treatments-xml/data/AF/1D/A8/AF1DA83BD458FFF0FF3AD91845017A6F.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3790426" ID-GBIF-Dataset="5101dbd6-06ac-4b6e-991e-b66eec55ca52" ID-GBIF-Taxon="163750796" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790426" ID-ZooBank="46085EC4-FB49-4272-803A-A0FF21EE52CD" LSID="urn:lsid:zoobank.org:act:29D10CFC-A8FB-431B-A61C-289B7619CF78" approvalRequired="22" approvalRequired_for_textStreams="22" checkinTime="1587986115529" checkinUser="plazi" docAuthor="Kraus, Fred &amp; Allison, Allen" docDate="2009" docId="AF1DA83BD458FFF0FF3AD91845017A6F" docLanguage="en" docName="ZK_article_2096.pdf" docOrigin="ZooKeys 26 (26)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@42210d27" docStyleName="zookeys.2008.journal_article" docTitle="Oreophryne anamiatoi Kraus &amp; Allison 2009, sp. n." docType="treatment" docUuid="29D10CFC-A8FB-431B-A61C-289B7619CF78" docUuidSource="ZooBank" docVersion="6" lastPageId="18" lastPageNumber="71" masterDocId="5324D043D455FFE2FFB6DC144745783C" masterDocTitle="New microhylid Frogs from the Muller Range, Papua New Guinea" masterLastPageNumber="76" masterPageNumber="53" pageId="13" pageNumber="66" updateTime="1643569204779" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New microhylid Frogs from the Muller Range, Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kraus, Fred</mods:namePart>
<mods:affiliation>University of Michigan, Ann Arbor, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Allison, Allen</mods:namePart>
<mods:affiliation>Bishop Museum ,,</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-10-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>26</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>26</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>53</mods:start>
<mods:end>76</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.26.258</mods:identifier>
<mods:identifier type="GBIF-Dataset">5101dbd6-06ac-4b6e-991e-b66eec55ca52</mods:identifier>
<mods:identifier type="ISSN">1313–2970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576562</mods:identifier>
<mods:identifier type="ZooBank">46085EC4-FB49-4272-803A-A0FF21EE52CD</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790426" ID-GBIF-Taxon="163750796" ID-Zenodo-Dep="3790426" LSID="urn:lsid:zoobank.org:act:29D10CFC-A8FB-431B-A61C-289B7619CF78" httpUri="http://treatment.plazi.org/id/AF1DA83BD458FFF0FF3AD91845017A6F" lastPageId="18" lastPageNumber="71" pageId="13" pageNumber="66">
<subSubSection pageId="13" pageNumber="66" type="nomenclature">
<paragraph blockId="13.[140,979,1292,1390]" box="[140,484,1292,1319]" pageId="13" pageNumber="66">
<heading box="[140,484,1292,1319]" fontSize="11" level="2" pageId="13" pageNumber="66" reason="3">
<emphasis bold="true" box="[140,484,1292,1319]" pageId="13" pageNumber="66">
<taxonomicName authority="Kraus &amp; Allison, 2009" authorityName="Kraus &amp; Allison" authorityYear="2009" box="[140,412,1292,1319]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="13" pageNumber="66" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
<emphasis bold="true" box="[140,412,1292,1319]" italics="true" pageId="13" pageNumber="66">
<emphasis box="[140,281,1292,1319]" inLineHeading="true" pageId="13" pageNumber="66" reason="1">Oreophryne</emphasis>
anamiatoi
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[419,484,1293,1319]" pageId="13" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="13.[140,979,1292,1390]" box="[140,979,1328,1354]" pageId="13" pageNumber="66">
<uri box="[140,979,1328,1354]" pageId="13" pageNumber="66">
urn:lsid:zoobank.org:act: 
<uuid box="[418,979,1328,1354]" pageId="13" pageNumber="66">29D10CFC-A8FB-431B-A61C-289B7619CF78</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection box="[140,263,1363,1390]" pageId="13" pageNumber="66" type="description">
<paragraph blockId="13.[140,979,1292,1390]" box="[140,263,1363,1390]" pageId="13" pageNumber="66">Fig. 1C, D</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="67" pageId="13" pageNumber="66" type="materials_examined">
<paragraph blockId="13.[140,1108,1434,1672]" pageId="13" pageNumber="66">
<materialsCitation ID-GBIF-Occurrence="2608940824" collectingDate="2009-03-27" collectionCode="BPBM, FK" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="Mt. Itukua" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="BPBM 33768, FK 12882" specimenCount="1" stateProvince="Southern Highlands" typeStatus="holotype">
<emphasis bold="true" box="[140,260,1434,1460]" pageId="13" pageNumber="66">
<emphasis box="[140,256,1434,1460]" inLineHeading="true" pageId="13" pageNumber="66" reason="1">
<typeStatus box="[140,256,1434,1460]" pageId="13" pageNumber="66">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode box="[267,425,1434,1460]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="66" type="Museum">BPBM 33768</specimenCode>
(field tag 
<specimenCode box="[538,657,1434,1460]" collectionCode="FK" pageId="13" pageNumber="66">FK 12882</specimenCode>
), collected by 
<collectorName box="[817,909,1434,1460]" pageId="13" pageNumber="66">F. Kraus</collectorName>
and 
<collectorName box="[965,1103,1434,1460]" pageId="13" pageNumber="66">J. Anamiato</collectorName>
, E slope 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D458FFEFFF5BD9AA46347DEB" box="[237,369,1469,1496]" country="Papua New Guinea" latitude="-5.66954" longLatPrecision="1" longitude="142.62334" name="Mt. Itukua" pageId="13" pageNumber="66" stateProvince="Southern Highlands">Mt. Itukua</location>
, 
<collectorName box="[385,546,1469,1496]" pageId="13" pageNumber="66">Muller Range</collectorName>
, 
<geoCoordinate box="[562,695,1469,1496]" degrees="5.66954" direction="south" orientation="latitude" pageId="13" pageNumber="66" precision="1" value="-5.66954">5.66954° S</geoCoordinate>
, 
<geoCoordinate box="[711,876,1469,1496]" degrees="142.62334" direction="east" orientation="longitude" pageId="13" pageNumber="66" precision="1" value="142.62334">142.62334° E</geoCoordinate>
, 
<quantity box="[891,987,1469,1496]" metricMagnitude="3" metricUnit="m" metricValue="2.177" pageId="13" pageNumber="66" unit="m" value="2177.0">
<elevation box="[891,987,1469,1496]" metricMagnitude="3" metricUnit="m" metricValue="2.177" pageId="13" pageNumber="66" unit="m" value="2177.0">2177 m</elevation>
</quantity>
, 
<collectingRegion country="Papua New Guinea" name="Southern Highlands" pageId="13" pageNumber="66">Southern Highlands Province</collectingRegion>
, 
<collectingCountry box="[377,599,1504,1531]" name="Papua New Guinea" pageId="13" pageNumber="66">Papua New Guinea</collectingCountry>
, 
<date box="[611,792,1504,1530]" pageId="13" pageNumber="66" value="2009-03-27">
<collectingDate box="[611,792,1504,1530]" pageId="13" pageNumber="66" value="2009-03-27">27 March 2009</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph blockId="13.[140,1108,1434,1672]" lastBlockId="14.[140,1108,167,1671]" lastPageId="14" lastPageNumber="67" pageId="13" pageNumber="66">
<materialsCitation ID-GBIF-Occurrence="2608940818" collectingDate="2009-03-27" collectionCode="BPBM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="BPBM 33763" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<emphasis bold="true" box="[185,422,1539,1565]" pageId="13" pageNumber="66">
<emphasis box="[185,300,1539,1565]" inLineHeading="true" pageId="13" pageNumber="66" reason="1">
<typeStatus box="[185,300,1539,1565]" pageId="13" pageNumber="66">Paratypes</typeStatus>
</emphasis>
(n = 20).
