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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.963.53500" ID-GBIF-Dataset="9bc27e02-8911-4afd-bd9a-586dcebbac08" ID-PMC="PMC7458947" ID-Pensoft-Pub="1313-2970-963-81" ID-Pensoft-UUID="5D9D4242A1F5518295D41AB07D79DD82" ID-PubMed="32922132" ID-ZooBank="7D9006BB83234F738A41D487EBEA297A" ModsDocID="1313-2970-963-81" checkinTime="1598316826408" checkinUser="pensoft" docAuthor="Rosas-Ramos, Natalia, Mas-Peinado, Paloma, Gil-Tapetado, Diego, Recuero, Ernesto, Ruiz, Jose L. &amp; Garcia-Paris, Mario" docDate="2020" docId="AF57D823BE125C5DB5EF0C611A597DA3" docLanguage="en" docName="ZooKeys 963: 81-129" docOrigin="ZooKeys 963" docSource="http://dx.doi.org/10.3897/zookeys.963.53500" docTitle="Misolampus goudotii Guerin-Meneville 1834" docType="treatment" docVersion="4" id="5D9D4242A1F5518295D41AB07D79DD82" lastPageNumber="81" masterDocId="5D9D4242A1F5518295D41AB07D79DD82" masterDocTitle="Catalogue, distribution, taxonomic notes, and conservation of the Western Palearctic endemic hunchback beetles (Tenebrionidae, Misolampus)" masterLastPageNumber="129" masterPageNumber="81" pageNumber="81" updateTime="1668169382325" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Catalogue, distribution, taxonomic notes, and conservation of the Western Palearctic endemic hunchback beetles (Tenebrionidae, Misolampus)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rosas-Ramos, Natalia</mods:namePart>
<mods:affiliation>Departamento de Biologia Animal (Area de Zoologia), Facultad de Biologia (Edificio de Farmacia, planta 5), Universidad de Salamanca, Campus Miguel de Unamuno s / n, 37007 Salamanca, Spain &amp; Departamento de Biodiversidad y Biologia Evolutiva. Museo Nacional de Ciencias Naturales, MNCN-CSIC. c / Jose Gutierrez Abascal, 2. 28006, Madrid, Spain</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8653-0306</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mas-Peinado, Paloma</mods:namePart>
<mods:affiliation>Departamento de Biodiversidad y Biologia Evolutiva. Museo Nacional de Ciencias Naturales, MNCN-CSIC. c / Jose Gutierrez Abascal, 2. 28006, Madrid, Spain &amp; Centro de Investigacion en Biodiversidad y Cambio Global CIBC-UAM, Facultad de Ciencias, Universidad Autonoma de Madrid, c / Darwin 2, 28049 - Madrid, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gil-Tapetado, Diego</mods:namePart>
<mods:affiliation>Departamento de Biodiversidad y Biologia Evolutiva. Museo Nacional de Ciencias Naturales, MNCN-CSIC. c / Jose Gutierrez Abascal, 2. 28006, Madrid, Spain &amp; Departamento de Biologia, Ecologia y Evolucion, Facultad de Ciencias Biologicas, Universidad Complutense de Madrid, c / Jose Antonio Novais, 12, 28040 - Madrid, Spain</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2147-4040</mods:nameIdentifier>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Recuero, Ernesto</mods:namePart>
<mods:affiliation>Departamento de Biodiversidad y Biologia Evolutiva. Museo Nacional de Ciencias Naturales, MNCN-CSIC. c / Jose Gutierrez Abascal, 2. 28006, Madrid, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ruiz, Jose L.</mods:namePart>
<mods:affiliation>Instituto de Estudios Ceuties. Paseo del Revellin, 30. 51001 Ceuta, Spain</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garcia-Paris, Mario</mods:namePart>
<mods:affiliation>Departamento de Biodiversidad y Biologia Evolutiva. Museo Nacional de Ciencias Naturales, MNCN-CSIC. c / Jose Gutierrez Abascal, 2. 28006, Madrid, Spain</mods:affiliation>
<mods:nameIdentifier type="email">mparis@mncn.csic.es</mods:nameIdentifier>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>963</mods:number>
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<mods:start>81</mods:start>
<mods:end>129</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.963.53500</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-963-81</mods:identifier>
<mods:identifier type="ZooBank">7D9006BB83234F738A41D487EBEA297A</mods:identifier>
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<treatment ID-GBIF-Taxon="167293874" LSID="urn:lsid:plazi:treatment:AF57D823BE125C5DB5EF0C611A597DA3" httpUri="http://treatment.plazi.org/id/AF57D823BE125C5DB5EF0C611A597DA3" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="AF57D823-BE12-5C5D-B5EF-0C611A597DA3" authority="Guerin-Meneville, 1834" authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
Misolampus goudotii
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
, 1834
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="reference_group">
<paragraph pageId="0" pageNumber="81">
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">Misolampus goudotii</taxonomicName>
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
, 1834: 28. Terra typica:
<normalizedToken originalValue="“trouvée">&quot;trouvee</normalizedToken>
<normalizedToken originalValue="à">a</normalizedToken>
Tanger... ...
<normalizedToken originalValue="à">a</normalizedToken>
trois lieues de Tanger, sur les bords
<normalizedToken originalValue="dune">d'une</normalizedToken>
<normalizedToken originalValue="rivière">riviere</normalizedToken>
, dans le tronc
<normalizedToken originalValue="dun">d'un</normalizedToken>
olivier.&quot; Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
(
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">1900</bibRefCitation>
),
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.12929" author="Reitter, E" journalOrPublisher="Wiener Entomologische Zeitung" pageId="0" pageNumber="81" pagination="129 - 150" refId="B137" refString="Reitter, E, 1917. Bestimmungstabelle der Cossyphini und Misolampini. (Tribus der Tenebrionidae). Wiener Entomologische Zeitung 36 (6-7): 129 - 150, DOI: https://doi.org/10.5962/bhl.part.12929" title="Bestimmungstabelle der Cossyphini und Misolampini. (Tribus der Tenebrionidae)." url="https://doi.org/10.5962/bhl.part.12929" volume="36" year="1917">Reitter (1917)</bibRefCitation>
,
<bibRefCitation author="Antoine, M" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles du Maroc" pageId="0" pageNumber="81" pagination="123 - 162" publicationUrl="https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" refId="B4" refString="Antoine, M, 1949. Notes d'entomologie marocaine. 46 Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc 27 (1947): 123 - 162, https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" title="Notes d'entomologie marocaine. 46 Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides)." url="https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" volume="27" year="1949">Antoine (1949)</bibRefCitation>
, and
<bibRefCitation author="Espanol, F" journalOrPublisher="Graellsia" pageId="0" pageNumber="81" pagination="1 - 7" refId="B41" refString="Espanol, F, 1949. Datos para el conocimiento de los tenebrionidos del Mediterraneo Occidental. VI. El genero Misolampus Latr. en Espana y Baleares. Graellsia 7: 1 - 7" title="Datos para el conocimiento de los tenebrionidos del Mediterraneo Occidental. VI. El genero Misolampus Latr. en Espana y Baleares." volume="7" year="1949">
<normalizedToken originalValue="Español">Espanol</normalizedToken>
(1949)
</bibRefCitation>
among others, wrote the species name with a single final -i.
<bibRefCitation author="Solier, AJJ" journalOrPublisher="Studi Entomologici" pageId="0" pageNumber="81" pagination="149 - 370" publicationUrl="https://www.biodiversitylibrary.org/page/40161343" refId="B161" refString="Solier, AJJ, 1848. Essai sur les Collapterides. 14e Tribu - Blapsites. Studi Entomologici 1: 149 - 370, https://www.biodiversitylibrary.org/page/40161343" title="Essai sur les Collapterides. 14 e Tribu - Blapsites." url="https://www.biodiversitylibrary.org/page/40161343" volume="1" year="1848">Solier (1848)</bibRefCitation>
and Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
(
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">1900</bibRefCitation>
) mentioned the unavailable name: &quot;
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus nigrita" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="nigrita">Misolampus nigrita</taxonomicName>
Dejean&quot;, and
<bibRefCitation author="Espanol, F" journalOrPublisher="Trabajos del Museo de Ciencias Naturales de Barcelona, Nueva Serie Zoologica" pageId="0" pageNumber="81" pagination="1 - 93" refId="B43" refString="Espanol, F, 1954a. Los tenebrionidos (Col.) de Baleares. Trabajos del Museo de Ciencias Naturales de Barcelona, Nueva Serie Zoologica 1 (5): 1 - 93" title="Los tenebrionidos (Col.) de Baleares." volume="1" year="1954 a">
<normalizedToken originalValue="Español">Espanol</normalizedToken>
(1954a)
</bibRefCitation>
: &quot;
<taxonomicName family="Tenebrionidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="M. moraguesi" order="Coleoptera" pageId="0" pageNumber="81" rank="species" species="moraguesi">M. moraguesi</taxonomicName>
&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">Misolampus goudotii</taxonomicName>
Erichson in Wagner, 1841: 184 (non
<bibRefCitation author="Guerin-Meneville, FE" journalOrPublisher="Magasin de Zoologie" pageId="0" pageNumber="81" pagination="1 - 39" publicationUrl="https://www.biodiversitylibrary.org/page/37085196" refId="B67" refString="Guerin-Meneville, FE, 1834. Materiaux pour une classification des melasomes. (Extraits d'une monographie de cette famille). Magasin de Zoologie 4: 1 - 39, https://www.biodiversitylibrary.org/page/37085196" title="Materiaux pour une classification des melasomes. (Extraits d'une monographie de cette famille)." url="https://www.biodiversitylibrary.org/page/37085196" volume="4" year="1834">
<normalizedToken originalValue="Guérin-Méneville">Guerin-Meneville</normalizedToken>
1834
</bibRefCitation>
). Terra typica: not indicated, but &quot;von Algier&quot; according to
<bibRefCitation author="Erichson, WF" editor="Wagner, MF" journalOrPublisher="Dritter Band. L. Voss, Leipzig" pageId="0" pageNumber="81" pagination="140 - 194" publicationUrl="https://www.biodiversitylibrary.org/page/33793901" refId="B39" refString="Erichson, WF, 1841. Ueber die Insecten von Algier mit besonderer Ruecksicht auf ihre geographische Verbreitung. In: Wagner, MF, Ed., Reisen in der Regentschaft Algier 1836, 1837 und 1838 nebst einem naturhistorischen Anhang un einem Kupferatlas. Dritter Band. L. Voss, Leipzig: 140 - 194, https://www.biodiversitylibrary.org/page/33793901" title="Ueber die Insecten von Algier mit besonderer Ruecksicht auf ihre geographische Verbreitung." url="https://www.biodiversitylibrary.org/page/33793901" volumeTitle="Reisen in der Regentschaft Algier 1836, 1837 und 1838 nebst einem naturhistorischen Anhang un einem Kupferatlas." year="1841">
<normalizedToken originalValue="Erichsons">Erichson's</normalizedToken>
(1841)
</bibRefCitation>
work title.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus erichsoni" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="erichsoni">Misolampus erichsoni</taxonomicName>
Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
, 1900: 674 syn. nov. Terra typica:
<normalizedToken originalValue="“Algérie">&quot;Algerie</normalizedToken>
: O., Oran...; Daya...; Tlemcen...; Mascara..., Ammi Moussa; A.: Blidah...; La Chiffa...; Margueritte...;
<normalizedToken originalValue="forêt">foret</normalizedToken>
de Boghar...; mont Ouarsenis...;
<normalizedToken originalValue="forêts">forets</normalizedToken>
de la Grande-Kabylie...&quot;.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus peyerimhoffi" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="peyerimhoffi">Misolampus peyerimhoffi</taxonomicName>
Antoine, 1926: 257 syn. nov. Terra typica: &quot;Grand Atlas,
<normalizedToken originalValue="région">region</normalizedToken>
du Glaoui: plateau des
<normalizedToken originalValue="Aït">Ait</normalizedToken>
Rba...&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="materials_examined">
<paragraph pageId="0" pageNumber="81">Studied material.</paragraph>
<paragraph pageId="0" pageNumber="81">
Algeria:
<normalizedToken originalValue="“Argelia”">&quot;Argelia&quot;</normalizedToken>
(Dufour leg.): 1 ex. Morocco - Marrakech-Tensift-Al Hauz: Toufliht, 1483 m,
<geoCoordinate degrees="31" direction="north" minutes="28" orientation="latitude" precision="1" seconds="34.6" value="31.476278">31°28'34.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="7" direction="west" minutes="26" orientation="longitude" precision="1" seconds="06.5" value="-7.435139">7°26'06.5&quot;W</geoCoordinate>
, 11-III-2013: 4 exx. -
<normalizedToken originalValue="Meknès">Meknes</normalizedToken>
- Tafilalet: Ain Leuh, 17-V-1925: 1 ex.; Azrou, 1900 m (Alluaud 79) (
<emphasis italics="true" pageId="0" pageNumber="81">
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudoti" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudoti">Misolampus goudoti</taxonomicName>
var.
