treatments-xml/data/CC/22/66/CC226640DA5DFFA3FF7879C2FA82BBB4.xml
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<document id="73F96A3FA02B6E870AF6D9B8A860D934" ID-DOI="10.5281/zenodo.182801" ID-GBIF-Dataset="5d4b0f48-7899-4938-bed3-377fdba78899" ID-ISSN="1175-5326" ID-Zenodo-Dep="182801" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459878118304" checkinUser="plazi" docAuthor="Last, Peter R. &amp; Yearsley, Stephen Mallick And Gordon K." docDate="2008" docId="CC226640DA5DFFA3FF7879C2FA82BBB4" docLanguage="en" docName="zt01812p045.pdf" docOrigin="Zootaxa 1812" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pavoraja umbrosa Last &amp; Yearsley, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="42" masterDocId="301B1E38DA79FF8AFFEF7955FFB9BA2B" masterDocTitle="A review of the Australian skate genus Pavoraja Whitley (Rajiformes: Arhynchobatidae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="37" updateTime="1698227224956" updateUser="plazi">
<mods:mods id="DD80C342CB03107E2557C646758806DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C285038D7E509F884C7E925251F5A167">A review of the Australian skate genus Pavoraja Whitley (Rajiformes: Arhynchobatidae)</mods:title>
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<mods:namePart id="ABD80B409FAC81D8555EF35AE64D948C">Last, Peter R.</mods:namePart>
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<mods:namePart id="0B61384846D38C83EC093FFCAA01102F">Yearsley, Stephen Mallick And Gordon K.</mods:namePart>
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<mods:title id="9A9FCE75445D52E8CF15B32CFB8BF5F4">Zootaxa</mods:title>
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<mods:date id="E4A5DFB92792E4A0C66CA5C84B22376F">2008</mods:date>
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<mods:number id="7FE71527F3E4CFCB37CE3A1A18A4414C">1812</mods:number>
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<treatment id="CC226640DA5DFFA3FF7879C2FA82BBB4" ID-DOI="http://doi.org/10.5281/zenodo.6235104" ID-GBIF-Taxon="119370398" ID-Zenodo-Dep="6235104" LSID="urn:lsid:plazi:treatment:CC226640DA5DFFA3FF7879C2FA82BBB4" httpUri="http://treatment.plazi.org/id/CC226640DA5DFFA3FF7879C2FA82BBB4" lastPageId="41" lastPageNumber="42" pageId="36" pageNumber="37">
<subSubSection id="0C9184DDDA5DFFAEFF7879C2FDE4BAFF" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="4434D756DA5DFFAEFF7879C2FE60BA9A" blockId="36.[151,605,151,212]" box="[151,473,151,177]" pageId="36" pageNumber="37">
<heading id="1F7C603ADA5DFFAEFF7879C2FE60BA9A" bold="true" box="[151,473,151,177]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="76FF0B44DA5DFFAEFF7879C2FE60BA9A" bold="true" box="[151,473,151,177]" pageId="36" pageNumber="37">
<taxonomicName id="838BACD5DA5DFFAEFF7879C2FECCBA9A" ID-CoL="75VPF" box="[151,373,151,177]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="umbrosa" status="sp. nov.">
<emphasis id="76FF0B44DA5DFFAEFF7879C2FECCBA9A" bold="true" box="[151,373,151,177]" italics="true" pageId="36" pageNumber="37">Pavoraja umbrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA5DFFAEFE9379C2FE60BA9A" box="[380,473,151,177]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4434D756DA5DFFAEFF7879EFFDE4BAFF" blockId="36.[151,605,151,212]" box="[151,605,186,212]" pageId="36" pageNumber="37">
(
<figureCitation id="DCB0CBD3DA5DFFAEFF7079EFFF5CBAFF" box="[159,229,186,212]" captionStart="FIGURE 1" captionStartId="5.[151,255,1649,1673]" captionTargetBox="[237,1390,619,1616]" captionTargetId="figure@5.[196,1392,594,1625]" captionTargetPageId="5" captionText="FIGURE 1. Lateral view of right clasper, partially expanded to show components: A, Pavoraja alleni, CSIRO CA 3929, 292 mm TL;" httpUri="https://zenodo.org/record/182802/files/figure.png" pageId="36" pageNumber="37">Figs 1</figureCitation>
f, 2f, 20df, 2123,
<tableCitation id="0909E2EDDA5DFFAEFE3C79EFFD8FBAFF" box="[467,566,186,212]" captionStart="TABLE 3" captionStartId="41.[151,239,151,175]" captionTargetBox="[153,1426,431,2014]" captionTargetPageId="41" captionText="TABLE 3. Proportional measurements of holotypes with ranges and means of other material of Pavoraja pseudonitida sp. nov. (holotype CSIRO H 438 01; 18 paratypes) and P. umbrosa sp. nov. (holotype CSIRO T 1367; 6 paratypes). Values are expressed as percentages of total length." httpUri="http://table.plazi.org/id/10F487DEDA50FFA3FF7879C2FD2FBADC" pageId="36" pageNumber="37" tableUuid="10F487DEDA50FFA3FF7879C2FD2FBADC">Tables 3</tableCitation>
,
<tableCitation id="0909E2EDDA5DFFAEFDAB79EFFDEDBAFF" box="[580,596,186,212]" captionStart="TABLE 4" captionStartId="42.[151,239,155,179]" captionTargetBox="[156,1414,218,1175]" captionTargetPageId="42" captionText="TABLE 4. Neurocranial measurements for Pavoraja species expressed as percentages of nasobasal length." httpUri="http://table.plazi.org/id/10F487DEDA53FFA0FF7879CEFAB2BA99" pageId="36" pageNumber="37" tableUuid="10F487DEDA53FFA0FF7879CEFAB2BA99">4</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0C9184DDDA5DFFAEFF78785FFAB7BB4A" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="4434D756DA5DFFAEFF78785FFAB7BB4A" blockId="36.[151,1436,266,353]" pageId="36" pageNumber="37">
<taxonomicName id="838BACD5DA5DFFAEFF78785FFF43BB0A" ID-CoL="63LD6" box="[151,250,266,289]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="genus">
<emphasis id="76FF0B44DA5DFFAEFF78785FFF43BB0A" box="[151,250,266,289]" italics="true" pageId="36" pageNumber="37">Pavoraja</emphasis>
</taxonomicName>
sp. F:
<bibRefCitation id="201AAAA7DA5DFFAEFEA7785FFDFFBB0A" author="Last" box="[328,582,266,289]" pageId="36" pageNumber="45" refString="Last, P. R. &amp; Stevens, J. D. (1994) Sharks and Rays of Australia. CSIRO Publishing, Melbourne, 513 p." type="book" year="1994">Last and Stevens, 1994</bibRefCitation>
,
<emphasis id="76FF0B44DA5DFFAEFDBC785FFC32BB0A" box="[595,907,266,289]" italics="true" pageId="36" pageNumber="37">
Sharks and rays of
<collectingCountry id="3C9C97C6DA5DFFAEFCC7785FFC32BB0A" box="[808,907,266,289]" name="Australia" pageId="36" pageNumber="37">Australia</collectingCountry>
</emphasis>
, pp 319, fig. 34.13, key fig. 42, pl. 46;
<bibRefCitation id="201AAAA7DA5DFFAEFAD0785FFE38BB6A" author="Last" pageId="36" pageNumber="45" refString="Last, P. R. &amp; Compagno, L. J. V. (1999) Arhynchobatidae, softnose skates, pp. 1457 - 1461. In: Carpenter, K. E. &amp; Niem, V. H. (e ds). FAO species identification guide for fisheries purposes. The marine living resources of the Western Central Pacific, FAO Rome." type="book chapter" year="1999">Last and Compagno, 1999</bibRefCitation>
,
<emphasis id="76FF0B44DA5DFFAEFE64787FFD57BB6A" box="[395,750,298,321]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="838BACD5DA5DFFAEFE64787FFDFCBB6A" ID-CoL="6PY" box="[395,581,298,321]" class="Elasmobranchii" family="Arhynchobatidae" kingdom="Animalia" order="Rajiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="family">Arhynchobatidae</taxonomicName>
, softnose skates
</emphasis>
. In: Carpenter and Niem (eds), 1999, FAO species identification guide for fisheries purposes. The marine living resources of the Western Central Pacific, pp 1461, figs.
