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<document id="E2B861BB647EFC808BA4ADBA79C4CEEA" ID-CLB-Dataset="3383" ID-DOI="10.1093/zoolinnean/zly037" ID-GBIF-Dataset="d90d12b0-99f7-4d30-8eea-1f5364bf490c" ID-ISSN="0024-4082" ID-Zenodo-Dep="5715063" ID-ZooBank="03DAADC2-E954-42C5-B44D-55927DF55837" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619628054066" checkinUser="carolina" docAuthor="Geroto, Caio Fabricio Cezar &amp; Bertini, Reinaldo J." docDate="2019" docId="03C35358FFC2936BFEA43CCCFB789425" docLanguage="en" docName="ZoolJLinnSoc.185.312-334.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 185" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Pepesuchinae Geroto &amp; Bertini 2019, SUBFAM. NOV." docType="treatment" docVersion="8" lastPageNumber="331" masterDocId="FFFA2B20FFD09378FF873E3FFFBE9013" masterDocTitle="New material of Pepesuchus (Crocodyliformes; Mesoeucrocodylia) from the Bauru Group: implications about its phylogeny and the age of the Adamantina Formation" masterLastPageNumber="334" masterPageNumber="312" pageNumber="330" updateTime="1732783493118" updateUser="ExternalLinkService">
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<mods:title id="932F797C855701EF59C471ECDC37171D">New material of Pepesuchus (Crocodyliformes; Mesoeucrocodylia) from the Bauru Group: implications about its phylogeny and the age of the Adamantina Formation</mods:title>
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<mods:namePart id="036AD870FF2352BD0B42AD0C0DF440D2">Geroto, Caio Fabricio Cezar</mods:namePart>
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<mods:namePart id="3FD8A463B522FBEDEADFE10BC37BC85E">Bertini, Reinaldo J.</mods:namePart>
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<mods:title id="719AC4B86D06BB96A6F775D0516247C1">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="09283C5894799DEC66D015956A3DD548">2019</mods:date>
<mods:detail id="43FF36AB10A8E7BA5610CD02557CA777" type="pubDate">
<mods:number id="0E3DA003FF94E468DE4F600F65EA92C0">2018-07-02</mods:number>
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<mods:detail id="2FCE7A006F86157C727C0816B3108D2E" type="volume">
<mods:number id="5245859843C6365194FE3AA952D50F0E">185</mods:number>
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<mods:classification id="0332FD32D5EE492E7647902E3962BC21">journal article</mods:classification>
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<mods:identifier id="3A2807F59A968DFCF86719C6EC8073FA" type="DOI">10.1093/zoolinnean/zly037</mods:identifier>
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<treatment id="03C35358FFC2936BFEA43CCCFB789425" ID-DOI="http://doi.org/10.5281/zenodo.5730280" ID-GBIF-Taxon="190310834" ID-Zenodo-Dep="5730280" LSID="urn:lsid:plazi:treatment:03C35358FFC2936BFEA43CCCFB789425" httpUri="http://treatment.plazi.org/id/03C35358FFC2936BFEA43CCCFB789425" lastPageId="19" lastPageNumber="331" pageId="18" pageNumber="330" scope_family="Peirosauridae" scope_order="Mesoeucrocodylia">
<subSubSection id="C370B1C5FFC2936AFEA43CCCFEE89343" pageId="18" pageNumber="330" type="nomenclature">
<paragraph id="8BD5E24EFFC2936AFEA43CCCFD349318" blockId="18.[291,650,755,779]" box="[291,650,755,779]" pageId="18" pageNumber="330">
<heading id="D09D5522FFC2936AFEA43CCCFD349318" box="[291,650,755,779]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="330" reason="3">
<taxonomicName id="4C6A99CDFFC2936AFEA43CCCFE639319" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[291,477,755,779]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">PEPESUCHINAE</taxonomicName>
<emphasis id="B91E3E5CFFC2936AFE633CC8FD349318" bold="true" box="[484,650,755,779]" pageId="18" pageNumber="330">
<taxonomicNameLabel id="A22D8327FFC2936AFE633CC8FD349318" box="[484,650,755,779]" pageId="18" pageNumber="330" rank="subFamily">SUBFAM. NOV.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BD5E24EFFC2936AFF243D23FEE89343" blockId="18.[163,779,796,848]" pageId="18" pageNumber="330">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C370B1C5FFC2936AFF243D5AFE9A94AC" pageId="18" pageNumber="330" type="diagnosis">
<paragraph id="8BD5E24EFFC2936AFF243D5AFEA2936F" blockId="18.[163,779,869,1168]" box="[163,284,869,892]" pageId="18" pageNumber="330">
<emphasis id="B91E3E5CFFC2936AFF243D5AFEA2936F" box="[163,284,869,892]" italics="true" pageId="18" pageNumber="330">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD5E24EFFC2936AFF243DBAFDA99483" blockId="18.[163,779,869,1168]" pageId="18" pageNumber="330">Crocodyliforms with an elongated platirostral rostrum, parallel lateral edges of the nasals and maxilla–lacrimal contact fully included in the antorbital fossa. The anterior wedge portion of the frontal region is elongated and inserted between the nasals; the nasals have a posterolateral process inserted between the prefrontals and lacrimals. The lateral aspect of the longitudinal axial inclination of the humeral shaft is slightly posterodorsally inclined.</paragraph>
<paragraph id="8BD5E24EFFC2936AFF243A98FE9A94AC" blockId="18.[163,778,1191,1276]" box="[163,292,1191,1215]" pageId="18" pageNumber="330">
