treatments-xml/data/54/69/8C/54698C42DFF65A60820498366B6ACE5A.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e84013" ID-Pensoft-Pub="1864-8312-80-575" ID-Pensoft-UUID="81B59BA932BC5DAF90815FD08F5820A6" ID-ZooBank="4A6669D57DDD4AE0A8B455724B9C594A" ModsDocID="1864-8312-80-575" checkinTime="1668813875820" checkinUser="pensoft" docAuthor="Linsky, Marek, Ciamporova-Zatovicova, Zuzana, Lassova, Kristina &amp; Ciampor Jr, Fedor" docDate="2022" docId="54698C42DFF65A60820498366B6ACE5A" docLanguage="en" docName="ArthropodSystPhyl 80: 575-602" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-11-18" docSource="http://dx.doi.org/10.3897/asp.80.e84013" docTitle="Rumilara riberai Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr 2022, sp. nov." docType="treatment" docUuid="7B43FDD9-535D-4D13-A19F-87573BF05C93" docUuidSource="ZooBank" docVersion="1" id="81B59BA932BC5DAF90815FD08F5820A6" lastPageNumber="575" masterDocId="81B59BA932BC5DAF90815FD08F5820A6" masterDocTitle="Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)" masterLastPageNumber="602" masterPageNumber="575" pageNumber="575" updateTime="1668813875820" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Linsky, Marek</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7206-2380</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ciamporova-Zatovicova, Zuzana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0506-6212</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lassova, Kristina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7175-2109</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ciampor Jr, Fedor</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6269-3592</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
<mods:nameIdentifier type="email">f.ciampor@savba.sk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>575</mods:start>
<mods:end>602</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e84013</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e84013</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-575</mods:identifier>
<mods:identifier type="ZooBank">4A6669D57DDD4AE0A8B455724B9C594A</mods:identifier>
<mods:identifier type="Pensoft-UUID">81B59BA932BC5DAF90815FD08F5820A6</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:7B43FDD9-535D-4D13-A19F-87573BF05C93" httpUri="http://treatment.plazi.org/id/54698C42DFF65A60820498366B6ACE5A" lastPageNumber="575" pageId="0" pageNumber="575">
<subSubSection pageId="0" pageNumber="575" type="nomenclature">
<paragraph pageId="0" pageNumber="575">
3.2.5. 
<taxonomicName LSID="https://zoobank.org/7B43FDD9-535D-4D13-A19F-87573BF05C93" authority="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr, 2022" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" family="Elmidae" genus="Rumilara" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rumilara riberai" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="riberai" status="sp. nov.">Rumilara riberai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="575">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">Figs 2c</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">, 3c</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">, 5c</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">, 6c</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">, 7c</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">, 8c</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">, 9c</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Larva of Rumilara obscura sp. nov. (a - d): a prosternum and mesosternum; b foreleg (without coxa); c midleg; d hindleg. Schematic illustrations: e midleg of Hexanchorus cordillierae (alp - anterior lateral pubescence; mst - mesotibial tubercle; plp - posterior lateral pubescence; vs - ventral seta); f aedeagus of Rumilara riberai sp. nov. (ap - apophyse; co - corona; df - dorsal fibula; en - endophallus; mi - fibula minor; mj - fibula major; ml - median lobe; pa - paramere; ph - phallobase). Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure10" httpUri="https://binary.pensoft.net/fig/770433" pageId="0" pageNumber="575">, 10f</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">, 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="materials_examined">
<paragraph pageId="0" pageNumber="575">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">
<emphasis italics="true" pageId="0" pageNumber="575">Type material</emphasis>
: 
<materialsCitation collectingDate="2013-08-11" country="Ecuador" elevation="917" latitude="-0.33325" location="Rio Toachi" longLatPrecision="1" longitude="-78.94072" municipality="Otongachi" specimenCount="♂" stateProvince="Santo Domingo de los Tsachilas prov." typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
(PUCE) &quot; 
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion country="Ecuador" name="Santo Domingo de los Tsachilas">
Santo Domingo de los 
<normalizedToken originalValue="Tsáchilas">Tsachilas</normalizedToken>
prov.
