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<document id="BAB4660CFEF03C7D19926A70E2AEDBC5" ID-CLB-Dataset="54010" ID-DOI="10.5852/ejt.2023.873.2133" ID-GBIF-Dataset="6b60ec14-d9a3-4400-b636-bb3ee16094b7" ID-ISSN="2118-9773" ID-Zenodo-Dep="8038022" ID-ZooBank="099F6A49-6516-4F39-9E20-A077A5D60535" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1686745931301" checkinUser="felipe" docAuthor="Pham, Nhi Thi, Pham, Phu Van, Matsumoto, Rikio, Shimizu, So &amp; Broad, Gavin R." docDate="2023" docId="03E3AC44FF9CFFEE9712FEE00686893E" docLanguage="en" docName="ejt-873_1-151.pdf" docOrigin="European Journal of Taxonomy 873 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.873.2133" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Enicospilus nigriventris Nikam 1975" docType="treatment" docVersion="8" lastPageNumber="101" masterDocId="FFDAD43CFFFFFF8A952BFFE8073A8A55" masterDocTitle="A review of the genus Enicospilus Stephens (Ichneumonidae: Ophioninae) from Vietnam, with descriptions of ten new species" masterLastPageNumber="151" masterPageNumber="1" pageNumber="100" updateTime="1699524284659" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EE671C6752F93D54440D4EB81C7B6E40">A review of the genus Enicospilus Stephens (Ichneumonidae: Ophioninae) from Vietnam, with descriptions of ten new species</mods:title>
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<mods:namePart id="A07E24CC9021E819189C0F5C7BB91DEB">Pham, Nhi Thi</mods:namePart>
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<mods:affiliation id="BB740E2067AD78C3DBF188F64C8E54E1">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. Osaka Museum of Natural History, Nagai Park 1 - 23, Higashisumiyoshi-ku, Osaka, Japan. Institute for Agro-Environmental Sciences, NARO, Kannondai 3 - 1 - 3, Tsukuba 305 - 8604, Japan. Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Kobe, Hyogo, Japan. The Natural History Museum, Cromwell Road, London SW 7 BD, UK.</mods:affiliation>
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<mods:affiliation id="69E201970291E9D5BFE1BD028D62CE73">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam. Osaka Museum of Natural History, Nagai Park 1 - 23, Higashisumiyoshi-ku, Osaka, Japan. Institute for Agro-Environmental Sciences, NARO, Kannondai 3 - 1 - 3, Tsukuba 305 - 8604, Japan. Laboratory of Insect Biodiversity and Ecosystem Science, Graduate School of Agricultural Science, Kobe University, Kobe, Hyogo, Japan. The Natural History Museum, Cromwell Road, London SW 7 BD, UK.</mods:affiliation>
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<taxonomicName id="4C4A66D1FF9CFFE99712FEE004C18B71" ID-CoL="6FCBN" authority="Nikam, 1975" authorityName="Nikam" authorityYear="1975" box="[569,1019,264,292]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="B93EC140FF9CFFE99712FEE004608B76" bold="true" box="[569,858,264,291]" italics="true" pageId="99" pageNumber="100">Enicospilus nigriventris</emphasis>
<bibRefCitation id="EFDB60A3FF9CFFE9964AFEE104C18B71" author="Nikam R. K." box="[865,1019,265,292]" pageId="99" pageNumber="100" pagination="193 - 202" refId="ref57945" refString="Nikam R. K. 1975. Studies on Indian Ichneumonidae. Four new species of Enicospilus Stephens (Ophioninae) from Marathwada. Marathwada University Journal of Science 14 (7): 193 - 202." type="journal article" year="1975">Nikam, 1975</bibRefCitation>
