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<document id="8851065D8AAC971EAEB273D6CD4B80A3" ID-CLB-Dataset="55812" ID-DOI="10.11646/zootaxa.5129.1.6" ID-GBIF-Dataset="a8ed59ae-958b-4979-9843-299d524c4a7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="6488262" ID-ZooBank="3FF266F0-BEDB-46AA-8E5D-AEA3434BA75E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1650960444504" checkinUser="plazi" docAuthor="Saigusa, Toyohei" docDate="2022" docId="03FDAA58FF85402FFF30263AFB0DD97B" docLanguage="en" docName="zootaxa.5129.1.6.pdf" docOrigin="Zootaxa 5129 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Empis (Sphenempis) bicoloriventris Saigusa 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="111" masterDocId="FFC4D220FF864029FFA7255FFFA1DE38" masterDocTitle="Sphenempis, a new Empis subgenus (Diptera, Empididae) from the Eastern Palaearctic Region" masterLastPageNumber="117" masterPageNumber="105" pageNumber="108" updateTime="1699329216215" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="DB99D2C36B5F911DBE2D295229398995" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="00E88E34245FA6D507E3B86BFC07CE71">
<mods:title id="36D2BDF341229DA715B7C2A4CCA19EC8">Sphenempis, a new Empis subgenus (Diptera, Empididae) from the Eastern Palaearctic Region</mods:title>
</mods:titleInfo>
<mods:name id="25AAEE12C70FAE4633561F2AECEB958D" type="personal">
<mods:role id="68B4D8767080336A65E3758D0798BE6F">
<mods:roleTerm id="BF4622118258098488995AB744B56C3C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="166C21AF357B735204FBFF110158A8DA">Saigusa, Toyohei</mods:namePart>
</mods:name>
<mods:typeOfResource id="69C174EAC5C22A900F7EA21A7327EE9F">text</mods:typeOfResource>
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<mods:titleInfo id="EA731F8792958A3A3DEB4A41A3911A5B">
<mods:title id="44E23CBD2EBAC707783530692E8365FA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="41AB25630BF7D3F7078915A52BE52075">
<mods:date id="7C0DD2A2C1B93D1C3A4E362697D1CD27">2022</mods:date>
<mods:detail id="A8F24ABE7E3FF1AA8062E6F72348A25D" type="pubDate">
<mods:number id="74AC53038CC0F2B8CCB1AEF0531AAF63">2022-04-26</mods:number>
</mods:detail>
<mods:detail id="EEFA41A7369FDC619354C3E709338F33" type="volume">
<mods:number id="AC665789137565B1B954B798B748E0EC">5129</mods:number>
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<mods:number id="6826F4F7C47101DDABBC4C2A305F2763">1</mods:number>
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<mods:classification id="17595F417048EB9D8A89D216918CAC26">journal article</mods:classification>
<mods:identifier id="519F689A73B9F2F5C6EC20729B47CCC1" type="CLB-Dataset">55812</mods:identifier>
<mods:identifier id="8B4810928E01871F68E5EB396A36B0F3" type="DOI">10.11646/zootaxa.5129.1.6</mods:identifier>
<mods:identifier id="DCAC000C2735E76800EE6BB658B0C5B5" type="GBIF-Dataset">a8ed59ae-958b-4979-9843-299d524c4a7a</mods:identifier>
<mods:identifier id="DEB7AF4F11E829F5C311A76A548BF77F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="48A6F98FCA2A4347272994D11A860FC4" type="Zenodo-Dep">6488262</mods:identifier>
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</mods:mods>
<treatment id="03FDAA58FF85402FFF30263AFB0DD97B" ID-DOI="http://doi.org/10.5281/zenodo.6502543" ID-GBIF-Taxon="194854847" ID-Zenodo-Dep="6502543" LSID="urn:lsid:plazi:treatment:03FDAA58FF85402FFF30263AFB0DD97B" httpUri="http://treatment.plazi.org/id/03FDAA58FF85402FFF30263AFB0DD97B" lastPageId="6" lastPageNumber="111" pageId="3" pageNumber="108">
<subSubSection id="C34E48C5FF85402AFF30263AFD0DDD47" box="[151,684,869,896]" pageId="3" pageNumber="108" type="nomenclature">
<paragraph id="8BEB1B4EFF85402AFF30263AFD0DDD47" blockId="3.[151,684,869,931]" box="[151,684,869,896]" pageId="3" pageNumber="108">
<heading id="D0A3AC22FF85402AFF30263AFD0DDD47" bold="true" box="[151,684,869,896]" fontSize="11" level="1" pageId="3" pageNumber="108" reason="1">
<emphasis id="B920C75CFF85402AFF30263AFD0DDD47" bold="true" box="[151,684,869,896]" pageId="3" pageNumber="108">
<taxonomicName id="4C5460CDFF85402AFF30263AFDE6DDB8" ID-CoL="9J2NN" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" box="[151,583,869,896]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="bicoloriventris" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF85402AFF30263AFF45DD47" bold="true" box="[151,228,869,895]" italics="true" pageId="3" pageNumber="108">Empis</emphasis>
(
<emphasis id="B920C75CFF85402AFF52263AFE29DDB8" bold="true" box="[245,392,869,896]" italics="true" pageId="3" pageNumber="108">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF85402AFE3E263AFDE6DDB8" bold="true" box="[409,583,869,896]" italics="true" pageId="3" pageNumber="108">bicoloriventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2137A27FF85402AFDE9263AFD0DDD47" box="[590,684,869,895]" pageId="3" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF85402AFF3026D6FE2DDD9B" box="[151,396,905,931]" pageId="3" pageNumber="108" type="description">
<paragraph id="8BEB1B4EFF85402AFF3026D6FE2DDD9B" blockId="3.[151,684,869,931]" box="[151,396,905,931]" pageId="3" pageNumber="108">
(