<emphasis id="B91E3E5CFFC2936AFF243A98FE9A94AC" box="[163,292,1191,1215]" italics="true" pageId="18" pageNumber="330">Etymology</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC2936AFF243AF7FD1A964D" pageId="18" pageNumber="330" type="reference_group">
<paragraph id="8BD5E24EFFC2936AFF243AF7FD0A94EF" blockId="18.[163,778,1191,1276]" pageId="18" pageNumber="330">
<taxonomicName id="4C6A99CDFFC2936AFF243AF7FE8194CD" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[163,319,1224,1246]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
was established as the name of the subfamily based on
<taxonomicName id="4C6A99CDFFC2936AFEDA3AD8FD0E94EF" authority="Campos et al., 2011" authorityName="Campos" authorityYear="2011" box="[349,688,1254,1276]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC2936AFEDA3AD8FE7694E8" box="[349,456,1254,1276]" italics="true" pageId="18" pageNumber="330">P. deiseae</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFE483AD8FD0E94EF" author="Campos DA &amp; Oliveira GR &amp; Figueiredo RG &amp; Riff D &amp; Azevedo SAK &amp; Carvalho LB &amp; Kellner AWA" box="[463,688,1254,1276]" pageId="18" pageNumber="330" pagination="317 - 327" refId="ref14304" refString="Campos DA, Oliveira GR, Figueiredo RG, Riff D, Azevedo SAK, Carvalho LB, Kellner AWA. 2011. On a new peirosaurid crocodyliform from the Upper Cretaceous, Bauru Group, southeastern Brazil. Anais da Academia Brasileira de Ciencias 83: 317 - 327." type="journal article" year="2011">
Campos
<emphasis id="B91E3E5CFFC2936AFDB63AD8FDD794E8" box="[561,617,1254,1276]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 2011
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD5E24EFFC2936AFF243B2CFE059538" blockId="18.[163,779,1299,1630]" box="[163,443,1299,1323]" pageId="18" pageNumber="330">
<emphasis id="B91E3E5CFFC2936AFF243B2CFE059538" box="[163,443,1299,1323]" italics="true" pageId="18" pageNumber="330">Phylogenetic definition</emphasis>
</paragraph>
<paragraph id="8BD5E24EFFC2936AFF243B0BFD1A964D" blockId="18.[163,779,1299,1630]" pageId="18" pageNumber="330">
<taxonomicName id="4C6A99CDFFC2936AFF243B0BFE2D955A" box="[163,403,1332,1353]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC2936AFF243B0BFE2D955A" box="[163,403,1332,1353]" italics="true" pageId="18" pageNumber="330">Pepesuchus deiseae</emphasis>
</taxonomicName>
and all Crocodyliformes that share a more recent common ancestor with
<taxonomicName id="4C6A99CDFFC2936AFD1A3B6CFCB49574" box="[669,778,1362,1384]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC2936AFD1A3B6CFCB49574" box="[669,778,1362,1384]" italics="true" pageId="18" pageNumber="330">P. deiseae</emphasis>
</taxonomicName>
than with
<taxonomicName id="4C6A99CDFFC2936AFEA73B4DFDC69594" authority="Woodward, 1896" authorityName="Woodward" authorityYear="1896" box="[288,632,1393,1415]" class="Reptilia" genus="Notosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="terrestris">
<emphasis id="B91E3E5CFFC2936AFEA73B4DFE139594" box="[288,429,1394,1415]" italics="true" pageId="18" pageNumber="330">N. terrestris</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFE333B4EFDC69594" author="Woodward AS" box="[436,632,1393,1415]" pageId="18" pageNumber="330" pagination="1 - 20" refId="ref16486" refString="Woodward AS. 1896. On two Mesozoic crocodilians, Notosuchus (genus novum) and Cynodontosuchus (genum novum) from the red sandstones of the Territory of Neuquen (Argentine Republic). Anales del Museo de La Plata, La Plata, Serie Paleontologia Argentina 4: 1 - 20." type="journal article" year="1896">Woodward, 1896</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFD023B4DFEC095B6" authority="Simpson, 1937" authorityName="Simpson" authorityYear="1937" class="Reptilia" family="Sebecidae" genus="Sebecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="icaeorhinus">
<emphasis id="B91E3E5CFFC2936AFD023B4DFF7395B6" italics="true" pageId="18" pageNumber="330">S. icaeorhinus</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFF533BAFFEC095B6" author="Simpson GG" box="[212,382,1424,1446]" pageId="18" pageNumber="330" pagination="1 - 20" refId="ref16226" refString="Simpson GG. 1937. An ancient eusuchian crocodile from Patagonia. American Museum Novitates. Published by the American Museum of Natural History of New York. 1 - 20." type="book chapter" year="1937">Simpson, 1937</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFE0C3BAFFD2B95B5" authority="Price, 1945" authorityName="Price" authorityYear="1945" box="[395,661,1424,1446]" class="Reptilia" family="Baurusuchidae" genus="Baurusuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="pachecoi">
<emphasis id="B91E3E5CFFC2936AFE0C3BAFFDB595B6" box="[395,523,1424,1445]" italics="true" pageId="18" pageNumber="330">B. pachecoi</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFD943BAFFD2B95B5" author="Price LI" box="[531,661,1424,1446]" pageId="18" pageNumber="330" pagination="1 - 9" refId="ref15697" refString="Price LI. 1945. A new reptil from the Cretaceous Brazil. Divisao de Geologia e Mineralogia Notas Preliminares e estudos 1 - 9." type="book chapter" year="1945">Price, 1945</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFD263BAFFE9A95D7" authority="Price, 1950" authorityName="Price" authorityYear="1950" class="Reptilia" family="Sphagesauridae" genus="Sphagesaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="huenei">