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<figureCitation id="137101D7FF9CFFE997DBFEC204798B10" box="[752,835,298,325]" captionStart="Fig" captionStartId="99.[189,231,1963,1989]" captionTargetBox="[189,1399,506,1924]" captionTargetId="figure-37@99.[189,1399,503,1928]" captionTargetPageId="99" captionText="Fig. 55. Enicospilus nigriventris Nikam, 1975, ♂ (IEBR). a. Habitus. b. Head, frontal view. c. Head, dorsal view. d. Head, lateral view. e. Mesosoma, lateral view. f. Central part of fore wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8038229" httpUri="https://zenodo.org/record/8038229/files/figure.png" pageId="99" pageNumber="100">Fig. 55</figureCitation>
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<emphasis id="B93EC140FF9CFFE99596FE82067C8BD1" box="[189,326,362,388]" italics="true" pageId="99" pageNumber="100">Enicospilus</emphasis>
(
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<emphasis id="B93EC140FF9CFFE994CDFE8305558BD1" box="[486,623,363,388]" italics="true" pageId="99" pageNumber="100">nigriventris</emphasis>
<bibRefCitation id="EFDB60A3FF9CFFE9975DFE8204778BD0" author="Nikam R. K." box="[630,845,362,389]" pageId="99" pageNumber="100" pagination="193 - 202" refId="ref57945" refString="Nikam R. K. 1975. Studies on Indian Ichneumonidae. Four new species of Enicospilus Stephens (Ophioninae) from Marathwada. Marathwada University Journal of Science 14 (7): 193 - 202." type="journal article" year="1975">Nikam, 1975: 195</bibRefCitation>
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;
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(
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<taxonomicName id="4C4A66D1FF9CFFE99596FE4106EC8B96" ID-CoL="6FCBN" authorityName="Nikam" authorityYear="1975" box="[189,470,425,451]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="B93EC140FF9CFFE99596FE4106EC8B96" box="[189,470,425,451]" italics="true" pageId="99" pageNumber="100">Enicospilus nigriventris</emphasis>
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<treatmentCitation id="0AEB3B43FF9CFFE994D8FE4105FE8B91" author="Nikam R. K." box="[499,708,425,452]" page="197" pageId="99" pageNumber="100" year="1980">
<bibRefCitation id="EFDB60A3FF9CFFE994D8FE4105FE8B91" author="Nikam R. K." box="[499,708,425,452]" pageId="99" pageNumber="100" pagination="131 - 219" refId="ref57983" refString="Nikam R. K. 1980. Studies on Indian species of Enicospilus Stephens (Hymenoptera: Ichneumonidae). Oriental Insects 14: 131 - 219. https: // doi. org / 10.1080 / 00305316.1980.10433632" type="journal article" year="1980">Nikam 1980: 197</bibRefCitation>
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.
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<paragraph id="8BF51D52FF9CFFE99596F843024D8DBC" blockId="99.[189,1399,1963,2025]" pageId="99" pageNumber="100">
<emphasis id="B93EC140FF9CFFE99596F843062F8D90" bold="true" box="[189,277,1963,1989]" pageId="99" pageNumber="100">Fig. 55.</emphasis>
<taxonomicName id="4C4A66D1FF9CFFE99430F84305F38D93" authority="Nikam, 1975" authorityName="Nikam" authorityYear="1975" box="[283,713,1963,1990]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="nigriventris">
<emphasis id="B93EC140FF9CFFE99430F84305178D90" box="[283,557,1963,1989]" italics="true" pageId="99" pageNumber="100">Enicospilus nigriventris</emphasis>
<bibRefCitation id="EFDB60A3FF9CFFE99719F84305F38D93" author="Nikam R. K." box="[562,713,1963,1990]" pageId="99" pageNumber="100" pagination="193 - 202" refId="ref57945" refString="Nikam R. K. 1975. Studies on Indian Ichneumonidae. Four new species of Enicospilus Stephens (Ophioninae) from Marathwada. Marathwada University Journal of Science 14 (7): 193 - 202." type="journal article" year="1975">Nikam, 1975</bibRefCitation>
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, ♂ (IEBR).