<figureCitation id="136F07CBFF85402AFF3826D6FEBCDD9B" box="[159,285,905,931]" captionStart="FIGURE 1" captionStartId="4.[151,250,1884,1908]" captionTargetBox="[157,1417,181,1859]" captionTargetId="figure-19@4.[155,1433,181,1859]" captionTargetPageId="4" captionText="FIGURE 1. Empis (Sphenempis) species. A, Empis (Sphenempis) bicoloriventris sp. nov., holotype male; B, ditto, anterodorsal aspect; C, Empis (Sphenempis) bicoloriventris sp. nov., paratype female; D, Empis (Sphenempis) amurensis, male; E, Empis (Sphenempis) oshimai sp. nov., holotype male. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6488266" httpUri="https://zenodo.org/record/6488266/files/figure.png" pageId="3" pageNumber="108">Figs 1A–C</figureCitation>
,
<figureCitation id="136F07CBFF85402AFE8E26D6FE98DD9B" box="[297,313,905,931]" captionStart="FIGURE 2" captionStartId="5.[151,250,1910,1934]" captionTargetBox="[227,1361,181,1886]" captionTargetId="figure-16@5.[225,1363,181,1886]" captionTargetPageId="5" captionText="FIGURE 2. Empis (Sphenempis) bicoloriventris sp. nov., paratype male from Okinawa-jima.A, head, lateral aspect; B, antenna; C, labium, lateral aspect; D, ditto, ventral aspect; E, labellum, lateral aspect; F, apex of labrum, anterior aspect; G, maxillary palpus. Scale bars = 0.5 mm (A, C. D), 0.2 mm (B, G), 0.1 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.6488270" httpUri="https://zenodo.org/record/6488270/files/figure.png" pageId="3" pageNumber="108">2</figureCitation>
,
<figureCitation id="136F07CBFF85402AFEE226D6FEC8DD9B" box="[325,361,905,931]" captionStart="FIGURE 3" captionStartId="6.[151,250,1114,1138]" captionTargetBox="[156,1431,309,1083]" captionTargetId="figure-73@6.[156,1431,309,1091]" captionTargetPageId="6" captionText="FIGURE 3. Wings of Empis species. A, Empis (Sphenempis) bicoloriventris sp. nov., paratype male from Okinawa-jima; B, Empis (Sphenempis) amurensis, male; C, Empis (Sphenempis) oshimai sp. nov., paratype male; D, Empis (Lissempis) nigritarsis, female; E, Empis sp. close to E. kasparyani, male from Kagoshima, Kyushu; F, Empis leptogastra Loew, male. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6488272" httpUri="https://zenodo.org/record/6488272/files/figure.png" pageId="3" pageNumber="108">3A</figureCitation>
,
<figureCitation id="136F07CBFF85402AFED226D6FE25DD9B" box="[373,388,905,931]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1916]" captionTargetBox="[217,1370,181,1864]" captionTargetId="figure-16@7.[217,1370,181,1869]" captionTargetPageId="7" captionText="FIGURE 4. Empis (Sphenempis) bicoloriventris sp. nov., paratype male from Okinawa-jima. A, hind femur, anterior aspect; B, hind tibia, anterior aspect; C, mid femur, anterior aspect; D, mid tibia, anterior aspect; E, distal part of hind femur, anterior aspect; F, distal part of mid femur, anterior aspect; G, terminalia, lateral aspect; H, ditto, dorsal aspect; I, ditto, dorsolateral aspect. Scale bars = 0.5 mm (A–F), 0.2 mm (G), 0.1 mm (H, I)." figureDoi="http://doi.org/10.5281/zenodo.6488274" httpUri="https://zenodo.org/record/6488274/files/figure.png" pageId="3" pageNumber="108">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF85402AFF30268EFAF2DB17" pageId="3" pageNumber="108" type="materials_examined">
<paragraph id="8BEB1B4EFF85402AFF30268EFAF2DB17" blockId="3.[151,1437,977,1982]" pageId="3" pageNumber="108">
<emphasis id="B920C75CFF85402AFF30268EFE59DDD3" bold="true" box="[151,504,977,1003]" pageId="3" pageNumber="108">
Type material.
<typeStatus id="54EFA5ECFF85402AFEFE268EFE59DDD3" box="[345,504,977,1003]" pageId="3" pageNumber="108" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="9D52D0C7FF85402AFDA2268DFDBDDDD4" box="[517,540,978,1004]" pageId="3" pageNumber="108" type="male">♂</specimenCount>
, labelled: [Mt. Nago-dake] alt.
<quantity id="4CACB6ABFF85402AFC3B268EFBAEDDD4" box="[924,1039,977,1004]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="3" pageNumber="108" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50-150 m</quantity>
/
<geoCoordinate id="EE607D89FF85402AFBB0268EFBC8DDD4" box="[1047,1129,977,1004]" degrees="26.35" direction="north" orientation="latitude" pageId="3" pageNumber="108" precision="555" value="26.35">26.35N</geoCoordinate>
<geoCoordinate id="EE607D89FF85402AFBD0268EFB77DDD4" box="[1143,1238,977,1004]" degrees="127.59" direction="east" orientation="longitude" pageId="3" pageNumber="108" precision="555" value="127.59">127.59E</geoCoordinate>
/Nago. Okinawahontô /
<collectingCountry id="F3435BDEFF85402AFF4026AAFE87DA37" box="[231,294,1013,1039]" name="Japan" pageId="3" pageNumber="108">Japan</collectingCountry>
/
<date id="FFEA3D8EFF85402AFE9126AAFE51DA37" box="[310,496,1013,1039]" pageId="3" pageNumber="108" value="2008-03-02">2nd March 2008</date>
/ H. Nakayama leg.; [
<typeStatus id="54EFA5ECFF85402AFD4426AAFCDDDA28" box="[739,892,1013,1040]" pageId="3" pageNumber="108" type="holotype">HOLOTYPE</typeStatus>
]/
<taxonomicName id="4C5460CDFF85402AFC3426A9FBD9DA37" authority="" authorityName="Saigusa" authorityYear="2022" box="[915,1144,1013,1039]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Sphenempis">
<emphasis id="B920C75CFF85402AFC3426A9FC78DA37" box="[915,985,1014,1039]" italics="true" pageId="3" pageNumber="108">Empis</emphasis>
(
<emphasis id="B920C75CFF85402AFC4C26AAFBD0DA37" box="[1003,1137,1013,1039]" italics="true" pageId="3" pageNumber="108">Sphenempis</emphasis>
)
</taxonomicName>
/
<taxonomicName id="4C5460CDFF85402AFB2E26AAFA8FDA37" authorityName="Saigusa" authorityYear="2022" box="[1161,1326,1013,1039]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="bicoloriventris">
<emphasis id="B920C75CFF85402AFB2E26AAFA8FDA37" box="[1161,1326,1013,1039]" italics="true" pageId="3" pageNumber="108">bicoloriventris</emphasis>
</taxonomicName>
/ Saigusa, 2022 (in Kyushu University Museum).
