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<document id="3275439BF4E77FD41C20F08E9E8F9E80" ID-DOI="10.11646/zootaxa.4804.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3927094" ID-ZooBank="8C1C1703-9BBC-4B33-8045-78BDD9738F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593672611012" checkinUser="plazi" docAuthor="Westrop, Madison Armstrong Stephen R. &amp; Eoff, Jennifer D." docDate="2020" docId="03F76D37FFA6FFC26BD2E65AFDAC9678" docLanguage="en" docName="zootaxa.4804.1.1.pdf" docOrigin="Zootaxa 4804 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Blountia mimula Walcott 1916" docType="treatment" docVersion="3" lastPageNumber="11" masterDocId="FFCE154FFFAFFFC86B45E438FF8E9206" masterDocTitle="Systematics of a survivor: the Cambrian kingstoniid trilobite Blountia Walcott, 1916 across the Marjuman-Steptoean (Guzhangian-Paibian) extinction interval in Laurentian North America" masterLastPageNumber="79" masterPageNumber="1" pageNumber="10" updateTime="1698776919363" updateUser="plazi">
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<mods:title id="7BE7268CE9F1FE2087236F7ADE9032A5">Systematics of a survivor: the Cambrian kingstoniid trilobite Blountia Walcott, 1916 across the Marjuman-Steptoean (Guzhangian-Paibian) extinction interval in Laurentian North America</mods:title>
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<mods:namePart id="341E96A5F8273A15999BCE9F5D798677">Westrop, Madison Armstrong Stephen R.</mods:namePart>
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<mods:namePart id="A82D9C6917923631F8585F008165C368">Eoff, Jennifer D.</mods:namePart>
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<treatment id="03F76D37FFA6FFC26BD2E65AFDAC9678" LSID="urn:lsid:plazi:treatment:03F76D37FFA6FFC26BD2E65AFDAC9678" httpUri="http://treatment.plazi.org/id/03F76D37FFA6FFC26BD2E65AFDAC9678" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<subSubSection id="C3448FAAFFA6FFC16BD2E65AFD96907A" box="[151,536,610,637]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BE1DC21FFA6FFC16BD2E65AFD96907A" blockId="9.[151,536,610,672]" box="[151,536,610,637]" pageId="9" pageNumber="10">
<heading id="D0A96B4DFFA6FFC16BD2E65AFD96907A" bold="true" box="[151,536,610,637]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="4C5EA7A2FFA6FFC16BD2E65AFD96907A" authority="Walcott, 1916" authorityName="Walcott" authorityYear="1916" box="[151,536,610,637]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA6FFC16BD2E65AFD96907A" bold="true" box="[151,536,610,637]" pageId="9" pageNumber="10">
<emphasis id="B92A0033FFA6FFC16BD2E65AFEEB907B" bold="true" box="[151,357,610,637]" italics="true" pageId="9" pageNumber="10">Blountia mimula</emphasis>
Walcott, 1916
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3448FAAFFA6FFC16BD2E6BEFBDC911C" pageId="9" pageNumber="10" type="description">
<paragraph id="8BE1DC21FFA6FFC16BD2E6BEFF6A90A6" blockId="9.[151,536,610,672]" box="[151,228,646,672]" pageId="9" pageNumber="10">
<figureCitation id="1365C0A4FFA6FFC16BD2E6BEFF6A90A6" box="[151,228,646,672]" captionStart="PLATE 1" captionStartId="26.[151,234,1839,1864]" captionText="PLATE 1. Blountia mimula Walcott, 1916 from the Nolichucky Formation, Tennessee. 14. Nearly complete exoskeleton, holotype, USNM 62781, dorsal, lateral, posterior and anterior views, x17, Shields Ridge, Jefferson County, Tennessee, USNM loc. 120 (illustrated by Walcott 1916, pl. 61, figs 4, 4). 57. Cranidium, paratype, USNM 62873, dorsal, lateral and anterior views, x18, 17.7 km [11 miles] northwest of Knoxville, Knox County, Tennessee, USNM loc. 107c (illustrated by Walcott 1916, pl. 61, figs 4b, 4b). 810. Pygidium, paratype, USNM 62782, lateral, dorsal and posterior views, x16, also from USNM loc. 107c (illustrated by Walcott 1916, pl. 61, fig. 4a)." pageId="9" pageNumber="10">Plate 1</figureCitation>
</paragraph>
<paragraph id="8BE1DC21FFA6FFC16BD2E6FDFD6490DB" blockId="9.[151,1106,709,795]" box="[151,746,709,734]" pageId="9" pageNumber="10">
1916
<taxonomicName id="4C5EA7A2FFA6FFC16B95E6FDFE5F90D8" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[208,465,709,734]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA6FFC16B95E6FDFEF490DB" box="[208,378,709,733]" italics="true" pageId="9" pageNumber="10">Blountia mimula</emphasis>
Walcott
</taxonomicName>
, p. 399, pl. 61, figs 4, 4ac.