</emphasis>
<specimenCode box="[434,599,1539,1566]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="66" type="Museum">BPBM 33763</specimenCode>
, same data as holotype except collected 17 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D458FFEFFF3ADA33479F7E7D" box="[140,218,1575,1601]" name="March" pageId="13" pageNumber="66">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940801" box="[232,983,1575,1601]" collectingDate="2009-03-27" collectionCode="BPBM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="BPBM 33764" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[232,396,1575,1601]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="66" type="Museum">BPBM 33764</specimenCode>
, same data as holotype except collected 22 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D458FFEFFC3FDA3344927E7D" box="[905,983,1575,1601]" name="March" pageId="13" pageNumber="66">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940812" collectingDate="2009-03-27" collectionCode="PNGNM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="PNGNM 24097" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode collectionCode="PNGNM" pageId="13" pageNumber="66">PNGNM 24097</specimenCode>
, same data as holotype except collected 25 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D458FFEFFD00DA5E44417E58" box="[694,772,1610,1636]" name="March" pageId="13" pageNumber="66">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940822" collectingDate="2009-03-27" collectionCode="BPBM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="BPBM 33765-66" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[782,987,1610,1637]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="66" type="Museum">BPBM 33765–66</specimenCode>
, same data as holotype except collected 26 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D458FFEFFE42DA7945077EBB" box="[500,578,1645,1671]" name="March" pageId="13" pageNumber="66">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940804" box="[590,1016,1645,1672]" collectingDate="2009-03-27" collectionCode="BPBM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="Mt. Itukua" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="BPBM 33767" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[590,753,1645,1672]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="66" type="Museum">BPBM 33767</specimenCode>
, same data as holotype
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940808" collectingDate="2009-03-27" collectionCode="BPBM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" lastPageId="14" lastPageNumber="67" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="13" pageNumber="66" specimenCode="BPBM 33769-71" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lastPageId="14" lastPageNumber="67" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="66" type="Museum">BPBM 33769–71</specimenCode>
, same data as holotype except collected 28 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D45BFFECFD44DCB3440578FD" box="[754,832,167,193]" name="March" pageId="14" pageNumber="67">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940807" collectingDate="2009-03-27" collectionCode="PNGNM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="14" pageNumber="67" specimenCode="PNGNM 24098" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[844,1039,167,193]" collectionCode="PNGNM" pageId="14" pageNumber="67">PNGNM 24098</specimenCode>
, same data as holotype except collected 29 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D45BFFECFD9CDCDE453D78D8" box="[554,632,202,228]" name="March" pageId="14" pageNumber="67">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940823" collectingDate="2009-03-27" collectionCode="BPBM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="14" pageNumber="67" specimenCode="BPBM 33772" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[644,806,202,228]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="67" type="Museum">BPBM 33772</specimenCode>
, same data as holotype except collected 30 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D45BFFECFEE5DCF946E4793B" box="[339,417,237,263]" name="March" pageId="14" pageNumber="67">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608939800" collectingDate="2009-03-27" collectionCode="PNGNM" collectorName="F. Kraus &amp; J. Anamiato &amp; Muller Range" country="Papua New Guinea" elevation="2177" latitude="-5.66954" location="March" longLatPrecision="1" longitude="142.62334" pageId="14" pageNumber="67" specimenCode="PNGNM 24099" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[429,624,237,263]" collectionCode="PNGNM" pageId="14" pageNumber="67">PNGNM 24099</specimenCode>
, same data as holotype except collected 31 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D45BFFECFF3ADD04479F7916" box="[140,218,272,298]" name="March" pageId="14" pageNumber="67">March</location>
</materialsCitation>
; 
<materialsCitation ID-GBIF-Occurrence="2608940815" collectingDate="2009-04-03" collectionCode="BPBM, PNGNM" country="Papua New Guinea" elevation="1874" latitude="-5.64509" location="Mt. Paramo" longLatPrecision="1" longitude="142.63618" municipality="Muller Range" pageId="14" pageNumber="67" specimenCode="BPBM 33773-79, PNGNM 24100" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
<specimenCode box="[229,436,272,299]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="67" type="Museum">BPBM 33773–79</specimenCode>
, 
<specimenCode box="[447,641,272,298]" collectionCode="PNGNM" pageId="14" pageNumber="67">PNGNM 24100</specimenCode>
, 
<location LSID="urn:lsid:plazi:treatment:AF1DA83BD458FFF0FF3AD91845017A6F:226B4FF6D45BFFECFD3BDD05445C7917" box="[653,793,273,299]" country="Papua New Guinea" latitude="-5.64509" longLatPrecision="1" longitude="142.63618" municipality="Muller Range" name="Mt. Paramo" pageId="14" pageNumber="67" stateProvince="Southern Highlands">Mt. Paramo</location>
, 
<collectingMunicipality box="[804,962,272,299]" pageId="14" pageNumber="67">Muller Range</collectingMunicipality>
, 
<geoCoordinate box="[974,1103,272,299]" degrees="5.64509" direction="south" orientation="latitude" pageId="14" pageNumber="67" precision="1" value="-5.64509">5.64509° S</geoCoordinate>
, 
<geoCoordinate box="[140,301,307,334]" degrees="142.63618" direction="east" orientation="longitude" pageId="14" pageNumber="67" precision="1" value="142.63618">142.63618° E</geoCoordinate>
, 
<quantity box="[312,404,308,334]" metricMagnitude="3" metricUnit="m" metricValue="1.874" pageId="14" pageNumber="67" unit="m" value="1874.0">
<elevation box="[312,404,308,334]" metricMagnitude="3" metricUnit="m" metricValue="1.874" pageId="14" pageNumber="67" unit="m" value="1874.0">1874 m</elevation>
</quantity>
, 
<date box="[416,565,307,334]" pageId="14" pageNumber="67" value="2009-04-03">
<collectingDate box="[416,565,307,334]" pageId="14" pageNumber="67" value="2009-04-03">3 April 2009</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="67" type="diagnosis">
<paragraph blockId="14.[140,1108,167,1671]" pageId="14" pageNumber="67">
<emphasis bold="true" box="[185,312,342,368]" pageId="14" pageNumber="67">
<emphasis box="[185,307,342,368]" inLineHeading="true" pageId="14" pageNumber="67" reason="1">Diagnosis</emphasis>
.
</emphasis>
A medium-sized species of 
<taxonomicName box="[640,766,343,369]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="genus">
<emphasis box="[640,766,343,369]" italics="true" pageId="14" pageNumber="67">Oreophryne</emphasis>
</taxonomicName>
(adult SV = 
<quantity box="[924,1100,343,369]" metricMagnitude="-2" metricUnit="m" metricValue="2.655" metricValueMax="2.9699999999999998" metricValueMin="2.34" pageId="14" pageNumber="67" unit="mm" value="26.549999999999997" valueMax="29.7" valueMin="23.4">23.4–29.7 mm</quantity>
) distinguished by its combination of a cartilaginous connection of the procoracoid to the scapula, no webbing between the toes, fifth toe longer than the third, relatively short snout (EN/SV = 
<geoCoordinate box="[412,479,448,475]" degrees="0.073" direction="north" orientation="latitude" pageId="14" pageNumber="67" precision="55" value="0.073">0.073</geoCoordinate>
<geoCoordinate box="[479,562,448,475]" degrees="0.086" direction="west" orientation="longitude" pageId="14" pageNumber="67" precision="55" value="-0.086">–0.086</geoCoordinate>
), dark face, venter with a dense array of darkbrown flecks, and call consisting of an extended multi-note chuckle or cackle.
</paragraph>
<paragraph blockId="14.[140,1108,167,1671]" pageId="14" pageNumber="67">
<emphasis bold="true" box="[185,580,518,545]" pageId="14" pageNumber="67">
<emphasis box="[185,344,518,544]" inLineHeading="true" pageId="14" pageNumber="67" reason="1">Comparisons</emphasis>
with other species.