<emphasis id="B91E3E5CFFC2936AFD263BAFFCB495B6" box="[673,778,1424,1445]" italics="true" pageId="18" pageNumber="330">S. huenei</emphasis>
Price, 1950
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFEA83B90FD8695D7" authority="Price, 1955" authorityName="Price" authorityYear="1955" box="[303,568,1454,1476]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC2936AFEA83B90FE1195D7" box="[303,431,1455,1476]" italics="true" pageId="18" pageNumber="330">P. torminni</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFE303B91FD8695D7" author="Price LI" box="[439,568,1454,1476]" pageId="18" pageNumber="330" refId="ref15723" refString="Price LI. 1955. Novos Crocodilideos dos Arenitos da Serie Bauru, Cretaceo do Estado de Minas Gerasi. Anais da Academia Brasileira de Ciencias 27: 487 - 453." type="journal volume" year="1955">Price, 1955</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFDC33B90FE7B95F0" authority="(Gasparini et al., 1991)" baseAuthorityName="Gasparini" baseAuthorityYear="1991" class="Reptilia" family="Peirosauridae" genus="Gasparinisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="peirosauroides">
<emphasis id="B91E3E5CFFC2936AFDC33B90FCB595D0" box="[580,779,1454,1476]" italics="true" pageId="18" pageNumber="330">G. peirosauroides</emphasis>
(
<bibRefCitation id="EFFB9FBFFFC2936AFF2C3BF2FE0595F0" author="Gasparini Z &amp; Chiappe LM &amp; Fernandez M" box="[171,443,1485,1507]" pageId="18" pageNumber="330" pagination="316 - 333" refId="ref14755" refString="Gasparini Z, Chiappe LM, Fernandez M. 1991. A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians. Journal of Vertebrate Paleontology 11: 316 - 333." type="journal article" year="1991">
Gasparini
<emphasis id="B91E3E5CFFC2936AFEB73BF1FED095F1" box="[304,366,1485,1507]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 1991
</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFE533BF1FE9B9612" authority="Gasparini et al., 1991" authorityName="Gasparini" authorityYear="1991" class="Reptilia" family="Peirosauridae" genus="Lomasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="palpebrosus">
<emphasis id="B91E3E5CFFC2936AFE533BF1FD3895F1" box="[468,646,1485,1507]" italics="true" pageId="18" pageNumber="330">L. palpebrosus</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFD163BF2FE9B9612" author="Gasparini Z &amp; Chiappe LM &amp; Fernandez M" pageId="18" pageNumber="330" pagination="316 - 333" refId="ref14755" refString="Gasparini Z, Chiappe LM, Fernandez M. 1991. A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians. Journal of Vertebrate Paleontology 11: 316 - 333." type="journal article" year="1991">
Gasparini
<emphasis id="B91E3E5CFFC2936AFF243BD3FF639612" box="[163,221,1516,1537]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 1991
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFEB63BD3FCB89612" authority="Carvalho et al., 2007" authorityName="Carvalho" authorityYear="2007" box="[305,774,1516,1538]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="arrudacamposi">
<emphasis id="B91E3E5CFFC2936AFEB63BD3FDB69612" box="[305,520,1516,1537]" italics="true" pageId="18" pageNumber="330">M. arrudacamposi</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFD963BD3FCB89612" author="Carvalho IS &amp; Vasconcellos FM &amp; Tavares SAS" box="[529,774,1516,1538]" pageId="18" pageNumber="330" pagination="35 - 46" refId="ref14427" refString="Carvalho IS, Vasconcellos FM, Tavares SAS. 2007. Montealtosuchus arrudacamposi, a new peirosaurid crocodile (Mesoeucrocodylia) from the Late Cretaceous Adamantina Formation of Brazil. Zootaxa 1607: 35 - 46." type="journal article" year="2007">
Carvalho
<emphasis id="B91E3E5CFFC2936AFD033BD3FD009612" box="[644,702,1516,1537]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 2007
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFF243834FE0E9633" authority="Price, 1959" authorityName="Price" authorityYear="1959" box="[163,432,1546,1568]" class="Reptilia" genus="Araripesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="gomesii">
<emphasis id="B91E3E5CFFC2936AFF243834FEA09633" box="[163,286,1547,1568]" italics="true" pageId="18" pageNumber="330">A. gomesii</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFEAF3835FE0E9633" author="Price LI" box="[296,432,1546,1568]" pageId="18" pageNumber="330" pagination="7 - 55" refId="ref15755" refString="Price LI. 1959. Sobre um novo crocodilideo notossuquido do Cretacico brasileiro. Boletim do Departamento Nacional de Producao Mineral, Divisao de Geologia e Mineralogia 188: 7 - 55." type="journal article" year="1959">Price, 1959</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFE393835FE71962D" authority="Gomani, 1997" authorityName="Gomani" authorityYear="1997" class="Reptilia" family="Itasuchidae" genus="Malawisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="mwakasyungutiensis">