</paragraph>
<paragraph id="8BE1DC21FFA6FFC16BD2E6DCFCAD90FA" blockId="9.[151,1106,709,795]" box="[151,803,739,764]" pageId="9" pageNumber="10">
1938
<taxonomicName id="4C5EA7A2FFA6FFC16B95E6DCFE5C90FA" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[208,466,739,764]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA6FFC16B95E6DCFEF490FA" box="[208,378,740,764]" italics="true" pageId="9" pageNumber="10">Blountia mimula</emphasis>
Walcott
</taxonomicName>
; Resser, p. 63, pl. 12, figs 1819.
</paragraph>
<paragraph id="8BE1DC21FFA6FFC16BD2E73BFBDC911C" blockId="9.[151,1106,709,795]" box="[151,1106,770,795]" pageId="9" pageNumber="10">
<emphasis id="B92A0033FFA6FFC16BD2E73BFF33911C" box="[151,189,771,794]" italics="true" pageId="9" pageNumber="10">non</emphasis>
1965
<taxonomicName id="4C5EA7A2FFA6FFC16BB8E73BFE70911D" authority="Walcott" authorityName="Walcott" authorityYear="1916" box="[253,510,770,795]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA6FFC16BB8E73BFE28911D" box="[253,422,771,795]" italics="true" pageId="9" pageNumber="10">Blountia mimula</emphasis>
Walcott
</taxonomicName>
; Rasetti, p. 59, pl. 10, figs 37 [=
<taxonomicName id="4C5EA7A2FFA6FFC16813E73BFB8A911D" authority="Westrop &amp; Eoff, 2020" authorityName="Westrop &amp; Eoff" authorityYear="2020" box="[854,1028,771,795]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="angelae" status="sp. nov.">
<emphasis id="B92A0033FFA6FFC16813E73BFB8A911D" box="[854,1028,771,795]" italics="true" pageId="9" pageNumber="10">Blountia angelae</emphasis>
</taxonomicName>
<emphasis id="B92A0033FFA6FFC16F4FE73BFBCD911D" bold="true" box="[1034,1091,771,795]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A219BD48FFA6FFC16F4FE73BFBCD911D" box="[1034,1091,771,795]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
].
</paragraph>
</subSubSection>
<subSubSection id="C3448FAAFFA6FFC16BD2E77BFA0A91CC" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BE1DC21FFA6FFC16BD2E77BFA0A91CC" blockId="9.[151,1437,835,2043]" pageId="9" pageNumber="10">
<emphasis id="B92A0033FFA6FFC16BD2E77BFE9C9158" bold="true" box="[151,274,835,862]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Frontal area slightly more that one-quarter (27%; 2529) cranidial length, with gently sloping preglabellar field and flat anterior border nearly equal in length (sag.); anterior border furrow well-defined. Moderately convex glabella with width at palpebral lobe equal to 75% of glabellar length. Seven thoracic segments. Where expressed, pygidial border 15% of total length of pygidium; border furrow barely perceptible to entirely effaced.