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="var. laevior" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="variety" species="goudoti" variety="laevior">laevior</taxonomicName>
</emphasis>
Alluaud): 1 ex.; Azrou, 19-V-1925: 1 ex.; Dj. [Yebel] Hebri, 20-V-1925: 1 ex.; Timadit [Timahdite], 21-V-1923: 1 ex. - Tanger -
<normalizedToken originalValue="Tétouan">Tetouan</normalizedToken>
: Rif: Beni Siyyel: Bab Ruadi: VI-1932 (C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 6 exx.; Tanger, 1897: 3 exx.; Tanger (M. Escalera leg.) (small square-label pinned): 36 exx., plus 3 exx. only square-labelled; 2 km al O de Bab Berret, 1318 m,
<geoCoordinate degrees="35" direction="north" minutes="00" orientation="latitude" precision="1" seconds="02.57" value="35.000713">35°00'02.57&quot;N</geoCoordinate>
,
<geoCoordinate degrees="4" direction="west" minutes="55" orientation="longitude" precision="1" seconds="31.91" value="-4.9255304">4°55'31.91&quot;W</geoCoordinate>
, 12-VI-2011: 3 exx.; Crtra. Zinat-Mulay Abdeselam, P-4702, Beni Aros, Yebala, 513 m,
<geoCoordinate degrees="35" direction="north" minutes="22" orientation="latitude" precision="15" seconds="04" value="35.367775">35°22'04&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="32" orientation="longitude" precision="15" seconds="17" value="-5.5380554">5°32'17&quot;W</geoCoordinate>
, 29-IV-2016: 5 exx.; Yebel Bou-Hachem, Beni Aros, Yebala, 1160 m,
<geoCoordinate degrees="35" direction="north" minutes="15" orientation="latitude" precision="15" seconds="31" value="35.25861">35°15'31&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="30" orientation="longitude" precision="15" seconds="49" value="-5.5136113">5°30'49&quot;W</geoCoordinate>
, 12-V-2012: 6 exx.; Crtra. Mulay Abdeselam-Al Hamra, P-4704, Beni Aros, 985 m,
<geoCoordinate degrees="35" direction="north" minutes="15" orientation="latitude" precision="15" seconds="50" value="35.26389">35°15'50&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="25" orientation="longitude" precision="15" seconds="36" value="-5.4266667">5°25'36&quot;W</geoCoordinate>
, 28-XI-2019: 2 exx.; Larache: Yebala: Beni
<normalizedToken originalValue="Arós">Aros</normalizedToken>
: Yebel Bou-Hachem,
<geoCoordinate degrees="35" direction="north" minutes="15" orientation="latitude" precision="15" seconds="31" value="35.25861">35°15'31&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="30" orientation="longitude" precision="15" seconds="49" value="-5.5136113">5°30'49&quot;W</geoCoordinate>
, 9-VI-2012: 2 exx.; Pinsapar del Talassemtane, 1900 m,
<geoCoordinate degrees="35" direction="north" minutes="08" orientation="latitude" precision="1" seconds="36.7" value="35.143528">35°08'36.7&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="08" orientation="longitude" precision="1" seconds="13.0" value="-5.1369443">5°08'13.0&quot;W</geoCoordinate>
, 11-VI-2011: 2 exx.; Bab Taza: Talassemtane, 1401 m,
<geoCoordinate degrees="35" direction="north" minutes="06" orientation="latitude" precision="1" seconds="10.9" value="35.103027">35°06'10.9&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="08" orientation="longitude" precision="1" seconds="21.3" value="-5.13925">5°08'21.3&quot;W</geoCoordinate>
, 27-VII-2013: 1 ex.; Bab Taza: Talassemtane: Plaza de
<normalizedToken originalValue="España">Espana</normalizedToken>
, 1667 m,
<geoCoordinate degrees="35" direction="north" minutes="09" orientation="latitude" precision="1" seconds="03.7" value="35.151028">35°09'03.7&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="08" orientation="longitude" precision="1" seconds="28.6" value="-5.141278">5°08'28.6&quot;W</geoCoordinate>
, 27-VII-2013: 1 ex.; Casa Forestal, Yebel Lekraa, P.N. Talasemtane, Chefchaouen,
<geoCoordinate degrees="35" direction="north" minutes="07" orientation="latitude" precision="15" seconds="56" value="35.13222">35°07'56&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="08" orientation="longitude" precision="15" seconds="11" value="-5.136389">5°08'11&quot;W</geoCoordinate>
, 1695 m, 7-VI-2008: 3 exx.; Yebel Talassemtane-vertiente sur, P.N. Talasemtane, Chefchaouen,
<geoCoordinate degrees="35" direction="north" minutes="07" orientation="latitude" precision="15" seconds="53" value="35.131386">35°07'53&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="08" orientation="longitude" precision="15" seconds="01" value="-5.133611">5°08'01&quot;W</geoCoordinate>
, 1650 m, 11-IV-2011: 4 exx.; P.N. Yebel Tazaot, Pinsapar, P.N. Talasemtane, Chefchaouen,
<geoCoordinate degrees="35" direction="north" minutes="15" orientation="latitude" precision="925" value="35.25">35°15'N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="07" orientation="longitude" precision="925" value="-5.116667">5°07'W</geoCoordinate>
, 1670 m, 7-V-2011: 2 exx.; Pista hacia Casa Forestal, Yebel Lekraa, P.N. Talasemtane, Chefchaouen,
<geoCoordinate degrees="35" direction="north" minutes="07" orientation="latitude" precision="15" seconds="45" value="35.129166">35°07'45&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="08" orientation="longitude" precision="15" seconds="09" value="-5.1358333">5°08'09&quot;W</geoCoordinate>
, 1700 m, 28-VII-2011: 1 ex.; E. de Yebel Talaousisse, P.N. Talasemtane, Chefchaouen,
<geoCoordinate degrees="35" direction="north" minutes="07" orientation="latitude" precision="15" seconds="33" value="35.12583">35°07'33&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="04" orientation="longitude" precision="15" seconds="03" value="-5.0675">5°04'03&quot;W</geoCoordinate>
, 1350 m, 1-XII-2018: 2 exx.; Pista hacia Haout Taznout, P.N. Talasemtane, Chefchaouen,
<geoCoordinate degrees="35" direction="north" minutes="08" orientation="latitude" precision="15" seconds="20" value="35.13889">35°08'20&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="07" orientation="longitude" precision="15" seconds="24" value="-5.1233335">5°07'24&quot;W</geoCoordinate>
, 1712 m, 27-IV-2019: 2 exx.; Yebel Tizirhen, Bab Berred, Rif Central, 1585 m,
<geoCoordinate degrees="35" direction="north" minutes="00" orientation="latitude" precision="15" seconds="54" value="35.015">35°00'54&quot;N</geoCoordinate>
,
<geoCoordinate degrees="4" direction="west" minutes="54" orientation="longitude" precision="15" seconds="57" value="-4.9158335">4°54'57&quot;W</geoCoordinate>
, 27-IV-2017: 3 exx.; Yebel Tizirhen, Bab Berred, Rif Central, 1570 m,
<geoCoordinate degrees="35" direction="north" minutes="00" orientation="latitude" precision="15" seconds="47" value="35.013054">35°00'47&quot;N</geoCoordinate>
,
<geoCoordinate degrees="4" direction="west" minutes="54" orientation="longitude" precision="15" seconds="03" value="-4.9008336">4°54'03&quot;W</geoCoordinate>
, 28-IV-2018: 1 ex.; Pista de Bab El Kar,
<normalizedToken originalValue="Montañas">Montanas</normalizedToken>
de Fifi, Rif, 1512 m,
<geoCoordinate degrees="34" direction="north" minutes="59" orientation="latitude" precision="15" seconds="13" value="34.986946">34°59'13&quot;N</geoCoordinate>
,
<geoCoordinate degrees="5" direction="west" minutes="11" orientation="longitude" precision="15" seconds="20" value="-5.188889">5°11'20&quot;W</geoCoordinate>
, 2-VI-2019: 2 exx. - Taza - Al Hoceima - Taounate: Iguermalen [Targuist]: Beni Mesdui, VI-1932 (M. Escalera leg.): 6 exx.; Rif: Beni Seddat: Imosiner: VI-1930 (exp. C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 3 exx.; Rif: Beni Seddat: Tizi Taka, VI-1932 (C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 4 exx.; Rif: Beni Seddat: Tizi Taka, VI-1932 (Exp. C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 1 ex.; Rif: Iguermalen (Targuist), VI-1930 (exp. C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 4 exx.; Rif: Ketama, Bab Chiquer, VI-1932 (C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 8 exx.; Rif: Ketama, Bab Chiquer, VI-1932 (M. Escalera leg.): 2 exx.; Rif: Ketama: Tainza, VI-1930 (exp. C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg): 3 exx.; Rif: Ketama: Tidiguin [Tidghine], VI-1930 (exp. C.