</paragraph>
</subSubSection>
<subSubSection id="0C9184DDDA5DFFAFFF7878C7FA87B919" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="4434D756DA5DFFAEFF7878C7FE2CBBFF" blockId="36.[151,1436,402,468]" pageId="36" pageNumber="37">
<bibRef id="9D2B43A0DA5DFFAEFF7878C7FE2CBBFF" pageId="36" pageNumber="37" type="journal volume">
<title id="368F3C60DA5DFFAEFF7878C7FEBABB80" box="[151,259,402,427]" jopScore="0.058884006" pageId="36" pageNumber="37">
<typeStatus id="9B3069F4DA5DFFAEFF7878C7FEBABB80" box="[151,259,402,427]" pageId="36" pageNumber="37" type="holotype">
<emphasis id="76FF0B44DA5DFFAEFF7878C7FEBABB80" box="[151,259,402,427]" italics="true" pageId="36" pageNumber="37">Holotype</emphasis>
</typeStatus>
</title>
.
<journalOrPublisher id="9E4ADD9ADA5DFFAEFEFC78C7FF5DBBFF" jopScore="2.237592" pageId="36" pageNumber="37">
<collectionCode id="229A4F93DA5DFFAEFEFC78C7FED0BB87" box="[275,361,402,428]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="36" pageNumber="37">CSIRO</collectionCode>
T
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TL, adult male, off Stradbroke Island, Queensland,
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,
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,
<quantity id="83737AB3DA5DFFAEFF7878EFFF5DBBFF" box="[151,228,442,468]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="36" pageNumber="37" unit="m" value="530.0">530 m</quantity>
</journalOrPublisher>
,
<part id="302F7DCBDA5DFFAEFF0078EFFEB5BBFF" box="[239,268,442,468]" pageId="36" pageNumber="37" type="volume">22</part>
Nov.
<year id="30239605DA5DFFAEFEBC78EFFE28BBFF" box="[339,401,442,468]" pageId="36" pageNumber="37">1982</year>
.
</bibRef>
</paragraph>
<caption id="10F487DEDA5DFFAEFF787E0DFF68BDB9" httpUri="https://zenodo.org/record/182821/files/figure.png" pageId="36" pageNumber="37" targetBox="[225,1410,606,1841]" targetPageId="36">
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<emphasis id="76FF0B44DA5DFFAEFF787E0DFE9EBD5B" bold="true" box="[151,295,1880,1904]" pageId="36" pageNumber="37">FIGURE 21.</emphasis>
<taxonomicName id="838BACD5DA5DFFAEFEC07E0CFE4EBD5B" box="[303,503,1881,1904]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="umbrosa" status="sp. nov.">
<emphasis id="76FF0B44DA5DFFAEFEC07E0CFE4EBD5B" box="[303,503,1881,1904]" italics="true" pageId="36" pageNumber="37">Pavoraja umbrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA5DFFAEFE107E0DFDECBD44" box="[511,597,1880,1903]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, holotype CSIRO T 1367 adult male 341 mm TL:
<emphasis id="76FF0B44DA5DFFAEFB947E0DFB37BD5B" bold="true" box="[1147,1166,1880,1904]" pageId="36" pageNumber="37">A</emphasis>
, Dorsal view;
<emphasis id="76FF0B44DA5DFFAEFAC17E0DFAF9BD5B" bold="true" box="[1326,1344,1880,1904]" pageId="36" pageNumber="37">B</emphasis>
, Ventral view.
</paragraph>
</caption>
<paragraph id="4434D756DA5CFFAFFF2979CDFA87B919" blockId="37.[151,1437,152,2018]" pageId="37" pageNumber="38">
<emphasis id="76FF0B44DA5CFFAFFF2979CDFEFCBA9A" box="[198,325,152,177]" italics="true" pageId="37" pageNumber="38">
<typeStatus id="9B3069F4DA5CFFAFFF2979CDFEF8BA9A" box="[198,321,152,177]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="528D1CDFDA5CFFAFFEA379CDFE52BA99" box="[332,491,152,178]" pageId="37" pageNumber="38" type="generic">28 specimens</specimenCount>
(
<quantity id="83737AB3DA5CFFAFFE1479CDFD33BA99" box="[507,650,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="2.26" metricValueMax="3.69" metricValueMin="0.83" pageId="37" pageNumber="38" unit="mm" value="226.0" valueMax="369.0" valueMin="83.0">83369 mm</quantity>
TL): AMS I 19093005 (
<specimenCount id="528D1CDFDA5CFFAFFC5279CDFBEABA99" box="[957,1107,152,178]" pageId="37" pageNumber="38" type="generic">5 specimens</specimenCount>
),
<quantity id="83737AB3DA5CFFAFFB8879CDFB03BA99" box="[1127,1210,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="37" pageNumber="38" unit="mm" value="83.0">83 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFF787995FF43BAF1" box="[151,250,192,218]" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" pageId="37" pageNumber="38" unit="mm" value="107.0">107 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFE787995FE40BAF1" box="[407,505,192,218]" metricMagnitude="-1" metricUnit="m" metricValue="1.13" pageId="37" pageNumber="38" unit="mm" value="113.0">113 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFD797995FD41BAF1" box="[662,760,192,218]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" pageId="37" pageNumber="38" unit="mm" value="115.0">115 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFC797995FC40BAF1" box="[918,1017,192,218]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="38" unit="mm" value="140.0">140 mm</quantity>
TL, female, off Wooli, New South
<collectingCountry id="3C9C97C6DA5CFFAFFF7879BDFF65BB29" box="[151,220,232,258]" name="United Kingdom" pageId="37" pageNumber="38">Wales</collectingCountry>
,
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,
<geoCoordinate id="21BFB191DA5CFFAFFE8E79BDFE6EBB29" box="[353,471,231,258]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.71666">153°43' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFE0B79BDFD8ABB29" box="[484,563,232,258]" metricMagnitude="2" metricUnit="m" metricValue="5.029999999999999" pageId="37" pageNumber="38" unit="m" value="503.0">503 m</quantity>
,
<date id="3035F196DA5CFFAFFDD079BDFD65BB29" box="[575,732,232,258]" pageId="37" pageNumber="38" value="1975-10-10">10 Oct. 1975</date>
; AMS I 20301023 (
<specimenCount id="528D1CDFDA5CFFAFFC3779BDFBC5BB29" box="[984,1148,232,258]" pageId="37" pageNumber="38" type="generic">11 specimens</specimenCount>
),