</paragraph>
</subSubSection>
<subSubSection id="C3448FAAFFA6FFC16B82E7EBFC65965C" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BE1DC21FFA6FFC16B82E7EBFC65965C" blockId="9.[151,1437,835,2043]" pageId="9" pageNumber="10">
<emphasis id="B92A0033FFA6FFC16B82E7EBFEB891E8" bold="true" box="[199,310,979,1006]" pageId="9" pageNumber="10">Material.</emphasis>
<materialsCitation id="3B36D67CFFA6FFC16A78E7ECFC66965C" collectionCode="USNM" county="Jefferson County" location="Knoxville" municipality="Nolichucky Formation" pageId="9" pageNumber="10" specimenCode="USNM 62781, USNM 62873, USNM 62872" specimenCount="2" stateProvince="Tennessee" typeStatus="holotype">
The
<typeStatus id="54E56283FFA6FFC16A35E7ECFE5C91E8" box="[368,466,980,1006]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
is a nearly complete exoskeleton (
<specimenCode id="DBF8745AFFA6FFC16813E7ECFC7791EB" box="[854,1017,979,1006]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM 62781</specimenCode>
; Pl. 1, figs 14) from the Nolichucky Formation at Shields Ridge,
<collectingCounty id="6280A4ADFFA6FFC16AA1E7C0FD269614" box="[484,680,1016,1042]" pageId="9" pageNumber="10">Jefferson County</collectingCounty>
,
<collectingRegion id="499A12C3FFA6FFC169F0E7C0FCA69614" box="[693,808,1016,1042]" country="United States of America" name="Tennessee" pageId="9" pageNumber="10">Tennessee</collectingRegion>
(
<collectionCode id="ED4F44E4FFA6FFC1687CE7C0FC029614" box="[825,908,1016,1042]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM</collectionCode>
loc. 120).
<typeStatus id="54E56283FFA6FFC16F48E7C0FBF49614" box="[1037,1146,1016,1042]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
are a cranidium (
<specimenCode id="DBF8745AFFA6FFC16E0CE7C0FF6F9633" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM 62873</specimenCode>
; Pl. 1, figs 57) and pygidium (
<specimenCode id="DBF8745AFFA6FFC1697AE024FD6C9633" box="[575,738,1051,1078]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM 62872</specimenCode>
; Pl. 1, figs 810) from the
<collectingMunicipality id="6B85465BFFA6FFC16F4EE024FA829630" box="[1035,1292,1052,1078]" pageId="9" pageNumber="10">Nolichucky Formation</collectingMunicipality>
, 17.7 km (
<quantity id="4CA671C4FFA6FFC16EC4E024FF5B965D" metricMagnitude="4" metricUnit="m" metricValue="1.7702784" pageId="9" pageNumber="10" unit="mi" value="11.0">11 miles</quantity>
) northwest of
<location id="8E818AFAFFA6FFC16A3DE078FE65965C" LSID="urn:lsid:plazi:treatment:03F76D37FFA6FFC26BD2E65AFDAC9678:8E818AFAFFA6FFC16A3DE078FE65965C" box="[376,491,1088,1114]" county="Jefferson County" municipality="Nolichucky Formation" name="Knoxville" pageId="9" pageNumber="10" stateProvince="Tennessee">Knoxville</location>
,
<location id="8E818AFAFFA6FFC16AB3E078FD01965C" LSID="urn:lsid:plazi:treatment:03F76D37FFA6FFC26BD2E65AFDAC9678:8E818AFAFFA6FFC16AB3E078FD01965C" box="[502,655,1088,1114]" county="Jefferson County" municipality="Nolichucky Formation" name="Knox County" pageId="9" pageNumber="10" stateProvince="Tennessee">Knox County</location>
,
<collectingRegion id="499A12C3FFA6FFC169DCE078FC82965C" box="[665,780,1088,1114]" country="United States of America" name="Tennessee" pageId="9" pageNumber="10">Tennessee</collectingRegion>
(
<collectionCode id="ED4F44E4FFA6FFC1685FE078FCE3965C" box="[794,877,1088,1114]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">USNM</collectionCode>
loc. 107c)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3448FAAFFA6FFC16B82E05AFD77954E" pageId="9" pageNumber="10" type="description">
<paragraph id="8BE1DC21FFA6FFC16B82E05AFBB49400" blockId="9.[151,1437,835,2043]" pageId="9" pageNumber="10">
<emphasis id="B92A0033FFA6FFC16B82E05AFED7967B" bold="true" box="[199,345,1122,1149]" pageId="9" pageNumber="10">Description.</emphasis>
Cranidium exclusive of posterolateral projection subrectangular in outline, width at palpebral lobe equal to 75% of length, with evenly rounded anterior margin. Frontal area long, occupying about one-quarter cranidial length (27%; 2529), preglabellar field comprises about 40% (39; 3642); gently slopes down with subtle increase in slope anteriorly to flat anterior border; anterior border furrow shallow. Moderately convex glabella subtrapezoidal in outline, narrowing slightly forward, with well-rounded anterior margin. Glabellar width at posterior end of palpebral lobe about 75% (74; 7078) its length. Glabella surface smooth, lacking lateral glabellar furrows and with only a hint of SO near axial furrow on some specimens (Pl. 1, fig. 1). Posterior margin of LO gently curved. Palpebral lobe short, equal to about one-tenth (12%; 1015) cranidial length, situated in front of glabellar mid-length. Anterior branches of facial suture diverge before curving along anterior cranidial margin; posterior branches extend at about 45 degrees, increasing curvature posteriorly. Posterolateral projections extend laterally with backwardly curved posterior margins, tapering to a sharp point. Posterior border furrow weakly expressed across two-thirds of posterior margin, becoming entirely effaced beyond that point.