<normalizedToken originalValue="Bolívar">Bolivar</normalizedToken>
leg.): 1 ex.; Rif: Ketama: Zoco Telata, VI-1932 (M. Escalera leg.): 7 exx.; Lurdeka [?]: 1 ex.; Yebel Tidighin, Azila, Rif central, 1705 m,
<geoCoordinate degrees="34" direction="north" minutes="51" orientation="latitude" precision="15" seconds="14" value="34.853886">34°51'14&quot;N</geoCoordinate>
,
<geoCoordinate degrees="4" direction="west" minutes="32" orientation="longitude" precision="15" seconds="19" value="-4.538611">4°32'19&quot;W</geoCoordinate>
, 29-XI-2019: 1 ex; Carretera P-5420, P.N. Tazzeka, Medio Atlas nororiental, 1000 m,
<geoCoordinate degrees="34" direction="north" minutes="03" orientation="latitude" precision="925" value="34.05">34°03'N</geoCoordinate>
,
<geoCoordinate degrees="4" direction="west" minutes="15" orientation="longitude" precision="925" value="-4.25">4°15'W</geoCoordinate>
, 25-XI-2004 (F.J.
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
leg.): 2 exx. -
<normalizedToken originalValue="Souss-Massa-Drâa">Souss-Massa-Draa</normalizedToken>
: Yebel Tual, 28-VII-1934: 1 ex.; Ifni: Yebel Tamarrut [25 km SE Ifni], I-1935 (F. Escalera leg.): 1 ex.; Sidi Ifni: Akarkor, Jbel Toual, 627 m,
<geoCoordinate degrees="29" direction="north" minutes="13" orientation="latitude" precision="1" seconds="48.9" value="29.230251">29°13'48.9&quot;N</geoCoordinate>
,
<geoCoordinate degrees="10" direction="west" minutes="00" orientation="longitude" precision="1" seconds="44.1" value="-10.01225">10°00'44.1&quot;W</geoCoordinate>
, 21-I-2020: 4 exx. Spain - Islas Baleares: Mallorca (Mas de Xaxars leg.) (
<taxonomicName class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus erichsoni" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="erichsoni">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus erichsoni</emphasis>
</taxonomicName>
): 2 exx.; Escorca, 26-III-1985, 1 ex.; Escorca, Coll de Femenia, 545 m,
<geoCoordinate degrees="39" direction="north" minutes="51" orientation="latitude" precision="1" seconds="33.68" value="39.859356">39°51'33.68&quot;N</geoCoordinate>
,
<geoCoordinate degrees="2" direction="west" minutes="54" orientation="longitude" precision="1" seconds="19.27" value="-2.9053528">2°54'19.27&quot;W</geoCoordinate>
, 25-III-2012: 10 exx.; Menorca (Cardona leg.): 2 exx. plus 6 exx. without data; 2 exx.; Menorca: 2 exx.; Algaiarens, 14 m,
<geoCoordinate degrees="40" direction="north" minutes="02" orientation="latitude" precision="1" seconds="28.3" value="40.041195">40°02'28.3&quot;N</geoCoordinate>
,
<geoCoordinate degrees="03" direction="west" minutes="55" orientation="longitude" precision="1" seconds="28.4" value="-3.9245555">03°55'28.4&quot;W</geoCoordinate>
, 27-IV-2006: 2 exx.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="diagnosis">
<paragraph pageId="0" pageNumber="81">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">
With a total length from 10 to 14 mm, this is the largest species of the genus (
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">
Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
1900
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.12929" author="Reitter, E" journalOrPublisher="Wiener Entomologische Zeitung" pageId="0" pageNumber="81" pagination="129 - 150" refId="B137" refString="Reitter, E, 1917. Bestimmungstabelle der Cossyphini und Misolampini. (Tribus der Tenebrionidae). Wiener Entomologische Zeitung 36 (6-7): 129 - 150, DOI: https://doi.org/10.5962/bhl.part.12929" title="Bestimmungstabelle der Cossyphini und Misolampini. (Tribus der Tenebrionidae)." url="https://doi.org/10.5962/bhl.part.12929" volume="36" year="1917">Reitter 1917</bibRefCitation>
;
<bibRefCitation author="Antoine, M" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles du Maroc" pageId="0" pageNumber="81" pagination="248 - 259" publicationUrl="https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" refId="B3" refString="Antoine, M, 1926. Notes d'entomologie marocaine. VI (1) Tenebrionides nouveaux et interessants. Bulletin de la Societe des Sciences Naturelles du Maroc 5: 248 - 259, https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" title="Notes d'entomologie marocaine. VI (1) Tenebrionides nouveaux et interessants." url="https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" volume="5" year="1926">Antoine 1926</bibRefCitation>
;
<bibRefCitation author="Espanol, F" journalOrPublisher="Graellsia" pageId="0" pageNumber="81" pagination="1 - 7" refId="B41" refString="Espanol, F, 1949. Datos para el conocimiento de los tenebrionidos del Mediterraneo Occidental. VI. El genero Misolampus Latr. en Espana y Baleares. Graellsia 7: 1 - 7" title="Datos para el conocimiento de los tenebrionidos del Mediterraneo Occidental. VI. El genero Misolampus Latr. en Espana y Baleares." volume="7" year="1949">
<normalizedToken originalValue="Español">Espanol</normalizedToken>
1949
</bibRefCitation>
). This species is well characterised and isolated within the genus
<taxonomicName authorityName="Latreille" authorityYear="1807" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus</emphasis>
</taxonomicName>
by the following traits: fore angles of the prothorax not protruding, almost rounded, forming an obtuse angle at apex; lateral surface of pronotum shallowly rugose, with the rugosity progressively erased towards the dorsal areas that appear smoother, propleural punctation fine and often erased; elytra with longitudinal series of small elongated tubercles, more apparent on the sides of the posterior half of the elytra (
<bibRefCitation author="Espanol, F" journalOrPublisher="Graellsia" pageId="0" pageNumber="81" pagination="1 - 7" refId="B41" refString="Espanol, F, 1949. Datos para el conocimiento de los tenebrionidos del Mediterraneo Occidental. VI. El genero Misolampus Latr. en Espana y Baleares. Graellsia 7: 1 - 7" title="Datos para el conocimiento de los tenebrionidos del Mediterraneo Occidental. VI. El genero Misolampus Latr. en Espana y Baleares." volume="7" year="1949">
<normalizedToken originalValue="Español">Espanol</normalizedToken>
1949
</bibRefCitation>
,
<bibRefCitation author="Espanol, F" journalOrPublisher="Trabajos del Museo de Ciencias Naturales de Barcelona, Nueva Serie Zoologica" pageId="0" pageNumber="81" pagination="1 - 93" refId="B43" refString="Espanol, F, 1954a. Los tenebrionidos (Col.) de Baleares. Trabajos del Museo de Ciencias Naturales de Barcelona, Nueva Serie Zoologica 1 (5): 1 - 93" title="Los tenebrionidos (Col.) de Baleares." volume="1" year="1954 a">1954a</bibRefCitation>
;
<bibRefCitation author="Palmer, M" journalOrPublisher="Bolleti de la Societat d' Historia Natural de les Balears" pageId="0" pageNumber="81" pagination="139 - 151" refId="B119" refString="Palmer, M, 1998. Phylogeny of the genus Misolampus (Coleoptera: Tenebrionidae). Bolleti de la Societat d' Historia Natural de les Balears 41: 139 - 151" title="Phylogeny of the genus Misolampus (Coleoptera: Tenebrionidae)." volume="41" year="1998">Palmer 1998</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3A-C</figureCitation>
). Female genitalia figured by
<bibRefCitation author="Palmer, M" journalOrPublisher="Bolleti de la Societat d' Historia Natural de les Balears" pageId="0" pageNumber="81" pagination="139 - 151" refId="B119" refString="Palmer, M, 1998. Phylogeny of the genus Misolampus (Coleoptera: Tenebrionidae). Bolleti de la Societat d' Historia Natural de les Balears 41: 139 - 151" title="Phylogeny of the genus Misolampus (Coleoptera: Tenebrionidae)." volume="41" year="1998">Palmer (1998)</bibRefCitation>
. Specimens from the Balearic Islands have been studied karyologically (Juan and Petipierre 1986, 1989;
<bibRefCitation DOI="https://doi.org/10.1007/BF00710770" author="Juan, C" journalOrPublisher="Chromosome Research" pageId="0" pageNumber="81" pagination="167 - 174" refId="B77" refString="Juan, C, Pons, J, Petitpierre, E, 1993. Localization of tandemly repeated DNA sequences in beetle chromosomes by fluorescent in situ hybridization. Chromosome Research 1 (3): 167 - 174, DOI: https://doi.org/10.1007/BF00710770" title="Localization of tandemly repeated DNA sequences in beetle chromosomes by fluorescent in situ hybridization." url="https://doi.org/10.1007/BF00710770" volume="1" year="1993">Juan et al. 1993</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1111/j.1601-5223.1993.00179.x" author="Pons, J" journalOrPublisher="Hereditas" pageId="0" pageNumber="81" pagination="179 - 185" refId="B134" refString="Pons, J, Petitpierre, E, Juan, C, 1993. Characterization of the heterochromatin of the darkling beetle Misolampus goudoti: cloning of two satellite DNA families and digestion of chromosomes with restriction enzymes. Hereditas 119 (2): 179 - 185, DOI: https://doi.org/10.1111/j.1601-5223.1993.00179.x" title="Characterization of the heterochromatin of the darkling beetle Misolampus goudoti: cloning of two satellite DNA families and digestion of chromosomes with restriction enzymes." url="https://doi.org/10.1111/j.1601-5223.1993.00179.x" volume="119" year="1993">Pons et al. 1993</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1139/g04-019" author="Pons, J" journalOrPublisher="Genome" pageId="0" pageNumber="81" pagination="769 - 774" refId="B133" refString="Pons, J, 2004. Cloning and characterization of a transposable-like repeat in the heterochromatin of the darkling beetle Misolampus goudoti. Genome 47 (4): 769 - 774, DOI: https://doi.org/10.1139/g04-019" title="Cloning and characterization of a transposable-like repeat in the heterochromatin of the darkling beetle Misolampus goudoti." url="https://doi.org/10.1139/g04-019" volume="47" year="2004">Pons 2004</bibRefCitation>
), presenting a chromosome number of 20 (2n) (
<bibRefCitation author="Juan, C" journalOrPublisher="Genetica Iberica" pageId="0" pageNumber="81" pagination="231 - 243" refId="B75" refString="Juan, C, Petitpierre, E, 1986. Karyological analyses on tenebrionid beetles from the Balearic Islands. Genetica Iberica 38 (3-4): 231 - 243" title="Karyological analyses on tenebrionid beetles from the Balearic Islands." volume="38" year="1986">Juan and Petitpierre 1986</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1139/g89-519" author="Juan, C" journalOrPublisher="Genetica Iberica" pageId="0" pageNumber="81" refId="B76" refString="Juan, C, Petitpierre, E, 1989. . C-banding and DNA content in seven species of Tenebrionidae (Coleoptera). Genome 32(5) 834-839. https://doi.org/10.1139/g89-519" title=". C-banding and DNA content in seven species of Tenebrionidae (Coleoptera). Genome 32 (5) 834 - 839." url="https://doi.org/10.1139/g89-519" year="1989">1989</bibRefCitation>