</emphasis>
The new species differs from all Papuan congeners except 
<taxonomicName authority="Zweifel, Cogger, and Richards" authorityName="Zweifel, Cogger, and Richards" box="[303,785,554,580]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="alticola">
<emphasis box="[303,421,554,580]" italics="true" pageId="14" pageNumber="67">O. alticola</emphasis>
Zweifel, Cogger, and Richards
</taxonomicName>
, 
<taxonomicName authority="Gunther" authorityName="Gunther" authorityYear="2003" box="[800,1064,554,580]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="asplenicola">
<emphasis box="[800,955,554,580]" italics="true" pageId="14" pageNumber="67">O. asplenicola</emphasis>
Günther
</taxonomicName>
, 
<taxonomicName authority="Zweifel" authorityName="Zweifel" authorityYear="1956" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="brevicrus">
<emphasis italics="true" pageId="14" pageNumber="67">O. brevicrus</emphasis>
Zweifel
</taxonomicName>
, 
<taxonomicName authority="Gunther" authorityName="Gunther" authorityYear="2003" box="[349,584,589,616]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="clamata">
<emphasis box="[349,474,589,616]" italics="true" pageId="14" pageNumber="67">O. clamata</emphasis>
Günther
</taxonomicName>
, 
<taxonomicName authority="(van Kampen)" baseAuthorityName="van Kampen" box="[599,912,589,616]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="crucifera">
<emphasis box="[599,732,589,616]" italics="true" pageId="14" pageNumber="67">O. crucifera</emphasis>
(van Kampen)
</taxonomicName>
, 
<taxonomicName authority="Parker" authorityName="Parker" authorityYear="1934" box="[927,1103,589,616]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="flava">
<emphasis box="[927,1019,589,616]" italics="true" pageId="14" pageNumber="67">O. flava</emphasis>
Parker
</taxonomicName>
, 
<taxonomicName authority="Zweifel, Cogger, and Richards" authorityName="Zweifel, Cogger, and Richards" box="[140,671,624,651]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="habbemensis">
<emphasis box="[140,312,624,651]" italics="true" pageId="14" pageNumber="67">O. habbemensis</emphasis>
Zweifel, Cogger, and Richards
</taxonomicName>
, 
<taxonomicName authority="Zweifel" authorityName="Zweifel" authorityYear="1956" box="[684,963,624,651]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="idenburgensis">
<emphasis box="[684,867,624,651]" italics="true" pageId="14" pageNumber="67">O. idenburgensis</emphasis>
Zweifel
</taxonomicName>
, 
<taxonomicName authority="Parker" authorityName="Parker" authorityYear="1934" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="kampeni">
<emphasis box="[976,1108,624,651]" italics="true" pageId="14" pageNumber="67">O. kampeni</emphasis>
Parker
</taxonomicName>
, 
<taxonomicName authorityName="Zweifel" authorityYear="2003" box="[226,333,660,686]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="notata">
<emphasis box="[226,333,660,686]" italics="true" pageId="14" pageNumber="67">O. notata</emphasis>
</taxonomicName>
, 
<taxonomicName authority="Gunther" authorityName="Gunther" authorityYear="2003" box="[350,671,659,686]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="pseudasplenicola">
<emphasis box="[350,564,659,686]" italics="true" pageId="14" pageNumber="67">O. pseudasplenicola</emphasis>
Günther
</taxonomicName>
, and 
<taxonomicName authority="Gunther" authorityName="Gunther" authorityYear="2003" box="[736,942,659,686]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="waira">
<emphasis box="[736,837,660,686]" italics="true" pageId="14" pageNumber="67">O. waira</emphasis>
Günther
</taxonomicName>
in having (vs. lacking) a procoracoid that reaches the scapula. It differs from 
<taxonomicName box="[865,996,695,721]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="crucifera">
<emphasis box="[865,996,695,721]" italics="true" pageId="14" pageNumber="67">O. crucifera</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Zweifel" authorityYear="1956" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="idenburgensis">
<emphasis italics="true" pageId="14" pageNumber="67">O. idenburgensis</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Parker" authorityYear="1934" box="[252,382,730,756]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="kampeni">
<emphasis box="[252,382,730,756]" italics="true" pageId="14" pageNumber="67">O. kampeni</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[446,545,730,756]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="waira">
<emphasis box="[446,545,730,756]" italics="true" pageId="14" pageNumber="67">O. waira</emphasis>
</taxonomicName>
in lacking (vs. having) webbing between the toes; and it differs from 
<taxonomicName box="[363,480,765,792]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="alticola">
<emphasis box="[363,480,765,792]" italics="true" pageId="14" pageNumber="67">O. alticola</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Zweifel" authorityYear="1956" box="[497,632,765,792]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="brevicrus">
<emphasis box="[497,632,765,792]" italics="true" pageId="14" pageNumber="67">O. brevicrus</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[649,774,765,792]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="clamata">
<emphasis box="[649,774,765,792]" italics="true" pageId="14" pageNumber="67">O. clamata</emphasis>
</taxonomicName>
, and 
<taxonomicName box="[843,1016,765,792]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="habbemensis">
<emphasis box="[843,1016,765,792]" italics="true" pageId="14" pageNumber="67">O. habbemensis</emphasis>
</taxonomicName>
in having the fifth toe longer than (vs. shorter than or subequal to) the third. 
<taxonomicName authorityName="Kraus &amp; Allison" authorityYear="2009" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="anamiatoi">
<emphasis italics="true" pageId="14" pageNumber="67">Oreophryne anamiatoi</emphasis>
</taxonomicName>
differs from the remaining species 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[661,815,835,862]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="asplenicola">
<emphasis box="[661,815,835,862]" italics="true" pageId="14" pageNumber="67">O. asplenicola</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Parker" authorityYear="1934" box="[833,924,835,862]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="flava">
<emphasis box="[833,924,835,862]" italics="true" pageId="14" pageNumber="67">O. flava</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Zweifel" authorityYear="2003" box="[941,1048,836,862]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="notata">
<emphasis box="[941,1048,836,862]" italics="true" pageId="14" pageNumber="67">O. notata</emphasis>
</taxonomicName>
, and 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[140,353,871,897]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="pseudasplenicola">
<emphasis box="[140,353,871,897]" italics="true" pageId="14" pageNumber="67">O. pseudasplenicola</emphasis>
</taxonomicName>
in its larger size (
<quantity box="[552,725,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="2.655" metricValueMax="2.9699999999999998" metricValueMin="2.34" pageId="14" pageNumber="67" unit="mm" value="26.549999999999997" valueMax="29.7" valueMin="23.4">23.4–29.7 mm</quantity>
vs. maximum of 
<quantity box="[925,1009,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="67" unit="mm" value="21.0">21 mm</quantity>
in those four species) in having a uniformly dark face, and in its call (call a series of peeps in the other species, but call unknown in 
<taxonomicName authorityName="Parker" authorityYear="1934" box="[585,676,941,968]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="flava">
<emphasis box="[585,676,941,968]" italics="true" pageId="14" pageNumber="67">O. flava</emphasis>
</taxonomicName>
); it further differs from 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[954,1107,941,968]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="asplenicola">
<emphasis box="[954,1107,941,968]" italics="true" pageId="14" pageNumber="67">O. asplenicola</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[189,400,976,1003]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="pseudasplenicola">
<emphasis box="[189,400,976,1003]" italics="true" pageId="14" pageNumber="67">O. pseudasplenicola</emphasis>
</taxonomicName>
in its shorter snout (EN/SV&gt; 
<quantity box="[747,843,976,1002]" metricMagnitude="-3" metricUnit="m" metricValue="2.1844" pageId="14" pageNumber="67" unit="in" value="0.086">0.086 in</quantity>
those species), from 
<taxonomicName authorityName="Zweifel" authorityYear="2003" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="notata">
<emphasis italics="true" pageId="14" pageNumber="67">O. notata</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Gunther" authorityYear="2003" box="[266,478,1011,1038]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="pseudasplenicola">
<emphasis box="[266,478,1011,1038]" italics="true" pageId="14" pageNumber="67">O. pseudasplenicola</emphasis>
</taxonomicName>
in lacking (vs. having) an inverted white U on the face, and from 
<taxonomicName authorityName="Parker" authorityYear="1934" box="[256,347,1047,1073]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="14" pageNumber="67" phylum="Chordata" rank="species" species="flava">
<emphasis box="[256,347,1047,1073]" italics="true" pageId="14" pageNumber="67">O. flava</emphasis>
</taxonomicName>
in having an abdomen with dark-brown flecking (vs. immaculate yellow-white), and lacking (vs. having) a dark W-shaped mark between the shoulders.
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="70" pageId="14" pageNumber="67" type="description">
<paragraph blockId="14.[140,1108,167,1671]" lastBlockId="15.[140,1108,167,1038]" lastPageId="15" lastPageNumber="68" pageId="14" pageNumber="67">
<emphasis bold="true" box="[185,488,1117,1143]" pageId="14" pageNumber="67">
<emphasis box="[185,331,1117,1143]" inLineHeading="true" pageId="14" pageNumber="67" reason="1">Description</emphasis>
of 
<typeStatus box="[372,483,1117,1143]" pageId="14" pageNumber="67">holotype</typeStatus>
.
</emphasis>
Adult female with incision on right side. Head wide (HW/SV = 0.40), with steep, slightly concave loreal region. Canthus rostralis round- ed, concave when viewed from above. Nostrils directed laterally, closer to tip of snout than to eyes. Internarial distance broader than distance from naris to eye (EN/IN = 0.80, IN/SV = 0.101, EN/SV = 0.081). Snout truncate when viewed from the side, shallowly angulate when viewed from above. Eyes moderately large (EY/SV = 0.11); eyelid approximately two-thirds width of interorbital distance. Tympanum distinct but small (TY/SV = 0.047). Dorsal skin granular with series of weakly raised parallel ridges and scattered small pustules; ventral surfaces coarsely granular. Supratympanic fold narrow. Fingers unwebbed, bearing discs with terminal grooves; relative lengths 3&gt;4&gt;2&gt;1. Finger discs approximately 3 times widths of penultimate phalanges, except for first finger, which is approximately twice width of penultimate phalanx. Subarticular tubercles well developed; inner metacarpal tubercle oval and low; outer rounded and obscure. Toes unwebbed, bearing discs with terminal grooves; relative lengths 4&gt;5&gt;3&gt;2&gt;1. Toe discs smaller than those of fingers (3 
<superScript attach="right" box="[860,874,1609,1623]" fontSize="6" pageId="14" pageNumber="67">rdF</superScript>
/4thT = 1.37), approximately twice width of penultimate phalanges. Subarticular tubercles low but distinct; inner metatarsal tubercle large, oval; outer absent. Hind legs of moderate length (TL/SV = 0.46).