</paragraph>
<paragraph id="8BE1DC21FFA6FFC16B82E228FADB9474" blockId="9.[151,1437,835,2043]" pageId="9" pageNumber="10">Thorax comprised of seven flat, transverse segments that taper to short, down-sloping pleural spines. Pleural furrow very faint but expressed near fulcrum. Axis is convex and raised above pleural lobes, occupies about one-third (31%) thoracic width at anterior, narrowing (tr.) posteriorly. Axial furrows shallow but clearly defined grooves.</paragraph>
<paragraph id="8BE1DC21FFA6FFC16B82E243FD77954E" blockId="9.[151,1437,835,2043]" pageId="9" pageNumber="10">Pygidium subcircular in outline, maximum length more than half of width (57%; 5064). Axis convex, raised above pleural field, width one-third (32%; 2133) maximum pygidial width (tr.). Axial furrows well-incised and shallow at axial tip. Faint axial ring furrows expressed at margin of axis in some individuals but effaced completely in others; at least seven axial rings present. Pleural field gently raised above down-sloping border, unfurrowed apart from barely perceptible pleural furrow at anterior. Lateral and posterior border furrows faint. Border length 15% maximum pygidial length. Entire exoskeleton smooth.</paragraph>
</subSubSection>
<subSubSection id="C3448FAAFFA6FFC26B82E369FDAC9678" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BE1DC21FFA6FFC16B82E369FD5295FD" blockId="9.[151,1437,835,2043]" pageId="9" pageNumber="10">
<emphasis id="B92A0033FFA6FFC16B82E369FEC2956A" bold="true" box="[199,332,1873,1900]" pageId="9" pageNumber="10">Discussion.</emphasis>
Walcotts (1916) types of
<taxonomicName id="4C5EA7A2FFA6FFC16932E36AFCBD956A" authorityName="Walcott" authorityYear="1916" box="[631,819,1874,1900]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA6FFC16932E36AFCBD956A" box="[631,819,1874,1900]" italics="true" pageId="9" pageNumber="10">Blountia mimula</emphasis>
</taxonomicName>
suggest that there may be size-related variation in glabellar width, which ranges from 64% of cranidial width at palpebral lobe in the
<typeStatus id="54E56283FFA6FFC16F49E34DFBE09589" box="[1036,1134,1909,1935]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
(Pl. 1, fig. 1) to 79% in the larger
<typeStatus id="54E56283FFA6FFC16B9AE3A2FEB195B2" box="[223,319,1946,1972]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
(Pl. 1. fig. 5). The anterior cranidial margin is less strongly curved in the
<typeStatus id="54E56283FFA6FFC16F30E3A2FB5795B2" box="[1141,1241,1946,1972]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
. The pygidia also show variation in proportions; the length of the pygidium of the
<typeStatus id="54E56283FFA6FFC16820E385FC4995D1" box="[869,967,1981,2007]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
is 50% of width (Pl. 1, figs 1, 3), whereas in the
<typeStatus id="54E56283FFA6FFC16B98E3D9FEB395FD" box="[221,317,2017,2043]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
(Pl. 1, fig. 9), the proportion is 64%.