). There is marked geographical variability on the sculpture and shape of pronotum and propleurae, and on the development of elytral tubercles (
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">
Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
1900
</bibRefCitation>
;
<bibRefCitation author="Antoine, M" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles du Maroc" pageId="0" pageNumber="81" pagination="123 - 162" publicationUrl="https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" refId="B4" refString="Antoine, M, 1949. Notes d'entomologie marocaine. 46 Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc 27 (1947): 123 - 162, https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" title="Notes d'entomologie marocaine. 46 Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides)." url="https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" volume="27" year="1949">Antoine 1949</bibRefCitation>
;
<bibRefCitation author="Espanol, F" journalOrPublisher="Trabajos del Museo de Ciencias Naturales de Barcelona, Nueva Serie Zoologica" pageId="0" pageNumber="81" pagination="1 - 93" refId="B43" refString="Espanol, F, 1954a. Los tenebrionidos (Col.) de Baleares. Trabajos del Museo de Ciencias Naturales de Barcelona, Nueva Serie Zoologica 1 (5): 1 - 93" title="Los tenebrionidos (Col.) de Baleares." volume="1" year="1954 a">
<normalizedToken originalValue="Español">Espanol</normalizedToken>
1954a
</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3A-C</figureCitation>
). Specimens from northern Morocco (excluding the Tingitane Peninsula), Algeria and the Balearic Islands, present a well-developed and evident elytral tuberculation that may form rugose ridges (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3A</figureCitation>
). On the other extreme, elytral tubercles are reduced in the Rif and Atlas populations (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3B</figureCitation>
), to become almost completely absent in the specimens from Sidi Ifni (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3C</figureCitation>
). Pronotum sculpture is formed by fine spaced punctures intermixed with granules, much denser on the sides in the Balearic Islands population (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3A</figureCitation>
); pronotal rugose areas are more marked and extended in the specimens from the High Atlas (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3B</figureCitation>
), and formed by sparse punctation, without granulose areas, in the specimens from Ifni (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3C</figureCitation>
). The anterior edge of the pronotum, in the Rif and Balearic populations, is straight at the middle, while it appears convex in the populations from the High Atlas (
<bibRefCitation author="Antoine, M" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles du Maroc" pageId="0" pageNumber="81" pagination="123 - 162" publicationUrl="https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" refId="B4" refString="Antoine, M, 1949. Notes d'entomologie marocaine. 46 Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides). Bulletin de la Societe des Sciences Naturelles du Maroc 27 (1947): 123 - 162, https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" title="Notes d'entomologie marocaine. 46 Materiaux pour l'etude des Helopinae du Maroc (Col. Tenebrionides)." url="https://bibdigital.rjb.csic.es/viewer/16061/?offset=page=137&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" volume="27" year="1949">Antoine 1949</bibRefCitation>
). The geographic distribution of this variability has been the subject of taxonomic discussion resulting in the proposal of different taxa, here formally synonymised (see synonymic list, and taxonomic discussion).
</paragraph>
<caption doi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 3.</emphasis>
Live specimens and habitat of
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="81">A-C</emphasis>
Live adult specimens of
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
from the Balearic Islands (
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
Cap Formentor, Mallorca) and Morocco (
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
Toufliht, High Atlas
<emphasis bold="true" pageId="0" pageNumber="81">C</emphasis>
Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species
<emphasis bold="true" pageId="0" pageNumber="81">D-F</emphasis>
A summary of the impressive habitat diversity used by
<taxonomicName lsidName="M. goudotii" pageId="0" pageNumber="81" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">M. goudotii</emphasis>
</taxonomicName>
from the Balearic Islands (
<emphasis bold="true" pageId="0" pageNumber="81">D</emphasis>
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus ilex" order="Fagales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis italics="true" pageId="0" pageNumber="81">Quercus ilex</emphasis>
</taxonomicName>
forest at Creu de Menut, Mallorca), to southwestern Morocco (
<emphasis bold="true" pageId="0" pageNumber="81">E</emphasis>
deep valleys in the Toubkal National Park, High Atlas
<emphasis bold="true" pageId="0" pageNumber="81">F</emphasis>
<taxonomicName authorityName="Skeels" authorityYear="1911" class="Insecta" family="Noctuidae" genus="Argania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argania spinosa" order="Lepidoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis italics="true" pageId="0" pageNumber="81">Argania spinosa</emphasis>
</taxonomicName>
open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Geographic distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">
Distributed throughout Morocco, northern Algeria and the Balearic Islands in Spain (
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<bibRefCitation author="Mouna, M" journalOrPublisher="Bulletin de l'Institut Scientifique, Rabat" pageId="0" pageNumber="81" pagination="133 - 142" refId="B110" refString="Mouna, M, Arahou, M, 1986. Comparaison des peuplements de Coleopteres frondicoles de deux essences forestieres resineuses au Maroc: Cedre (Cedrus atlantica Man) et Thuya (Tetraclinis articulata Vahl et Benth). Bulletin de l'Institut Scientifique, Rabat 1: 133 - 142" title="Comparaison des peuplements de Coleopteres frondicoles de deux essences forestieres resineuses au Maroc: Cedre (Cedrus atlantica Man) et Thuya (Tetraclinis articulata Vahl et Benth)." volume="1" year="1986">Mouna and Arahou 1986</bibRefCitation>
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,
<bibRefCitation DOI="https://doi.org/10.12976/jib/2015.4.13" author="Benyahia, Y" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="81" pagination="1 - 30" refId="B12" refString="Benyahia, Y, Brustel, H, El Antry, S, Courtin, O, Maatouf, N, Valladares, L, Rohi, L, 2016. Preliminary list of Coleoptera heritage species of the Talassemtane National Park, Morocco. Journal of Insect Biodiversity 4 (13): 1 - 30, DOI: https://doi.org/10.12976/jib/2015.4.13" title="Preliminary list of Coleoptera heritage species of the Talassemtane National Park, Morocco." url="https://doi.org/10.12976/jib/2015.4.13" volume="4" year="2016">2016</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.31167/csef.v0i42.17493" author="Nunez, L" journalOrPublisher="Cuadernos de la Sociedad Espanola de Ciencias Forestales" pageId="0" pageNumber="81" pagination="401 - 408" refId="B115" refString="Nunez, L, Closa, S, Gonzalez, E, Lencina, JL, Gallego, D, 2016. Metodologia para la determinacion de niveles de infestacion por Cerambyx cerdo (Linnaeus, 1758), desarrollo de tecnicas de captura y estudio de fauna saproxilica acompanante en los encinares de Mallorca. Cuadernos de la Sociedad Espanola de Ciencias Forestales 43: 401 - 408, DOI: https://doi.org/10.31167/csef.v0i42.17493" title="Metodologia para la determinacion de niveles de infestacion por Cerambyx cerdo (Linnaeus, 1758), desarrollo de tecnicas de captura y estudio de fauna saproxilica acompanante en los encinares de Mallorca." url="https://doi.org/10.31167/csef.v0i42.17493" volume="43" year="2016">
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;
<bibRefCitation author="Chavanon, G" journalOrPublisher="Travaux de l'Institut Scientifique, Serie Zoologie" pageId="0" pageNumber="81" pagination="1 - 192" refId="B27" refString="Chavanon, G, 2020. Catalogue des Coleopteres de la region orientale du Maroc (Province de Guercif exceptee). Travaux de l'Institut Scientifique, Serie Zoologie 57 (2018): 1 - 192" title="Catalogue des Coleopteres de la region orientale du Maroc (Province de Guercif exceptee)." volume="57" year="2020">Chavanon 2020</bibRefCitation>
) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Geographic distribution of Misolampus goudotii. Map for the distribution range of Misolampus goudotii (pale blue spot). Blue dots correspond to the species records, including both recent and old, as well as previously published data. The population from Ifni remains isolated from the main distribution range, by a distance of ca. 250 km B potential geographic distribution of Misolampus goudotii: Red indicates high suitable areas, and blue, areas of low suitability. Species distribution model was generated using MaxEnt v 3.4.1 (Elith et al. 2006) and the set of WorldClim v 2.0 (Fick and Hijmans 2017) environmental variables." figureDoi="10.3897/zookeys.963.53500.figure4" httpUri="https://binary.pensoft.net/fig/444545" pageId="0" pageNumber="81">4</figureCitation>
). The record from Ceuta, Spain (
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">
Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
1900
</bibRefCitation>
) corresponds to the mountain Yebel Musa (just 1.5 km west of Ceuta), currently in Moroccan territory (region of
<normalizedToken originalValue="Tanger-Tétouan">Tanger-Tetouan</normalizedToken>
).