</paragraph>
<paragraph blockId="15.[140,1108,167,1038]" pageId="15" pageNumber="68">In preservative, dorsum medium brown with small black flecks scattered throughout, these concentrated dorsolaterally, mid-dorsally, and on top of head. Entire face to posterior of jaw angle uniform dark brown. Dark-brown postocular stripe extends along ventral side of supratympanic ridge. Rear and front of thighs uniform medium brown. Tops of wrist, hand, and first three fingers boldly marked with dark brown. Sides darker brown with very pale straw flecks. Venter very pale straw with bold, large, dark-brown flecks, these dense on chin, throat, and under legs, sparse on abdomen. Palmar and plantar surfaces dark brown spotted with very pale straw. Iris dark brown minutely flecked with silver.</paragraph>
<paragraph blockId="15.[140,1108,167,1038]" pageId="15" pageNumber="68">
<emphasis bold="true" box="[185,304,553,579]" pageId="15" pageNumber="68">
<emphasis box="[185,298,553,579]" inLineHeading="true" pageId="15" pageNumber="68" reason="1">Variation</emphasis>
.
</emphasis>
Mensural variation for the 
<typeStatus box="[625,673,554,580]" pageId="15" pageNumber="68">type</typeStatus>
series is shown in 
<tableCitation box="[891,979,554,581]" captionStart="Table 5" captionStartId="15.[140,200,1082,1104]" captionText="Table 5. Mensural data for type series of Oreophryne anamiatoi sp. n. Data include only adult animals." pageId="15" pageNumber="68">Table 5</tableCitation>
. Sexual dimorphism is slight except that females clearly attain larger size than males and may have slightly narrower snouts (reflected in EN/IN values).
</paragraph>
<paragraph blockId="15.[140,1108,167,1038]" pageId="15" pageNumber="68">Dorsal ground color varies from light brown to dark brown, and pattern varies from virtually absent to moderately well stippled and streaked with black. Black stippling may be uniformly distributed, or concentrated laterally or dorsolaterally. Two specimens have a narrow tan vertebral stripe margined with black stippling. The dark face and postocular bar are present in all. Venters of all specimens are boldly spotted with dark brown on white, but two specimens have the abdomen clear and a few others have it less spotted than the chin and throat.</paragraph>
<paragraph blockId="15.[140,1108,167,1038]" lastBlockId="16.[140,1108,166,1038]" lastPageId="16" lastPageNumber="69" pageId="15" pageNumber="68">
<emphasis bold="true" box="[185,346,905,931]" pageId="15" pageNumber="68">
<emphasis box="[185,255,905,931]" inLineHeading="true" pageId="15" pageNumber="68" reason="1">Color</emphasis>
in life.
</emphasis>
BPBM 33764 (Fig. 1C): “Dorsum medium brown with a slight russet cast. Face dark brown; short dark-brown postocular dash; and few dark-brown lateral flecks. Rear of thighs brown, slightly darker than dorsum. Venter pale yellow heavily flecked with dark gray. Iris bronze with narrow red rim around pupil. Upper arms and tarsi burnt orange.” BPBM 33765 was uniform brown dorsally with a dark face mask and postocular stripe (Fig. 1D); rear and front of thighs, and groin, uniform brown, slightly darker than dorsum; venter dirty cream flecked with gray; iris dark brown. BPBM 33766 had black flecks on sides and venter brighter yellow but with fewer gray flecks and with white chromatophores; BPBM 33767 had the dorsum and rear of thighs burnt orange and venter pale yellow; BPBM 33769 and 33771 had a tan vertebral line. BPBM 33774 had a metallic green sheen on eyelids and top of snout; BPBM 33776 same but less extensive. PNGNM 24097 was light brown with straw yellow on sides, with dark-brown spots dorsally and laterally, denser on sides; face to rictus and short postocular stripe dark brown; rear of thighs brown like dorsum but unspotted; venter pale yellow spotted with dark-gray flecks; iris brown.
</paragraph>
<caption box="[140,1094,1082,1105]" pageId="15" pageNumber="68">
<paragraph blockId="15.[140,1094,1082,1105]" box="[140,1094,1082,1105]" pageId="15" pageNumber="68">
<emphasis bold="true" box="[140,227,1082,1104]" pageId="15" pageNumber="68">Table 5.</emphasis>
Mensural data for type series of 
<taxonomicName authority="Kraus &amp; Allison, 2009" authorityName="Kraus &amp; Allison" authorityYear="2009" box="[528,728,1083,1104]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="15" pageNumber="68" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
<emphasis box="[528,728,1083,1104]" italics="true" pageId="15" pageNumber="68">
Oreophryne 
<taxonomicName box="[637,728,1083,1104]" pageId="15" pageNumber="68" rank="species" species="anamiatoi" status="sp. n.">anamiatoi</taxonomicName>
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[734,785,1083,1104]" pageId="15" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
Data include only adult animals.
</paragraph>
</caption>
<paragraph pageId="15" pageNumber="68">
<table box="[182,1079,1127,1669]" gridcols="5" gridrows="15" pageId="15" pageNumber="68">
<tr box="[182,1079,1127,1186]" gridrow="0" pageId="15" pageNumber="68">
<th box="[182,306,1127,1186]" gridcol="0" gridrow="0" pageId="15" pageNumber="68">Character</th>
<th box="[413,699,1127,1186]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="15" pageNumber="68">Males (n = 11) mean range</th>
<th box="[793,1079,1127,1186]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="15" pageNumber="68">Females (n = 4) Mean range</th>
</tr>
<tr box="[182,1079,1196,1221]" gridrow="1" pageId="15" pageNumber="68">
<th box="[182,306,1196,1221]" gridcol="0" gridrow="1" pageId="15" pageNumber="68">SV (mm)</th>
<td box="[413,473,1196,1221]" gridcol="1" gridrow="1" pageId="15" pageNumber="68">25.2</td>
<td box="[566,699,1196,1221]" gridcol="2" gridrow="1" pageId="15" pageNumber="68">23.4–26.7</td>
<td box="[793,853,1196,1221]" gridcol="3" gridrow="1" pageId="15" pageNumber="68">29.0</td>
<td box="[946,1079,1196,1221]" gridcol="4" gridrow="1" pageId="15" pageNumber="68">27.4–29.7</td>
</tr>
<tr box="[182,1079,1231,1255]" gridrow="2" pageId="15" pageNumber="68">
<th box="[182,306,1231,1255]" gridcol="0" gridrow="2" pageId="15" pageNumber="68">TL/SV</th>
<td box="[413,473,1231,1255]" gridcol="1" gridrow="2" pageId="15" pageNumber="68">0.45</td>
<td box="[566,699,1231,1255]" gridcol="2" gridrow="2" pageId="15" pageNumber="68">0.43–0.49</td>
<td box="[793,853,1231,1255]" gridcol="3" gridrow="2" pageId="15" pageNumber="68">0.47</td>
<td box="[946,1079,1231,1255]" gridcol="4" gridrow="2" pageId="15" pageNumber="68">0.45–0.48</td>
</tr>
<tr box="[182,1079,1265,1290]" gridrow="3" pageId="15" pageNumber="68">
<th box="[182,306,1265,1290]" gridcol="0" gridrow="3" pageId="15" pageNumber="68">EN/SV</th>
<td box="[413,473,1265,1290]" gridcol="1" gridrow="3" pageId="15" pageNumber="68">0.077</td>
<td box="[566,699,1265,1290]" gridcol="2" gridrow="3" pageId="15" pageNumber="68">0.073–0.082</td>
<td box="[793,853,1265,1290]" gridcol="3" gridrow="3" pageId="15" pageNumber="68">0.079</td>
<td box="[946,1079,1265,1290]" gridcol="4" gridrow="3" pageId="15" pageNumber="68">0.075–0.084</td>