</paragraph>
<paragraph id="8BE1DC21FFA5FFC26B82E4A1FDFB938C" blockId="10.[151,1437,152,1150]" pageId="10" pageNumber="11">
Sclerites in
<collectingRegion id="499A12C3FFA5FFC26A15E4A1FE4D92B5" box="[336,451,153,179]" country="United States of America" name="Tennessee" pageId="10" pageNumber="11">Tennessee</collectingRegion>
identified by
<bibRefCitation id="EFCFA1D0FFA5FFC26922E4A1FC8F92B5" author="Rasetti, F." box="[615,769,152,179]" pageId="10" pageNumber="11" pagination="1 - 127" refId="ref23892" refString="Rasetti, F. (1965) Upper Cambrian trilobite faunas of Northeastern Tennessee. Smithsonian Miscellaneous Collections, 148, 1 - 127." type="journal article" year="1965">Rasetti (1965</bibRefCitation>
, pl. 10, figs 37) as
<taxonomicName id="4C5EA7A2FFA5FFC268B1E4A1FB3C92B5" authorityName="Walcott" authorityYear="1916" box="[1012,1202,153,179]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC268B1E4A1FB3C92B5" box="[1012,1202,153,179]" italics="true" pageId="10" pageNumber="11">Blountia mimula</emphasis>
</taxonomicName>
do not appear to be conspecific with the
<typeStatus id="54E56283FFA5FFC26A38E485FE5192D1" box="[381,479,189,215]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
and represent a new species,
<taxonomicName id="4C5EA7A2FFA5FFC26860E486FC1292D1" authorityName="Westrop &amp; Eoff" authorityYear="2020" box="[805,924,189,215]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="angelae">
<emphasis id="B92A0033FFA5FFC26860E486FCB892D1" box="[805,822,190,215]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26806E485FC1292D1" box="[835,924,189,215]" italics="true" pageId="10" pageNumber="11">angelae</emphasis>
</taxonomicName>
(Pl. 15). These species are differentiated most clearly by pygidial morphology: the pygidium of
<taxonomicName id="4C5EA7A2FFA5FFC26997E4D9FCC292FC" authorityName="Westrop &amp; Eoff" authorityYear="2020" box="[722,844,224,251]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="angelae">
<emphasis id="B92A0033FFA5FFC26997E4D9FD6D92FC" box="[722,739,225,250]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC269B6E4D8FCC292FC" box="[755,844,224,250]" italics="true" pageId="10" pageNumber="11">angelae</emphasis>
</taxonomicName>
has a clearly defined border furrow on both testate surfaces (Pl. 15, fig. 6) and internal moulds (Pl. 15, figs 79), whereas
<taxonomicName id="4C5EA7A2FFA5FFC268F6E53DFBA89318" authorityName="Walcott" authorityYear="1916" box="[947,1062,260,287]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC268F6E53DFC4A9318" box="[947,964,261,286]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26897E53CFBA89318" box="[978,1062,260,286]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
is effaced (Pl. 1, figs 13. 9, 10). Cranidia of
<taxonomicName id="4C5EA7A2FFA5FFC26A58E511FE1B9344" authorityName="Westrop &amp; Eoff" authorityYear="2020" box="[285,405,296,322]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="angelae">
<emphasis id="B92A0033FFA5FFC26A58E511FEA09344" box="[285,302,297,322]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26A79E510FE1B9344" box="[316,405,296,322]" italics="true" pageId="10" pageNumber="11">angelae</emphasis>
</taxonomicName>
differ from
<taxonomicName id="4C5EA7A2FFA5FFC2695BE511FD1F9344" authorityName="Walcott" authorityYear="1916" box="[542,657,296,322]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC2695BE511FDA19344" box="[542,559,297,322]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26978E510FD1F9344" box="[573,657,296,322]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
in frontal area proportions: the anterior border is noticeably longer in the former (compare Pl. 1, figs 1, 5 with Pl. 15, figs 1, 4). In addition, the palpebral areas are wider in
<taxonomicName id="4C5EA7A2FFA5FFC26E61E574FA139360" authorityName="Westrop &amp; Eoff" authorityYear="2020" box="[1316,1437,332,358]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="angelae">
<emphasis id="B92A0033FFA5FFC26E61E574FABB9363" box="[1316,1333,332,357]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26E01E574FA139360" box="[1348,1437,332,358]" italics="true" pageId="10" pageNumber="11">angelae</emphasis>
</taxonomicName>
(compare Pl. 1, fig. 5 and Pl. 15, figs 1, 4).