</paragraph>
<caption doi="10.3897/zookeys.963.53500.figure4" httpUri="https://binary.pensoft.net/fig/444545" pageId="0" pageNumber="81" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 4.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
Geographic distribution of
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
. Map for the distribution range of
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
(pale blue spot). Blue dots correspond to the species records, including both recent and old, as well as previously published data. The population from Ifni remains isolated from the main distribution range, by a distance of ca. 250 km
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
potential geographic distribution of
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
: Red indicates high suitable areas, and blue, areas of low suitability. Species distribution model was generated using MaxEnt v 3.4.1 (
<bibRefCitation DOI="https://doi.org/10.1111/j.2006.0906-7590.04596.x" author="Elith, J" journalOrPublisher="Ecography" pageId="0" pageNumber="81" pagination="129 - 151" refId="B37" refString="Elith, J, Graham, CH, Anderson, RP, Dudik, M, Ferrier, S, Guisan, A, Hijmans, RJ, Huettmann, F, Leathwick, JR, Lehmann, A, Li, J, Lohmann, LG, Loiselle, BA, Manion, G, Moritz, C, Nakamura, M, Nakazawa, Y, Overton, JMcCM, Peterson, AT, Phillips, SJ, Richardson, K, Scachetti-Pereira, R, Schapire, RE, Soberon, J, Williams, S, Wisz, MS, Zimmermann, NE, 2006. Novel methods improve prediction of species' distributions from occurrence data. Ecography 29: 129 - 151, DOI: https://doi.org/10.1111/j.2006.0906-7590.04596.x" title="Novel methods improve prediction of species' distributions from occurrence data." url="https://doi.org/10.1111/j.2006.0906-7590.04596.x" volume="29" year="2006">Elith et al. 2006</bibRefCitation>
) and the set of WorldClim v 2.0 (
<bibRefCitation DOI="https://doi.org/10.1002/joc.5086" author="Fick, SE" journalOrPublisher="International Journal of Climatology," pageId="0" pageNumber="81" pagination="4302 - 4315" refId="B52" refString="Fick, SE, Hijmans, RJ, 2017. WorldClim 2: new 1-km spatial resolution climate surfaces for global land areas. International Journal of Climatology, 37 (12): 4302 - 4315, DOI: https://doi.org/10.1002/joc.5086" title="WorldClim 2: new 1 - km spatial resolution climate surfaces for global land areas." url="https://doi.org/10.1002/joc.5086" volume="37" year="2017">Fick and Hijmans 2017</bibRefCitation>
) environmental variables.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
The studied materials include recent and old records of populations from the Balearic Islands (Mallorca and Menorca) and from the Moroccan regions of
<normalizedToken originalValue="Meknès-Tafilalet">Meknes-Tafilalet</normalizedToken>
,
<normalizedToken originalValue="Souss-Massa-Drâa">Souss-Massa-Draa</normalizedToken>
,
<normalizedToken originalValue="Tanger-Tétouan">Tanger-Tetouan</normalizedToken>
, and Taza-Al Hoceima-Taounate. Recent data are available from all four regions, with a large number of localities from the Rif, and less numerous in the Middle and High Atlas. Among these records, we emphasise the re-discovery of the population from the province of Sidi Ifni, in January-2020, 85 years after its original finding, by F.
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
de la Escalera in 1934 and 1935 (
<bibRefCitation author="Paris Garcia, M" journalOrPublisher="Tamuda" pageId="0" pageNumber="81" refId="B123" refString="Paris Garcia, M, Blay Goicoechea, A, Hitado Morales, M, 2011. Las &quot;especies&quot; no descritas de M. Martinez de la Escalera. In: Martin Albaladejo, Izquierdo Moya (Eds) Manuel Martinez de la Escalera. Biografia y Publicaciones. DVD. Museo Nacional de Ciencias Naturales (CSIC) and Asociacion espanola de Entomologia." title="Las &quot; especies &quot; no descritas de M. Martinez de la Escalera. In: Martin Albaladejo, Izquierdo Moya (Eds) Manuel Martinez de la Escalera. Biografia y Publicaciones. DVD. Museo Nacional de Ciencias Naturales (CSIC) and Asociacion espanola de Entomologia." year="2011">
<normalizedToken originalValue="París">Paris</normalizedToken>
<normalizedToken originalValue="García">Garcia</normalizedToken>
et al. 2011
</bibRefCitation>
). The latter is a singular population, apparently isolated in the arid mountains near Ifni; its closest known population is located in the Western High Atlas, ca. 250 km to the northeast (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Geographic distribution of Misolampus goudotii. Map for the distribution range of Misolampus goudotii (pale blue spot). Blue dots correspond to the species records, including both recent and old, as well as previously published data. The population from Ifni remains isolated from the main distribution range, by a distance of ca. 250 km B potential geographic distribution of Misolampus goudotii: Red indicates high suitable areas, and blue, areas of low suitability. Species distribution model was generated using MaxEnt v 3.4.1 (Elith et al. 2006) and the set of WorldClim v 2.0 (Fick and Hijmans 2017) environmental variables." figureDoi="10.3897/zookeys.963.53500.figure4" httpUri="https://binary.pensoft.net/fig/444545" pageId="0" pageNumber="81">4A</figureCitation>
). The potential distribution map locates high suitable areas for this species along the mountain ranges of northwestern Africa, the coastal and mountain areas in the Tingitane peninsula, and along the coast of
<normalizedToken originalValue="Rabat-Salé-Kénitra">Rabat-Sale-Kenitra</normalizedToken>
region. It also identifies areas where the species does not occur as high suitable, including sothwestern Iberia, the Balearic Islands and Sardinia. The Ifni population is located in a very fragmented area of high suitability, suggesting a possible Pleistocene relict status for this population (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Geographic distribution of Misolampus goudotii. Map for the distribution range of Misolampus goudotii (pale blue spot). Blue dots correspond to the species records, including both recent and old, as well as previously published data. The population from Ifni remains isolated from the main distribution range, by a distance of ca. 250 km B potential geographic distribution of Misolampus goudotii: Red indicates high suitable areas, and blue, areas of low suitability. Species distribution model was generated using MaxEnt v 3.4.1 (Elith et al. 2006) and the set of WorldClim v 2.0 (Fick and Hijmans 2017) environmental variables." figureDoi="10.3897/zookeys.963.53500.figure4" httpUri="https://binary.pensoft.net/fig/444545" pageId="0" pageNumber="81">4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="notes on natural history">
<paragraph pageId="0" pageNumber="81">Notes on natural history.</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
is widely distributed over northwestern Africa, though restricted to mountain ranges and adjacent areas: Rif, Middle Atlas, western High Atlas, Beni Snassen mountains, southwestern foothills of the Anti-Atlas (Morocco) and Tellian Atlas (Algerie) (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Geographic distribution of Misolampus goudotii. Map for the distribution range of Misolampus goudotii (pale blue spot). Blue dots correspond to the species records, including both recent and old, as well as previously published data. The population from Ifni remains isolated from the main distribution range, by a distance of ca. 250 km B potential geographic distribution of Misolampus goudotii: Red indicates high suitable areas, and blue, areas of low suitability. Species distribution model was generated using MaxEnt v 3.4.1 (Elith et al. 2006) and the set of WorldClim v 2.0 (Fick and Hijmans 2017) environmental variables." figureDoi="10.3897/zookeys.963.53500.figure4" httpUri="https://binary.pensoft.net/fig/444545" pageId="0" pageNumber="81">4</figureCitation>
). Altitudinal range in the Maghreb from 2 to 2064 m a.s.l., with 70.5% of records above 800 m of altitude (62% above 1000 m). In the Balearic Islands its altitudinal range is lower, between 15 and 718 m a.s.l., but the species is found mainly in areas of mountainous topography (e.g., Serra de Tramuntana in Mallorca). It inhabits a wide range of geological substrates, both acid and basic, from plutonic and metamorphic types to calcareous and dolomitic rocks (see
<bibRefCitation author="Michard, A" journalOrPublisher="Notes et Memories Service Geologique du Maroc" pageId="0" pageNumber="81" pagination="1 - 408" refId="B105" refString="Michard, A, 1976. Elements de Geologie marocaine. Notes et Memories Service Geologique du Maroc 252: 1 - 408" title="Elements de Geologie marocaine." volume="252" year="1976">Michard 1976</bibRefCitation>
;
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;
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,
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).