</tr>
<tr box="[182,1079,1300,1324]" gridrow="4" pageId="15" pageNumber="68">
<th box="[182,306,1300,1324]" gridcol="0" gridrow="4" pageId="15" pageNumber="68">IN/SV</th>
<td box="[413,473,1300,1324]" gridcol="1" gridrow="4" pageId="15" pageNumber="68">0.103</td>
<td box="[566,699,1300,1324]" gridcol="2" gridrow="4" pageId="15" pageNumber="68">0.100–0.107</td>
<td box="[793,853,1300,1324]" gridcol="3" gridrow="4" pageId="15" pageNumber="68">0.100</td>
<td box="[946,1079,1300,1324]" gridcol="4" gridrow="4" pageId="15" pageNumber="68">0.098–0.102</td>
</tr>
<tr box="[182,1079,1334,1358]" gridrow="5" pageId="15" pageNumber="68">
<th box="[182,306,1334,1358]" gridcol="0" gridrow="5" pageId="15" pageNumber="68">SN/SV</th>
<td box="[413,473,1334,1358]" gridcol="1" gridrow="5" pageId="15" pageNumber="68">0.14</td>
<td box="[566,699,1334,1358]" gridcol="2" gridrow="5" pageId="15" pageNumber="68">0.13–0.15</td>
<td box="[793,853,1334,1358]" gridcol="3" gridrow="5" pageId="15" pageNumber="68">0.13</td>
<td box="[946,1079,1334,1358]" gridcol="4" gridrow="5" pageId="15" pageNumber="68">0.13–0.14</td>
</tr>
<tr box="[182,1079,1369,1393]" gridrow="6" pageId="15" pageNumber="68">
<th box="[182,306,1369,1393]" gridcol="0" gridrow="6" pageId="15" pageNumber="68">TY/SV</th>
<td box="[413,473,1369,1393]" gridcol="1" gridrow="6" pageId="15" pageNumber="68">0.045</td>
<td box="[566,699,1369,1393]" gridcol="2" gridrow="6" pageId="15" pageNumber="68">0.039–0.056</td>
<td box="[793,853,1369,1393]" gridcol="3" gridrow="6" pageId="15" pageNumber="68">0.049</td>
<td box="[946,1079,1369,1393]" gridcol="4" gridrow="6" pageId="15" pageNumber="68">0.047–0.051</td>
</tr>
<tr box="[182,1079,1403,1427]" gridrow="7" pageId="15" pageNumber="68">
<th box="[182,306,1403,1427]" gridcol="0" gridrow="7" pageId="15" pageNumber="68">EY/SV</th>
<td box="[413,473,1403,1427]" gridcol="1" gridrow="7" pageId="15" pageNumber="68">0.12</td>
<td box="[566,699,1403,1427]" gridcol="2" gridrow="7" pageId="15" pageNumber="68">0.11–0.13</td>
<td box="[793,853,1403,1427]" gridcol="3" gridrow="7" pageId="15" pageNumber="68">0.12</td>
<td box="[946,1079,1403,1427]" gridcol="4" gridrow="7" pageId="15" pageNumber="68">0.11–0.12</td>
</tr>
<tr box="[182,1079,1438,1462]" gridrow="8" pageId="15" pageNumber="68">
<th box="[182,306,1438,1462]" gridcol="0" gridrow="8" pageId="15" pageNumber="68">HW/SV</th>
<td box="[413,473,1438,1462]" gridcol="1" gridrow="8" pageId="15" pageNumber="68">0.39</td>
<td box="[566,699,1438,1462]" gridcol="2" gridrow="8" pageId="15" pageNumber="68">0.38–0.40</td>
<td box="[793,853,1438,1462]" gridcol="3" gridrow="8" pageId="15" pageNumber="68">0.39</td>
<td box="[946,1079,1438,1462]" gridcol="4" gridrow="8" pageId="15" pageNumber="68">0.38–0.40</td>
</tr>
<tr box="[182,1079,1472,1496]" gridrow="9" pageId="15" pageNumber="68">
<th box="[182,306,1472,1496]" gridcol="0" gridrow="9" pageId="15" pageNumber="68">HL/SV</th>
<td box="[413,473,1472,1496]" gridcol="1" gridrow="9" pageId="15" pageNumber="68">0.33</td>
<td box="[566,699,1472,1496]" gridcol="2" gridrow="9" pageId="15" pageNumber="68">0.32–0.35</td>
<td box="[793,853,1472,1496]" gridcol="3" gridrow="9" pageId="15" pageNumber="68">0.33</td>
<td box="[946,1079,1472,1496]" gridcol="4" gridrow="9" pageId="15" pageNumber="68">0.32–0.34</td>
</tr>
<tr box="[182,1079,1506,1531]" gridrow="10" pageId="15" pageNumber="68">
<th box="[182,306,1506,1531]" gridcol="0" gridrow="10" pageId="15" pageNumber="68">3rdF/SV</th>
<td box="[413,473,1506,1531]" gridcol="1" gridrow="10" pageId="15" pageNumber="68">0.071</td>
<td box="[566,699,1506,1531]" gridcol="2" gridrow="10" pageId="15" pageNumber="68">0.062–0.083</td>
<td box="[793,853,1506,1531]" gridcol="3" gridrow="10" pageId="15" pageNumber="68">0.074</td>
<td box="[946,1079,1506,1531]" gridcol="4" gridrow="10" pageId="15" pageNumber="68">0.065–0.082</td>
</tr>
<tr box="[182,1079,1541,1565]" gridrow="11" pageId="15" pageNumber="68">
<th box="[182,306,1541,1565]" gridcol="0" gridrow="11" pageId="15" pageNumber="68">4thT/SV</th>
<td box="[413,473,1541,1565]" gridcol="1" gridrow="11" pageId="15" pageNumber="68">0.056</td>
<td box="[566,699,1541,1565]" gridcol="2" gridrow="11" pageId="15" pageNumber="68">0.050–0.062</td>
<td box="[793,853,1541,1565]" gridcol="3" gridrow="11" pageId="15" pageNumber="68">0.058</td>
<td box="[946,1079,1541,1565]" gridcol="4" gridrow="11" pageId="15" pageNumber="68">0.055–0.062</td>
</tr>
<tr box="[182,1079,1575,1600]" gridrow="12" pageId="15" pageNumber="68">
<th box="[182,306,1575,1600]" gridcol="0" gridrow="12" pageId="15" pageNumber="68">EN/IN</th>
<td box="[413,473,1575,1600]" gridcol="1" gridrow="12" pageId="15" pageNumber="68">0.74</td>
<td box="[566,699,1575,1600]" gridcol="2" gridrow="12" pageId="15" pageNumber="68">0.72–0.77</td>
<td box="[793,853,1575,1600]" gridcol="3" gridrow="12" pageId="15" pageNumber="68">0.79</td>
<td box="[946,1079,1575,1600]" gridcol="4" gridrow="12" pageId="15" pageNumber="68">0.75–0.86</td>
</tr>
<tr box="[182,1079,1610,1634]" gridrow="13" pageId="15" pageNumber="68">
<th box="[182,306,1610,1634]" gridcol="0" gridrow="13" pageId="15" pageNumber="68">3rd F/4th T</th>
<td box="[413,473,1610,1634]" gridcol="1" gridrow="13" pageId="15" pageNumber="68">1.28</td>
<td box="[566,699,1610,1634]" gridcol="2" gridrow="13" pageId="15" pageNumber="68">1.20–1.40</td>
<td box="[793,853,1610,1634]" gridcol="3" gridrow="13" pageId="15" pageNumber="68">1.27</td>
<td box="[946,1079,1610,1634]" gridcol="4" gridrow="13" pageId="15" pageNumber="68">1.18–1.37</td>
</tr>
<tr box="[182,1079,1644,1669]" gridrow="14" pageId="15" pageNumber="68">
<th box="[182,306,1644,1669]" gridcol="0" gridrow="14" pageId="15" pageNumber="68">HL/HW</th>
<td box="[413,473,1644,1669]" gridcol="1" gridrow="14" pageId="15" pageNumber="68">0.85</td>
<td box="[566,699,1644,1669]" gridcol="2" gridrow="14" pageId="15" pageNumber="68">0.81–0.89</td>
<td box="[793,853,1644,1669]" gridcol="3" gridrow="14" pageId="15" pageNumber="68">0.85</td>
<td box="[946,1079,1644,1669]" gridcol="4" gridrow="14" pageId="15" pageNumber="68">0.82–0.86</td>
</tr>
</table>
</paragraph>
<paragraph blockId="16.[140,1108,166,1038]" pageId="16" pageNumber="69">
<emphasis bold="true" box="[185,242,553,579]" pageId="16" pageNumber="69">
<emphasis box="[185,237,553,579]" inLineHeading="true" pageId="16" pageNumber="69" reason="1">Call</emphasis>
.
</emphasis>
Animals called during the first few hours of darkness. We recorded ten calls from two individuals (
<tableCitation box="[390,477,589,616]" captionStart="Table 6" captionStartId="16.[140,200,1095,1117]" captionText="Table 6. Call statistics of Oreophryne anamiatoi sp. n. recorded on Mt. Paramo, Muller Range, Southern Highlands Province, Papua New Guinea on 3 April 2009. BPBM 33774 was recorded at 2000 h and BPBM 33775 at 2100 h. Air temperatures were 17.0 and 17.7 °C, respectively." pageId="16" pageNumber="69">Table 6</tableCitation>
). Th e calls from both were similar, so we combined data from both for analysis.