<quantity id="83737AB3DA5CFFAFFB7F79BDFB5ABB29" box="[1168,1251,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="38" unit="mm" value="90.0">90 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFA9179BDFF7CBB01" metricMagnitude="-2" metricUnit="m" metricValue="9.4" pageId="37" pageNumber="38" unit="mm" value="94.0">94 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFE597845FDB2BB01" box="[438,523,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="9.9" pageId="37" pageNumber="38" unit="mm" value="99.0">99 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFD127845FCD9BB01" box="[765,864,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="1.28" pageId="37" pageNumber="38" unit="mm" value="128.0">128 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFBEF7845FBDDBB01" box="[1024,1124,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="2.33" pageId="37" pageNumber="38" unit="mm" value="233.0">233 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFAEB7845FADEBB01" box="[1284,1383,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="2.41" pageId="37" pageNumber="38" unit="mm" value="241.0">241 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFEAA786DFE1EBB79" box="[325,423,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="2.7" pageId="37" pageNumber="38" unit="mm" value="270.0">270 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFD60786DFD48BB79" box="[655,753,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="3.02" pageId="37" pageNumber="38" unit="mm" value="302.0">302 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFC65786DFC55BB79" box="[906,1004,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="3.06" pageId="37" pageNumber="38" unit="mm" value="306.0">306 mm</quantity>
TL, female;
<quantity id="83737AB3DA5CFFAFFB6A786DFB5FBB79" box="[1157,1254,312,338]" metricMagnitude="-1" metricUnit="m" metricValue="3.06" pageId="37" pageNumber="38" unit="mm" value="306.0">306 mm</quantity>
TL, adolescent male;
<quantity id="83737AB3DA5CFFAFFF0F7835FEFABB51" box="[224,323,352,378]" metricMagnitude="-1" metricUnit="m" metricValue="3.34" pageId="37" pageNumber="38" unit="mm" value="334.0">334 mm</quantity>
TL, female, off Wooli, New South
<collectingCountry id="3C9C97C6DA5CFFAFFD037835FC8FBB51" box="[748,822,352,378]" name="United Kingdom" pageId="37" pageNumber="38">Wales</collectingCountry>
,
<geoCoordinate id="21BFB191DA5CFFAFFCAC7835FC11BB51" box="[835,936,351,378]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-29.883333">29°53' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFC587835FB97BB51" box="[951,1070,351,378]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.7">153°42' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFBD47835FB33BB51" box="[1083,1162,352,378]" metricMagnitude="2" metricUnit="m" metricValue="5.02" pageId="37" pageNumber="38" unit="m" value="502.0">502 m</quantity>
,
<date id="3035F196DA5CFFAFFB787835FA84BB51" box="[1175,1341,352,378]" pageId="37" pageNumber="38" value="1977-08-23">23 Aug. 1977</date>
; AMS I 21776006,
<quantity id="83737AB3DA5CFFAFFEC378DDFE29BB89" box="[300,400,392,418]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="37" pageNumber="38" unit="mm" value="250.0">250 mm</quantity>
TL, female, off Danger Point, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFC4778DDFBA9BB89" box="[936,1040,391,418]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-28.033333">28°02' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFBF078DDFB21BB89" box="[1055,1176,391,418]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.95">153°57' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFB4778DDFB41BB89" box="[1192,1272,392,418]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="37" pageNumber="38" unit="m" value="410.0">410 m</quantity>
,
<date id="3035F196DA5CFFAFFAE978DDFA2EBB89" box="[1286,1431,392,418]" pageId="37" pageNumber="38" value="1978-06-01">1 Jun. 1978</date>
; AMS I 21777004 (
<specimenCount id="528D1CDFDA5CFFAFFE7E78E5FD93BBE1" box="[401,554,432,458]" pageId="37" pageNumber="38" type="generic">3 specimens</specimenCount>
),
<quantity id="83737AB3DA5CFFAFFDAD78E5FD11BBE1" box="[578,680,432,458]" metricMagnitude="-1" metricUnit="m" metricValue="2.06" pageId="37" pageNumber="38" unit="mm" value="206.0">206 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFC4E78E5FBBEBBE1" box="[929,1031,432,458]" metricMagnitude="-1" metricUnit="m" metricValue="2.7199999999999998" pageId="37" pageNumber="38" unit="mm" value="272.0">272 mm</quantity>
TL, juvenile male;
<quantity id="83737AB3DA5CFFAFFAEF78E5FADCBBE1" box="[1280,1381,432,458]" metricMagnitude="-1" metricUnit="m" metricValue="3.35" pageId="37" pageNumber="38" unit="mm" value="335.0">335 mm</quantity>
TL, female, off Danger Point, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFD8B788DFD73BBD9" box="[612,714,471,498]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-28.1">28°06' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFD38788DFCF7BBD9" box="[727,846,471,498]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.96666">153°58' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFCB4788DFC10BBD9" box="[859,937,472,498]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="37" pageNumber="38" unit="m" value="410.0">410 m</quantity>
,
<date id="3035F196DA5CFFAFFC5A788DFBFBBBD9" box="[949,1090,472,498]" pageId="37" pageNumber="38" value="1978-06-01">1 Jun. 1978</date>
; AMS I 21789003,
<quantity id="83737AB3DA5CFFAFFAD5788DFA25BBD9" box="[1338,1436,472,498]" metricMagnitude="-1" metricUnit="m" metricValue="3.02" pageId="37" pageNumber="38" unit="mm" value="302.0">302 mm</quantity>
TL, juvenile male, off Danger Point, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFD017B55FCECB831" box="[750,853,511,538]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-27.916666">27°55' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFC8C7B55FC63B831" box="[867,986,511,538]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="154.05">154°03' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFC067B55FB8EB831" box="[1001,1079,512,538]" metricMagnitude="2" metricUnit="m" metricValue="5.4799999999999995" pageId="37" pageNumber="38" unit="m" value="548.0">548 m</quantity>
,
<date id="3035F196DA5CFFAFFBAA7B55FB62B831" box="[1093,1243,512,538]" pageId="37" pageNumber="38" value="1978-11-06">6 Nov. 1978</date>
; AMS I 21795 0
<quantity id="83737AB3DA5CFFAFFF487B7DFE8BB869" box="[167,306,552,578]" metricMagnitude="1" metricUnit="m" metricValue="1.1329" pageId="37" pageNumber="38" unit="mm" value="11329.0">11, 329 mm</quantity>
TL, female, off Danger Point, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFCD67B7DFC27B869" box="[825,926,551,578]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-28.05">28°03' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFC437B7DFB98B869" box="[940,1057,551,578]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="154.06667">154°04' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFBC17B7DFBC2B869" box="[1070,1147,552,578]" metricMagnitude="2" metricUnit="m" metricValue="7.31" pageId="37" pageNumber="38" unit="m" value="731.0">731 m</quantity>
