA4E0F545FFED2DDEFC8DFD8CFCE6" box="[448,541,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2A</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2E37FC8CFDDDFCE6" box="[553,588,834,861]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5A</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets laterally on sclerotized plate (arrow ep in
<figureCitation id="F1BEA4E0F545FFED2DBDFCA9FE79FC3A" box="[419,488,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2</figureCitation>
A–B, 5A–B), 0.21 apart. Vulva (
<figureCitation id="F1BEA4E0F545FFED2F4BFCA9FC20FC3A" box="[853,945,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2B</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2FA3FCA8FC4EFC3A" box="[957,991,870,897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5B</figureCitation>
) with a pair of long, narrow pore plates (arrow pp in
<figureCitation id="F1BEA4E0F545FFED2D37FC45FE12FC1E" box="[297,387,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2B</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED2D91FC44FE23FC1E" box="[399,434,906,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5B</figureCitation>
), without serrated sclerites, and with short scape (arrow s in
<figureCitation id="F1BEA4E0F545FFED2847FC45FB25FC1E" box="[1113,1204,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1898]" captionTargetBox="[226,1361,181,1851]" captionTargetId="figure-17@3.[226,1361,181,1851]" captionTargetPageId="3" captionText="FIGURE 2. Belisana bubeng sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view, arrow points at long distal spine on embolus; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate, s = scape. Scale bars: 0.05 (A–D), 0.30 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.4556724" httpUri="https://zenodo.org/record/4556724/files/figure.png" pageId="4" pageNumber="561">Figs 2B</figureCitation>
,
<figureCitation id="F1BEA4E0F545FFED28A1FC44FB72FC1E" box="[1215,1251,906,933]" captionStart="FIGURE 5" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[230,1359,188,1912]" captionTargetId="figure-17@7.[220,1368,181,1919]" captionTargetPageId="7" captionText="FIGURE 5. Drawings of female genitalia in ventral and dorsal views and male left procursus in prolateral and retrolateral views (arrows point at same structures as in photos of each species).A–D. B. bubeng sp. nov.; E–F. B. xiaolongha sp. nov. ep = frontal “epigynal” pocket, f = flap, pp = pore plate, pr = procursus, s = scape. Scale bars: 0.10 (A–B), 0.05 (C–F)." figureDoi="http://doi.org/10.5281/zenodo.4556735" httpUri="https://zenodo.org/record/4556735/files/figure.png" pageId="4" pageNumber="561">5B</figureCitation>
).
</paragraph>
<paragraph id="693AB865F545FFED2CD9FC61FA98FC72" blockId="4.[151,1437,150,1042]" box="[199,1289,942,969]" pageId="4" pageNumber="561">
<emphasis id="5BF16477F545FFED2CD9FC61FE14FC73" bold="true" box="[199,389,942,969]" pageId="4" pageNumber="561">Natural history.</emphasis>
The species was found in leaf litter and collected by small plot direct searching.
</paragraph>
<paragraph id="693AB865F545FFED2CD9FC1CFC15FC56" blockId="4.[151,1437,150,1042]" box="[199,900,978,1005]" pageId="4" pageNumber="561">
<emphasis id="5BF16477F545FFED2CD9FC1CFED0FC57" bold="true" box="[199,321,978,1004]" pageId="4" pageNumber="561">Variation:</emphasis>
<materialsCitation id="D9EDB238F545FFED2D56FC1DFC10FC56" ID-GBIF-Occurrence="3302850306" box="[328,897,979,1005]" collectionCode="IZCAS" collectorName="Tibia I" pageId="4" pageNumber="561" specimenCount="1" typeStatus="paratype">
<collectorName id="C470DDB3F545FFED2D56FC1DFE03FC56" box="[328,402,979,1005]" pageId="4" pageNumber="561">Tibia I</collectorName>
in female
<typeStatus id="B63E06C7F545FFED2E17FC1DFDF8FC56" box="[521,617,979,1005]" pageId="4" pageNumber="561" type="paratype">paratype</typeStatus>
(
<collectionCode id="0F9420A0F545FFED2E66FC1DFD5AFC56" box="[632,715,979,1005]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="561">IZCAS</collectionCode>
Ar41823): 2.24
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="219FEBEEF545FFED2CD9FC38FCD3FBA9" box="[199,834,1014,1042]" pageId="4" pageNumber="561" type="distribution">
<paragraph id="693AB865F545FFED2CD9FC38FCD3FBA9" blockId="4.[151,1437,150,1042]" box="[199,834,1014,1042]" pageId="4" pageNumber="561">
<emphasis id="5BF16477F545FFED2CD9FC38FEF0FBAB" bold="true" box="[199,353,1014,1040]" pageId="4" pageNumber="561">Distribution.</emphasis>
<collectingCountry id="1192F8F5F545FFED2D79FC39FE3BFBAA" box="[359,426,1015,1041]" name="China" pageId="4" pageNumber="561">China</collectingCountry>
(
<collectingRegion id="AB417687F545FFED2DA7FC39FD82FBAA" box="[441,531,1015,1041]" country="China" name="Yunnan" pageId="4" pageNumber="561">Yunnan</collectingRegion>
,
<typeStatus id="B63E06C7F545FFED2E00FC39FDDFFBAA" box="[542,590,1015,1041]" pageId="4" pageNumber="561">type</typeStatus>
locality only;
<figureCitation id="F1BEA4E0F545FFED2EEFFC39FCA5FBA9" box="[753,820,1015,1042]" captionStart="FIGURE 6" captionStartId="8.[151,250,1217,1241]" captionTargetBox="[151,1435,151,1190]" captionTargetId="figure-197@8.[151,1435,151,1190]" captionTargetPageId="8" captionText="FIGURE 6. Distribution records of new species of the genus Belisana. 1. B. bubeng sp. nov.; 2. B. xiaolongha sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4556737" httpUri="https://zenodo.org/record/4556737/files/figure.png" pageId="4" pageNumber="561">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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