<emphasis id="B920C75CFF85402AFDFD2146FCA6DA0B" bold="true" box="[602,775,1049,1075]" pageId="3" pageNumber="108">
<typeStatus id="54EFA5ECFF85402AFDFD2146FCA3DA0B" box="[602,770,1049,1075]" pageId="3" pageNumber="108" type="paratype">PARATYPES</typeStatus>
:
</emphasis>
<specimenCount id="9D52D0C7FF85402AFCB72146FC95DA0B" box="[784,820,1049,1075]" pageId="3" pageNumber="108" type="male">1♂</specimenCount>
, [
<collectingRegion id="4990D5ACFF85402AFCEE2146FC72DA0C" box="[841,979,1049,1076]" country="Japan" name="Okinawa" pageId="3" pageNumber="108">OKINAWA</collectingRegion>
]/ Yona/ Okinawa-jima/
<date id="FFEA3D8EFF85402AFB432145FAD4DA0B" box="[1252,1397,1049,1076]" pageId="3" pageNumber="108" value="2006-12-06">Dec. 6, 2006</date>
/
<collectionCode id="ED45838BFF85402AFA212145FA39DA0C" box="[1414,1432,1050,1076]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. SAIGUSA leg.;
<specimenCount id="9D52D0C7FF85402AFEEA2162FED9DA6F" box="[333,376,1085,1111]" pageId="3" pageNumber="108" type="male">2♂</specimenCount>
<specimenCount id="9D52D0C7FF85402AFEDF2162FE35DA6F" box="[376,404,1085,1111]" pageId="3" pageNumber="108" type="female">1♀</specimenCount>
, RYUKYUS /OKINAWA- /
<collectionCode id="ED45838BFF85402AFD772162FC95DA6F" box="[720,820,1085,1111]" pageId="3" pageNumber="108">HONTO</collectionCode>
, Kudeken/
<date id="FFEA3D8EFF85402AFC162162FB8FDA60" box="[945,1070,1085,1112]" pageId="3" pageNumber="108" value="1954-03-20">20.III.1954</date>
/ Syusiro Ito, Japan-U.S./ Co-op. Sci. / Programme;
<specimenCount id="9D52D0C7FF85402AFEC9213EFE33DA43" box="[366,402,1121,1147]" pageId="3" pageNumber="108" type="female">3♀</specimenCount>
, [OKINAWA-jima]/ Tanagawagumui/ Anba, Kunigami/
<date id="FFEA3D8EFF85402AFBB2213EFB1ADA43" box="[1045,1211,1121,1147]" pageId="3" pageNumber="108" value="1997-03-03">March 3, 1997</date>
/
<collectionCode id="ED45838BFF85402AFB6B213DFB7FDA44" box="[1228,1246,1122,1148]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. SAIGUSA col.;
<specimenCount id="9D52D0C7FF85402AFF3021DAFF1BDAA7" box="[151,186,1157,1183]" pageId="3" pageNumber="108" type="female">1♀</specimenCount>
, [OKINAWA-jima]/ Ryukyu Univ. Exp./ Forest Yona/
<date id="FFEA3D8EFF85402AFC8621DAFC62DAA7" box="[801,963,1157,1183]" pageId="3" pageNumber="108" value="1997-03-03">March 3, 1997</date>
/
<collectionCode id="ED45838BFF85402AFC7421D9FC44DA98" box="[979,997,1158,1184]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. SAIGUSA col.;
<specimenCount id="9D52D0C7FF85402AFB0121DAFB68DAA7" box="[1190,1225,1157,1183]" pageId="3" pageNumber="108" type="female">1♀</specimenCount>
, same data,
<date id="FFEA3D8EFF85402AFAF421DAFF4EDAFB" pageId="3" pageNumber="108" value="1997-03-05">March 5, 1997</date>
;
<specimenCount id="9D52D0C7FF85402AFF5B21F6FE81DAFB" box="[252,288,1193,1219]" pageId="3" pageNumber="108" type="male">1♂</specimenCount>
, Ishigaki-jima/ Banna-dake (foot)/
<date id="FFEA3D8EFF85402AFD1721F6FC8FDAFB" box="[688,814,1193,1219]" pageId="3" pageNumber="108" value="2002-03-18">18.III.2002</date>
/M. SUGIMOTO col.;
<specimenCount id="9D52D0C7FF85402AFB9D21F6FBFCDAFB" box="[1082,1117,1193,1219]" pageId="3" pageNumber="108" type="female">1♀</specimenCount>
, [
<collectingRegion id="4990D5ACFF85402AFBD521F6FB5DDAFC" box="[1138,1276,1193,1220]" country="Japan" name="Okinawa" pageId="3" pageNumber="108">OKINAWA</collectingRegion>
] /Omotodake/ Ishigakijima /
<date id="FFEA3D8EFF85402AFE932192FE46DADF" box="[308,487,1229,1256]" pageId="3" pageNumber="108" value="2004-03-25">March 25, 2004</date>
/
<collectionCode id="ED45838BFF85402AFE502191FDA8DAD0" box="[503,521,1230,1256]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. Nakamura col.;
<specimenCount id="9D52D0C7FF85402AFD6B2192FD50DAD0" box="[716,753,1229,1256]" pageId="3" pageNumber="108" type="male">5♂</specimenCount>
, [ISHIGAKI-JIMA]/ small torrent south of Golf field/
<quantity id="4CACB6ABFF85402AFAC42192FA39DAD0" box="[1379,1432,1229,1256]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="108" unit="m" value="95.0">95m</quantity>
, Hirae-Ômata/
<date id="FFEA3D8EFF85402AFE9B21ADFDA3DB33" box="[316,514,1265,1292]" pageId="3" pageNumber="108" value="2019-01-29">January 29, 2019</date>
/
<collectionCode id="ED45838BFF85402AFDB321ADFD87DB34" box="[532,550,1266,1292]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. SAIGUSA col.;
<specimenCount id="9D52D0C7FF85402AFD5521AEFCB7DB33" box="[754,790,1265,1291]" pageId="3" pageNumber="108" type="female">3♀</specimenCount>
, [ISHIGAKI-JIMA]/ North of Golf field/
<quantity id="4CACB6ABFF85402AFAA321AEFAE7DB33" box="[1284,1350,1265,1291]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="108" unit="m" value="100.0">100m</quantity>
, Hirae- Ômata /
<date id="FFEA3D8EFF85402AFF5F2049FE16DB17" box="[248,439,1301,1328]" pageId="3" pageNumber="108" value="2019-01-28">January 28, 2019</date>
/
<collectionCode id="ED45838BFF85402AFE612049FE79DB08" box="[454,472,1302,1328]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. SAIGUSA col.;
<specimenCount id="9D52D0C7FF85402AFD06204AFD67DB17" box="[673,710,1301,1327]" pageId="3" pageNumber="108" type="male">2♂</specimenCount>
,
<date id="FFEA3D8EFF85402AFD75204AFCE1DB08" box="[722,832,1301,1328]" pageId="3" pageNumber="108" value="1953-01-19">I-19-1953</date>
/ Ryukyu Is./
<collectionCode id="ED45838BFF85402AFC7E2049FC4ADB08" box="[985,1003,1302,1328]" name="Tavera, Department of Geology and Geophysics" pageId="3" pageNumber="108">T</collectionCode>
. SHIRAKI, Iriomote Is./ Ohara.