,
<date id="3035F196DA5CFFAFFB677B7DFAA5B869" box="[1160,1308,552,578]" pageId="37" pageNumber="38" value="1978-11-06">6 Nov. 1978</date>
;
<collectionCode id="229A4F93DA5CFFAFFAC77B7DFAC7B869" box="[1320,1406,552,578]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38">CSIRO</collectionCode>
H 15701,
<quantity id="83737AB3DA5CFFAFFF117B05FED9B841" box="[254,352,592,618]" metricMagnitude="-1" metricUnit="m" metricValue="3.4899999999999998" pageId="37" pageNumber="38" unit="mm" value="349.0">349 mm</quantity>
TL, adult male, east of Wooli, New South
<collectingCountry id="3C9C97C6DA5CFFAFFCB37B05FC1FB841" box="[860,934,592,618]" name="United Kingdom" pageId="37" pageNumber="38">Wales</collectingCountry>
,
<geoCoordinate id="21BFB191DA5CFFAFFC5D7B05FBA1B841" box="[946,1048,591,618]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-29.833334">29°50' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFBCA7B05FB25B841" box="[1061,1180,591,618]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.68333">153°41' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFB467B05FB4FB841" box="[1193,1270,592,618]" metricMagnitude="2" metricUnit="m" metricValue="3.6" pageId="37" pageNumber="38" unit="m" value="360.0">360 m</quantity>
,
<date id="3035F196DA5CFFAFFAEC7B05FA2EB841" box="[1283,1431,592,618]" pageId="37" pageNumber="38" value="1982-07-18">18 Jul. 1982</date>
;
<collectionCode id="229A4F93DA5CFFAFFF787B2DFF57B8B9" box="[151,238,632,658]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38">CSIRO</collectionCode>
T
<quantity id="83737AB3DA5CFFAFFEE27B2DFE0EB8B9" box="[269,439,632,658]" metricMagnitude="3" metricUnit="m" metricValue="1.3633689999999998" pageId="37" pageNumber="38" unit="mm" value="1363369.0">1363, 369 mm</quantity>
TL, adult male;
<collectionCode id="229A4F93DA5CFFAFFD967B2DFD76B8B9" box="[633,719,632,658]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38">CSIRO</collectionCode>
T
<quantity id="83737AB3DA5CFFAFFD007B2DFC20B8B9" box="[751,921,632,658]" metricMagnitude="3" metricUnit="m" metricValue="1.364309" pageId="37" pageNumber="38" unit="mm" value="1364309.0">1364, 309 mm</quantity>
TL, female, east of Moreton Island, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFF0C7BF5FEF1B891" box="[227,328,671,698]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-27.2">27°12' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFEBA7BF5FE73B891" box="[341,458,671,698]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.86667">153°52' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFE397BF5FD9DB891" box="[470,548,672,698]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="37" pageNumber="38" unit="m" value="555.0">555 m</quantity>
,
<date id="3035F196DA5CFFAFFDC07BF5FD77B891" box="[559,718,672,698]" pageId="37" pageNumber="38" value="1983-05-10">10 May 1983</date>
;
<collectionCode id="229A4F93DA5CFFAFFD357BF5FC89B891" box="[730,816,672,698]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38">CSIRO</collectionCode>
T
<quantity id="83737AB3DA5CFFAFFCBF7BF5FC42B891" box="[848,1019,672,698]" metricMagnitude="3" metricUnit="m" metricValue="1.3653" pageId="37" pageNumber="38" unit="mm" value="1365300.0">1365, 300 mm</quantity>
TL, juvenile male, off North Stradbroke Island, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFE207B9DFD8AB8C9" box="[463,563,711,738]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-27.583334">27°35' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFDAF7B9DFD0CB8C9" box="[576,693,711,738]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.95">153°57' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFD2E7B9DFCB7B8C9" box="[705,782,712,738]" metricMagnitude="2" metricUnit="m" metricValue="5.45" pageId="37" pageNumber="38" unit="m" value="545.0">545 m</quantity>
,
<date id="3035F196DA5CFFAFFCF67B9DFC00B8C9" box="[793,953,712,738]" pageId="37" pageNumber="38" value="1983-03-31">31 Mar. 1983</date>
;
<collectionCode id="229A4F93DA5CFFAFFC2B7B9DFBA3B8C9" box="[964,1050,712,738]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38">CSIRO</collectionCode>
T
<quantity id="83737AB3DA5CFFAFFBD57B9DFB5DB8C9" box="[1082,1252,712,738]" metricMagnitude="3" metricUnit="m" metricValue="1.366319" pageId="37" pageNumber="38" unit="mm" value="1366319.0">1366, 319 mm</quantity>
TL, female, off North Stradbroke Island, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFDB87BA5FD05B921" box="[599,700,751,778]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-27.316668">27°19' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFD277BA5FC84B921" box="[712,829,751,778]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.88333">153°53' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFCA67BA5FC2FB921" box="[841,918,752,778]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="38" unit="m" value="600.0">600 m</quantity>
,
<date id="3035F196DA5CFFAFFC4E7BA5FB86B921" box="[929,1087,752,778]" pageId="37" pageNumber="38" value="1983-05-10">10 May 1983</date>
;
<collectionCode id="229A4F93DA5CFFAFFBA47BA5FB18B921" box="[1099,1185,752,778]" httpUri="http://grbio.org/cool/0k2v-y6wz" name="Australian National Fish Collection" pageId="37" pageNumber="38">CSIRO</collectionCode>
T
<quantity id="83737AB3DA5CFFAFFB2E7BA5FAD3B921" box="[1217,1386,752,778]" metricMagnitude="3" metricUnit="m" metricValue="1.368304" pageId="37" pageNumber="38" unit="mm" value="1368304.0">1368, 304 mm</quantity>
TL, adolescent male, off North Stradbroke Island, Queensland,
<geoCoordinate id="21BFB191DA5CFFAFFCA37A4DFC08B919" box="[844,945,791,818]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="925" value="-27.583334">27°35' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5CFFAFFC517A4DFB8DB919" box="[958,1076,791,818]" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="925" value="153.95">153°57' E</geoCoordinate>
,
<quantity id="83737AB3DA5CFFAFFBAE7A4DFB37B919" box="[1089,1166,792,818]" metricMagnitude="2" metricUnit="m" metricValue="5.45" pageId="37" pageNumber="38" unit="m" value="545.0">545 m</quantity>
,
<date id="3035F196DA5CFFAFFB767A4DFA83B919" box="[1177,1338,792,818]" pageId="37" pageNumber="38" value="1983-03-31">31 Mar. 1983</date>
.