<taxonomicName authorityName="Guerin-Meneville" authorityYear="1834" class="Insecta" family="Tenebrionidae" genus="Misolampus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Misolampus goudotii" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">Misolampus goudotii</emphasis>
</taxonomicName>
is a euryecious species that occurs at infra-, thermo-, meso- and supra-Mediterranean thermoclimatic belts, in regions with ombrotypes from arid to hyperhumid (
<bibRefCitation author="Benabid, A" journalOrPublisher="Foret mediterraneenne" pageId="0" pageNumber="81" pagination="53 - 64" refId="B9" refString="Benabid, A, 1985. Les ecosystemes forestiers, preforestiers et presteppiques du Maroc: diversite, repartition biogeographique et problemes poses par leur amenagement. Foret mediterraneenne 7 (1): 53 - 64" title="Les ecosystemes forestiers, preforestiers et presteppiques du Maroc: diversite, repartition biogeographique et problemes poses par leur amenagement." volume="7" year="1985">Benabid 1985</bibRefCitation>
;
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<normalizedToken originalValue="Rivas-Martínez">Rivas-Martinez</normalizedToken>
1987
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3406/ecmed.1989.1643" author="Le Houerou, HN" journalOrPublisher="Amand Koenig, Parisiis et Argentoratii" pageId="0" pageNumber="81" refId="B87" refString="Le Houerou, HN, 1989. Classification ecoclimatique des zones arides (s.l.) de l'Afrique du Nord. Ecologia mediterranea 15(3/4): 95-144. [7 pls] https://doi.org/10.3406/ecmed.1989.1643" title="Classification ecoclimatique des zones arides (s. l.) de l'Afrique du Nord. Ecologia mediterranea 15 (3 / 4): 95 - 144. [7 pls]" url="https://doi.org/10.3406/ecmed.1989.1643" year="1989">Le Houerou 1989</bibRefCitation>
;
<bibRefCitation author="Rivas-Martinez, S" journalOrPublisher="Itinera Geobotanica" pageId="0" pageNumber="81" pagination="5 - 432" refId="B142" refString="Rivas-Martinez, S, Diaz, TE, Fernandez-Gonzalez, F, Izco, J, Loidi, J, Lousa, M, Penas, A, 2002. Vascular plant communities of Spain and Portugal. Addenda to the syntaxonomical checklist of 2001. Part I. Itinera Geobotanica 15: 5 - 432" title="Vascular plant communities of Spain and Portugal. Addenda to the syntaxonomical checklist of 2001. Part I." volume="15" year="2002">
<normalizedToken originalValue="Rivas-Martínez">Rivas-Martinez</normalizedToken>
et al. 2002
</bibRefCitation>
;
<bibRefCitation author="Rivas-Martinez, S" journalOrPublisher="Itinera Geobotanica" pageId="0" pageNumber="81" pagination="1 - 436" refId="B140" refString="Rivas-Martinez, S, 2007. Mapa de series, geoseries y geopermaseries de vegetacion de Espana [Memoria del Mapa de Vegetacion Potencial de Espana. Parte I]. Itinera Geobotanica 17 (1): 1 - 436" title="Mapa de series, geoseries y geopermaseries de vegetacion de Espana [Memoria del Mapa de Vegetacion Potencial de Espana. Parte I]." volume="17" year="2007">
<normalizedToken originalValue="Rivas-Martínez">Rivas-Martinez</normalizedToken>
2007
</bibRefCitation>
;
<bibRefCitation author="Sebbar, A" journalOrPublisher="Coleccion Monografica Tierras del Sur, Universidad de Granada, Granada" pageId="0" pageNumber="81" refId="B157" refString="Sebbar, A, Hsaine, M, Fougrach, H, Badri, W, 2013. Carte des precipitations annuelles au Maroc (1935/2006). 26eme Colloque de l'Association Internationale de Climatologie, Cotonou (Benin), September 2013: 37-43." title="Carte des precipitations annuelles au Maroc (1935 / 2006). 26 eme Colloque de l'Association Internationale de Climatologie, Cotonou (Benin), September 2013: 37 - 43." year="2013">Sebbar et al. 2013</bibRefCitation>
), and occupies a wide variety of forest formations, both coniferous [
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Tetraclinis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetraclinis articulata" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="articulata">
<emphasis italics="true" pageId="0" pageNumber="81">Tetraclinis articulata</emphasis>
</taxonomicName>
(Vahl) Mast.,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Abies maroccana" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="maroccana">
<emphasis italics="true" pageId="0" pageNumber="81">Abies maroccana</emphasis>
</taxonomicName>
Trab.,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Cedrus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cedrus atlantica" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="atlantica">
<emphasis italics="true" pageId="0" pageNumber="81">Cedrus atlantica</emphasis>
</taxonomicName>
(Endl.) Manetti ex
<normalizedToken originalValue="Carrière">Carriere</normalizedToken>
,
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus phoenicea" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="phoenicea">
<emphasis italics="true" pageId="0" pageNumber="81">Juniperus phoenicea</emphasis>
</taxonomicName>
L.,
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Juniperus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Juniperus thurifera" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="thurifera">
<emphasis italics="true" pageId="0" pageNumber="81">Juniperus thurifera</emphasis>
</taxonomicName>
L.,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus nigra" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="81">Pinus nigra</emphasis>
</taxonomicName>
J.F. Arnold,
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus halepensis" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="halepensis">
<emphasis italics="true" pageId="0" pageNumber="81">Pinus halepensis</emphasis>
</taxonomicName>
Mill.,
<taxonomicName lsidName="P. pinaster" pageId="0" pageNumber="81" rank="species" species="pinaster">
<emphasis italics="true" pageId="0" pageNumber="81">P. pinaster</emphasis>
</taxonomicName>
] and broadleaved [deciduous:
<taxonomicName authorityName="Willd" authorityYear="1809" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus canariensis" order="Fagales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="canariensis">
<emphasis italics="true" pageId="0" pageNumber="81">Quercus canariensis</emphasis>
</taxonomicName>
Willd.,
<taxonomicName authorityName="Pomel" authorityYear="1875" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus afares" order="Fagales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="afares">
<emphasis italics="true" pageId="0" pageNumber="81">Quercus afares</emphasis>
</taxonomicName>
Pomel,
<taxonomicName lsidName="Q. faginea" pageId="0" pageNumber="81" rank="species" species="faginea">
<emphasis italics="true" pageId="0" pageNumber="81">Q. faginea</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. pyrenaica" pageId="0" pageNumber="81" rank="species" species="pyrenaica">
<emphasis italics="true" pageId="0" pageNumber="81">Q. pyrenaica</emphasis>
</taxonomicName>
; perennial:
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus ilex" order="Fagales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis italics="true" pageId="0" pageNumber="81">Quercus ilex</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. suber" pageId="0" pageNumber="81" rank="species" species="suber">
<emphasis italics="true" pageId="0" pageNumber="81">Q. suber</emphasis>
</taxonomicName>
,
<taxonomicName class="Magnoliopsida" family="Oleaceae" genus="Olea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Olea europaea subsp. var. var. sylvestris" order="Lamiales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="variety" species="europaea" subSpecies="var." variety="sylvestris">Olea europaea var. sylvestris</taxonomicName>
(Mill.) Lehr] (see
<bibRefCitation author="Benabid, A" journalOrPublisher="Universite de Droit, d'Economie et des Sciences d'Aix-Marseille" pageId="0" pageNumber="81" refId="B7" refString="Benabid, A, 1982. Etudes phytoecologique, biogeographique et dinamique des associations et series sylvatiques du Rif occidental (Maroc). These Doct. es-Sc. Faculte des Sciences et Techniques St. Jerome. Universite de Droit, d'Economie et des Sciences d'Aix-Marseille" title="Etudes phytoecologique, biogeographique et dinamique des associations et series sylvatiques du Rif occidental (Maroc). These Doct. es-Sc. Faculte des Sciences et Techniques St. Jerome." year="1982">Benabid 1982</bibRefCitation>
,
<bibRefCitation author="Benabid, A" journalOrPublisher="Travaux de l'Institut Scientifique, Rabat, serie Botanique" pageId="0" pageNumber="81" pagination="1 - 64" refId="B8" refString="Benabid, A, 1984. Etude phytoecologique des peuplements forestiers et preforestiers du Rif centro-occidental (Maroc). Travaux de l'Institut Scientifique, Rabat, serie Botanique 34: 1 - 64" title="Etude phytoecologique des peuplements forestiers et preforestiers du Rif centro-occidental (Maroc)." volume="34" year="1984">1984</bibRefCitation>
,
<bibRefCitation author="Benabid, A" journalOrPublisher="Foret mediterraneenne" pageId="0" pageNumber="81" pagination="53 - 64" refId="B9" refString="Benabid, A, 1985. Les ecosystemes forestiers, preforestiers et presteppiques du Maroc: diversite, repartition biogeographique et problemes poses par leur amenagement. Foret mediterraneenne 7 (1): 53 - 64" title="Les ecosystemes forestiers, preforestiers et presteppiques du Maroc: diversite, repartition biogeographique et problemes poses par leur amenagement." volume="7" year="1985">1985</bibRefCitation>
;
<bibRefCitation author="Benabid, A" journalOrPublisher="Lazaroa" pageId="0" pageNumber="81" pagination="21 - 97" refId="B11" refString="Benabid, A, Fennane, M, 1994. Connaissances sur la vegetation du Maroc: Phytogeographie, phytosociologie et series de vegetation. Lazaroa 14: 21 - 97" title="Connaissances sur la vegetation du Maroc: Phytogeographie, phytosociologie et series de vegetation." volume="14" year="1994">Benabid and Fennane 1994</bibRefCitation>
;
<bibRefCitation author="Bolos de, O" journalOrPublisher="Institut d'Estudis Catalans, Arxius de les Seccions de Ciencies 114 [1996]" pageId="0" pageNumber="81" refId="B14" refString="Bolos de, O, 1997. La vegetacio de les Illes Balears. Comunitat des plantes. Institut d'Estudis Catalans, Arxius de les Seccions de Ciencies 114 [1996]" title="La vegetacio de les Illes Balears. Comunitat des plantes." year="1997">
<normalizedToken originalValue="Bolòs">Bolos</normalizedToken>
1997
</bibRefCitation>
;
<bibRefCitation author="Charco, J" journalOrPublisher="Agencia Espanola de Cooperacion Internacional, Madrid" pageId="0" pageNumber="81" refId="B26" refString="Charco, J, 1999. El bosque mediterraneo en el norte de Africa. Biodiversidad y lucha contra la desertificacion. Agencia Espanola de Cooperacion Internacional, Madrid" title="El bosque mediterraneo en el norte de Africa. Biodiversidad y lucha contra la desertificacion." year="1999">Charco 1999</bibRefCitation>
;
<bibRefCitation author="Benabid, A" journalOrPublisher="Ibis Press, Paris" pageId="0" pageNumber="81" refId="B10" refString="Benabid, A, 2000. Flore et ecosystemes du Maroc. Evaluation et preservation de la biodiversite. Ed. Ibis Press, Paris" title="Flore et ecosystemes du Maroc. Evaluation et preservation de la biodiversite. Ed." year="2000">Benabid 2000</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1007/978-3-319-93704-5" author="Taleb, MS" editor="Pedrotti, F" journalOrPublisher="Basics, Methods and Case Studies. Springer Nature Switzerland AG. Cham." pageId="0" pageNumber="81" pagination="1 - 161" refId="B163" refString="Taleb, MS, Fennane, M, 2019. Vascular Plant Communities of Morocco. Phytosociology, Ecology and Geography. In: Pedrotti, F, Ed., Geobotany Studies. Basics, Methods and Case Studies. Springer Nature Switzerland AG. Cham.: 1 - 161, DOI: https://doi.org/10.1007/978-3-319-93704-5" title="Vascular Plant Communities of Morocco. Phytosociology, Ecology and Geography." url="https://doi.org/10.1007/978-3-319-93704-5" volumeTitle="Geobotany Studies." year="2019">Taleb and Fennane 2019</bibRefCitation>