</collectingRegion>
, 
<collectingMunicipality>Otongachi</collectingMunicipality>
env., tributary of 
<location LSID="urn:lsid:plazi:treatment:54698C42DFF65A60820498366B6ACE5A:38F2CFA0E3E769995561541B0FAE3062" country="Ecuador" latitude="-0.33325" longLatPrecision="1" longitude="-78.94072" municipality="Otongachi" name="Rio Toachi" stateProvince="Santo Domingo de los Tsachilas prov.">
<normalizedToken originalValue="Río">Rio</normalizedToken>
Toachi
</location>
, 
<geoCoordinate degrees="00" direction="south" minutes="19" orientation="latitude" precision="1" seconds="59.7" value="-0.33325">00°19′59.7″ S</geoCoordinate>
, 
<geoCoordinate degrees="78" direction="west" minutes="56" orientation="longitude" precision="1" seconds="26.6" value="-78.94072">78°56′26.6″ W</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.17" unit="m" value="917.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.17" unit="m" value="917.0">917m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="2013-08-11">11.8.2013</collectingDate>
, stream ca 
<quantity metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="1.0" unit="m" value="10.0">10m</elevation>
</quantity>
wide, with sand, gravel, boulders, 
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
</emphasis>
; 
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Ecuador" elevation="917" latitude="-0.33325" location="Rio Toachi" longLatPrecision="1" longitude="-78.94072" municipality="Otongachi" specimenCount="10" specimenCount-female="1" specimenCount-male="6" stateProvince="Santo Domingo de los Tsachilas" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="575">
<typeStatus>Paratypes</typeStatus>
</emphasis>
<specimenCount type="male">6♂♂</specimenCount>
, 
<specimenCount type="female">1♀</specimenCount>
(CCB), 
<specimenCount type="generic">3 ex</specimenCount>
with the same data as the holotype
</materialsCitation>
; 
<materialsCitation collectingDate="2013-08-11" country="Ecuador" elevation="914" latitude="-0.33069444" location="Rio Toachi" longLatPrecision="1" longitude="-78.94489" municipality="Otongachi" specimenCount="10" specimenCount-female="4" stateProvince="Santo Domingo de los Tsachilas prov." typeStatus="paratype">
<specimenCount type="female">4♀♀</specimenCount>
, 
<specimenCount type="generic">6 ex</specimenCount>
&quot; 
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion country="Ecuador" name="Santo Domingo de los Tsachilas">
Santo Domingo de los 
<normalizedToken originalValue="Tsáchilas">Tsachilas</normalizedToken>
prov.
</collectingRegion>
, 
<collectingMunicipality>Otongachi</collectingMunicipality>
env., next to the tributary of 
<location LSID="urn:lsid:plazi:treatment:54698C42DFF65A60820498366B6ACE5A:05E53D56956678430F93E0331C1354A3" country="Ecuador" latitude="-0.33069444" longLatPrecision="1" longitude="-78.94489" municipality="Otongachi" name="Rio Toachi" stateProvince="Santo Domingo de los Tsachilas prov.">
<normalizedToken originalValue="Río">Rio</normalizedToken>
Toachi
</location>
, 
<geoCoordinate degrees="00" direction="south" minutes="19" orientation="latitude" precision="1" seconds="50.5" value="-0.33069444">00°19′50.5″ S</geoCoordinate>
, 
<geoCoordinate degrees="78" direction="west" minutes="56" orientation="longitude" precision="1" seconds="41.6" value="-78.94489">78°56′41.6″ W</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">914m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="2013-08-11">11.8.2013</collectingDate>
, very shallow stream running along a rock covered by periphyton, 
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Ecuador" elevation="917" latitude="-0.33325" location="Rio Toachi" longLatPrecision="1" longitude="-78.94072" municipality="Otongachi" specimenCount="13" specimenCount-larva="13" stateProvince="Santo Domingo de los Tsachilas" typeStatus="Other material">
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">
<typeStatus isOtherTypeStatus="true">Other material</typeStatus>
</emphasis>
: 
<specimenCount type="larva">13 larvae</specimenCount>
with the same data as the holotype
</materialsCitation>
; 
<materialsCitation collectingDate="2013-08-11" country="Ecuador" elevation="914" latitude="-0.33069444" location="Rio Toachi" longLatPrecision="1" longitude="-78.94489" municipality="Otongachi" specimenCount="6" specimenCount-larva="6" stateProvince="Santo Domingo de los Tsachilas prov.">
<specimenCount type="larva">6 larvae</specimenCount>
&quot; 
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion country="Ecuador" name="Santo Domingo de los Tsachilas">
Santo Domingo de los 
<normalizedToken originalValue="Tsáchilas">Tsachilas</normalizedToken>
prov.