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF85402AFF602066FE03DB4F" pageId="3" pageNumber="108" type="etymology">
<paragraph id="8BEB1B4EFF85402AFF602066FE03DB4F" blockId="3.[151,1437,977,1982]" pageId="3" pageNumber="108">
<emphasis id="B920C75CFF85402AFF602066FEEFDB6B" bold="true" box="[199,334,1337,1363]" pageId="3" pageNumber="108">Etymology.</emphasis>
The specific name
<taxonomicName id="4C5460CDFF85402AFD9F2066FD7CDB6B" authorityName="Saigusa" authorityYear="2022" box="[568,733,1337,1363]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="108" phylum="Arthropoda" rank="species" species="bicoloriventris">
<emphasis id="B920C75CFF85402AFD9F2066FD7CDB6B" box="[568,733,1337,1363]" italics="true" pageId="3" pageNumber="108">bicoloriventris</emphasis>
</taxonomicName>
is based on the bicolored male abdomen, white basally and blackish brown distally.
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF85402AFF6020DEFC02DB87" pageId="3" pageNumber="108" type="diagnosis">
<paragraph id="8BEB1B4EFF85402AFF6020DEFC02DB87" blockId="3.[151,1437,977,1982]" pageId="3" pageNumber="108">
<emphasis id="B920C75CFF85402AFF6020DEFEE3DBA3" bold="true" box="[199,322,1409,1435]" pageId="3" pageNumber="108">Diagnosis.</emphasis>
Mesonotum densely light brownish pollinose; laterotergal brush of whitish setae; male abdomen whitish on anterior half, blackish brown on posterior half; halter pale.
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF85402FFF602096FBD8DBAB" lastPageId="6" lastPageNumber="111" pageId="3" pageNumber="108" type="description">
<paragraph id="8BEB1B4EFF85402AFF602096FD5DD957" blockId="3.[151,1437,977,1982]" pageId="3" pageNumber="108">
<emphasis id="B920C75CFF85402AFF602096FE3DDBDB" bold="true" box="[199,412,1481,1507]" pageId="3" pageNumber="108">Description. Male</emphasis>
(
<figureCitation id="136F07CBFF85402AFE0E2096FD84DBDC" box="[425,549,1481,1508]" captionStart="FIGURE 1" captionStartId="4.[151,250,1884,1908]" captionTargetBox="[157,1417,181,1859]" captionTargetId="figure-19@4.[155,1433,181,1859]" captionTargetPageId="4" captionText="FIGURE 1. Empis (Sphenempis) species. A, Empis (Sphenempis) bicoloriventris sp. nov., holotype male; B, ditto, anterodorsal aspect; C, Empis (Sphenempis) bicoloriventris sp. nov., paratype female; D, Empis (Sphenempis) amurensis, male; E, Empis (Sphenempis) oshimai sp. nov., holotype male. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6488266" httpUri="https://zenodo.org/record/6488266/files/figure.png" pageId="3" pageNumber="108">Figs 1A, B</figureCitation>
). Body length
<quantity id="4CACB6ABFF85402AFD6F2096FCF3DBDC" box="[712,850,1481,1508]" metricMagnitude="-3" metricUnit="m" metricValue="3.6999999999999997" metricValueMax="4.1" metricValueMin="3.3" pageId="3" pageNumber="108" unit="mm" value="3.6999999999999997" valueMax="4.1" valueMin="3.3">3.3–4.1 mm</quantity>
; wing length
<quantity id="4CACB6ABFF85402AFC412096FBCEDBDC" box="[998,1135,1481,1508]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="3" pageNumber="108" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
. Coloration. Head blackish brown, densely light grey pollinose. Antenna with pedicel and scape brown, postpedicel black with extreme base brown. Palpus pale brown; labrum yellow, dark brown at extreme tip; labium pale brown, with pale yellowishbrown labella. Thorax blackish brown, densely light grey pollinose, postpronotum polished brown on lateral part; viewed from above pollinosity of mesoscutum more brownish on area surrounding posterior part of dorsocentrals; prescutellar area and scutellum ash grey pollinose. Abdomen bicolored, anterior 4 segments yellowish white, thinly white pollinose, clothed with short white setae, posterior 4 segments and genitalia polished blackish brown, with darker setae. Wing uniformly infuscate to light brown, with membrane somewhat polished, pterostigma indistinct; veins dark brown; squama whitish with pale fine setae. Halter yellowish white, apical half of knob dark in specimens from Okinawa-jima, entirely white in specimens from Ishigaki-jima and Iriomote-jima. Legs yellow; coxae thinly light grey pollinose; mid and hind coxae somewhat infuscate; trochanters black apically; femora and tibiae polished yellow, somewhat darkened on apical 1/2; tarsi brown.
</paragraph>
<paragraph id="8BEB1B4EFF85402FFF602225FDB8DF25" blockId="3.[151,1437,977,1982]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="111" pageId="3" pageNumber="108">
Most structural and chaetotaxy characters as in subgenus description. Wing (
<figureCitation id="136F07CBFF85402AFBBE2226FBD0D9AB" box="[1049,1137,1913,1939]" captionStart="FIGURE 3" captionStartId="6.[151,250,1114,1138]" captionTargetBox="[156,1431,309,1083]" captionTargetId="figure-73@6.[156,1431,309,1091]" captionTargetPageId="6" captionText="FIGURE 3. Wings of Empis species. A, Empis (Sphenempis) bicoloriventris sp. nov., paratype male from Okinawa-jima; B, Empis (Sphenempis) amurensis, male; C, Empis (Sphenempis) oshimai sp. nov., paratype male; D, Empis (Lissempis) nigritarsis, female; E, Empis sp. close to E. kasparyani, male from Kagoshima, Kyushu; F, Empis leptogastra Loew, male. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6488272" httpUri="https://zenodo.org/record/6488272/files/figure.png" pageId="3" pageNumber="108">Fig. 3A</figureCitation>
) about 3.5 times as long as wide: cell r
<subScript id="17D0190BFF85402AFEB222EFFE8FD986" attach="left" box="[277,302,1968,1982]" fontSize="6" pageId="3" pageNumber="108">2+3</subScript>
almost as wide as cell r