). It also occurs in areas reforested with pines (pers. obs.) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3D, E</figureCitation>
). The population of Ifni inhabits mountains (620-1225 m of altitude) at the infra-Mediterranean thermoclimatic belt, probably affected by the proximity to the Atlantic Ocean and consequently by the presence of some degree of cryptic precipitation (
<bibRefCitation author="Gehu, JM" journalOrPublisher="Acta Botanica Barcinonensia" pageId="0" pageNumber="81" pagination="439 - 453" refId="B58" refString="Gehu, JM, Biondi, E, 1998. Nature et limites de quelques vegetations littorales de type macaronesien sur les cotes sud occidentales du Maroc. Acta Botanica Barcinonensia 45: 439 - 453" title="Nature et limites de quelques vegetations littorales de type macaronesien sur les cotes sud occidentales du Maroc." volume="45" year="1998">
<normalizedToken originalValue="Géhu">Gehu</normalizedToken>
and Biondi 1998
</bibRefCitation>
). The vegetation of the area is dominated by open forest of
<taxonomicName authorityName="Skeels" authorityYear="1911" class="Insecta" family="Noctuidae" genus="Argania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argania spinosa" order="Lepidoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis italics="true" pageId="0" pageNumber="81">Argania spinosa</emphasis>
</taxonomicName>
(L.) Skeels, with sparse cactiform and arbustive
<taxonomicName class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Euphorbia" order="Malpighiales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Euphorbia</emphasis>
</taxonomicName>
L. (
<bibRefCitation DOI="https://doi.org/10.1023/A:1009775327616" author="Medail, F" journalOrPublisher="Plant Ecology" pageId="0" pageNumber="81" pagination="221 - 244" refId="B103" refString="Medail, F, Quezel, P, 1999. The phytogeographical significance of S.W. Morocco compared to the Canary Islands. Plant Ecology 140: 221 - 244, DOI: https://doi.org/10.1023/A:1009775327616" title="The phytogeographical significance of S. W. Morocco compared to the Canary Islands." url="https://doi.org/10.1023/A:1009775327616" volume="140" year="1999">
<normalizedToken originalValue="Médail">Medail</normalizedToken>
and
<normalizedToken originalValue="Quézel">Quezel</normalizedToken>
1999
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3989/graellsia.2015.v71.118" author="Ruiz, JL" journalOrPublisher="Boletin de la Asociacion espanola de Entomologia" pageId="0" pageNumber="81" refId="B148" refString="Ruiz, JL, Garcia-Paris, M, 2015. Una nueva especie de Meloe Linnaeus, 1758 del grupo de M. (Eurymeloe) rugosus Marsham, 1802 (Coleoptera: Meloidae) del suroeste de Marruecos. Graellsia 71(1): e018. https://doi.org/10.3989/graellsia.2015.v71.118" title="Una nueva especie de Meloe Linnaeus, 1758 del grupo de M. (Eurymeloe) rugosus Marsham, 1802 (Coleoptera: Meloidae) del suroeste de Marruecos. Graellsia 71 (1): e 018." url="https://doi.org/10.3989/graellsia.2015.v71.118" year="2015">
Ruiz and
<normalizedToken originalValue="García-París">Garcia-Paris</normalizedToken>
2015
</bibRefCitation>
), and large areas covered by formerly cultivated
<taxonomicName class="Magnoliopsida" family="Cactaceae" genus="Opuntia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Opuntia" order="Caryophyllales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Opuntia</emphasis>
</taxonomicName>
Mill (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3F</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
In the Moroccan Rif,
<taxonomicName lsidName="M. goudotii" pageId="0" pageNumber="81" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">M. goudotii</emphasis>
</taxonomicName>
is often encountered under bark, inside fallen logs or stumps, and at the base of dead old oaks (perennial:
<taxonomicName lsidName="Q. ilex" pageId="0" pageNumber="81" rank="species" species="ilex">
<emphasis italics="true" pageId="0" pageNumber="81">Q. ilex</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. suber" pageId="0" pageNumber="81" rank="species" species="suber">
<emphasis italics="true" pageId="0" pageNumber="81">Q. suber</emphasis>
</taxonomicName>
; deciduous:
<taxonomicName lsidName="Q. canariensis" pageId="0" pageNumber="81" rank="species" species="canariensis">
<emphasis italics="true" pageId="0" pageNumber="81">Q. canariensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. faginea" pageId="0" pageNumber="81" rank="species" species="faginea">
<emphasis italics="true" pageId="0" pageNumber="81">Q. faginea</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="Q. pyrenaica" pageId="0" pageNumber="81" rank="species" species="pyrenaica">
<emphasis italics="true" pageId="0" pageNumber="81">Q. pyrenaica</emphasis>
</taxonomicName>
), arbutus trees (
<taxonomicName class="Magnoliopsida" family="Ericaceae" genus="Arbutus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Arbutus unedo" order="Ericales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="unedo">
<emphasis italics="true" pageId="0" pageNumber="81">Arbutus unedo</emphasis>
</taxonomicName>
L.), wild olive trees (
<taxonomicName lsidName="O. europaea subsp. var. var. sylvestris" pageId="0" pageNumber="81" rank="variety" species="europaea" subSpecies="var." variety="sylvestris">O. europaea var. sylvestris</taxonomicName>
), pines (
<taxonomicName lsidName="P. nigra" pageId="0" pageNumber="81" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="81">P. nigra</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. pinaster" pageId="0" pageNumber="81" rank="species" species="pinaster">
<emphasis italics="true" pageId="0" pageNumber="81">P. pinaster</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="P. halepensis" pageId="0" pageNumber="81" rank="species" species="halepensis">
<emphasis italics="true" pageId="0" pageNumber="81">P. halepensis</emphasis>
</taxonomicName>
), firs (
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Abies maroccana" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="maroccana">
<emphasis italics="true" pageId="0" pageNumber="81">Abies maroccana</emphasis>
</taxonomicName>
), and cedars (
<taxonomicName class="Pinopsida" family="Pinaceae" genus="Cedrus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cedrus atlantica" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="atlantica">
<emphasis italics="true" pageId="0" pageNumber="81">Cedrus atlantica</emphasis>
</taxonomicName>
), as already reported partially by Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
(
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">1900</bibRefCitation>
),
<bibRefCitation author="Cobos, A" journalOrPublisher="Archivos del Instituto de Aclimatacion de Almeria" pageId="0" pageNumber="81" pagination="1 - 9" refId="B29" refString="Cobos, A, 1955. Prospeccion entomologica en la zona espanola del Protectorado en Marruecos. Archivos del Instituto de Aclimatacion de Almeria 4: 1 - 9" title="Prospeccion entomologica en la zona espanola del Protectorado en Marruecos." volume="4" year="1955">Cobos (1955</bibRefCitation>
,
<bibRefCitation author="Cobos, A" journalOrPublisher="Archivos del Instituto de Aclimatacion de Almeria" pageId="0" pageNumber="81" pagination="63 - 98" refId="B30" refString="Cobos, A, 1961. Exploracion entomologica del cedral de Tizi-Ifri (macizo del Iguelmalet, Rif central, Marruecos). Coleopteros. Archivos del Instituto de Aclimatacion de Almeria 10: 63 - 98" title="Exploracion entomologica del cedral de Tizi-Ifri (macizo del Iguelmalet, Rif central, Marruecos). Coleopteros." volume="10" year="1961">1961</bibRefCitation>
), and
<bibRefCitation DOI="https://doi.org/10.15560/11.5.1778" author="Benyahia, Y" journalOrPublisher="Check List" pageId="0" pageNumber="81" pagination=": 1778" refId="B13" refString="Benyahia, Y, Soldati, F, Rohi, L, Valladares, L, Maatouf, N, Courtin, O, El Antry, S, Brustel, H, 2015. First survey of darkling beetles (Coleoptera, Tenebrionidae) of Talassemtane National Park, Western Rif, Morocco. Check List 11 (5): 1778, DOI: https://doi.org/10.15560/11.5.1778" title="First survey of darkling beetles (Coleoptera, Tenebrionidae) of Talassemtane National Park, Western Rif, Morocco." url="https://doi.org/10.15560/11.5.1778" volume="11" year="2015">Benyahia et al. (2015</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.12976/jib/2015.4.13" author="Benyahia, Y" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="81" pagination="1 - 30" refId="B12" refString="Benyahia, Y, Brustel, H, El Antry, S, Courtin, O, Maatouf, N, Valladares, L, Rohi, L, 2016. Preliminary list of Coleoptera heritage species of the Talassemtane National Park, Morocco. Journal of Insect Biodiversity 4 (13): 1 - 30, DOI: https://doi.org/10.12976/jib/2015.4.13" title="Preliminary list of Coleoptera heritage species of the Talassemtane National Park, Morocco." url="https://doi.org/10.12976/jib/2015.4.13" volume="4" year="2016">2016</bibRefCitation>
). They can also be found under bark of standing dead trees (
<taxonomicName lsidName="A. maroccana" pageId="0" pageNumber="81" rank="species" species="maroccana">
<emphasis italics="true" pageId="0" pageNumber="81">A. maroccana</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="C. atlantica" pageId="0" pageNumber="81" rank="species" species="atlantica">
<emphasis italics="true" pageId="0" pageNumber="81">C. atlantica</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. suber" pageId="0" pageNumber="81" rank="species" species="suber">
<emphasis italics="true" pageId="0" pageNumber="81">Q. suber</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. pyrenaica" pageId="0" pageNumber="81" rank="species" species="pyrenaica">
<emphasis italics="true" pageId="0" pageNumber="81">Q. pyrenaica</emphasis>
</taxonomicName>
). In the Middle and High Atlas, it is usually found under bark and inside large decaying logs of
<taxonomicName lsidName="Q. ilex" pageId="0" pageNumber="81" rank="species" species="ilex">
<emphasis italics="true" pageId="0" pageNumber="81">Q. ilex</emphasis>
</taxonomicName>
(
<bibRefCitation author="Antoine, M" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles du Maroc" pageId="0" pageNumber="81" pagination="248 - 259" publicationUrl="https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" refId="B3" refString="Antoine, M, 1926. Notes d'entomologie marocaine. VI (1) Tenebrionides nouveaux et interessants. Bulletin de la Societe des Sciences Naturelles du Maroc 5: 248 - 259, https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" title="Notes d'entomologie marocaine. VI (1) Tenebrionides nouveaux et interessants." url="https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" volume="5" year="1926">Antoine 1926</bibRefCitation>
), but also in old decomposing logs of
<taxonomicName lsidName="P. nigra" pageId="0" pageNumber="81" rank="species" species="nigra">
<emphasis italics="true" pageId="0" pageNumber="81">P. nigra</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. atlantica" pageId="0" pageNumber="81" rank="species" species="atlantica">
<emphasis italics="true" pageId="0" pageNumber="81">C. atlantica</emphasis>
</taxonomicName>
.