<emphasis id="B93EC140FF9CFFE9967BF84304648D90" bold="true" box="[848,862,1963,1989]" pageId="99" pageNumber="100">a</emphasis>
. Habitus.
<emphasis id="B93EC140FF9CFFE996F9F84304D88D90" bold="true" box="[978,994,1963,1989]" pageId="99" pageNumber="100">b</emphasis>
. Head, frontal view.
<emphasis id="B93EC140FF9CFFE991E6F84303E08D90" bold="true" box="[1229,1242,1963,1989]" pageId="99" pageNumber="100">c</emphasis>
. Head, dorsal view.
<emphasis id="B93EC140FF9CFFE99429F82606288DBD" bold="true" box="[258,274,1998,2024]" pageId="99" pageNumber="100">d</emphasis>
. Head, lateral view.
<emphasis id="B93EC140FF9CFFE9972BF82605378DBD" bold="true" box="[512,525,1998,2024]" pageId="99" pageNumber="100">e</emphasis>
. Mesosoma, lateral view.
<emphasis id="B93EC140FF9CFFE99617F826047C8DBD" bold="true" box="[828,838,1998,2024]" pageId="99" pageNumber="100">f</emphasis>
. Central part of fore wing. Scale bars = 1.0 mm.
</paragraph>
</caption>
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<emphasis id="B93EC140FF9BFFEE9596FEE0060C8B77" bold="true" box="[189,310,264,290]" pageId="100" pageNumber="101">Diagnosis</emphasis>
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<paragraph id="8BF51D52FF9BFFEE9596FED1024D8B91" blockId="100.[189,1399,313,453]" pageId="100" pageNumber="101">Interocellar area reddish brown; clypeus moderately convex, ventral margin acute; mandible twisted ca 75°, outer surface without a diagonal setose groove; body reddish with black markings, fore wing fenestra with proximal, central and distal sclerites present, central sclerite large, C-shaped, formed from the antero-distal periphery to almost ventral margin of an extensive quadra; mesoscutum posteriorly black.</paragraph>
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<emphasis id="B93EC140FF9BFFEE9596FE060690885D" bold="true" box="[189,426,494,520]" pageId="100" pageNumber="101">Material examined</emphasis>
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</paragraph>
<paragraph id="8BF51D52FF9BFFEE9596FDC805018884" blockId="100.[189,1399,543,721]" pageId="100" pageNumber="101">
<materialsCitation id="3B22170FFF9BFFEE9596FDC8068E8835" ID-GBIF-Occurrence="4130444614" collectingDate="2017-09-21" collectingMethod="light trap" collectionCode="IEBR" collectorName="Nguyen Q. C. &amp; Nguyen T. P. L." country="Vietnam" location="Kon Chu Rang NP" pageId="100" pageNumber="101" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai Province">
<collectingCountry id="F35D5DC2FF9BFFEE9596FDC8067E886F" box="[189,324,544,570]" name="Vietnam" pageId="100" pageNumber="101">VIETNAM</collectingCountry>
<specimenCount id="9D4CD6DBFF9BFFEE9477FDC806B2886F" box="[348,392,544,570]" pageId="100" pageNumber="101" type="female">1 ♀</specimenCount>
;
<collectingRegion id="498ED3B0FF9BFFEE94BDFDF7055A886C" box="[406,608,543,569]" country="Vietnam" name="Gia Lai" pageId="100" pageNumber="101">Gia Lai Province</collectingRegion>
,
<location id="8E954B89FF9BFFEE9740FDC80473886F" LSID="urn:lsid:plazi:treatment:03E3AC44FF9CFFEE9712FEE00686893E:8E954B89FF9BFFEE9740FDC80473886F" box="[619,841,543,570]" country="Vietnam" name="Kon Chu Rang NP" pageId="100" pageNumber="101" stateProvince="Gia Lai Province">Kon Chu Rang NP</location>