<subScript id="17D0190BFF85402AFD9C22EFFDF4D986" attach="left" box="[571,597,1968,1982]" fontSize="6" pageId="3" pageNumber="108">4+5</subScript>
at level of tip of vein R
<subScript id="17D0190BFF85402AFCC722EFFCD8D986" attach="left" box="[864,889,1968,1982]" fontSize="6" pageId="3" pageNumber="108">2+3</subScript>
; discal cell half as long as vein M
<subScript id="17D0190BFF85402AFB5F22EFFAA1D986" attach="left" box="[1272,1280,1968,1982]" fontSize="6" pageId="3" pageNumber="108">1</subScript>
; axillary lobe narrow, widest at middle, with evenly curved margin. Legs (
<figureCitation id="136F07CBFF80402FFC9425C8FC10DE89" box="[819,945,151,177]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1916]" captionTargetBox="[217,1370,181,1864]" captionTargetId="figure-16@7.[217,1370,181,1869]" captionTargetPageId="7" captionText="FIGURE 4. Empis (Sphenempis) bicoloriventris sp. nov., paratype male from Okinawa-jima. A, hind femur, anterior aspect; B, hind tibia, anterior aspect; C, mid femur, anterior aspect; D, mid tibia, anterior aspect; E, distal part of hind femur, anterior aspect; F, distal part of mid femur, anterior aspect; G, terminalia, lateral aspect; H, ditto, dorsal aspect; I, ditto, dorsolateral aspect. Scale bars = 0.5 mm (A–F), 0.2 mm (G), 0.1 mm (H, I)." figureDoi="http://doi.org/10.5281/zenodo.6488274" httpUri="https://zenodo.org/record/6488274/files/figure.png" pageId="6" pageNumber="111">Figs 4A–F</figureCitation>
): Mid femur with 3–4 anteroventral and 1–2 posteroventral spines, hind femur 12.5 times as long as thick, with 3–4 anteroventral spines, without posteroventral spines. Terminalia (
<figureCitation id="136F07CBFF80402FFED12580FE4EDEC1" box="[374,495,223,249]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1916]" captionTargetBox="[217,1370,181,1864]" captionTargetId="figure-16@7.[217,1370,181,1869]" captionTargetPageId="7" captionText="FIGURE 4. Empis (Sphenempis) bicoloriventris sp. nov., paratype male from Okinawa-jima. A, hind femur, anterior aspect; B, hind tibia, anterior aspect; C, mid femur, anterior aspect; D, mid tibia, anterior aspect; E, distal part of hind femur, anterior aspect; F, distal part of mid femur, anterior aspect; G, terminalia, lateral aspect; H, ditto, dorsal aspect; I, ditto, dorsolateral aspect. Scale bars = 0.5 mm (A–F), 0.2 mm (G), 0.1 mm (H, I)." figureDoi="http://doi.org/10.5281/zenodo.6488274" httpUri="https://zenodo.org/record/6488274/files/figure.png" pageId="6" pageNumber="111">Figs 4G–I</figureCitation>
): Palm-like process with two moderately long finger-like lobes and short proximal lobe; cercus not raised posteriorly.
</paragraph>
<caption id="DF2B4BC6FF82402DFF302203FC9CD984" ID-DOI="http://doi.org/10.5281/zenodo.6488266" ID-Zenodo-Dep="6488266" httpUri="https://zenodo.org/record/6488266/files/figure.png" pageId="4" pageNumber="109" startId="4.[151,250,1884,1908]" targetBox="[157,1417,181,1859]" targetPageId="4">
<paragraph id="8BEB1B4EFF82402DFF302203FC9CD984" blockId="4.[151,1436,1884,1981]" pageId="4" pageNumber="109">
<emphasis id="B920C75CFF82402DFF302203FEB3D94C" bold="true" box="[151,274,1884,1908]" pageId="4" pageNumber="109">FIGURE 1.</emphasis>
<taxonomicName id="4C5460CDFF82402DFEBF2202FE47D94D" authority="" authorityName="Saigusa" authorityYear="2022" box="[280,486,1885,1909]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="subGenus" subGenus="Sphenempis">
<emphasis id="B920C75CFF82402DFEBF2202FEF6D94C" box="[280,343,1885,1908]" italics="true" pageId="4" pageNumber="109">Empis</emphasis>
(
<emphasis id="B920C75CFF82402DFEC22202FE7FD94D" box="[357,478,1885,1909]" italics="true" pageId="4" pageNumber="109">Sphenempis</emphasis>
)
</taxonomicName>
species.
<emphasis id="B920C75CFF82402DFDE52203FDFBD94C" bold="true" box="[578,602,1884,1908]" pageId="4" pageNumber="109">A,</emphasis>
<taxonomicName id="4C5460CDFF82402DFDC72202FC68D94D" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" box="[608,969,1885,1909]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="bicoloriventris" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF82402DFDC72202FD3ED94C" box="[608,671,1885,1908]" italics="true" pageId="4" pageNumber="109">Empis</emphasis>
(
<emphasis id="B920C75CFF82402DFD0A2202FC87D94D" box="[685,806,1885,1909]" italics="true" pageId="4" pageNumber="109">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF82402DFC932202FC68D94D" box="[820,969,1885,1909]" italics="true" pageId="4" pageNumber="109">bicoloriventris</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF82402DFC682202FBBED94D" bold="true" box="[975,1055,1885,1909]" pageId="4" pageNumber="109">
<taxonomicNameLabel id="A2137A27FF82402DFC682202FBBED94D" box="[975,1055,1885,1909]" pageId="4" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male;
<emphasis id="B920C75CFF82402DFB6F2202FB7ED94D" bold="true" box="[1224,1247,1885,1909]" pageId="4" pageNumber="109">B,</emphasis>
ditto, anterodorsal aspect;
<emphasis id="B920C75CFF82402DFF4222DFFF5FD9A0" bold="true" box="[229,254,1920,1944]" pageId="4" pageNumber="109">C,</emphasis>
<taxonomicName id="4C5460CDFF82402DFEA122DEFDD5D9A1" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" box="[262,628,1921,1945]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="bicoloriventris" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF82402DFEA122DEFEE4D9A0" box="[262,325,1921,1944]" italics="true" pageId="4" pageNumber="109">Empis</emphasis>
(
<emphasis id="B920C75CFF82402DFEF222DEFE6FD9A1" box="[341,462,1921,1945]" italics="true" pageId="4" pageNumber="109">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF82402DFE7822DEFDD5D9A1" box="[479,628,1921,1945]" italics="true" pageId="4" pageNumber="109">bicoloriventris</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF82402DFDDB22DEFD6FD9A1" bold="true" box="[636,718,1921,1945]" pageId="4" pageNumber="109">