</paragraph>
<paragraph blockId="16.[140,1108,166,1038]" pageId="16" pageNumber="69">
The call consists of 17–22 pulsed notes (
<figureCitation box="[654,755,659,686]" captionStart="Figure 8" captionStartId="17.[140,212,578,600]" captionTargetBox="[142,1068,170,497]" captionTargetId="figure@17.[140,1108,165,559]" captionTargetPageId="17" captionText="Figure 8. A Waveform, B power spectrum, and C spectrogram of Call “D” of Oreophryne anamiatoi sp. n. (BPBM 33774) recorded at Mt. Paramo, Muller Range, Southern Highlands Province, Papua New Guinea on 3 April 2009 at 2000 h. Air temperature 17.7 °C." figureDoi="http://doi.org/10.5281/zenodo.3769065" httpUri="https://zenodo.org/record/3769065/files/figure.png" pageId="16" pageNumber="69">Figs. 8A</figureCitation>
, 
<figureCitation box="[766,805,660,686]" captionStart="Figure 9" captionStartId="17.[140,212,1161,1183]" captionTargetBox="[175,1085,739,1077]" captionTargetId="figure@17.[140,1108,694,1142]" captionTargetPageId="17" captionText="Figure 9. Detail of the middle three notes of call “D” of Oreophryne anamiatoi sp. n. (BPBM 33774). A Waveform, B power spectrum, and C spectrogram. Note the pulsing of the notes." figureDoi="http://doi.org/10.5281/zenodo.3769067" httpUri="https://zenodo.org/record/3769067/files/figure.png" pageId="16" pageNumber="69">9A</figureCitation>
). Note duration was similar over the course of the call and averaged 
<geoCoordinate box="[644,745,694,721]" degrees="0.0569" direction="south" orientation="latitude" pageId="16" pageNumber="69" precision="5" value="-0.0569">0.0569 s</geoCoordinate>
(range 
<geoCoordinate box="[834,921,695,722]" degrees="0.0266" direction="north" orientation="latitude" pageId="16" pageNumber="69" precision="5" value="0.0266">0.0266</geoCoordinate>
<geoCoordinate box="[920,1021,695,722]" degrees="0.0681" direction="west" orientation="longitude" pageId="16" pageNumber="69" precision="5" value="-0.0681">–0.0681</geoCoordinate>
). Internote intervals were also similar throughout the call and were similar in duration to the notes, averaging 
<geoCoordinate box="[422,525,765,792]" degrees="0.0563" direction="south" orientation="latitude" pageId="16" pageNumber="69" precision="5" value="-0.0563">0.0563 s</geoCoordinate>
(range 
<geoCoordinate box="[619,705,765,791]" degrees="0.0390" direction="north" orientation="latitude" pageId="16" pageNumber="69" precision="5" value="0.039">0.0390</geoCoordinate>
<geoCoordinate box="[704,806,765,791]" degrees="0.0942" direction="west" orientation="longitude" pageId="16" pageNumber="69" precision="5" value="-0.0942">–0.0942</geoCoordinate>
). Mean call duration was 
<geoCoordinate box="[140,202,800,826]" degrees="2.10" direction="south" orientation="latitude" pageId="16" pageNumber="69" precision="555" value="-2.1">2.10s</geoCoordinate>
(range 1.77–2.33), and notes were delivered at a mean repetition rate of 18.5 notes/s (range 17.3–20.3). Number of pulses/note ranged from 3–11 (
<figureCitation captionStart="Figure 9" captionStartId="17.[140,212,1161,1183]" captionTargetBox="[175,1085,739,1077]" captionTargetId="figure@17.[140,1108,694,1142]" captionTargetPageId="17" captionText="Figure 9. Detail of the middle three notes of call “D” of Oreophryne anamiatoi sp. n. (BPBM 33774). A Waveform, B power spectrum, and C spectrogram. Note the pulsing of the notes." figureDoi="http://doi.org/10.5281/zenodo.3769067" httpUri="https://zenodo.org/record/3769067/files/figure.png" pageId="16" pageNumber="69">Fig. 9A</figureCitation>
), with a strong tendency to decrease over the course of the call (Fig 10). Modal number of pulses in the first three notes in each call ranged from 10–11, producing a mean rate for those notes of 200.9 pulses/s, while modal number of pulses in the last three notes ranged from 7–9, producing a mean rate for those notes of 146.4 pulses/s (
<tableCitation box="[244,335,1011,1038]" captionStart="Table 7" captionStartId="18.[140,200,846,868]" captionText="Table 7. Pulse rates of the calls of Oreophryne anamiatoi sp. n. See Table 6 for additional details." pageId="16" pageNumber="69">Table 7</tableCitation>
). Notes are finely tuned, with a mean dominant frequency of 2490
</paragraph>
<caption pageId="16" pageNumber="69">
<paragraph blockId="16.[140,1108,1095,1181]" pageId="16" pageNumber="69">
<emphasis bold="true" box="[140,227,1095,1117]" pageId="16" pageNumber="69">Table 6.</emphasis>
Call statistics of 
<taxonomicName authority="Kraus &amp; Allison, 2009" authorityName="Kraus &amp; Allison" authorityYear="2009" box="[383,583,1095,1117]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="16" pageNumber="69" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
<emphasis box="[383,583,1095,1117]" italics="true" pageId="16" pageNumber="69">Oreophryne anamiatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[589,640,1095,1116]" pageId="16" pageNumber="69" rank="species">sp. n.</taxonomicNameLabel>
recorded on Mt. Paramo, Muller Range, Southern Highlands Province, Papua New Guinea on 3 April 2009. BPBM 33774 was recorded at 2000 h and BPBM 33775 at 2100 h. Air temperatures were 17.0 and 17.7 °C, respectively.
</paragraph>
</caption>
<paragraph pageId="16" pageNumber="69">
<table box="[153,1097,1204,1669]" gridcols="9" gridrows="11" pageId="16" pageNumber="69" rowsContinueIn="18.[262,1029,891,1359]">
<tr box="[153,1097,1204,1324]" gridrow="0" pageId="16" pageNumber="69">
<th box="[153,298,1204,1324]" gridcol="0" gridrow="0" pageId="16" pageNumber="69">Specimen</th>
<th box="[319,362,1204,1324]" gridcol="1" gridrow="0" pageId="16" pageNumber="69">Call</th>
<th box="[379,433,1204,1324]" gridcol="2" gridrow="0" pageId="16" pageNumber="69">Total notes</th>
<th box="[451,540,1204,1324]" gridcol="3" gridrow="0" pageId="16" pageNumber="69">Call duration (s)</th>
<th box="[560,637,1204,1324]" gridcol="4" gridrow="0" pageId="16" pageNumber="69">Notes/s</th>
<th box="[659,748,1204,1324]" gridcol="5" gridrow="0" pageId="16" pageNumber="69">Mean note duration (s)</th>
<th box="[770,864,1204,1324]" gridcol="6" gridrow="0" pageId="16" pageNumber="69">Mean internote duration (s)</th>
<th box="[885,970,1204,1324]" gridcol="7" gridrow="0" pageId="16" pageNumber="69">No. pulses per note</th>
<th box="[989,1097,1204,1324]" gridcol="8" gridrow="0" pageId="16" pageNumber="69">Dominant frequency (Hz)</th>
</tr>
<tr box="[153,1097,1334,1359]" gridrow="1" pageId="16" pageNumber="69">
<th box="[153,298,1334,1359]" gridcol="0" gridrow="1" pageId="16" pageNumber="69">BPBM 33774</th>
<td box="[319,362,1334,1359]" gridcol="1" gridrow="1" pageId="16" pageNumber="69">A</td>
<td box="[379,433,1334,1359]" gridcol="2" gridrow="1" pageId="16" pageNumber="69">19</td>
<td box="[451,540,1334,1359]" gridcol="3" gridrow="1" pageId="16" pageNumber="69">2.15</td>
<td box="[560,637,1334,1359]" gridcol="4" gridrow="1" pageId="16" pageNumber="69">17.27</td>
<td box="[659,748,1334,1359]" gridcol="5" gridrow="1" pageId="16" pageNumber="69">0.0611</td>
<td box="[770,864,1334,1359]" gridcol="6" gridrow="1" pageId="16" pageNumber="69">0.0551</td>
<td box="[885,970,1334,1359]" gridcol="7" gridrow="1" pageId="16" pageNumber="69">8–11</td>
<td box="[989,1097,1334,1359]" gridcol="8" gridrow="1" pageId="16" pageNumber="69">2540</td>
</tr>
<tr box="[153,1097,1369,1393]" gridrow="2" pageId="16" pageNumber="69">
<th box="[153,298,1369,1393]" gridcol="0" gridrow="2" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1369,1393]" gridcol="1" gridrow="2" pageId="16" pageNumber="69">B</td>
<td box="[379,433,1369,1393]" gridcol="2" gridrow="2" pageId="16" pageNumber="69">20</td>
<td box="[451,540,1369,1393]" gridcol="3" gridrow="2" pageId="16" pageNumber="69">2.33</td>
<td box="[560,637,1369,1393]" gridcol="4" gridrow="2" pageId="16" pageNumber="69">17.86</td>
<td box="[659,748,1369,1393]" gridcol="5" gridrow="2" pageId="16" pageNumber="69">0.0596</td>
<td box="[770,864,1369,1393]" gridcol="6" gridrow="2" pageId="16" pageNumber="69">0.0598</td>
<td box="[885,970,1369,1393]" gridcol="7" gridrow="2" pageId="16" pageNumber="69">8–12</td>
<td box="[989,1097,1369,1393]" gridcol="8" gridrow="2" pageId="16" pageNumber="69">2520</td>
</tr>
<tr box="[153,1097,1403,1427]" gridrow="3" pageId="16" pageNumber="69">
<th box="[153,298,1403,1427]" gridcol="0" gridrow="3" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1403,1427]" gridcol="1" gridrow="3" pageId="16" pageNumber="69">C</td>
<td box="[379,433,1403,1427]" gridcol="2" gridrow="3" pageId="16" pageNumber="69">19</td>
<td box="[451,540,1403,1427]" gridcol="3" gridrow="3" pageId="16" pageNumber="69">2.14</td>
<td box="[560,637,1403,1427]" gridcol="4" gridrow="3" pageId="16" pageNumber="69">17.67</td>