</paragraph>
</subSubSection>
<subSubSection id="0C9184DDDA5CFFAFFF297A15FDD0BEE9" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="4434D756DA5CFFAFFF297A15FDD0BEE9" blockId="37.[151,1437,152,2018]" pageId="37" pageNumber="38">
<emphasis id="76FF0B44DA5CFFAFFF297A15FEFCB972" box="[198,325,832,857]" italics="true" pageId="37" pageNumber="38">Diagnosis.</emphasis>
A species of
<taxonomicName id="838BACD5DA5CFFAFFE047A15FDE1B972" box="[491,600,832,857]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
<emphasis id="76FF0B44DA5CFFAFFE047A15FDE1B972" box="[491,600,832,857]" italics="true" pageId="37" pageNumber="38">Pavoraja</emphasis>
</taxonomicName>
with: with a relatively long, narrow tail, length 5660% TL, width at midlength 1.31.8% TL; large orbits, diameter 4.35.4% TL; broad nasal curtain, total width 6.77.6% TL; orbital thorns large, mostly 3 on posteromedial margin; interorbital usually absent; spiracular and scapular thorns absent; nuchal pore patch indistinct, not preceded by a nuchal thorn; thorns of tail series normally absent near first dorsal fin, lateral series often absent; dorsal fin placement variable, fins confluent to widely spaced; epichordal lobe of caudal fin not confluent with second dorsal fin, base of lobe equal to in length or shorter than dorsal-fin bases; tooth rows in lower jaw 3642; predorsal caudal centra 7483; interdorsal vertebrae 1013; pectoral radials 6871; greyish brown, often with darker dusky blotches, lacking pattern of white spots; dorsal fins uniformly greyish to blackish; epichordal lobe pale to dusky, not strongly demarcated from dorsal fins; ventral surface mostly dark.
</paragraph>
</subSubSection>
<subSubSection id="0C9184DDDA5CFFADFF297D85FE44BCE1" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="4434D756DA5CFFAFFF297D85FC55BC89" blockId="37.[151,1437,152,2018]" pageId="37" pageNumber="38">
<emphasis id="76FF0B44DA5CFFAFFF297D85FEE1BEC2" box="[198,344,1232,1257]" italics="true" pageId="37" pageNumber="38">Description.</emphasis>
Disc 1.15 (1.071.19) times as broad as long; maximum angle in front of spiracles 103° (94 101°); anterior margin weakly convex to double concave in females and immature males, deeply concave opposite spiracles in males; posterior margin strongly convex; outer corners broadly rounded. Snout width at axis through anterior border of orbits 60% (6365% in mature male, 71% in female
<typeStatus id="9B3069F4DA5CFFAFFB797C1DFAB1BF49" box="[1174,1288,1352,1378]" pageId="37" pageNumber="38" type="paratype">paratypes</typeStatus>
) of distance from tip of snout to axil of pectoral fins. Pelvic-fin anterior margin 72% (7488%) of distance from origin of anterior lobe to posterior extremity of fin. Tail length 1.25 (1.101.37) times disc length; widths at midlength and at axils of pelvic fins 32% (2534%) and 84% (7384%) of orbit diameter respectively; skin fold very narrow anteriorly, extending along ventrolateral surface from near pelvic-fin tip to near origin of hypochordal lobe of caudal fin, much wider beneath dorsal fins (subequal or slightly exceeding height of epichordal lobe of caudal fin). Interdorsal distance variable, interdorsal membrane continuous or fins separated by as much as epichordal lobe length; epichordal caudal-fin lobe separated slightly from base of second dorsal fin (by a distance of up to base length of lobe), base shorter than bases of dorsal fins.
</paragraph>
<paragraph id="4434D756DA5CFFAFFF297FE5FE2FBD41" blockId="37.[151,1437,152,2018]" pageId="37" pageNumber="38">Preocular length 2.08 (2.092.53) times longer than orbit diameter; preoral length 2.40 (2.192.55) times internarial distance. Orbit diameter 1.57 (1.661.95) times interorbital distance; 2.12 (1.692.18) times length of spiracles. Internarial distance 0.44 (0.400.48) in distance between first gill slits; 0.78 (0.660.85) in distance between fifth gill slits. Length of first gill slit 1.41 (1.261.50) times length of fifth gill slit; 0.18 (0.17 0.20) in mouth width.</paragraph>
<paragraph id="4434D756DA5CFFADFF297E2DFC4FB8C9" blockId="37.[151,1437,152,2018]" lastBlockId="39.[151,1437,152,2018]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
Dorsal surfaces of disc, tail and posterior lobes of pelvic fins densely covered with granular denticles. Dorsal fins denticulate. Claspers, anterior lobes of pelvic fins, skin folds on tail, caudal lobes and entire ventral surface naked. Orbit with 24 thorns on anteromedial margin, 14 (mainly 3) on posteromedial margin (often damaged); one interorbital thorn rarely present; interspiracular and scapular thorns absent. Prenuchal and nuchal thorns 04 (
<typeStatus id="9B3069F4DA5FFFACFE477995FDA9BAF1" box="[424,528,192,218]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
damaged, mainly 2 or
<quantity id="83737AB3DA5FFFACFCF27995FCF0BAF1" box="[797,841,192,218]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="38" pageNumber="39" unit="in" value="3.0">3 in</quantity>
<typeStatus id="9B3069F4DA5FFFACFCBF7995FC7CBAF1" box="[848,965,192,218]" pageId="38" pageNumber="39" type="paratype">paratypes</typeStatus>
), situated posterior to pale, pored prenuchal area; thorns well developed, prominent when undamaged. Tail with 13 (mainly 3) series of thorns; dorsolateral series short, variable (usually absent in specimens &lt;
<quantity id="83737AB3DA5FFFACFC877845FC73BB01" box="[872,970,272,298]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="38" pageNumber="39" unit="mm" value="240.0">240 mm</quantity>
TL), rarely extending onto disc, sparse or absent on posterior half of tail, thorns often staggered; median series more regular, never extending onto disc, rudimentary or absent before dorsal fins; interdorsal thorns never present. Adult males with 23 irregular rows of small non-retractable alar thorns; alar thorns sometimes paired, not forming a dense patch; malar thorns much smaller, situated anterolaterally to alar thorns, patch extending anteriorly well short of greatest concavity of anterior margin of disc, to about level of pectoral-fin apex. Smallest juvenile (AMS I 19093 005): disc uniformly granular, no obvious orbital or nuchal thorns; tail thorns in a linear median row, equally Claspers slender (
<figureCitation id="DCB0CBD3DA5EFFADFE7579CDFE4EBA99" box="[410,503,152,178]" captionStart="FIGURE 20" captionStartId="34.[151,255,1854,1878]" captionTargetBox="[155,1434,770,1766]" captionTargetId="figure@34.[155,1436,748,1827]" captionTargetPageId="34" captionText="FIGURE 20. Right clasper cartilages of Pavoraja pseudonitida sp. nov., holotype CSIRO H 438 01, adult male 372 mm TL: A, Lateral view, partially expanded with dorsal and ventral terminal cartilages shown separately; B, Dorsal view; C, Ventral view. Right clasper cartilages of P. umbrosa sp. nov., paratype CSIRO T 1363, adult male 369 mm TL; D, Lateral view, partially expanded with dorsal and ventral terminal cartilages shown separately; E, Dorsal view; F, Ventral view. Abbreviations as in Fig. 6." httpUri="https://zenodo.org/record/182820/files/figure.png" pageId="39" pageNumber="40">Figs 20</figureCitation>
df); pseudorhipidion continuing distally under sentinel as fold of integument, finally becoming fused to glans floor; spur well developed though proximal section poorly defined, distal tip slightly recurved, lying inside glans margin; rhipidion moderately well developed, extending from middle to distal one-third of glans, proximal section very thin, thicker distal section lying over base of sentinel; sentinel well developed, oval, covered by integument which is thicker and more fleshy over lateral region, extending from level of slit to past tip of glans; spike hidden within sentina; axial cartilage straight, slender; dorsal marginal not expanded distally; pseudorhipidion rod-shaped though thickened before thinning distally to a point; ventral marginal with an evenly convex distal margin; dorsal terminal 1 and ventral terminal cartilages narrowly joined on ventral aspect of glans or connected by tissue, together forming sheath-like covering over central half of glans; dorsal terminal 1 membranous; ventral terminal mostly membranous, with thickened area immediately distal to short proximal arm, penetrating glans and lying against dorsal margin of proximal extension of accessory terminal 1; dorsal terminal 2 with poorly defined cartilaginous projection joined to axial near level of accessory terminal 2 base by connective tissue (incomplete terminal bridge), lateral margin irregular; dorsal terminal 3 moderately large; accessory terminal 1 with a bow-shaped distal extension forming sentinel, extending slightly past distal margin of accessory terminal 2.