<taxonomicNameLabel id="A2137A27FF82402DFDDB22DEFD6FD9A1" box="[636,718,1921,1945]" pageId="4" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype female;
<emphasis id="B920C75CFF82402DFC2822DEFC06D9A1" bold="true" box="[911,935,1921,1945]" pageId="4" pageNumber="109">D,</emphasis>
<taxonomicName id="4C5460CDFF82402DFC1722DEFB4ED9A0" authorityName="Shamshev" authorityYear="1998" box="[944,1263,1921,1945]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="amurensis" subGenus="Sphenempis">
<emphasis id="B920C75CFF82402DFC1722DEFC4ED9A0" box="[944,1007,1921,1944]" italics="true" pageId="4" pageNumber="109">Empis</emphasis>
(
<emphasis id="B920C75CFF82402DFC5822DEFBD9D9A1" box="[1023,1144,1921,1945]" italics="true" pageId="4" pageNumber="109">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF82402DFB2F22DEFB4ED9A0" box="[1160,1263,1921,1944]" italics="true" pageId="4" pageNumber="109">amurensis</emphasis>
</taxonomicName>
, male;
<emphasis id="B920C75CFF82402DFA9922DEFAF4D9A1" bold="true" box="[1342,1365,1921,1945]" pageId="4" pageNumber="109">E,</emphasis>
<taxonomicName id="4C5460CDFF82402DFAFA22DEFED6D985" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="109" phylum="Arthropoda" rank="species" species="oshimai" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF82402DFAFA22DEFA3DD9A0" box="[1373,1436,1921,1944]" italics="true" pageId="4" pageNumber="109">Empis</emphasis>
(
<emphasis id="B920C75CFF82402DFF0722FAFEB8D985" box="[160,281,1957,1981]" italics="true" pageId="4" pageNumber="109">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF82402DFE8022FAFED6D985" box="[295,375,1957,1981]" italics="true" pageId="4" pageNumber="109">oshimai</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF82402DFED922FAFE6FD985" bold="true" box="[382,462,1957,1981]" pageId="4" pageNumber="109">
<taxonomicNameLabel id="A2137A27FF82402DFED922FAFE6FD985" box="[382,462,1957,1981]" pageId="4" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF2B4BC6FF83402CFF302229FCFED9EE" ID-DOI="http://doi.org/10.5281/zenodo.6488270" ID-Zenodo-Dep="6488270" httpUri="https://zenodo.org/record/6488270/files/figure.png" pageId="5" pageNumber="110" startId="5.[151,250,1910,1934]" targetBox="[227,1361,181,1886]" targetPageId="5">
<paragraph id="8BEB1B4EFF83402CFF302229FCFED9EE" blockId="5.[151,1436,1910,2006]" pageId="5" pageNumber="110">
<emphasis id="B920C75CFF83402CFF302229FEB0D9B6" bold="true" box="[151,273,1910,1934]" pageId="5" pageNumber="110">FIGURE 2.</emphasis>
<taxonomicName id="4C5460CDFF83402CFEB12229FDDFD9B6" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" box="[278,638,1910,1934]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="110" phylum="Arthropoda" rank="species" species="bicoloriventris" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF83402CFEB12229FEF4D9B5" box="[278,341,1910,1933]" italics="true" pageId="5" pageNumber="110">Empis</emphasis>
(
<emphasis id="B920C75CFF83402CFEC42229FE7DD9B6" box="[355,476,1910,1934]" italics="true" pageId="5" pageNumber="110">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF83402CFE4E2229FDDFD9B6" box="[489,638,1910,1934]" italics="true" pageId="5" pageNumber="110">bicoloriventris</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF83402CFD242229FD73D9B6" bold="true" box="[643,722,1910,1934]" pageId="5" pageNumber="110">
<taxonomicNameLabel id="A2137A27FF83402CFD242229FD73D9B6" box="[643,722,1910,1934]" pageId="5" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male from Okinawa-jima.
<emphasis id="B920C75CFF83402CFBE52229FBFAD9B6" bold="true" box="[1090,1115,1910,1934]" pageId="5" pageNumber="110">A,</emphasis>
head, lateral aspect;
<emphasis id="B920C75CFF83402CFA8D2229FAE0D9B6" bold="true" box="[1322,1345,1910,1934]" pageId="5" pageNumber="110">B,</emphasis>
antenna;
<emphasis id="B920C75CFF83402CFF3022C5FF11D98A" bold="true" box="[151,176,1946,1970]" pageId="5" pageNumber="110">C,</emphasis>
labium, lateral aspect;
<emphasis id="B920C75CFF83402CFE0722C5FE19D98A" bold="true" box="[416,440,1946,1970]" pageId="5" pageNumber="110">D,</emphasis>
ditto, ventral aspect;
<emphasis id="B920C75CFF83402CFD3F22C5FD0ED98A" bold="true" box="[664,687,1946,1970]" pageId="5" pageNumber="110">E,</emphasis>
labellum, lateral aspect;
<emphasis id="B920C75CFF83402CFC1622C5FC64D98A" bold="true" box="[945,965,1946,1970]" pageId="5" pageNumber="110">F,</emphasis>
apex of labrum, anterior aspect;
<emphasis id="B920C75CFF83402CFABD22C5FA92D98A" bold="true" box="[1306,1331,1946,1970]" pageId="5" pageNumber="110">G,</emphasis>
maxillary palpus. Scale bars = 0.5 mm (A, C. D), 0.2 mm (B, G), 0.1 mm (E, F).
</paragraph>
</caption>
<caption id="DF2B4BC6FF80402FFF302105FF67DAE6" ID-DOI="http://doi.org/10.5281/zenodo.6488272" ID-Zenodo-Dep="6488272" httpUri="https://zenodo.org/record/6488272/files/figure.png" pageId="6" pageNumber="111" startId="6.[151,250,1114,1138]" targetBox="[156,1431,309,1083]" targetPageId="6">
<paragraph id="8BEB1B4EFF80402FFF302105FF67DAE6" blockId="6.[151,1437,1114,1246]" pageId="6" pageNumber="111">
<emphasis id="B920C75CFF80402FFF302105FEB5DA4A" bold="true" box="[151,276,1114,1138]" pageId="6" pageNumber="111">FIGURE 3.</emphasis>
Wings of
<taxonomicName id="4C5460CDFF80402FFE262104FD93DA4A" box="[385,562,1114,1138]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="undefined-a">
<emphasis id="B920C75CFF80402FFE262104FE61DA4A" box="[385,448,1115,1138]" italics="true" pageId="6" pageNumber="111">Empis</emphasis>
species.