<td box="[659,748,1403,1427]" gridcol="5" gridrow="3" pageId="16" pageNumber="69">0.0596</td>
<td box="[770,864,1403,1427]" gridcol="6" gridrow="3" pageId="16" pageNumber="69">0.0560</td>
<td box="[885,970,1403,1427]" gridcol="7" gridrow="3" pageId="16" pageNumber="69">7–11</td>
<td box="[989,1097,1403,1427]" gridcol="8" gridrow="3" pageId="16" pageNumber="69">2520</td>
</tr>
<tr box="[153,1097,1437,1462]" gridrow="4" pageId="16" pageNumber="69">
<th box="[153,298,1437,1462]" gridcol="0" gridrow="4" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1437,1462]" gridcol="1" gridrow="4" pageId="16" pageNumber="69">D</td>
<td box="[379,433,1437,1462]" gridcol="2" gridrow="4" pageId="16" pageNumber="69">19</td>
<td box="[451,540,1437,1462]" gridcol="3" gridrow="4" pageId="16" pageNumber="69">2.13</td>
<td box="[560,637,1437,1462]" gridcol="4" gridrow="4" pageId="16" pageNumber="69">18.12</td>
<td box="[659,748,1437,1462]" gridcol="5" gridrow="4" pageId="16" pageNumber="69">0.0580</td>
<td box="[770,864,1437,1462]" gridcol="6" gridrow="4" pageId="16" pageNumber="69">0.0569</td>
<td box="[885,970,1437,1462]" gridcol="7" gridrow="4" pageId="16" pageNumber="69">7–11</td>
<td box="[989,1097,1437,1462]" gridcol="8" gridrow="4" pageId="16" pageNumber="69">2480</td>
</tr>
<tr box="[153,1097,1472,1496]" gridrow="5" pageId="16" pageNumber="69">
<th box="[153,298,1472,1496]" gridcol="0" gridrow="5" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1472,1496]" gridcol="1" gridrow="5" pageId="16" pageNumber="69">E</td>
<td box="[379,433,1472,1496]" gridcol="2" gridrow="5" pageId="16" pageNumber="69">16</td>
<td box="[451,540,1472,1496]" gridcol="3" gridrow="5" pageId="16" pageNumber="69">1.77</td>
<td box="[560,637,1472,1496]" gridcol="4" gridrow="5" pageId="16" pageNumber="69">17.75</td>
<td box="[659,748,1472,1496]" gridcol="5" gridrow="5" pageId="16" pageNumber="69">0.0572</td>
<td box="[770,864,1472,1496]" gridcol="6" gridrow="5" pageId="16" pageNumber="69">0.0574</td>
<td box="[885,970,1472,1496]" gridcol="7" gridrow="5" pageId="16" pageNumber="69">4–12</td>
<td box="[989,1097,1472,1496]" gridcol="8" gridrow="5" pageId="16" pageNumber="69">2480</td>
</tr>
<tr box="[153,1097,1506,1531]" gridrow="6" pageId="16" pageNumber="69">
<th box="[153,298,1506,1531]" gridcol="0" gridrow="6" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1506,1531]" gridcol="1" gridrow="6" pageId="16" pageNumber="69">F</td>
<td box="[379,433,1506,1531]" gridcol="2" gridrow="6" pageId="16" pageNumber="69">18</td>
<td box="[451,540,1506,1531]" gridcol="3" gridrow="6" pageId="16" pageNumber="69">1.96</td>
<td box="[560,637,1506,1531]" gridcol="4" gridrow="6" pageId="16" pageNumber="69">18.68</td>
<td box="[659,748,1506,1531]" gridcol="5" gridrow="6" pageId="16" pageNumber="69">0.0564</td>
<td box="[770,864,1506,1531]" gridcol="6" gridrow="6" pageId="16" pageNumber="69">0.0553</td>
<td box="[885,970,1506,1531]" gridcol="7" gridrow="6" pageId="16" pageNumber="69">3–11</td>
<td box="[989,1097,1506,1531]" gridcol="8" gridrow="6" pageId="16" pageNumber="69">2480</td>
</tr>
<tr box="[153,1097,1541,1565]" gridrow="7" pageId="16" pageNumber="69">
<th box="[153,298,1541,1565]" gridcol="0" gridrow="7" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1541,1565]" gridcol="1" gridrow="7" pageId="16" pageNumber="69">G</td>
<td box="[379,433,1541,1565]" gridcol="2" gridrow="7" pageId="16" pageNumber="69">17</td>
<td box="[451,540,1541,1565]" gridcol="3" gridrow="7" pageId="16" pageNumber="69">1.94</td>
<td box="[560,637,1541,1565]" gridcol="4" gridrow="7" pageId="16" pageNumber="69">17.85</td>
<td box="[659,748,1541,1565]" gridcol="5" gridrow="7" pageId="16" pageNumber="69">0.0584</td>
<td box="[770,864,1541,1565]" gridcol="6" gridrow="7" pageId="16" pageNumber="69">0.0592</td>
<td box="[885,970,1541,1565]" gridcol="7" gridrow="7" pageId="16" pageNumber="69">8–12</td>
<td box="[989,1097,1541,1565]" gridcol="8" gridrow="7" pageId="16" pageNumber="69">2460</td>
</tr>
<tr box="[153,1097,1575,1600]" gridrow="8" pageId="16" pageNumber="69">
<th box="[153,298,1575,1600]" gridcol="0" gridrow="8" pageId="16" pageNumber="69">BPBM 33775</th>
<td box="[319,362,1575,1600]" gridcol="1" gridrow="8" pageId="16" pageNumber="69">A</td>
<td box="[379,433,1575,1600]" gridcol="2" gridrow="8" pageId="16" pageNumber="69">22</td>
<td box="[451,540,1575,1600]" gridcol="3" gridrow="8" pageId="16" pageNumber="69">2.31</td>
<td box="[560,637,1575,1600]" gridcol="4" gridrow="8" pageId="16" pageNumber="69">20.29</td>
<td box="[659,748,1575,1600]" gridcol="5" gridrow="8" pageId="16" pageNumber="69">0.0516</td>
<td box="[770,864,1575,1600]" gridcol="6" gridrow="8" pageId="16" pageNumber="69">0.0558</td>
<td box="[885,970,1575,1600]" gridcol="7" gridrow="8" pageId="16" pageNumber="69">6–12</td>
<td box="[989,1097,1575,1600]" gridcol="8" gridrow="8" pageId="16" pageNumber="69">2460</td>
</tr>
<tr box="[153,1097,1610,1634]" gridrow="9" pageId="16" pageNumber="69">
<th box="[153,298,1610,1634]" gridcol="0" gridrow="9" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1610,1634]" gridcol="1" gridrow="9" pageId="16" pageNumber="69">B</td>
<td box="[379,433,1610,1634]" gridcol="2" gridrow="9" pageId="16" pageNumber="69">21</td>
<td box="[451,540,1610,1634]" gridcol="3" gridrow="9" pageId="16" pageNumber="69">2.21</td>
<td box="[560,637,1610,1634]" gridcol="4" gridrow="9" pageId="16" pageNumber="69">19.86</td>
<td box="[659,748,1610,1634]" gridcol="5" gridrow="9" pageId="16" pageNumber="69">0.0532</td>
<td box="[770,864,1610,1634]" gridcol="6" gridrow="9" pageId="16" pageNumber="69">0.0545</td>
<td box="[885,970,1610,1634]" gridcol="7" gridrow="9" pageId="16" pageNumber="69">7–10</td>
<td box="[989,1097,1610,1634]" gridcol="8" gridrow="9" pageId="16" pageNumber="69">2460</td>
</tr>
<tr box="[153,1097,1644,1669]" gridrow="10" pageId="16" pageNumber="69">
<th box="[153,298,1644,1669]" gridcol="0" gridrow="10" pageId="16" pageNumber="69">“</th>
<td box="[319,362,1644,1669]" gridcol="1" gridrow="10" pageId="16" pageNumber="69">C</td>
<td box="[379,433,1644,1669]" gridcol="2" gridrow="10" pageId="16" pageNumber="69">20</td>
<td box="[451,540,1644,1669]" gridcol="3" gridrow="10" pageId="16" pageNumber="69">2.08</td>
<td box="[560,637,1644,1669]" gridcol="4" gridrow="10" pageId="16" pageNumber="69">19.80</td>
<td box="[659,748,1644,1669]" gridcol="5" gridrow="10" pageId="16" pageNumber="69">0.0531</td>
<td box="[770,864,1644,1669]" gridcol="6" gridrow="10" pageId="16" pageNumber="69">0.0536</td>
<td box="[885,970,1644,1669]" gridcol="7" gridrow="10" pageId="16" pageNumber="69">7–11</td>
<td box="[989,1097,1644,1669]" gridcol="8" gridrow="10" pageId="16" pageNumber="69">2460</td>
</tr>
</table>
</paragraph>
<paragraph blockId="17.[140,1108,505,664]" box="[437,977,505,527]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[437,977,505,527]" pageId="17" pageNumber="70">0.5 ļ.0 ļ.5 2.0 2.5</emphasis>
</paragraph>
<paragraph blockId="17.[140,1108,505,664]" box="[656,737,536,558]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[656,737,536,558]" pageId="17" pageNumber="70">Time (s)</emphasis>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769065" ID-Zenodo-Dep="3769065" httpUri="https://zenodo.org/record/3769065/files/figure.png" pageId="17" pageNumber="70" startId="17.[140,212,578,600]" targetBox="[142,1068,170,497]" targetPageId="17">
<paragraph blockId="17.[140,1108,505,664]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[140,266,578,600]" pageId="17" pageNumber="70">Figure 8. A</emphasis>
Waveform, 
<emphasis bold="true" box="[383,400,578,600]" pageId="17" pageNumber="70">B</emphasis>
power spectrum, and 
<emphasis bold="true" box="[614,633,578,600]" pageId="17" pageNumber="70">C</emphasis>
spectrogram of Call “D” of 
<taxonomicName authority="Kraus &amp; Allison, 2009" authorityName="Kraus &amp; Allison" authorityYear="2009" box="[905,1107,578,600]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="17" pageNumber="70" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
<emphasis box="[905,1107,578,600]" italics="true" pageId="17" pageNumber="70">
<taxonomicName box="[905,1008,578,599]" class="Amphibia" family="Microhylidae" genus="Oreophryne" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="17" pageNumber="70" phylum="Chordata" rank="genus">Oreophryne</taxonomicName>
anamiatoi
</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[140,190,611,632]" pageId="17" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
(BPBM 33774) recorded at Mt. Paramo, Muller Range, Southern Highlands Province, Papua New Guinea on 3 April 2009 at 2000 h. Air temperature 17.7 °C.