<emphasis id="B91E3E5CFFC2936AFE393835FEAD962C" italics="true" pageId="18" pageNumber="330">Malawisuchus mwakasyungutiensis</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFE993816FE71962D" author="Gomani EM" box="[286,463,1577,1599]" pageId="18" pageNumber="330" pagination="280 - 294" refId="ref14984" refString="Gomani EM. 1997. A crocodyliform from the Early Cretaceous dinosaur beds, northern Malawi. Journal of Vertebrate Paleontology 17: 280 - 294." type="journal article" year="1997">Gomani, 1997</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFE583815FE84964E" authority="Carvalho &amp; Bertini, 1999" authorityName="Carvalho &amp; Bertini" authorityYear="1999" class="Reptilia" family="Candidodontidae" genus="Mariliasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="amarali">
<emphasis id="B91E3E5CFFC2936AFE583815FDD0962D" box="[479,622,1577,1599]" italics="true" pageId="18" pageNumber="330">M. amarali</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFDFE3816FE84964E" author="Carvalho IS &amp; Bertini RJ" pageId="18" pageNumber="330" pagination="83 - 105" refId="ref14355" refString="Carvalho IS, Bertini RJ. 1999. Mariliasuchus: um novo Crocodylomorpha (Notosuchia) do Cretaceo da Bacia Bauru, Brasil. Geologia Colombiana 24: 83 - 105." type="journal article" year="1999">Carvalho &amp; Bertini, 1999</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6A99CDFFC2936AFEF13877FD1E964D" authority="Laurent, 1768" authorityName="Laurent" authorityYear="1768" box="[374,672,1608,1630]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="niloticus">
<emphasis id="B91E3E5CFFC2936AFEF13877FE48964E" box="[374,502,1608,1629]" italics="true" pageId="18" pageNumber="330">C. niloticus</emphasis>
Laurent, 1768
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC2936AFF24384AFAAC967C" pageId="18" pageNumber="330" type="discussion">
<paragraph id="8BD5E24EFFC2936AFF24384AFE96969F" blockId="18.[163,779,1653,1891]" box="[163,296,1653,1676]" pageId="18" pageNumber="330">
<emphasis id="B91E3E5CFFC2936AFF24384AFE96969F" box="[163,296,1653,1676]" italics="true" pageId="18" pageNumber="330">Discussion</emphasis>
</paragraph>
<paragraph id="8BD5E24EFFC2936AFF2438AAFB2B9180" blockId="18.[163,779,1653,1891]" lastBlockId="18.[827,1444,197,863]" pageId="18" pageNumber="330">
This phylogenetic analysis revealed a monophyletic clade comprising
<taxonomicName id="4C6A99CDFFC2936AFEFC388BFE4096DA" baseAuthorityName="Kellner" baseAuthorityYear="1987" box="[379,510,1716,1737]" class="Reptilia" genus="Caririsuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="camposi">
<emphasis id="B91E3E5CFFC2936AFEFC388BFE4096DA" box="[379,510,1716,1737]" italics="true" pageId="18" pageNumber="330">C. camposi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFD88388AFD1896DA" authorityName="Iori &amp; Carvalho" authorityYear="2012" box="[527,678,1716,1738]" class="Reptilia" family="Trematochampsidae" genus="Barreirosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="franciscoi">
<emphasis id="B91E3E5CFFC2936AFD88388AFD1896DA" box="[527,678,1716,1738]" italics="true" pageId="18" pageNumber="330">B. franciscoi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFD30388AFF6D96FB" authorityName="Price" authorityYear="1955" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="jesuinoi">
<emphasis id="B91E3E5CFFC2936AFD30388AFF6D96FB" italics="true" pageId="18" pageNumber="330">I. jesuinoi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFF6138ECFEDE96F4" box="[230,352,1746,1768]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC2936AFF6138ECFEDE96F4" box="[230,352,1746,1768]" italics="true" pageId="18" pageNumber="330">P. deiseae</emphasis>
</taxonomicName>
and MCT 1723-R, and excluding
<taxonomicName id="4C6A99CDFFC2936AFF2438CEFE1E9715" authorityName="Buffetaut and Taqueti" authorityYear="1979" box="[163,416,1777,1798]" class="Reptilia" family="Trematochampsidae" genus="Miadanasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="oblita">
<emphasis id="B91E3E5CFFC2936AFF2438CEFE1E9715" box="[163,416,1777,1798]" italics="true" pageId="18" pageNumber="330">Miadanasuchus oblita</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC2936AFE5F38CEFD849714" authorityName="Chiappe" authorityYear="1988" box="[472,570,1777,1799]" class="Reptilia" family="Trematochampsidae" genus="Amargasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="minor">
<emphasis id="B91E3E5CFFC2936AFE5F38CEFD849714" box="[472,570,1777,1799]" italics="true" pageId="18" pageNumber="330">A. minor</emphasis>
</taxonomicName>
, making the clade
<taxonomicName id="4C6A99CDFFC2936AFF24392FFE269735" authorityName="Buffetaut" authorityYear="1974" box="[163,408,1808,1830]" class="Reptilia" family="Trematochampsidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="family">Trematochampsidae</taxonomicName>
paraphyletic,
<emphasis id="B91E3E5CFFC2936AFDCD392FFD329736" box="[586,652,1808,1829]" italics="true" pageId="18" pageNumber="330">sensu</emphasis>
<bibRefCitation id="EFFB9FBFFFC2936AFD10392FFF509756" author="Buffetaut E" pageId="18" pageNumber="330" pagination="348 - 350" refId="ref14170" refString="Buffetaut E. 1991. Itasuchus Price 1955. In: Maisey J, ed. Santana fossils: an illustrated atlas. Neptune City, NJ: T. F. H. Publications, 348 - 350." type="book chapter" year="1991">Buffetaut (1991)</bibRefCitation>
.