</paragraph>
<caption id="10F487DEDA5FFFACFF787EFCFBFCBDC8" httpUri="https://zenodo.org/record/182822/files/figure.png" pageId="38" pageNumber="39" targetBox="[153,1389,573,1909]" targetPageId="38">
<paragraph id="4434D756DA5FFFACFF787EFCFBFCBDC8" blockId="38.[151,1434,1961,2019]" pageId="38" pageNumber="39">
<emphasis id="76FF0B44DA5FFFACFF787EFCFE93BDEA" bold="true" box="[151,298,1961,1985]" pageId="38" pageNumber="39">FIGURE 22.</emphasis>
<taxonomicName id="838BACD5DA5FFFACFEDB7EFFFE44BDEA" box="[308,509,1962,1985]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="umbrosa" status="sp. nov.">
<emphasis id="76FF0B44DA5FFFACFEDB7EFFFE44BDEA" box="[308,509,1962,1985]" italics="true" pageId="38" pageNumber="39">Pavoraja umbrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA5FFFACFDE87EFCFDE6BDEB" box="[519,607,1961,1984]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="76FF0B44DA5FFFACFD827EFCFD39BDEA" bold="true" box="[621,640,1961,1985]" pageId="38" pageNumber="39">A</emphasis>
, paratype CSIRO T 1366, female 319 mm TL, dorsal view;
<emphasis id="76FF0B44DA5FFFACFA907EFCFA28BDEA" bold="true" box="[1407,1425,1961,1985]" pageId="38" pageNumber="39">B</emphasis>
, paratype AMS I 19093005 (1 of 5 specimens), juvenile male 83 mm TL, dorsal view.
</paragraph>
</caption>
<paragraph id="4434D756DA5EFFADFF297BA5FBDEB971" blockId="39.[151,1437,152,2018]" pageId="39" pageNumber="40">Neurocranium of CSIRO T 1364 typical of genus; with very small accessory lateral cartilages; 2 foramina near apex of rostral node; rostral shaft relatively long, longer than length of basal fenestra and extending about halfway to rostral base; posterior fontanelle slightly longer than anterior fontanelle.</paragraph>
<paragraph id="4434D756DA5EFFADFF297A3DFECCB981" blockId="39.[151,1437,152,2018]" pageId="39" pageNumber="40">Scapulocoracoid expanded anteriorly, with 2 postventral foramina; extra condyle sometimes present on neopterygial ridge.</paragraph>
<paragraph id="4434D756DA5EFFADFF297AEDFC76BE59" blockId="39.[151,1437,152,2018]" pageId="39" pageNumber="40">
<emphasis id="76FF0B44DA5EFFADFF297AEDFE8AB9FA" box="[198,307,952,977]" italics="true" pageId="39" pageNumber="40">Meristics</emphasis>
(n=6). Tooth rows in upper jaw 41 (3646), in lower jaw 42 (3642). Vertebral counts: monospondylous centra 28 (2730), predorsal caudal centra 80 (7483), interdorsal centra 12 (113), diplospondylous centra 116 (106117), total centra 144 (134147). Pectoral-fin counts: propterygial radials 32 (3133), mesopterygial radials 12 (1114), metapterygial radials 26 (2327), total radials 70 (6871). Pelvic-fin counts: males 4 (34) + 19 (1719) radials, females 3 + 1920 radials.
</paragraph>
<paragraph id="4434D756DA5EFFADFF297DD5FD7FBC51" blockId="39.[151,1437,152,2018]" pageId="39" pageNumber="40">
<emphasis id="76FF0B44DA5EFFADFF297DD5FEF7BEB2" box="[198,334,1152,1177]" italics="true" pageId="39" pageNumber="40">Coloration.</emphasis>
Preserved and live colours similar. Dorsal surface of disc, tail and posterior lobes of pelvic fins medium brown to dark brown, usually slightly mottled, with large, diffuse-edged dark brown blotches, no light-coloured spots but sometimes with indistinct paler blotches; membrane covering orbit semi-translucent to slightly pigmented; mid area of snout and outer part of disc only slightly paler than central part of disc; nuchal pore patch indistinct, not demarcated by a pale blotch; lateral skin fold dusky to almost black distally. Claspers brownish dorsally with blackish lateral margin, posterolateral extremity of glans white, dusky ventrally. Dorsal fins generally uniformly greyish to blackish; epichordal lobe of caudal fin dusky to pale, not usually strongly demarcated from dorsal fins. Ventral surface relatively dark, dark brownish to greyish; paler areas on snout, oronasal region, gill slits, cloaca and sometimes basal half of anterior pelvic-fin lobe; darkest areas between gills and around pectoral-fin corners; tail variably coloured with white and dark areas. Smallest juveniles (&lt;
<quantity id="83737AB3DA5EFFADFECC7F45FE3EBC01" box="[291,391,1552,1578]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="40" unit="mm" value="100.0">100 mm</quantity>
TL) with fine black spots on disc (plain coloured in one
<typeStatus id="9B3069F4DA5EFFADFBD77F45FB24BC01" box="[1080,1181,1552,1578]" pageId="39" pageNumber="40" type="paratype">paratype</typeStatus>
from AMS I 19093 0 0 5,&gt;
<quantity id="83737AB3DA5EFFADFF0B7F6DFEF1BC79" box="[228,328,1592,1618]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="39" pageNumber="40" unit="mm" value="110.0">110 mm</quantity>
TL); predorsal tail with irregular spotting and indistinct bands; prominent dark band below dorsal fins; ventral surface translucent or white.