</collectingRegion>
, 
<collectingMunicipality>Otongachi</collectingMunicipality>
env., next to the tributary of 
<location LSID="urn:lsid:plazi:treatment:54698C42DFF65A60820498366B6ACE5A:2BDF7D21F6C307E96522C2BA40E3BCD0" country="Ecuador" latitude="-0.33069444" longLatPrecision="1" longitude="-78.94489" municipality="Otongachi" name="Rio Toachi" stateProvince="Santo Domingo de los Tsachilas prov.">
<normalizedToken originalValue="Río">Rio</normalizedToken>
Toachi
</location>
, 
<geoCoordinate degrees="00" direction="south" minutes="19" orientation="latitude" precision="1" seconds="50.5" value="-0.33069444">00°19′50.5″ S</geoCoordinate>
, 
<geoCoordinate degrees="78" direction="west" minutes="56" orientation="longitude" precision="1" seconds="41.6" value="-78.94489">78°56′41.6″ W</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">914m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="2013-08-11">11.8.2013</collectingDate>
, very shallow stream running along a rock covered by periphyton, 
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
; 
<materialsCitation collectingDate="2013-08-11" country="Ecuador" elevation="852" latitude="-0.3261111" location="Rio Toachi" longLatPrecision="1" longitude="-78.94975" municipality="Otongachi" specimenCount="1" specimenCount-larva="1" stateProvince="Santo Domingo de los Tsachilas prov.">
<specimenCount type="larva">1 larva</specimenCount>
&quot; 
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
, 
<collectingRegion country="Ecuador" name="Santo Domingo de los Tsachilas">
Santo Domingo de los 
<normalizedToken originalValue="Tsáchilas">Tsachilas</normalizedToken>
prov.
</collectingRegion>
, 
<collectingMunicipality>Otongachi</collectingMunicipality>
env., tributary of 
<location LSID="urn:lsid:plazi:treatment:54698C42DFF65A60820498366B6ACE5A:24772860EA6DDA9D1A35DC5B6A6462DC" country="Ecuador" latitude="-0.3261111" longLatPrecision="1" longitude="-78.94975" municipality="Otongachi" name="Rio Toachi" stateProvince="Santo Domingo de los Tsachilas prov.">
<normalizedToken originalValue="Río">Rio</normalizedToken>
Toachi
</location>
, 
<geoCoordinate degrees="00" direction="south" minutes="19" orientation="latitude" precision="1" seconds="34.0" value="-0.3261111">00°19′34.0″ S</geoCoordinate>
, 
<geoCoordinate degrees="78" direction="west" minutes="56" orientation="longitude" precision="1" seconds="59.1" value="-78.94975">78°56′59.1″ W</geoCoordinate>
, 
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.52" unit="m" value="852.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.52" unit="m" value="852.0">852m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="2013-08-11">11.8.2013</collectingDate>
, wild river ca 
<quantity metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="2.0" unit="m" value="20.0">20m</elevation>
</quantity>
wide, fast flowing, with large boulders, 
<normalizedToken originalValue="Čiampor">Ciampor</normalizedToken>
Jr &amp; 
<normalizedToken originalValue="Čiamporová-Zaťovičová">Ciamporova-Zatovicova</normalizedToken>
lgt.&quot;
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/asp.80.e84013.figure15" httpUri="https://binary.pensoft.net/fig/770438" pageId="0" pageNumber="575" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" pageId="0" pageNumber="575">Figure 15.</emphasis>
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Deleve" baseAuthorityYear="1968" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara leleupi" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara leleupi</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="575">comb. nov.</emphasis>
: 
<emphasis bold="true" pageId="0" pageNumber="575">a</emphasis>
aedeagus, left lateral view; 
<emphasis bold="true" pageId="0" pageNumber="575">b</emphasis>
aedeagus, ventral view; 
<emphasis bold="true" pageId="0" pageNumber="575">c</emphasis>
aedeagus, right lateral view; 
<emphasis bold="true" pageId="0" pageNumber="575">d</emphasis>
male sternite VIII. Scale: 
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.1mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="adult diagnosis">
<paragraph pageId="0" pageNumber="575">Adult diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="575">The species can be distinguished by a combination of following characters: 1) head only partially clothed by pubescence, clypeus bare; 2) femora bare or with a row of dark, long hairs on dorsal margin; 3) elytra without dense pubescence; 4) aedeagus with rod-like sclerite; 5) parameres only feebly asymmetrical.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Adult diagnostic description.</paragraph>
<paragraph pageId="0" pageNumber="575">
Male. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Colour</emphasis>
: Body (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">2c</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3c</figureCitation>