;
<date id="FFF43B92FF9BFFEE967EFDC804CB886F" box="[853,1009,544,570]" pageId="100" pageNumber="101" value="2017-09-21">
<collectingDate id="EFB0C27AFF9BFFEE967EFDC804CB886F" box="[853,1009,544,570]" pageId="100" pageNumber="101" value="2017-09-21">21 Sep. 2017</collectingDate>
</date>
;
<collectorName id="26BF7884FF9BFFEE96D7FDC803AD886F" box="[1020,1175,544,570]" pageId="100" pageNumber="101">Nguyen Q.C.</collectorName>
and
<collectorName id="26BF7884FF9BFFEE91E4FDC8024D886F" box="[1231,1399,544,570]" pageId="100" pageNumber="101">Nguyen T.P.L.</collectorName>
leg.;
<collectingMethod id="520B6545FF9BFFEE95DCFDAD065F8835" box="[247,357,581,608]" pageId="100" pageNumber="101">light trap</collectingMethod>
;
<collectionCode id="ED5B8597FF9BFFEE945AFDAE068E8835" box="[369,436,582,608]" pageId="100" pageNumber="101">IEBR</collectionCode>
</materialsCitation>
<materialsCitation id="3B22170FFF9BFFEE94E6FDAD037888D0" ID-GBIF-Occurrence="4130444774" collectingDate="2021-06-09" collectingMethod="light trap" collectionCode="IEBR" collectorName="Pham T. N. &amp; Hoang V. T. &amp; Pham V. P." country="Vietnam" elevation="95" latitude="11.347466" location="Phu Ly" longLatPrecision="1" longitude="107.1419" municipality="Vinh Cuu" pageId="100" pageNumber="101" specimenCount="1" specimenCount-male="1" stateProvince="Dong Nai Province">
<specimenCount id="9D4CD6DBFF9BFFEE94E6FDAD06C18835" box="[461,507,581,608]" pageId="100" pageNumber="101" type="male">1 ♂</specimenCount>
;
<collectingRegion id="498ED3B0FF9BFFEE9723FDAE05D6880A" box="[520,748,581,608]" country="Vietnam" name="Dong Nai" pageId="100" pageNumber="101">Dong Nai Province</collectingRegion>
,
<collectingMunicipality id="6B918728FF9BFFEE97D3FDAD0456880A" box="[760,876,581,607]" pageId="100" pageNumber="101">Vinh Cuu</collectingMunicipality>
,
<location id="8E954B89FF9BFFEE965CFDAD04F58835" LSID="urn:lsid:plazi:treatment:03E3AC44FF9CFFEE9712FEE00686893E:8E954B89FF9BFFEE965CFDAD04F58835" box="[887,975,581,608]" country="Vietnam" latitude="11.347466" longLatPrecision="1" longitude="107.1419" municipality="Vinh Cuu" name="Phu Ly" pageId="100" pageNumber="101" stateProvince="Dong Nai Province">Phu Ly</location>
;
<geoCoordinate id="EE7E7B95FF9BFFEE96F1FDAB03428835" box="[986,1144,579,608]" degrees="11" direction="north" minutes="20.848" orientation="latitude" pageId="100" pageNumber="101" precision="1" value="11.347466">11°20.848 N</geoCoordinate>
,
<geoCoordinate id="EE7E7B95FF9BFFEE91AFFDAB02158835" box="[1156,1327,579,608]" degrees="107" direction="east" minutes="08.514" orientation="longitude" pageId="100" pageNumber="101" precision="1" value="107.1419">107°08.514 E</geoCoordinate>
;
<elevation id="0067FA61FF9BFFEE9010FDAD07CF88D0" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="100" pageNumber="101" unit="m" value="95.0">
<quantity id="4CB2B0B7FF9BFFEE9010FDAD024D8835" box="[1339,1399,581,608]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="100" pageNumber="101" unit="m" value="95.0">95 m</quantity>
a.s.l.