</paragraph>
<paragraph id="8BE1DC21FFA5FFC26B82E5ACFEB990C8" blockId="10.[151,1437,152,1150]" pageId="10" pageNumber="11">
Cranidia of
<taxonomicName id="4C5EA7A2FFA5FFC26A13E5ACFD1093A8" authority="Resser, 1938" authorityName="Resser" authorityYear="1938" box="[342,670,404,430]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
<emphasis id="B92A0033FFA5FFC26A13E5ACFE7593A8" box="[342,507,404,430]" italics="true" pageId="10" pageNumber="11">B. bristolensis</emphasis>
<bibRefCitation id="EFCFA1D0FFA5FFC26943E5ACFD1093A8" author="Resser, C. E." box="[518,670,404,430]" pageId="10" pageNumber="11" pagination="1 - 140" refId="ref23917" refString="Resser, C. E. (1938) The Cambrian (Restricted) of the southern Appalachians. Geological Society of America Special Papers, 15, 1 - 140." type="journal article" year="1938">Resser, 1938</bibRefCitation>
</taxonomicName>
, including its synonyms
<taxonomicName id="4C5EA7A2FFA5FFC26883E5ACFAE693A8" authority="Resser, 1938" authorityName="Resser" authorityYear="1938" box="[966,1384,404,430]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
<emphasis id="B92A0033FFA5FFC26883E5ACFB4693A8" box="[966,1224,404,430]" italics="true" pageId="10" pageNumber="11">Maryvillia bristolensis</emphasis>
<bibRefCitation id="EFCFA1D0FFA5FFC26F91E5ACFAE693A8" author="Resser, C. E." box="[1236,1384,404,430]" pageId="10" pageNumber="11" pagination="1 - 140" refId="ref23917" refString="Resser, C. E. (1938) The Cambrian (Restricted) of the southern Appalachians. Geological Society of America Special Papers, 15, 1 - 140." type="journal article" year="1938">Resser, 1938</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C5EA7A2FFA5FFC26BD2E580FCCB93D4" authority="Lochman, in Lochman &amp; Duncan, 1944" authorityName="Lochman, in Lochman &amp; Duncan" authorityYear="1944" box="[151,837,440,466]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="nixonensis">
<emphasis id="B92A0033FFA5FFC26BD2E580FEF893D7" box="[151,374,440,466]" italics="true" pageId="10" pageNumber="11">Blountia nixonensis</emphasis>
Lochman, in
<bibRefCitation id="EFCFA1D0FFA5FFC26953E580FCCB93D4" author="Lochman, C. &amp; Duncan, D." box="[534,837,440,466]" pageId="10" pageNumber="11" pagination="1 - 172" refId="ref23573" refString="Lochman, C. &amp; Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman &amp; Duncan, 1944</bibRefCitation>
</taxonomicName>
(see below), have anterior branches of the facial sutures that are subparallel, or nearly so (e.g., Pl. 2, Pl. 3, fig. 5), whereas as those of
<taxonomicName id="4C5EA7A2FFA5FFC26F13E5E4FB4593F0" authorityName="Walcott" authorityYear="1916" box="[1110,1227,476,502]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26F13E5E4FBE993F3" box="[1110,1127,476,501]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26F32E5E4FB4593F0" box="[1143,1227,476,502]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
are more strongly diveregent (e.g., Pl. 1, fig. 5). The frontal area of
<taxonomicName id="4C5EA7A2FFA5FFC26984E638FCBA901C" authorityName="Walcott" authorityYear="1916" box="[705,820,512,538]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26984E638FD5C901F" box="[705,722,512,537]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC269A5E638FCBA901C" box="[736,820,512,538]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
is longer (29% total cranidial length compared to 23% in
<taxonomicName id="4C5EA7A2FFA5FFC26BF1E61CFEDB9038" authorityName="Resser" authorityYear="1938" box="[180,341,548,574]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
<emphasis id="B92A0033FFA5FFC26BF1E61CFF4B903B" box="[180,197,548,573]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26B91E61CFEDB9038" box="[212,341,548,574]" italics="true" pageId="10" pageNumber="11">bristolensis</emphasis>
</taxonomicName>
), and the palpebral area of the fixigena is distinctly narrower (compare Pls 2, 4 with Pl. 1, figs 1, 5). Pygidial contrasts are even stronger, with the firmly impressed border furrows of
<taxonomicName id="4C5EA7A2FFA5FFC26F0FE670FB649064" authorityName="Resser" authorityYear="1938" box="[1098,1258,584,610]" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