<bibRefCitation author="Mouna, M" journalOrPublisher="Bulletin de l'Institut Scientifique, Rabat" pageId="0" pageNumber="81" pagination="133 - 142" refId="B110" refString="Mouna, M, Arahou, M, 1986. Comparaison des peuplements de Coleopteres frondicoles de deux essences forestieres resineuses au Maroc: Cedre (Cedrus atlantica Man) et Thuya (Tetraclinis articulata Vahl et Benth). Bulletin de l'Institut Scientifique, Rabat 1: 133 - 142" title="Comparaison des peuplements de Coleopteres frondicoles de deux essences forestieres resineuses au Maroc: Cedre (Cedrus atlantica Man) et Thuya (Tetraclinis articulata Vahl et Benth)." volume="1" year="1986">Mouna and Arahou (1986)</bibRefCitation>
collected the species on thuya (
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Tetraclinis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tetraclinis articulata" order="Pinales" pageId="0" pageNumber="81" phylum="Tracheophyta" rank="species" species="articulata">
<emphasis italics="true" pageId="0" pageNumber="81">Tetraclinis articulata</emphasis>
</taxonomicName>
) in the Korifla Valley (northwestern Morocco). Sidi Ifni specimens were found within crevices in old dead logs of
<taxonomicName authorityName="Skeels" authorityYear="1911" class="Insecta" family="Noctuidae" genus="Argania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Argania spinosa" order="Lepidoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis italics="true" pageId="0" pageNumber="81">Argania spinosa</emphasis>
</taxonomicName>
, almost buried on the ground of a steep slope (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3F</figureCitation>
). Nearby standing dead trunks were occupied by
<taxonomicName class="Insecta" family="Asilidae" genus="Nesotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesotes tuberculipennis subsp. villarubiai" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="tuberculipennis" subSpecies="villarubiai">
<emphasis italics="true" pageId="0" pageNumber="81">Nesotes tuberculipennis villarubiai</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Español">Espanol</normalizedToken>
, 1943) as described by
<bibRefCitation DOI="https://doi.org/10.23885/1814-3326-2015-11-1-33-37" author="Nabozhenko, MV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="0" pageNumber="81" pagination="33 - 37" refId="B111" refString="Nabozhenko, MV, 2015. To the knowledge of Helopini (Coleoptera: Tenebrionidae) of Morocco. Caucasian Entomological Bulletin 11 (1): 33 - 37, DOI: https://doi.org/10.23885/1814-3326-2015-11-1-33-37" title="To the knowledge of Helopini (Coleoptera: Tenebrionidae) of Morocco." url="https://doi.org/10.23885/1814-3326-2015-11-1-33-37" volume="11" year="2015">Nabozhenko (2015)</bibRefCitation>
. In Algeria, they have been found under bark of fallen pines (
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">
Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
1900
</bibRefCitation>
). In Menorca, it has been found in oak forests of
<taxonomicName lsidName="Q. ilex" pageId="0" pageNumber="81" rank="species" species="ilex">
<emphasis italics="true" pageId="0" pageNumber="81">Q. ilex</emphasis>
</taxonomicName>
, under bark or under stones and leaf litter (
<bibRefCitation author="Cardona Orfila, F" journalOrPublisher="Imprenta de Miguel Parpal, Mao" pageId="0" pageNumber="81" publicationUrl="https://issuu.com/bibliomao/docs/doscientos_cole__pteros_mas_de_meno" refId="B24" refString="Cardona Orfila, F, 1875. Doscientos coleopteros mas de Menorca. Imprenta de Miguel Parpal, Mao, https://issuu.com/bibliomao/docs/doscientos_cole__pteros_mas_de_meno" title="Doscientos coleopteros mas de Menorca." url="https://issuu.com/bibliomao/docs/doscientos_cole__pteros_mas_de_meno" year="1875">Cardona Orfila 1875</bibRefCitation>
), and in Mallorca it is frequent in decaying wood of fallen pines (
<taxonomicName lsidName="P. pinea" pageId="0" pageNumber="81" rank="species" species="pinea">
<emphasis italics="true" pageId="0" pageNumber="81">P. pinea</emphasis>
</taxonomicName>
) and old oaks (
<taxonomicName lsidName="Q. ilex" pageId="0" pageNumber="81" rank="species" species="ilex">
<emphasis italics="true" pageId="0" pageNumber="81">Q. ilex</emphasis>
</taxonomicName>
) (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Live specimens and habitat of Misolampus goudotii A-C Live adult specimens of Misolampus goudotii from the Balearic Islands (A Cap Formentor, Mallorca) and Morocco (B Toufliht, High Atlas C Akarkor, Sidi Ifni); the specimens selected represent the diversity of sculptural patterns in elytra and pronotum reported for the species D-F A summary of the impressive habitat diversity used by M. goudotii from the Balearic Islands (D Quercus ilex forest at Creu de Menut, Mallorca), to southwestern Morocco (E deep valleys in the Toubkal National Park, High Atlas F Argania spinosa open forests at Jbel Toual in Sidi Ifni). Photographs by MGP and NRR." figureDoi="10.3897/zookeys.963.53500.figure3" httpUri="https://binary.pensoft.net/fig/444544" pageId="0" pageNumber="81">3D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
Adult specimens are often found in aggregations. We found aggregations of approximately 15 specimens close together in a single large rotting pine log in Mallorca. We also found aggregations of
<taxonomicName lsidName="M. goudotii" pageId="0" pageNumber="81" rank="species" species="goudotii">
<emphasis italics="true" pageId="0" pageNumber="81">M. goudotii</emphasis>
</taxonomicName>
together with
<taxonomicName class="Insecta" family="Leiodidae" genus="Helops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helops insignis subsp. maroccanus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="subSpecies" species="insignis" subSpecies="maroccanus">
<emphasis italics="true" pageId="0" pageNumber="81">Helops insignis maroccanus</emphasis>
</taxonomicName>
(Fairmaire, 1873) (
<taxonomicName authorityName="Latreille" authorityYear="1802" family="Tenebrionidae" lsidName="" pageId="0" pageNumber="81" rank="family">Tenebrionidae</taxonomicName>
,
<taxonomicName lsidName="" pageId="0" pageNumber="81" rank="subFamily" subFamily="Helopinae">Helopinae</taxonomicName>
) under bark of dead trees of
<taxonomicName lsidName="Q. suber" pageId="0" pageNumber="81" rank="species" species="suber">
<emphasis italics="true" pageId="0" pageNumber="81">Q. suber</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="A. maroccana" pageId="0" pageNumber="81" rank="species" species="maroccana">
<emphasis italics="true" pageId="0" pageNumber="81">A. maroccana</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. atlantica" pageId="0" pageNumber="81" rank="species" species="atlantica">
<emphasis italics="true" pageId="0" pageNumber="81">C. atlantica</emphasis>
</taxonomicName>
in the Rif Mountains.
<bibRefCitation author="Whitehead, PF" journalOrPublisher="Bolleti de la Societat d'Historia Natural de les Balears" pageId="0" pageNumber="81" pagination="45 - 56" refId="B180" refString="Whitehead, PF, 1993. Observations on Coleoptera of Mallorca, Balearic Islands. Bolleti de la Societat d'Historia Natural de les Balears 36: 45 - 56" title="Observations on Coleoptera of Mallorca, Balearic Islands." volume="36" year="1993">Whitehead (1993)</bibRefCitation>
relates the finding on two occasions of groups of individuals between the annual rings of dead pines (
<taxonomicName lsidName="P. halepensis" pageId="0" pageNumber="81" rank="species" species="halepensis">
<emphasis italics="true" pageId="0" pageNumber="81">P. halepensis</emphasis>
</taxonomicName>
) in active colonies of ants of the genus
<taxonomicName authorityName="Forel" authorityYear="1890" class="Insecta" family="Formicidae" genus="Messor" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Messor" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Messor</emphasis>
</taxonomicName>
Forel, 1890 and of the species
<taxonomicName authorityName="Emery" authorityYear="1877" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium bicolor" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="81">Monomorium bicolor</emphasis>
</taxonomicName>
Emery, 1877 (probably another species of
<taxonomicName authorityName="Mayr" authorityYear="1855" class="Insecta" family="Formicidae" genus="Monomorium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monomorium" order="Hymenoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Monomorium</emphasis>
</taxonomicName>
Mayr, 1855, since the invasive
<taxonomicName lsidName="M. bicolor" pageId="0" pageNumber="81" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="81">M. bicolor</emphasis>
</taxonomicName>
is not present in Balearic Islands;
<bibRefCitation author="Salata, S" journalOrPublisher="North-Western Journal of Zoology" pageId="0" pageNumber="81" pagination="13 - 23" refId="B150" refString="Salata, S, Georgiadis, C, Borowiec, L, 2019. Invasive ant species (Hymenoptera: Formicidae) of Greece and Cyprus. North-Western Journal of Zoology 15 (1): 13 - 23" title="Invasive ant species (Hymenoptera: Formicidae) of Greece and Cyprus." volume="15" year="2019">Salata et al. 2019</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">
Adults are present all year round, but they are more commonly seen in winter and spring in middle and low elevations (
<bibRefCitation author="Vauloger de, Beaupre" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="81" pagination="669 - 722" publicationUrl="https://www.biodiversitylibrary.org/page/8431237" refId="B170" refString="Vauloger de, Beaupre, 1900. Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini. Annales de la Societe Entomologique de France 68 (1899): 669 - 722, https://www.biodiversitylibrary.org/page/8431237" title="Contribution au catalogue des coleopteres du Nord de l'Afrique. Helopini." url="https://www.biodiversitylibrary.org/page/8431237" volume="68" year="1900">
Vauloger de
<normalizedToken originalValue="Beaupré">Beaupre</normalizedToken>
1900
</bibRefCitation>
;
<bibRefCitation author="Espanol, F" journalOrPublisher="Publicaciones del Instituto de Biologia Aplicada" pageId="0" pageNumber="81" pagination="17 - 47" refId="B49" refString="Espanol, F, 1967. Mision entomologica Hakan Lindberg y M. Meinander a Marruecos. Col. Tenebrionidae. Publicaciones del Instituto de Biologia Aplicada 42: 17 - 47" title="Mision entomologica Hakan Lindberg y M. Meinander a Marruecos. Col. Tenebrionidae." volume="42" year="1967">
<normalizedToken originalValue="Español">Espanol</normalizedToken>
1967
</bibRefCitation>
; pers. obs.), and in summer at higher altitude (
<bibRefCitation author="Antoine, M" journalOrPublisher="Bulletin de la Societe des Sciences Naturelles du Maroc" pageId="0" pageNumber="81" pagination="248 - 259" publicationUrl="https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" refId="B3" refString="Antoine, M, 1926. Notes d'entomologie marocaine. VI (1) Tenebrionides nouveaux et interessants. Bulletin de la Societe des Sciences Naturelles du Maroc 5: 248 - 259, https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" title="Notes d'entomologie marocaine. VI (1) Tenebrionides nouveaux et interessants." url="https://bibdigital.rjb.csic.es/viewer/12177/?offset=page=302&amp;viewer=picture&amp;o=bookmark&amp;n=0&amp;q=" volume="5" year="1926">Antoine 1926</bibRefCitation>
), however,
<bibRefCitation author="Moragues, F" journalOrPublisher="Anales de la Sociedad Espanola de Historia Natural" pageId="0" pageNumber="81" pagination="11 - 34" publicationUrl="https://www.biodiversitylibrary.org/page/8157089" refId="B109" refString="Moragues, F, 1889. Coleopteros de Mallorca. Anales de la Sociedad Espanola de Historia Natural 18 (1888): 11 - 34, https://www.biodiversitylibrary.org/page/8157089" title="Coleopteros de Mallorca." url="https://www.biodiversitylibrary.org/page/8157089" volume="18" year="1889">Moragues (1889)</bibRefCitation>
mentioned collections during the summer in Mallorca.
</paragraph>
</subSubSection>
</treatment>
</document>