<taxonomicName id="4C6A99CDFFC2936AFF793911FE1C9757" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[254,418,1838,1860]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
has phylogenetic stability as a name for this group and was recovered in further analyses as a polytomy (
<bibRefCitation id="EFFB9FBFFFC2936AFBCB3EFAFAEE90C9" author="Montefeltro FC &amp; Larsson HCE &amp; De Franca MAG &amp; Langer MC" box="[1100,1360,197,219]" pageId="18" pageNumber="330" pagination="835 - 841" refId="ref15584" refString="Montefeltro FC, Larsson HCE, De Franca MAG, Langer MC. 2013. A new neosuchian with Asian affinities from the Jurassic of northeastern Brazil. Naturwissenschaften 100: 835 - 841." type="journal article" year="2013">
Montefeltro
<emphasis id="B91E3E5CFFC2936AFB503EF9FAB090C9" box="[1239,1294,197,219]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 2013
</bibRefCitation>
); internal relationships were recovered and solved. The clade definition contains
<taxonomicName id="4C6A99CDFFC2936AFB9A3F3CFB329104" box="[1053,1164,258,280]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC2936AFB9A3F3CFB329104" box="[1053,1164,258,280]" italics="true" pageId="18" pageNumber="330">P. deiseae</emphasis>
</taxonomicName>
as an internal specifier because the taxon is more preserved than
<taxonomicName id="4C6A99CDFFC2936AFAAD3F1DFA229124" authorityName="Price" authorityYear="1955" box="[1322,1436,290,311]" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="jesuinoi">
<emphasis id="B91E3E5CFFC2936AFAAD3F1DFA229124" box="[1322,1436,290,311]" italics="true" pageId="18" pageNumber="330">I. jesuinoi</emphasis>
</taxonomicName>
, with
<taxonomicName id="4C6A99CDFFC2936AFCF13F7FFC489146" authorityName="Price" authorityYear="1955" box="[886,1014,320,341]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC2936AFCF13F7FFC489146" box="[886,1014,320,341]" italics="true" pageId="18" pageNumber="330">P. torminni</emphasis>
</taxonomicName>
as an external specifier, allowing the clade to remain stable even with changes in the phylogenetic position of
<taxonomicName id="4C6A99CDFFC2936AFB973F41FB309180" authorityName="Price" authorityYear="1955" box="[1040,1166,382,403]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC2936AFB973F41FB309180" box="[1040,1166,382,403]" italics="true" pageId="18" pageNumber="330">P. torminni</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD5E24EFFC2936AFCD43FA3FAED934C" blockId="18.[827,1444,197,863]" pageId="18" pageNumber="330">This group is supported by ten synapomorphies as follows: posterolateral nasal process inserted between the prefrontal and lacrimal (54.1); a longer total lacrimal length relative to total prefrontal length (57.0); a rod-shaped jugal below the laterotemporal fenestra (65.1); an anterior margin of the prefrontals at the same level as the frontal anterior margin (70.1); a slightly depressed nasal–frontal contact area (80.1); an anterior process of the frontal extending between the prefrontals (82.1); a postorbital bar inclined medially and posteroventrally (95.1); mandibular branching with anteroposterior axis twice the medial–lateral width in the dorsal view (that forms a Y shape) (187.2); an anterior dentary direction of the teeth that is slightly procumbent (240.1); no internarial bar (250.2).</paragraph>
<paragraph id="8BD5E24EFFC2936AFCFD3DB1FADD93B5" blockId="18.[890,1379,910,935]" box="[890,1379,910,935]" pageId="18" pageNumber="330">
THE AGE OF THE
<taxonomicName id="4C6A99CDFFC2936AFBBA3DB1FADD93B5" authority="FORMATION" box="[1085,1379,910,935]" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">ADAMANTINA FORMATION</taxonomicName>
</paragraph>
<paragraph id="8BD5E24EFFC2936AFCBC3D89FBDF958B" blockId="18.[827,1444,950,1892]" pageId="18" pageNumber="330">
Time calibration of the phylogenetic hypotheses described above implies that
<taxonomicName id="4C6A99CDFFC2936AFB293DEAFAE893F8" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[1198,1366,981,1003]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
originated from an Early Cretaceous ancestry. The presence of early members of the clade from the Araripe Basin suggests an origin there. A subsequent cladogenic event split this group into two lineages, the first giving rise to
<taxonomicName id="4C6A99CDFFC2936AFB903A50FB2B9497" baseAuthorityName="Kellner" baseAuthorityYear="1987" box="[1047,1173,1135,1156]" class="Reptilia" genus="Caririsuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="camposi">
<emphasis id="B91E3E5CFFC2936AFB903A50FB2B9497" box="[1047,1173,1135,1156]" italics="true" pageId="18" pageNumber="330">C. camposi</emphasis>
</taxonomicName>
, and the other spreading to the Bauru Group and giving rise to some of its taxa (