) black; lateral sides of femora and tibiae brown; tarsal claws pale brown with reddish tincture. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Pubescence</emphasis>
: Head behind eyes clothed with short, prone pubescence. Elytra very readily covered by short, recumbent pubescence, and with numerous moderately long, curved, semi-erect, hair-like setae. Femora with long, dark, densely set, hair-like setae near dorsal margin; pro- and metatibiae apically with a fringe of long, dark, hair-like setae on inner margin; mesotibiae without pubescence. Metaventrite readily clothed with short, prone pubescence. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Surface</emphasis>
: Head, clypeus and pronotum densely punctate. Elytra with rows of punctures moderately deeply impressed; third interval slightly elevated in anterior 1/5; remaining intervals nearly flat. Prosternal process (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">7c</figureCitation>
) moderately wide, concave between procoxae, then subtriangular with rounded apex. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Aedeagus</emphasis>
(Figs 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Rumilara riberai sp. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d ovipositor; e male sternite VIII; f female sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure13" httpUri="https://binary.pensoft.net/fig/770436" pageId="0" pageNumber="575">13a-c</figureCitation>
) with long, dorsal, rod-like sclerite on penis; parameres asymmetrical, left one slightly longer. - Female externally similar to male, except slightly larger.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="575">♂ - CL: 2.16-2.53 mm; PL: 0.56-0.65 mm; PW: 0.71-0.79 mm; EL: 1.60-1.88 mm; EW: 0.83-0.97 mm. ♀ - CL: 2.68-2.69 mm; PL: 0.68-0.70 mm; PW: 0.82-0.85 mm; EL: 1.99-2.00 mm; EW: 1.04-1.06 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="variation">
<paragraph pageId="0" pageNumber="575">Variation.</paragraph>
<paragraph pageId="0" pageNumber="575">The greatest differences were observed in distribution of pubescence (mainly on legs, around eyes, partly on pronotum and elytra) due to its often removal.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="etymology">
<paragraph pageId="0" pageNumber="575">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="575">
Named after Ignacio Ribera 
<normalizedToken originalValue="Galán">Galan</normalizedToken>
a great expert on systematics, phylogeny, evolution, biogeography, and conservation of water beetles, who untimely passed away.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="larva">
<paragraph pageId="0" pageNumber="575">Larva</paragraph>
<paragraph pageId="0" pageNumber="575">
(Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Larvae of Rumilara gen. nov. species: a R. obscura sp. nov., dorsal; b R. suppressa sp. nov., dorsal; c R. riberai sp. nov., dorsal; d R. paterna sp. nov., dorsal; e R. obscura sp. nov., ventral; f R. suppressa sp. nov., ventral. Scale: 1 mm." figureDoi="10.3897/asp.80.e84013.figure9" httpUri="https://binary.pensoft.net/fig/770432" pageId="0" pageNumber="575">9c</figureCitation>
) with length 2.92-2.96 mm, greatest width 0.96-0.99 mm. Pleurites on abdominal segments 1-5. Abdominal segments 1-6 explanate. Surface around the frontal pair of pronotal gibbosities of the same colour as the rest of pronotum. Tergum of abdominal segments 1-7 with a pair of distinct, longitudinally arranged small gibbosities.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="distribution">
<paragraph pageId="0" pageNumber="575">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="575">
Known from three localities in Santo Domingo de los 
<normalizedToken originalValue="Tsáchilas">Tsachilas</normalizedToken>
Province (Figs 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Sampling sites of Rumilara gen. nov. in Ecuador: a Cotopaxi prov., Otonga env., Rio Esmeralda; b Cotopaxi prov., Artos env., stream above confluence with Rio Esmeralda; c Santo Domingo de los Tsachilas prov., Otongachi env., tributary of Rio Toachi; d Santo Domingo de los Tsachilas prov., thin water layer on rock, Otongachi env.; e Morona-Santiago prov., small stream near Indanza; f Morona-Santiago prov., Santa Rosa de Mamanguy env., Cascada la Encanada; g fast-flowing stream in Napo prov.; h Morona-Santiago prov., tributary of Rio Tinajillas; i Morona-Santiago prov., tributary of Rio Tinajillas; (localities of R. obscura - a, b; R. riberai - c, d; R. paterna - e, f; R. suppressa - g - i)." figureDoi="10.3897/asp.80.e84013.figure16" httpUri="https://binary.pensoft.net/fig/770439" pageId="0" pageNumber="575">16c-d</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="remarks">
<paragraph pageId="0" pageNumber="575">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="575">Several paratypes are markedly smaller than the rest (including holotype), however, they agree in all diagnostic characters.</paragraph>
</subSubSection>
</treatment>
</document>