</paragraph>
<paragraph id="4434D756DA5EFFADFF297FDDFE44BCE1" blockId="39.[151,1437,152,2018]" pageId="39" pageNumber="40">
<emphasis id="76FF0B44DA5EFFADFF297FDDFF44BC8A" box="[198,253,1672,1697]" italics="true" pageId="39" pageNumber="40">Size.</emphasis>
At least
<quantity id="83737AB3DA5EFFADFE817FDDFE6BBC89" box="[366,466,1672,1698]" metricMagnitude="-1" metricUnit="m" metricValue="3.69" pageId="39" pageNumber="40" unit="mm" value="369.0">369 mm</quantity>
TL and about
<quantity id="83737AB3DA5EFFADFD657FDDFD57BC89" box="[650,750,1672,1698]" metricMagnitude="-1" metricUnit="m" metricValue="1.91" pageId="39" pageNumber="40" unit="mm" value="191.0">191 mm</quantity>
disc width. Smallest mature male
<quantity id="83737AB3DA5EFFADFB7C7FDDFB41BC89" box="[1171,1272,1672,1698]" metricMagnitude="-1" metricUnit="m" metricValue="3.4899999999999998" pageId="39" pageNumber="40" unit="mm" value="349.0">349 mm</quantity>
TL. Smallest post-natal juvenile
<quantity id="83737AB3DA5EFFADFE987FE5FE73BCE1" box="[375,458,1712,1738]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="39" pageNumber="40" unit="mm" value="83.0">83 mm</quantity>
TL.
</paragraph>
</subSubSection>
<subSubSection id="0C9184DDDA5EFFADFF297F8DFA2FBD31" pageId="39" pageNumber="40" type="distribution">
<paragraph id="4434D756DA5EFFADFF297F8DFA2FBD31" blockId="39.[151,1437,152,2018]" pageId="39" pageNumber="40">
<emphasis id="76FF0B44DA5EFFADFF297F8DFEE4BCDA" box="[198,349,1752,1777]" italics="true" pageId="39" pageNumber="40">Distribution.</emphasis>
Upper continental slope off eastern
<collectingCountry id="3C9C97C6DA5EFFADFCCF7F8DFC37BCD9" box="[800,910,1752,1778]" name="Australia" pageId="39" pageNumber="40">Australia</collectingCountry>
, between east of Wooli, New South
<collectingCountry id="3C9C97C6DA5EFFADFABA7F8DFA25BCD9" box="[1365,1436,1752,1778]" name="United Kingdom" pageId="39" pageNumber="40">Wales</collectingCountry>
(
<geoCoordinate id="21BFB191DA5EFFADFF707E55FEBCBD31" box="[159,261,1791,1818]" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="-29.883333">29°53' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5EFFADFEFD7E55FE3EBD31" box="[274,391,1791,1818]" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="153.7">153°42' E</geoCoordinate>
) and east of Moreton Island, Queensland (
<geoCoordinate id="21BFB191DA5EFFADFC957E55FC59BD31" box="[890,992,1791,1818]" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="-27.2">27°12' S</geoCoordinate>
,
<geoCoordinate id="21BFB191DA5EFFADFC027E55FBDABD31" box="[1005,1123,1791,1818]" direction="east" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="153.86667">153°52' E</geoCoordinate>
), in depths of
<quantity id="83737AB3DA5EFFADFAE57E55FA2BBD31" box="[1290,1426,1792,1818]" metricMagnitude="2" metricUnit="m" metricValue="5.455" metricValueMax="7.31" metricValueMin="3.6" pageId="39" pageNumber="40" unit="m" value="545.5" valueMax="731.0" valueMin="360.0">
360
<specimenCount id="528D1CDFDA5EFFADFAA97E55FA2BBD31" box="[1350,1426,1792,1818]" pageId="39" pageNumber="40" type="generic">731 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="0C9184DDDA5EFFA2FF297E7DFA30BAF1" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="etymology">
<paragraph id="4434D756DA5EFFADFF297E7DFA23BD69" blockId="39.[151,1437,152,2018]" box="[198,1434,1832,1858]" pageId="39" pageNumber="40">
<emphasis id="76FF0B44DA5EFFADFF297E7DFEF3BD6A" box="[198,330,1832,1857]" italics="true" pageId="39" pageNumber="40">Etymology.</emphasis>
From the Latin
<emphasis id="76FF0B44DA5EFFADFDE77E7DFDC0BD6A" box="[520,633,1832,1857]" italics="true" pageId="39" pageNumber="40">umbrosus</emphasis>
(shady) in allusion to the uniform greyish dorsal surface of the disc.
</paragraph>
<paragraph id="4434D756DA5EFFA2FF297E05FA30BAF1" blockId="39.[151,1437,152,2018]" lastBlockId="40.[151,1436,152,218]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="76FF0B44DA5EFFADFF297E05FDE8BD42" box="[198,593,1872,1897]" italics="true" pageId="39" pageNumber="40">
Comparisons.
<taxonomicName id="838BACD5DA5EFFADFE9B7E05FDE8BD42" box="[372,593,1872,1897]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="umbrosa">Pavoraja umbrosa</taxonomicName>
</emphasis>
is similar morphometrically and meristically to its eastern Australian congener,
<taxonomicName id="838BACD5DA5EFFADFEFF7E2DFE7DBDBA" box="[272,452,1912,1937]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="pseudonitida">
<emphasis id="76FF0B44DA5EFFADFEFF7E2DFE7DBDBA" box="[272,452,1912,1937]" italics="true" pageId="39" pageNumber="40">P. pseudonitida</emphasis>
</taxonomicName>
, but lacks interorbital thorns (sometimes present in
<taxonomicName id="838BACD5DA5EFFADFBC87E2DFB62BDBA" box="[1063,1243,1912,1937]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="pseudonitida">
<emphasis id="76FF0B44DA5EFFADFBC87E2DFB62BDBA" box="[1063,1243,1912,1937]" italics="true" pageId="39" pageNumber="40">P. pseudonitida</emphasis>
</taxonomicName>
), the dorsal surface is greyish without small white spots (otherwise spotted), the ventral surface is dusky (rather than pale), and the epichordal lobe of caudal fin is mainly dusky and is not usually strongly demarcated from the dorsal fins (in
<taxonomicName id="838BACD5DA51FFA2FF1C79CDFE11BA9A" box="[243,424,152,177]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="pseudonitida">
<emphasis id="76FF0B44DA51FFA2FF1C79CDFE11BA9A" box="[243,424,152,177]" italics="true" pageId="40" pageNumber="41">P. pseudonitida</emphasis>
</taxonomicName>
the dorsal fins are strongly contrasted against a pale epichordal lobe).
<taxonomicName id="838BACD5DA51FFA2FB0479CDFF70BAF2" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="umbrosa">
<emphasis id="76FF0B44DA51FFA2FB0479CDFF70BAF2" italics="true" pageId="40" pageNumber="41">Pavoraja umbrosa</emphasis>
</taxonomicName>
differs from other members of the genus by combinations of characters given in the key and diagnoses.