<taxonomicName id="4C6A99CDFFC2936AFC233A93FBAB94D2" box="[932,1045,1196,1217]" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC2936AFC233A93FBAB94D2" box="[932,1045,1196,1217]" italics="true" pageId="18" pageNumber="330">Itasuchus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6A99CDFFC2936AFBA43A93FB1994D2" box="[1059,1191,1196,1217]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC2936AFBA43A93FB1994D2" box="[1059,1191,1196,1217]" italics="true" pageId="18" pageNumber="330">Pepesuchus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC2936AFB653A93FA2A94D2" authorityName="Iori &amp; Garcia" authorityYear="2012" box="[1250,1428,1196,1217]" class="Reptilia" family="Trematochampsidae" genus="Barreirosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC2936AFB653A93FA2A94D2" box="[1250,1428,1196,1217]" italics="true" pageId="18" pageNumber="330">Barreirosuchus</emphasis>
</taxonomicName>
). A palaeobiogeographical link has been suggested between the Araripe and
<collectingRegion id="49AE2CACFFC2936AFBF03AD6FB7094EC" box="[1143,1230,1257,1279]" country="Brazil" name="Parana" pageId="18" pageNumber="330">Paraná</collectingRegion>
basins, although these crocodyliform taxa cannot be used to correlate these localities. Except for
<taxonomicName id="4C6A99CDFFC2936AFB073B18FABE952F" baseAuthorityName="Kellner" baseAuthorityYear="1987" box="[1152,1280,1319,1340]" class="Reptilia" genus="Caririsuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="camposi">
<emphasis id="B91E3E5CFFC2936AFB073B18FABE952F" box="[1152,1280,1319,1340]" italics="true" pageId="18" pageNumber="330">C. camposi</emphasis>
</taxonomicName>
, found in the Lower Cretaceous of the Araripe Basin, all other taxa included in
<taxonomicName id="4C6A99CDFFC2936AFB8B3B5BFB0F9569" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[1036,1201,1380,1402]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
occur in the Upper Cretaceous Bauru Group.
</paragraph>
<paragraph id="8BD5E24EFFC2936AFCD43B9EFAAC967C" blockId="18.[827,1444,950,1892]" pageId="18" pageNumber="330">
The phylogenetic results show that diversification within itasuchine taxa from the Bauru Group occurred throughout the Santonian, matching the stage during which depositions and basin border rising occurred in the region (
<bibRefCitation id="EFFB9FBFFFC2936AFC673823FB269622" author="Riccomini C" box="[992,1176,1564,1586]" pageId="18" pageNumber="330" pagination="153 - 162" refId="ref15811" refString="Riccomini C. 1997. Arcabouco estrutural e aspectos do tectonismo gerador e deformador da Bacia Bauru no Estado de Sao Paulo. Revista Brasileira de Geociencias 27: 153 - 162." type="journal article" year="1997">Riccomini, 1997</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC2936AFB213823FA2A9622" author="Batezelli A &amp; Saad AR &amp; Etchebehere MLC &amp; Perinotto JAJ &amp; Fulfaro VJ" box="[1190,1428,1564,1586]" pageId="18" pageNumber="330" pagination="5 - 19" refId="ref13823" refString="Batezelli A, Saad AR, Etchebehere MLC, Perinotto JAJ, Fulfaro VJ. 2003. Analise estratigrafica aplicada a Formacao Aracatuba (Grupo Bauru - KS) no Centro-Oeste do Estado de Sao Paulo. Geociencias 22: 5 - 19." type="journal article" year="2003">
Batezelli
<emphasis id="B91E3E5CFFC2936AFA923823FAF19622" box="[1301,1359,1564,1585]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 2003
</bibRefCitation>
). Both
<taxonomicName id="4C6A99CDFFC2936AFCFE3805FBA69643" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[889,1048,1594,1616]" class="Reptilia" family="Peirosauridae" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Peirosaurinae">Peirosaurinae</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC2936AFBD63805FB539643" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[1105,1261,1594,1616]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
can be found in the Adamantina and Marília Formations.
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC2936BFCD43847FEF6923F" lastPageId="19" lastPageNumber="331" pageId="18" pageNumber="330" type="reference_group">
<paragraph id="8BD5E24EFFC2936BFCD43847FD1D90EA" blockId="18.[827,1444,950,1892]" lastBlockId="19.[145,762,197,1660]" lastPageId="19" lastPageNumber="331" pageId="18" pageNumber="330">
<taxonomicName id="4C6A99CDFFC2936AFCD43847FB90969E" authorityName="Price" authorityYear="1955" box="[851,1070,1656,1677]" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="jesuinoi">
<emphasis id="B91E3E5CFFC2936AFCD43847FB90969E" box="[851,1070,1656,1677]" italics="true" pageId="18" pageNumber="330">Itasuchus jesuinoi</emphasis>
</taxonomicName>
occurs in two localities and in distinct geological units: the
<taxonomicName id="4C6A99CDFFC2936AFB0E38A9FC3D96D9" authority="(Mezzalira, 1989)" baseAuthorityName="Mezzalira" baseAuthorityYear="1989" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
Adamantina (
<bibRefCitation id="EFFB9FBFFFC2936AFAAD38A9FCC796D9" author="Mezzalira S" pageId="18" pageNumber="330" refId="ref15543" refString="Mezzalira S. 1989. Os fosseis do Estado de Sao Paulo. Sao Paulo: Secretaria do Meio Ambiente / Instituto Geologico." type="book" year="1989">Mezzalira, 1989</bibRefCitation>
)
</taxonomicName>
and Marília Formations (
<bibRefCitation id="EFFB9FBFFFC2936AFB71388AFA2E96D8" author="Price LI" box="[1270,1424,1717,1739]" pageId="18" pageNumber="330" refId="ref15723" refString="Price LI. 1955. Novos Crocodilideos dos Arenitos da Serie Bauru, Cretaceo do Estado de Minas Gerasi. Anais da Academia Brasileira de Ciencias 27: 487 - 453." type="journal volume" year="1955">Price, 1955</bibRefCitation>
).