<emphasis id="B920C75CFF80402FFDB82105FD93DA4A" bold="true" box="[543,562,1114,1138]" pageId="6" pageNumber="111">A</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF80402FFD952105FD96DA4A" bold="true" box="[562,567,1114,1138]" pageId="6" pageNumber="111">,</emphasis>
<taxonomicName id="4C5460CDFF80402FFD982104FC0CDA4A" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" box="[575,941,1114,1139]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="bicoloriventris" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF80402FFD982104FDDFDA4A" box="[575,638,1115,1138]" italics="true" pageId="6" pageNumber="111">Empis</emphasis>
(
<emphasis id="B920C75CFF80402FFD282105FCA9DA4A" box="[655,776,1114,1138]" italics="true" pageId="6" pageNumber="111">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF80402FFCBF2105FC0CDA4A" box="[792,941,1114,1138]" italics="true" pageId="6" pageNumber="111">bicoloriventris</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF80402FFC122104FBA7DA4B" bold="true" box="[949,1030,1115,1139]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="A2137A27FF80402FFC122104FBA7DA4B" box="[949,1030,1115,1139]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male from Okinawa-jima;
<emphasis id="B920C75CFF80402FFA222104FA3DDA4B" bold="true" box="[1413,1436,1115,1139]" pageId="6" pageNumber="111">B,</emphasis>
<taxonomicName id="4C5460CDFF80402FFF302120FE70DAAE" authorityName="Shamshev" authorityYear="1998" box="[151,465,1150,1175]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="amurensis" subGenus="Sphenempis">
<emphasis id="B920C75CFF80402FFF302120FF77DAAE" box="[151,214,1151,1174]" italics="true" pageId="6" pageNumber="111">Empis</emphasis>
(
<emphasis id="B920C75CFF80402FFF432121FEFCDAAE" box="[228,349,1150,1174]" italics="true" pageId="6" pageNumber="111">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF80402FFECD2120FE70DAAE" box="[362,465,1151,1174]" italics="true" pageId="6" pageNumber="111">amurensis</emphasis>
</taxonomicName>
, male;
<emphasis id="B920C75CFF80402FFDBE2121FD93DAAE" bold="true" box="[537,562,1150,1174]" pageId="6" pageNumber="111">C,</emphasis>
<taxonomicName id="4C5460CDFF80402FFD912120FCF8DAAE" authority="Saigusa, 2022" authorityName="Saigusa" authorityYear="2022" box="[566,857,1150,1175]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="oshimai" status="sp. nov." subGenus="Sphenempis">
<emphasis id="B920C75CFF80402FFD912120FDD4DAAE" box="[566,629,1151,1174]" italics="true" pageId="6" pageNumber="111">Empis</emphasis>
(
<emphasis id="B920C75CFF80402FFD242121FD5DDAAE" box="[643,764,1150,1174]" italics="true" pageId="6" pageNumber="111">Sphenempis</emphasis>
)
<emphasis id="B920C75CFF80402FFCAE2121FCF8DAAE" box="[777,857,1150,1174]" italics="true" pageId="6" pageNumber="111">oshimai</emphasis>
</taxonomicName>
<emphasis id="B920C75CFF80402FFCF92120FC0CDAAF" bold="true" box="[862,941,1151,1175]" pageId="6" pageNumber="111">
<taxonomicNameLabel id="A2137A27FF80402FFCF92120FC0CDAAF" box="[862,941,1151,1175]" pageId="6" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype male;
<emphasis id="B920C75CFF80402FFBF62120FBCBDAAF" bold="true" box="[1105,1130,1151,1175]" pageId="6" pageNumber="111">D,</emphasis>
<taxonomicName id="4C5460CDFF80402FFBC82120FA37DAAE" authorityName="Meigen" authorityYear="1804" box="[1135,1430,1151,1175]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="nigritarsis" subGenus="Lissempis">
<emphasis id="B920C75CFF80402FFBC82120FB0FDAAE" box="[1135,1198,1151,1174]" italics="true" pageId="6" pageNumber="111">Empis</emphasis>
(
<emphasis id="B920C75CFF80402FFB1C2120FABEDAAE" box="[1211,1311,1151,1174]" italics="true" pageId="6" pageNumber="111">Lissempis</emphasis>
)
<emphasis id="B920C75CFF80402FFA8B2120FA37DAAE" box="[1324,1430,1151,1174]" italics="true" pageId="6" pageNumber="111">nigritarsis</emphasis>
</taxonomicName>
, female;
<emphasis id="B920C75CFF80402FFF4D21FCFEA0DA83" bold="true" box="[234,257,1187,1211]" pageId="6" pageNumber="111">E,</emphasis>
<taxonomicName id="4C5460CDFF80402FFEAF21FCFECADA82" box="[264,363,1187,1210]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B920C75CFF80402FFEAF21FCFEE6DA82" box="[264,327,1187,1210]" italics="true" pageId="6" pageNumber="111">Empis</emphasis>
sp.
</taxonomicName>
close to
<taxonomicName id="4C5460CDFF80402FFE6021FCFDF5DA82" authorityName="Shamshev" authorityYear="1998" box="[455,596,1186,1210]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="kasparyani">
<emphasis id="B920C75CFF80402FFE6021FCFDF5DA82" box="[455,596,1186,1210]" italics="true" pageId="6" pageNumber="111">E. kasparyani</emphasis>
</taxonomicName>
, male from Kagoshima, Kyushu;
<emphasis id="B920C75CFF80402FFC0921FCFC63DA83" bold="true" box="[942,962,1187,1211]" pageId="6" pageNumber="111">F,</emphasis>
<taxonomicName id="4C5460CDFF80402FFC6F21FCFB65DA82" authority="Loew" authorityName="Loew" authorityYear="1863" box="[968,1220,1186,1210]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="leptogastra">
<emphasis id="B920C75CFF80402FFC6F21FCFB20DA82" box="[968,1153,1186,1210]" italics="true" pageId="6" pageNumber="111">Empis leptogastra</emphasis>
Loew
</taxonomicName>
, male. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BEB1B4EFF80402FFF602052FBD8DBAB" blockId="6.[151,1437,1292,1859]" pageId="6" pageNumber="111">
<emphasis id="B920C75CFF80402FFF602052FEBFDB1F" bold="true" box="[199,286,1293,1319]" pageId="6" pageNumber="111">Female</emphasis>
(
<figureCitation id="136F07CBFF80402FFE882052FE26DB1F" box="[303,391,1293,1319]" captionStart="FIGURE 1" captionStartId="4.[151,250,1884,1908]" captionTargetBox="[157,1417,181,1859]" captionTargetId="figure-19@4.[155,1433,181,1859]" captionTargetPageId="4" captionText="FIGURE 1. Empis (Sphenempis) species. A, Empis (Sphenempis) bicoloriventris sp. nov., holotype male; B, ditto, anterodorsal aspect; C, Empis (Sphenempis) bicoloriventris sp. nov., paratype female; D, Empis (Sphenempis) amurensis, male; E, Empis (Sphenempis) oshimai sp. nov., holotype male. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6488266" httpUri="https://zenodo.org/record/6488266/files/figure.png" pageId="6" pageNumber="111">Fig. 1C</figureCitation>
). Similar to male. Body length
<quantity id="4CACB6ABFF80402FFD482052FCDBDB1F" box="[751,890,1293,1319]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.1" metricValueMin="3.7" pageId="6" pageNumber="111" unit="mm" value="3.9" valueMax="4.1" valueMin="3.7">3.7–4.1 mm</quantity>
; wing length
<quantity id="4CACB6ABFF80402FFBB22053FB01DB1F" box="[1045,1184,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.5" metricValueMin="3.6" pageId="6" pageNumber="111" unit="mm" value="4.05" valueMax="4.5" valueMin="3.6">3.6–4.5 mm</quantity>
. Frons almost as wide as ocellar tubercle (distance between posterior ocelli); mesoscutum more brownish pollinose, particularly along dorsocentral setae; wings clear, not infuscate; abdomen with polished brown terga and yellowish brown sterna, sparsely clothed with short brown setae; cercus slender, as long as abdominal segment 8.