</elevation>
;
<date id="FFF43B92FF9BFFEE942AFD8306B388D0" box="[257,393,619,645]" pageId="100" pageNumber="101" value="2021-06-09">
<collectingDate id="EFB0C27AFF9BFFEE942AFD8306B388D0" box="[257,393,619,645]" pageId="100" pageNumber="101" value="2021-06-09">9 Jun. 2021</collectingDate>
</date>
;
<collectorName id="26BF7884FF9BFFEE94BFFD83052B88D0" box="[404,529,619,645]" pageId="100" pageNumber="101">Pham T.N.</collectorName>
,
<collectorName id="26BF7884FF9BFFEE9737FD83059A88D0" box="[540,672,619,645]" pageId="100" pageNumber="101">Hoang V.T.</collectorName>
and
<collectorName id="26BF7884FF9BFFEE97FEFD83047088D0" box="[725,842,619,645]" pageId="100" pageNumber="101">Pham V.P.</collectorName>
leg.;
<collectingMethod id="520B6545FF9BFFEE96A2FD8304CF88D0" box="[905,1013,619,645]" pageId="100" pageNumber="101">light trap</collectingMethod>
;
<collectionCode id="ED5B8597FF9BFFEE96D4FD83037888D0" box="[1023,1090,619,645]" pageId="100" pageNumber="101">IEBR</collectionCode>
</materialsCitation>
<materialsCitation id="3B22170FFF9BFFEE9173FD83050D8884" ID-GBIF-Occurrence="4130444550" collectingDate="2021-06-11" collectingMethod="light trap" collectionCode="IEBR" collectorName="Pham T. N. &amp; Hoang V. T. &amp; Pham V. P." country="Vietnam" elevation="178" latitude="11.44195" location="Phu Ly" longLatPrecision="1" longitude="107.103035" municipality="Vinh Cuu" pageId="100" pageNumber="101" specimenCount="1" specimenCount-female="1" stateProvince="Dong Nai Province">
<specimenCount id="9D4CD6DBFF9BFFEE9173FD8303B988D0" box="[1112,1155,619,645]" pageId="100" pageNumber="101" type="female">1 ♀</specimenCount>
;
<collectingRegion id="498ED3B0FF9BFFEE91BBFD83024888D0" box="[1168,1394,619,645]" country="Vietnam" name="Dong Nai" pageId="100" pageNumber="101">Dong Nai Province</collectingRegion>
,
<collectingMunicipality id="6B918728FF9BFFEE9596FD78060B88FE" box="[189,305,656,683]" pageId="100" pageNumber="101">Vinh Cuu</collectingMunicipality>
,
<location id="8E954B89FF9BFFEE9417FD7806A988FE" LSID="urn:lsid:plazi:treatment:03E3AC44FF9CFFEE9712FEE00686893E:8E954B89FF9BFFEE9417FD7806A988FE" box="[316,403,656,683]" country="Vietnam" latitude="11.44195" longLatPrecision="1" longitude="107.103035" municipality="Vinh Cuu" name="Phu Ly" pageId="100" pageNumber="101" stateProvince="Dong Nai Province">Phu Ly</location>
;
<geoCoordinate id="EE7E7B95FF9BFFEE94B5FD67050688FE" box="[414,572,655,683]" degrees="11" direction="north" minutes="26.517" orientation="latitude" pageId="100" pageNumber="101" precision="1" value="11.44195">11°26.517 N</geoCoordinate>
,
<geoCoordinate id="EE7E7B95FF9BFFEE976CFD6705CB88FE" box="[583,753,655,683]" degrees="107" direction="east" minutes="06.182" orientation="longitude" pageId="100" pageNumber="101" precision="1" value="107.103035">107°06.182 E</geoCoordinate>
;
<elevation id="0067FA61FF9BFFEE97D6FD7904BC88FF" box="[765,902,656,683]" metricMagnitude="2" metricUnit="m" metricValue="1.78" pageId="100" pageNumber="101" unit="m" value="178.0">
<quantity id="4CB2B0B7FF9BFFEE97D6FD79047D88FE" box="[765,839,657,683]" metricMagnitude="2" metricUnit="m" metricValue="1.78" pageId="100" pageNumber="101" unit="m" value="178.0">178 m</quantity>
a.s.l.