</paragraph>
</subSubSection>
<caption id="10F487DEDA51FFA2FF787FCBFB9ABCF3" httpUri="https://zenodo.org/record/182823/files/figure.png" pageId="40" pageNumber="41" targetBox="[214,1361,374,1635]" targetPageId="40">
<paragraph id="4434D756DA51FFA2FF787FCBFB9ABCF3" blockId="40.[151,1435,1694,1753]" pageId="40" pageNumber="41">
<emphasis id="76FF0B44DA51FFA2FF787FCBFE90BC9D" bold="true" box="[151,297,1694,1718]" pageId="40" pageNumber="41">FIGURE 23.</emphasis>
Neurocranium of
<taxonomicName id="838BACD5DA51FFA2FE197FCAFD06BC9D" box="[502,703,1695,1718]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="umbrosa" status="sp. nov.">
<emphasis id="76FF0B44DA51FFA2FE197FCAFD06BC9D" box="[502,703,1695,1718]" italics="true" pageId="40" pageNumber="41">Pavoraja umbrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA51FFA2FD277FCAFC99BC9D" box="[712,800,1695,1718]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, paratype CSIRO T 1364, female 309 mm TL:
<emphasis id="76FF0B44DA51FFA2FADE7FCBFAFDBC9D" bold="true" box="[1329,1348,1694,1718]" pageId="40" pageNumber="41">A</emphasis>
, Dorsal view;
<emphasis id="76FF0B44DA51FFA2FF367F94FF52BCF2" bold="true" box="[217,235,1729,1753]" pageId="40" pageNumber="41">B</emphasis>
, Posterior view;
<emphasis id="76FF0B44DA51FFA2FE4E7F94FE0DBCF2" bold="true" box="[417,436,1729,1753]" pageId="40" pageNumber="41">C</emphasis>
, Ventral view;
<emphasis id="76FF0B44DA51FFA2FDB87F94FDD3BCF2" bold="true" box="[599,618,1729,1753]" pageId="40" pageNumber="41">D</emphasis>
, Lateral view. Abbreviations as in Fig. 5.
</paragraph>
</caption>
<subSubSection id="0C9184DDDA51FFA3FF297E59FA82BBB4" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="discussion">
<paragraph id="4434D756DA51FFA2FF297E59FB5BBD65" blockId="40.[151,1435,1804,1870]" pageId="40" pageNumber="41">
<emphasis id="76FF0B44DA51FFA2FF297E59FDA1BD0E" box="[198,536,1804,1829]" italics="true" pageId="40" pageNumber="41">
Remarks.
<taxonomicName id="838BACD5DA51FFA2FED27E59FDA1BD0E" box="[317,536,1804,1829]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="umbrosa">Pavoraja umbrosa</taxonomicName>
</emphasis>
belongs to the Central Eastern marine biogeographic province of
<collectingCountry id="3C9C97C6DA51FFA2FADF7E59FA22BD0D" box="[1328,1435,1804,1830]" name="Australia" pageId="40" pageNumber="41">Australia</collectingCountry>
where it is found primarily in the upper slope and midupper slope biomes (
<bibRefCitation id="201AAAA7DA51FFA2FBFE7E61FB6FBD65" author="Last" box="[1041,1238,1844,1870]" pageId="40" pageNumber="45" refString="Last, P. R., Lyne, V., Yearsley, G., Gledhill, D., Gomon, M., Rees, T. &amp; White, W. (2005) Validation of national fish datasets for the regionalisation of the Australian continental slope and shelf (&gt; 40 m depth). National Oceans Office, Hobart, 99 p." type="book" year="2005">
Last
<emphasis id="76FF0B44DA51FFA2FBA27E61FB31BD66" box="[1101,1160,1844,1869]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
, 2005
</bibRefCitation>
).
</paragraph>
<caption id="10F487DEDA50FFA3FF7879C2FD2FBADC" ID-Table-UUID="10F487DEDA50FFA3FF7879C2FD2FBADC" httpUri="http://table.plazi.org/id/10F487DEDA50FFA3FF7879C2FD2FBADC" pageId="41" pageNumber="42" targetBox="[153,1426,431,2014]" targetIsTable="true" targetPageId="41">
<paragraph id="4434D756DA50FFA3FF7879C2FD2FBADC" blockId="41.[151,1436,151,247]" pageId="41" pageNumber="42">
<emphasis id="76FF0B44DA50FFA3FF7879C2FEB3BA84" bold="true" box="[151,266,151,175]" pageId="41" pageNumber="42">TABLE 3.</emphasis>
Proportional measurements of holotypes with ranges and means of other material of
<taxonomicName id="838BACD5DA50FFA3FB4879CDFA25BA84" box="[1191,1436,152,175]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="pseudonitida" status="sp. nov.">
<emphasis id="76FF0B44DA50FFA3FB4879CDFA25BA84" box="[1191,1436,152,175]" italics="true" pageId="41" pageNumber="42">Pavoraja pseudonitida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA50FFA3FF7879EFFF50BAFA" box="[151,233,186,209]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
(holotype CSIRO H 43801; 18 paratypes) and
<taxonomicName id="838BACD5DA50FFA3FD0279EFFCDABAFA" box="[749,867,186,209]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="umbrosa" status="sp. nov.">
<emphasis id="76FF0B44DA50FFA3FD0279EFFCDABAFA" box="[749,867,186,209]" italics="true" pageId="41" pageNumber="42">P. umbrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA50FFA3FC8579EFFC04BAFA" box="[874,957,186,209]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
(holotype CSIRO T 1367; 6 paratypes). Values are expressed as percentages of total length.
</paragraph>
</caption>
<paragraph id="4434D756DA50FFA3FD5F7842FA8ABB07" blockId="41.[153,1425,279,415]" box="[688,1331,279,300]" pageId="41" pageNumber="42">
<taxonomicName id="838BACD5DA50FFA3FD5F7842FCFEBB07" box="[688,839,279,300]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="pseudonitida" status="sp. nov.">
<emphasis id="76FF0B44DA50FFA3FD5F7842FCFEBB07" bold="true" box="[688,839,279,300]" italics="true" pageId="41" pageNumber="42">P. pseudonitida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA50FFA3FCA37842FC2EBB07" box="[844,919,279,300]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="838BACD5DA50FFA3FB9B7842FB5ABB07" box="[1140,1251,279,300]" class="Elasmobranchii" family="Arhynchobatidae" genus="Pavoraja" kingdom="Animalia" order="Rajiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="umbrosa" status="sp. nov.">
<emphasis id="76FF0B44DA50FFA3FB9B7842FB5ABB07" bold="true" box="[1140,1251,279,300]" italics="true" pageId="41" pageNumber="42">P. umbrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DCCB63FDA50FFA3FB077842FA8ABB07" box="[1256,1331,279,300]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="4434D756DA50FFA3FD87786AFAFABB7F" blockId="41.[153,1425,279,415]" box="[616,1347,319,340]" pageId="41" pageNumber="42">
<typeStatus id="9B3069F4DA50FFA3FD87786AFD7BBB7F" box="[616,706,319,340]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
<typeStatus id="9B3069F4DA50FFA3FCD6786AFC2FBB7F" box="[825,918,319,340]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
<typeStatus id="9B3069F4DA50FFA3FBCF786AFBC0BB7F" box="[1056,1145,319,340]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
<typeStatus id="9B3069F4DA50FFA3FB09786AFAFABB7F" box="[1254,1347,319,340]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
</paragraph>
<paragraph id="4434D756DA50FFA3FD1B7830FA28BB51" blockId="41.[153,1425,279,415]" box="[756,1425,357,378]" pageId="41" pageNumber="42">Min. Max. Mean Min. Max. Mean</paragraph>
<paragraph id="4434D756DA50FFA3FF7678DFFA82BBB4" blockId="41.[153,1425,279,415]" box="[153,1339,394,415]" pageId="41" pageNumber="42">Total length (mm) 372 250 370 351 300 369</paragraph>
</subSubSection>
</treatment>
</document>