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF80402FFF6020C3FB87DB8F" box="[199,1062,1436,1464]" pageId="6" pageNumber="111" type="distribution">
<paragraph id="8BEB1B4EFF80402FFF6020C3FB87DB8F" blockId="6.[151,1437,1292,1859]" box="[199,1062,1436,1464]" pageId="6" pageNumber="111">
<emphasis id="B920C75CFF80402FFF6020C3FEC0DB8E" bold="true" box="[199,353,1436,1462]" pageId="6" pageNumber="111">Distribution.</emphasis>
Ryukyus (Okinawa-jima, Ishigaki-jima, Iriomote-jima),
<collectingCountry id="F3435BDEFF80402FFC7820C2FB83DB8F" box="[991,1058,1437,1463]" name="Japan" pageId="6" pageNumber="111">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF80402FFF60209EFBFDDBE3" box="[199,1116,1472,1499]" pageId="6" pageNumber="111" type="materials_examined">
<paragraph id="8BEB1B4EFF80402FFF60209EFBFDDBE3" blockId="6.[151,1437,1292,1859]" box="[199,1116,1472,1499]" pageId="6" pageNumber="111">
<materialsCitation id="3B3C1113FF80402FFF60209EFBF6DBE3" ID-GBIF-Occurrence="3757641303" box="[199,1111,1472,1499]" country="Japan" county="Ryukyus" latitude="26.35" location="Mt. Nago-dake" longLatPrecision="745" longitude="127.59" municipality="Okinawa-jima" pageId="6" pageNumber="111" specimenCount="1" typeStatus="holotype">
<emphasis id="B920C75CFF80402FFF60209EFEC7DBE2" bold="true" box="[199,358,1472,1499]" pageId="6" pageNumber="111">
<typeStatus id="54EFA5ECFF80402FFF60209EFEA3DBE3" box="[199,258,1473,1499]" pageId="6" pageNumber="111">Type</typeStatus>
locality.
</emphasis>
<location id="8E8B4D95FF80402FFECB209EFDB7DBE3" LSID="urn:lsid:plazi:treatment:03FDAA58FF85402FFF30263AFB0DD97B:8E8B4D95FF80402FFECB209EFDB7DBE3" box="[364,534,1473,1499]" country="Japan" county="Ryukyus" latitude="26.35" longLatPrecision="745" longitude="127.59" municipality="Okinawa-jima" name="Mt. Nago-dake" pageId="6" pageNumber="111">Mt. Nago-dake</location>
(
<geoCoordinate id="EE607D89FF80402FFD83209FFDDBDBE3" box="[548,634,1472,1499]" degrees="26.35" direction="north" orientation="latitude" pageId="6" pageNumber="111" precision="555" value="26.35">26.35N</geoCoordinate>
<geoCoordinate id="EE607D89FF80402FFD27209FFD42DBE3" box="[640,739,1472,1499]" degrees="127.59" direction="east" orientation="longitude" pageId="6" pageNumber="111" precision="555" value="127.59">127.59E</geoCoordinate>
),
<collectingMunicipality id="6B8F8134FF80402FFD51209EFC38DBE3" box="[758,921,1473,1499]" pageId="6" pageNumber="111">Okinawa-jima</collectingMunicipality>
,
<collectingCounty id="628A63C2FF80402FFC03209EFBA8DBE3" box="[932,1033,1473,1499]" pageId="6" pageNumber="111">Ryukyus</collectingCounty>
,
<collectingCountry id="F3435BDEFF80402FFBB3209EFBF6DBE3" box="[1044,1111,1473,1499]" name="Japan" pageId="6" pageNumber="111">Japan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E48C5FF80402FFF6020BAFB0DD97B" pageId="6" pageNumber="111" type="discussion">
<paragraph id="8BEB1B4EFF80402FFF6020BAFEA4D88B" blockId="6.[151,1437,1292,1859]" pageId="6" pageNumber="111">
<emphasis id="B920C75CFF80402FFF6020BAFE9ADBC7" bold="true" box="[199,315,1509,1535]" pageId="6" pageNumber="111">Remarks.</emphasis>
The male of this species is easily distinguished from
<taxonomicName id="4C5460CDFF80402FFC1220BAFBE8DBC6" authorityName="Shamshev" authorityYear="1998" box="[949,1097,1509,1534]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="B920C75CFF80402FFC1220BAFBE8DBC6" box="[949,1097,1509,1534]" italics="true" pageId="6" pageNumber="111">E. amurensis</emphasis>
</taxonomicName>
by the pale (almost white in fresh material) anterior half of the abdomen. The female also may be distinguished from unknown females of the following two species by the basally white halter. This species is not rare in the Ryukyus. The color difference in halteres between the males from Okinawa-jima and the Yaeyama islands (Ishigaki-jima, Iriomote-jima) is found, but there are no differences in the male genitalia between these two populations. This species appears from January to March.
</paragraph>
<paragraph id="8BEB1B4EFF80402FFF6023E2FB0DD97B" blockId="6.[151,1437,1292,1859]" pageId="6" pageNumber="111">
The white colored male abdomen is similar to that of the western North American species,
<taxonomicName id="4C5460CDFF80402FFB7B23E2FECCD8C3" authority="Arnaud &amp; Birchim" authorityName="Arnaud &amp; Birchim" authorityYear="1966" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="111" phylum="Arthropoda" rank="species" species="melanderi">
<emphasis id="B920C75CFF80402FFB7B23E2FA3CD8EF" box="[1244,1437,1725,1751]" italics="true" pageId="6" pageNumber="111">Empis melanderi</emphasis>
Arnaud &amp; Birchim
</taxonomicName>
(
<bibRefCitation id="EFC566BFFF80402FFEDB23BEFD37D8C3" author="Arnaud, P. H. &amp; Birchim, J. D." box="[380,662,1761,1787]" pageId="6" pageNumber="111" pagination="332 - 336" refId="ref6270" refString="Arnaud, P. H. &amp; Birchim, J. D. (1966) A new brumal Empis from the San Francisco Bay Area, California (Diptera: Empididae). The Pan-Pacific Entomologist, 42 (4), 332 - 336." type="journal article" year="1966">Arnaud &amp; Birchim 1966</bibRefCitation>
, fig. 1). This species lacks acrostichals as in
<taxonomicName id="4C5460CDFF80402FFB3423BEFAB8D8C3" authorityName="Saigusa" authorityYear="2022" box="[1171,1305,1761,1787]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="108" phylum="Arthropoda" rank="subGenus" subGenus="Sphenempis">
<emphasis id="B920C75CFF80402FFB3423BEFAB8D8C3" box="[1171,1305,1761,1787]" italics="true" pageId="6" pageNumber="111">Sphenempis</emphasis>
</taxonomicName>
, but differs from the latter in the wing shape, apically weakened vein CuA+CuP, simple mid and hind femora without strong spine-like setae and the male terminalia with well-developed cerci and no palm-like sclerites.
</paragraph>
</subSubSection>
</treatment>
</document>
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