<emphasis id="B92A0033FFA5FFC26F0FE670FBD59067" box="[1098,1115,584,609]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26F2CE670FB649064" box="[1129,1258,584,610]" italics="true" pageId="10" pageNumber="11">bristolensis</emphasis>
</taxonomicName>
(Pl. 3, figs 1, 2, Pl. 5) clearly differing from the effaced condition of
<taxonomicName id="4C5EA7A2FFA5FFC2699AE654FCDF9080" authorityName="Walcott" authorityYear="1916" box="[735,849,620,646]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC2699AE654FD7E9083" box="[735,752,620,645]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC269B8E654FCDF9080" box="[765,849,620,646]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
. The articulated specimens of both
<taxonomicName id="4C5EA7A2FFA5FFC26F98E654FAC19080" authorityName="Walcott" authorityYear="1916" box="[1245,1359,620,646]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26F98E654FB609083" box="[1245,1262,620,645]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26FBEE654FAC19080" box="[1275,1359,620,646]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5EA7A2FFA5FFC26EC1E654FE9690AC" authorityName="Resser" authorityYear="1938" class="Trilobita" family="Kingstoniidae" genus="Maryvillia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bristolensis">
<emphasis id="B92A0033FFA5FFC26EC1E654FA1B9083" box="[1412,1429,620,645]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26BD2E6A8FE9690AC" box="[151,280,656,682]" italics="true" pageId="10" pageNumber="11">bristolensis</emphasis>
</taxonomicName>
have seven thoracic segments, but the latter species has a wider axis (38% of thorax width versus 30% in
<taxonomicName id="4C5EA7A2FFA5FFC26BF1E68CFEA990C8" authorityName="Walcott" authorityYear="1916" box="[180,295,692,718]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26BF1E68CFF4B90CB" box="[180,197,692,717]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26B96E68CFEA990C8" box="[211,295,692,718]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BE1DC21FFA5FFC26B82E6E0FE3A9614" blockId="10.[151,1437,152,1150]" pageId="10" pageNumber="11">
Pygidia of
<taxonomicName id="4C5EA7A2FFA5FFC26A06E6E0FCEE90F4" authority="Lochman, in Lochman &amp; Duncan, 1944" authorityName="Lochman, in Lochman &amp; Duncan" authorityYear="1944" box="[323,864,728,754]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cora">
<emphasis id="B92A0033FFA5FFC26A06E6E0FEDA90F7" box="[323,340,728,753]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26A27E6E0FE1B90F7" box="[354,405,728,753]" italics="true" pageId="10" pageNumber="11">cora</emphasis>
Lochman, in
<bibRefCitation id="EFCFA1D0FFA5FFC26977E6E0FCEE90F4" author="Lochman, C. &amp; Duncan, D." box="[562,864,728,754]" pageId="10" pageNumber="11" pagination="1 - 172" refId="ref23573" refString="Lochman, C. &amp; Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman &amp; Duncan, 1944</bibRefCitation>
</taxonomicName>
, from the Pilgrim Formation of
<collectingRegion id="499A12C3FFA5FFC26F8CE6E0FAA190F4" box="[1225,1327,728,754]" country="United States of America" name="Montana" pageId="10" pageNumber="11">Montana</collectingRegion>
, show effacement of the border furrow that resembles the condition in
<taxonomicName id="4C5EA7A2FFA5FFC2680BE6C4FC4F9110" authorityName="Walcott" authorityYear="1916" box="[846,961,764,790]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC2680BE6C4FCD19113" box="[846,863,764,789]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26828E6C4FC4F9110" box="[877,961,764,790]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
(compare Pl. 1, figs 3, 9 with Pl. 6, figs 8, 9 and Pl. 7, figs 9, 10, 14). If the
<typeStatus id="54E56283FFA5FFC2694AE718FDF4913C" box="[527,634,800,826]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
are assigned correctly, the pygidium of
<taxonomicName id="4C5EA7A2FFA5FFC26F05E718FB1C913F" authorityName="Lochman, in Lochman &amp; Duncan" authorityYear="1944" box="[1088,1170,800,826]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cora">
<emphasis id="B92A0033FFA5FFC26F05E718FBDF913F" box="[1088,1105,800,825]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26F1AE718FB1C913F" box="[1119,1170,800,825]" italics="true" pageId="10" pageNumber="11">cora</emphasis>