<taxonomicName id="4C6A99CDFFC2936AFCBC38EBFBA896FA" box="[827,1046,1748,1769]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC2936AFCBC38EBFBA896FA" box="[827,1046,1748,1769]" italics="true" pageId="18" pageNumber="330">Pepesuchus deiseae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC2936AFBD638EBFB6096FA" authorityName="Iori &amp; Carvalho" authorityYear="2012" box="[1105,1246,1748,1769]" class="Reptilia" family="Trematochampsidae" genus="Barreirosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="species" species="franciscoi">
<emphasis id="B91E3E5CFFC2936AFBD638EBFB6096FA" box="[1105,1246,1748,1769]" italics="true" pageId="18" pageNumber="330">B. franciscoi</emphasis>
</taxonomicName>
also occur in the
<taxonomicName id="4C6A99CDFFC2936AFCBC38CDFBEC971B" authority="Formation" authorityName="Formation" box="[827,1106,1778,1800]" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">Adamantina Formation</taxonomicName>
(
<bibRefCitation id="EFFB9FBFFFC2936AFBE438CDFAF29714" author="Campos DA &amp; Oliveira GR &amp; Figueiredo RG &amp; Riff D &amp; Azevedo SAK &amp; Carvalho LB &amp; Kellner AWA" box="[1123,1356,1778,1800]" pageId="18" pageNumber="330" pagination="317 - 327" refId="ref14304" refString="Campos DA, Oliveira GR, Figueiredo RG, Riff D, Azevedo SAK, Carvalho LB, Kellner AWA. 2011. On a new peirosaurid crocodyliform from the Upper Cretaceous, Bauru Group, southeastern Brazil. Anais da Academia Brasileira de Ciencias 83: 317 - 327." type="journal article" year="2011">
Campos
<emphasis id="B91E3E5CFFC2936AFB4E38CCFABB9714" box="[1225,1285,1778,1800]" italics="true" pageId="18" pageNumber="330">et al.</emphasis>
, 2011
</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC2936AFADC38CDFC719735" author="Iori FV &amp; Garcia KL" pageId="18" pageNumber="330" pagination="397 - 410" refId="ref15011" refString="Iori FV, Garcia KL. 2012. Barreirosuchus franciscoi, um novo Crocodylomorpha Trematochampsidae da Bacia Bauru, Brasil. Revista Brasileira de Geociencias 42: 397 - 410." type="journal article" year="2012">Iori &amp; Garcia, 2012</bibRefCitation>
).
<taxonomicName id="4C6A99CDFFC2936AFC61392EFB369734" authorityName="GASPARINI" authorityYear="1982" box="[998,1160,1809,1831]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="family">Peirosauridae</taxonomicName>
occurs in outcrops from the Adamantina Formation, considering the presence of
<taxonomicName id="4C6A99CDFFC2936AFC133971FAD79777" authority="(Carvalho et al., 2007)" baseAuthorityName="Carvalho" baseAuthorityYear="2007" box="[916,1385,1870,1892]" class="Reptilia" family="Peirosauridae" genus="Montealtosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="18" pageNumber="330" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC2936AFC133971FBE79770" box="[916,1113,1870,1891]" italics="true" pageId="18" pageNumber="330">Montealtosuchus</emphasis>
(
<bibRefCitation id="EFFB9FBFFFC2936AFBED3971FADE9770" author="Carvalho IS &amp; Vasconcellos FM &amp; Tavares SAS" box="[1130,1376,1870,1892]" pageId="18" pageNumber="330" pagination="35 - 46" refId="ref14427" refString="Carvalho IS, Vasconcellos FM, Tavares SAS. 2007. Montealtosuchus arrudacamposi, a new peirosaurid crocodile (Mesoeucrocodylia) from the Late Cretaceous Adamantina Formation of Brazil. Zootaxa 1607: 35 - 46." type="journal article" year="2007">Carvalho et al., 2007</bibRefCitation>
)
</taxonomicName>
, and in the Marília Formation, considering the presence of
<taxonomicName id="4C6A99CDFFC3936BFF163EDAFEB190EA" authorityName="Price" authorityYear="1955" box="[145,271,228,250]" class="Reptilia" family="Peirosauridae" genus="Peirosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="torminni">
<emphasis id="B91E3E5CFFC3936BFF163EDAFEB190EA" box="[145,271,228,250]" italics="true" pageId="19" pageNumber="331">P. torminni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFB9FBFFFC3936BFE993EDBFE2290E9" author="Price LI" box="[286,412,228,250]" pageId="19" pageNumber="331" refId="ref15723" refString="Price LI. 1955. Novos Crocodilideos dos Arenitos da Serie Bauru, Cretaceo do Estado de Minas Gerasi. Anais da Academia Brasileira de Ciencias 27: 487 - 453." type="journal volume" year="1955">Price, 1955</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC3936BFE2E3EDBFD2A90EA" author="Carvalho IS &amp; Ribeiro LCB &amp; Avilla LS" box="[425,660,228,250]" pageId="19" pageNumber="331" pagination="975 - 1002" refId="ref14387" refString="Carvalho IS, Ribeiro LCB, Avilla LS. 2004. Uberabasuchus terrificus sp. nov., a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil. Gondwana Research 7: 975 - 1002." type="journal article" year="2004">