</elevation>
;
<date id="FFF43B92FF9BFFEE96B8FD79031188FE" box="[915,1067,657,683]" pageId="100" pageNumber="101" value="2021-06-11">
<collectingDate id="EFB0C27AFF9BFFEE96B8FD79031188FE" box="[915,1067,657,683]" pageId="100" pageNumber="101" value="2021-06-11">11 Jun. 2021</collectingDate>
</date>
;
<collectorName id="26BF7884FF9BFFEE911DFD78038F88FE" box="[1078,1205,656,683]" pageId="100" pageNumber="101">Pham T.N.</collectorName>
,
<collectorName id="26BF7884FF9BFFEE91EAFD79027C88FE" box="[1217,1350,657,683]" pageId="100" pageNumber="101">Hoang V.T.</collectorName>
and
<collectorName id="26BF7884FF9BFFEE9596FD5E060E8884" box="[189,308,694,721]" pageId="100" pageNumber="101">Pham V.P.</collectorName>
leg.;
<collectingMethod id="520B6545FF9BFFEE945DFD5E06DE8884" box="[374,484,694,721]" pageId="100" pageNumber="101">light trap</collectingMethod>
;
<collectionCode id="ED5B8597FF9BFFEE94DBFD5F050D8884" box="[496,567,695,721]" pageId="100" pageNumber="101">IEBR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3504ED9FF9BFFEE9596FD120686893E" pageId="100" pageNumber="101" type="distribution">
<paragraph id="8BF51D52FF9BFFEE9596FD12066C8941" blockId="100.[189,342,762,788]" box="[189,342,762,788]" pageId="100" pageNumber="101">
<heading id="D0BDAA3EFF9BFFEE9596FD12066C8941" bold="true" box="[189,342,762,788]" fontSize="11" level="3" pageId="100" pageNumber="101" reason="3">
<emphasis id="B93EC140FF9BFFEE9596FD12066C8941" bold="true" box="[189,342,762,788]" pageId="100" pageNumber="101">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BF51D52FF9BFFEE9596FCC30686893E" blockId="100.[189,1398,811,875]" pageId="100" pageNumber="101">
Previously known from
<collectingCountry id="F35D5DC2FF9BFFEE94F2FCC305168910" box="[473,556,811,837]" name="Brunei" pageId="100" pageNumber="101">Brunei</collectingCountry>
,
<collectingCountry id="F35D5DC2FF9BFFEE971CFCC3054C8910" box="[567,630,811,837]" name="India" pageId="100" pageNumber="101">India</collectingCountry>
, and
<collectingCountry id="F35D5DC2FF9BFFEE9798FCC3040D8910" box="[691,823,811,837]" name="Philippines" pageId="100" pageNumber="101">Philippines</collectingCountry>
(
<bibRefCitation id="EFDB60A3FF9BFFEE966EFCC304CB8910" author="Yu D. S. &amp; van Achterberg C. &amp; Horstmann K." box="[837,1009,811,837]" pageId="100" pageNumber="101" refId="ref59419" refString="Yu D. S., van Achterberg C. &amp; Horstmann K. 2016. Taxapad 2016, Ichneumonoidea 2015. Database on flash-drive. Taxapad, Ottawa, Ontario." type="book" year="2016">
Yu
<emphasis id="B93EC140FF9BFFEE9644FCC404908910" box="[879,938,811,837]" italics="true" pageId="100" pageNumber="101">et al.</emphasis>
2016
</bibRefCitation>
). These are the first record of this species from
<collectingCountry id="F35D5DC2FF9BFFEE9472FCB90686893E" box="[345,444,849,875]" name="Vietnam" pageId="100" pageNumber="101">Vietnam</collectingCountry>
</paragraph>
</subSubSection>
</treatment>
</document>