</taxonomicName>
lengthens considerably during holaspid ontogeny (e.g. compare Pl. 6, fig. 9 and Pl. 7, fig. 9). At a similar size, pygidia of
<taxonomicName id="4C5EA7A2FFA5FFC26FB8E77CFAFC9158" authorityName="Walcott" authorityYear="1916" box="[1277,1394,836,862]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26FB8E77CFA80915B" box="[1277,1294,836,861]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26E5BE77CFAFC9158" box="[1310,1394,836,862]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
are relatively longer (e.g., compare Pl. 1, fig. 3 and Pl. 6, fig. 9). Cranidia of
<taxonomicName id="4C5EA7A2FFA5FFC2688EE750FB939187" authorityName="Lochman, in Lochman &amp; Duncan" authorityYear="1944" box="[971,1053,872,898]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cora">
<emphasis id="B92A0033FFA5FFC2688EE750FC529187" box="[971,988,872,897]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC268AFE750FB939187" box="[1002,1053,872,897]" italics="true" pageId="10" pageNumber="11">cora</emphasis>
</taxonomicName>
also show substantial ontogenetic changes in proportions, with strong positive allometry of the frontal area (e.g., compare Pl. 6, figs 17 and Pl. 7, figs 1113). However similarly-sized cranidia of
<taxonomicName id="4C5EA7A2FFA5FFC269D5E788FD6D91CF" authorityName="Lochman, in Lochman &amp; Duncan" authorityYear="1944" box="[656,739,944,969]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cora">
<emphasis id="B92A0033FFA5FFC269D5E788FD6D91CF" box="[656,739,944,969]" italics="true" pageId="10" pageNumber="11">B. cora</emphasis>
</taxonomicName>
(Pl. 6, figs 4, 6) and
<taxonomicName id="4C5EA7A2FFA5FFC26896E788FBC991CC" authorityName="Walcott" authorityYear="1916" box="[979,1095,944,970]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26896E788FC6A91CF" box="[979,996,944,969]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC268B6E788FBC991CC" box="[1011,1095,944,970]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
(Pl. 1, fig. 5) are dramatically different, with the latter having a frontal area that is less than half of the relative length of the former, and a much shorter preglabellar field.
</paragraph>
<paragraph id="8BE1DC21FFA5FFC26B82E024FDAC9678" blockId="10.[151,1437,152,1150]" pageId="10" pageNumber="11">
Pygidial anatomy also separates
<taxonomicName id="4C5EA7A2FFA5FFC2697FE024FBEB9630" authority="Lochman, in Lochman &amp; Duncan, 1944" authorityName="Lochman, in Lochman &amp; Duncan" authorityYear="1944" box="[570,1125,1052,1078]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="janei">
<emphasis id="B92A0033FFA5FFC2697FE024FDC59633" box="[570,587,1052,1077]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC2691EE024FD1D9633" box="[603,659,1052,1077]" italics="true" pageId="10" pageNumber="11">janei</emphasis>
Lochman, in
<bibRefCitation id="EFCFA1D0FFA5FFC26876E024FBEB9630" author="Lochman, C. &amp; Duncan, D." box="[819,1125,1052,1078]" pageId="10" pageNumber="11" pagination="1 - 172" refId="ref23573" refString="Lochman, C. &amp; Duncan, D. (1944) Early Upper Cambrian faunas of Central Montana. Geological Society of America Special Papers, 54, 1 - 172. https: // doi. org / 10.1130 / SPE 54 - p 1" type="journal article" year="1944">Lochman &amp; Duncan, 1944</bibRefCitation>
</taxonomicName>
, also from the Pilgrim Formation, from
<taxonomicName id="4C5EA7A2FFA5FFC26A71E078FE27965C" authorityName="Walcott" authorityYear="1916" box="[308,425,1088,1114]" class="Trilobita" family="Kingstoniidae" genus="Blountia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ptychopariida" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="mimula">
<emphasis id="B92A0033FFA5FFC26A71E078FECB965F" box="[308,325,1088,1113]" italics="true" pageId="10" pageNumber="11">B</emphasis>
.
<emphasis id="B92A0033FFA5FFC26A10E078FE27965C" box="[341,425,1088,1114]" italics="true" pageId="10" pageNumber="11">mimula</emphasis>
</taxonomicName>
. The pygidium of the former is proportionately longer and has a shorter axis throughout holaspid ontogeny (Pl. 7, figs 18).
</paragraph>
</subSubSection>
</treatment>
</document>