</paragraph>
</caption>
<paragraph blockId="17.[140,1108,1085,1215]" box="[441,1012,1085,1107]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[441,1012,1085,1107]" pageId="17" pageNumber="70">0.05 0.ļ0 0.ļ5 0.20 0.25 0.30</emphasis>
</paragraph>
<paragraph blockId="17.[140,1108,1085,1215]" box="[666,747,1116,1138]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[666,747,1116,1138]" pageId="17" pageNumber="70">Time (s)</emphasis>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3769067" ID-Zenodo-Dep="3769067" httpUri="https://zenodo.org/record/3769067/files/figure.png" pageId="17" pageNumber="70" startId="17.[140,212,1161,1183]" targetBox="[175,1085,739,1077]" targetPageId="17">
<paragraph blockId="17.[140,1108,1085,1215]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[140,238,1161,1183]" pageId="17" pageNumber="70">Figure 9.</emphasis>
Detail of the middle three notes of call “D” of 
<taxonomicName authority="Kraus &amp; Allison, 2009" authorityName="Kraus &amp; Allison" authorityYear="2009" box="[671,871,1161,1183]" class="Amphibia" family="Microhylidae" genus="Oreophryne" kingdom="Animalia" order="Anura" pageId="17" pageNumber="70" phylum="Chordata" rank="species" species="anamiatoi" status="sp. n.">
<emphasis box="[671,871,1161,1183]" italics="true" pageId="17" pageNumber="70">Oreophryne anamiatoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[877,927,1162,1183]" pageId="17" pageNumber="70" rank="species">sp. n.</taxonomicNameLabel>
(BPBM 33774). 
<emphasis bold="true" box="[1089,1108,1161,1183]" pageId="17" pageNumber="70">A</emphasis>
Waveform, 
<emphasis bold="true" box="[248,265,1193,1215]" pageId="17" pageNumber="70">B</emphasis>
power spectrum, and 
<emphasis bold="true" box="[471,490,1193,1215]" pageId="17" pageNumber="70">C</emphasis>
spectrogram. Note the pulsing of the notes.
</paragraph>
</caption>
<paragraph blockId="17.[140,1108,1293,1672]" pageId="17" pageNumber="70">
Hz (range 2460–2540) (
<figureCitation box="[428,495,1293,1319]" captionStart="Figure 8" captionStartId="17.[140,212,578,600]" captionTargetBox="[142,1068,170,497]" captionTargetId="figure@17.[140,1108,165,559]" captionTargetPageId="17" captionText="Figure 8. A Waveform, B power spectrum, and C spectrogram of Call “D” of Oreophryne anamiatoi sp. n. (BPBM 33774) recorded at Mt. Paramo, Muller Range, Southern Highlands Province, Papua New Guinea on 3 April 2009 at 2000 h. Air temperature 17.7 °C." figureDoi="http://doi.org/10.5281/zenodo.3769065" httpUri="https://zenodo.org/record/3769065/files/figure.png" pageId="17" pageNumber="70">Fig. 8</figureCitation>
B–C, 9B–C) and no frequency modulation over the duration of the note.
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="71" pageId="17" pageNumber="70" type="etymology">
<paragraph blockId="17.[140,1108,1293,1672]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[185,320,1363,1389]" pageId="17" pageNumber="70">
<emphasis box="[185,315,1363,1389]" inLineHeading="true" pageId="17" pageNumber="70" reason="1">Etymology</emphasis>
.
</emphasis>
The species is named for Jim Anamiato of the 
<collectingCountry box="[840,1058,1363,1390]" name="Papua New Guinea" pageId="17" pageNumber="70">Papua New Guinea</collectingCountry>
National Museum for his considerable assistance on several of our expeditions, including the one during which this frog was discovered.
</paragraph>
<paragraph blockId="17.[140,1108,1293,1672]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[185,268,1469,1495]" pageId="17" pageNumber="70">
<emphasis box="[185,263,1469,1495]" inLineHeading="true" pageId="17" pageNumber="70" reason="1">Range</emphasis>
.
</emphasis>
Known only from the vicinity of the 
<typeStatus box="[725,773,1470,1496]" pageId="17" pageNumber="70">type</typeStatus>
locality on the northeastern slopes of the Muller Range, 
<collectingRegion box="[450,785,1504,1530]" country="Papua New Guinea" name="Southern Highlands" pageId="17" pageNumber="70">Southern Highlands Province</collectingRegion>
, 
<collectingCountry box="[795,1012,1504,1531]" name="Papua New Guinea" pageId="17" pageNumber="70">Papua New Guinea</collectingCountry>
(
<figureCitation box="[1026,1093,1504,1530]" captionStart="Figure 4" captionStartId="7.[140,212,1622,1644]" captionTargetBox="[140,1108,787,1602]" captionTargetId="figure@7.[140,1109,787,1602]" captionTargetPageId="7" captionText="Figure 4. Map showing known distribution of three new species of microhylid frogs from along the eastern slope of Mt. Itukua, Muller Range, Southern Highlands Province, Papua New Guinea (star)." figureDoi="http://doi.org/10.5281/zenodo.3769057" httpUri="https://zenodo.org/record/3769057/files/figure.png" pageId="17" pageNumber="70">Fig. 4</figureCitation>
).
</paragraph>
<paragraph blockId="17.[140,1108,1293,1672]" pageId="17" pageNumber="70">
<emphasis bold="true" box="[185,386,1539,1565]" pageId="17" pageNumber="70">
<emphasis box="[185,309,1539,1565]" inLineHeading="true" pageId="17" pageNumber="70" reason="1">Ecological</emphasis>
notes.
</emphasis>
Animals inhabited both primary and secondary rainforest at elevations ranging from 
<quantity box="[380,545,1575,1601]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.18" metricValueMin="1.8699999999999999" pageId="17" pageNumber="70" unit="m" value="2025.0" valueMax="2180.0" valueMin="1870.0">1870–2180 m</quantity>
. Most animals were found in the immediate vicinity of stream banks, but a few were found several meters away in adjacent forest. Males called from dense moss mats on standing or fallen trees from 
<quantity box="[824,898,1645,1672]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="17" pageNumber="70" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">1–5 m</quantity>
above the ground,
</paragraph>
<paragraph blockId="18.[224,580,542,595]" box="[301,580,573,595]" pageId="18" pageNumber="71">
<emphasis bold="true" box="[301,580,573,595]" pageId="18" pageNumber="71">2 4 6 8 ļ0</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>