Carvalho
<emphasis id="B91E3E5CFFC3936BFD9F3EDAFDEE90EA" box="[536,592,228,250]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2004
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD5E24EFFC3936BFF2E3F3CFEF6923F" blockId="19.[145,762,197,1660]" pageId="19" pageNumber="331">
The morphological similarity between
<taxonomicName id="4C6A99CDFFC3936BFF163F1DFEFA9124" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[145,324,289,311]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
taxa from the Adamantina and Marília Formations, and the occurrence of the same species in both, reveals a close chronology for these geological units. The age of the Marília Formation is usually considered Maastrichtian owing to the presence of charophytes, ostracods, turtles and dinosaurs (
<bibRefCitation id="EFFB9FBFFFC3936BFF1E3FE6FE2691FD" author="Dias-Brito D &amp; Musacchio EA &amp; Castro JC &amp; Maranhao MSAS &amp; Suarez JM &amp; Rodrigues R" box="[153,408,473,495]" pageId="19" pageNumber="331" pagination="245 - 304" refId="ref14621" refString="Dias-Brito D, Musacchio EA, Castro JC, Maranhao MSAS, Suarez JM, Rodrigues R. 2001. Grupo Bauru: uma unidade continental do Cretaceo no Brasil. Revue de Paleobiologie 20: 245 - 304." type="journal article" year="2001">
Dias-Brito
<emphasis id="B91E3E5CFFC3936BFE9D3FE5FEED91FD" box="[282,339,473,495]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2001
</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC3936BFE213FE6FD4C91FD" author="Gobbo-Rodrigues SR &amp; Santucci RM &amp; Bertini RJ" box="[422,754,473,495]" pageId="19" pageNumber="331" pagination="48" refId="ref14901" refString="Gobbo-Rodrigues SR, Santucci RM, Bertini RJ. 2001. Consideracoes sobre a idade da Formacao Marilia (Grupo Bauru, Cretaceo Superior) na regiao de Peiropolis, Estado de Minas Gerais. Boletim de Resumos do 17 º Congresso Brasileiro de Paleontologia Rio Branco: Universidade Federal do Acre, 48." type="journal article" year="2001">
Gobbo-Rodrigues
<emphasis id="B91E3E5CFFC3936BFDF33FE5FD1391FD" box="[628,685,473,495]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2001
</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC3936BFF163FC7FE6E921E" author="Santucci RM &amp; Bertini RJ" box="[145,464,504,526]" pageId="19" pageNumber="331" pagination="307 - 314" refId="ref15906" refString="Santucci RM, Bertini RJ. 2001. Distribuicao paleogeografica e biocronologica dos titanossauros (Saurischia, Sauropoda) do Grupo Bauru, Cretaceo Superior do Sudeste brasileiro. Revista Brasileira de Geociencias 31: 307 - 314." type="journal article" year="2001">Santucci &amp; Bertini, 2001</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC3936BFE643FC7FE89923F" author="Menegazzo MC &amp; Bertini RJ &amp; Manzini FF" pageId="19" pageNumber="331" pagination="18 - 32" refId="ref15417" refString="Menegazzo MC, Bertini RJ, Manzini FF. 2015. A new turtle from the Upper Cretaceous Bauru Group of Brazil, updated phylogeny and implications for age of the Santo Anastacio Formation. Journal of South American Earth Science 58: 18 - 32." type="journal article" year="2015">Menegazzo, Bertini &amp; Manzini, 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C370B1C5FFC3936BFF2E3C0AFB789425" pageId="19" pageNumber="331" type="description">
<paragraph id="8BD5E24EFFC3936BFF2E3C0AFEB19310" blockId="19.[145,762,197,1660]" pageId="19" pageNumber="331">
Another faunal correlation can be determined between the Adamantina and Santo Anastácio Formations. The presence of a podocnemidid in both geological units derived from Early Cretaceous taxa (
<bibRefCitation id="EFFB9FBFFFC3936BFF1D3C8FFE0892D5" author="Menegazzo MC &amp; Bertini RJ &amp; Manzini FF" box="[154,438,688,710]" pageId="19" pageNumber="331" pagination="18 - 32" refId="ref15417" refString="Menegazzo MC, Bertini RJ, Manzini FF. 2015. A new turtle from the Upper Cretaceous Bauru Group of Brazil, updated phylogeny and implications for age of the Santo Anastacio Formation. Journal of South American Earth Science 58: 18 - 32." type="journal article" year="2015">
Menegazzo
<emphasis id="B91E3E5CFFC3936BFEAC3C8FFED492D6" box="[299,362,688,709]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2015
</bibRefCitation>
).
<bibRefCitation id="EFFB9FBFFFC3936BFE563C8FFD4792D5" author="Menegazzo MC &amp; Catuneanu O &amp; Chang HK" box="[465,761,688,710]" pageId="19" pageNumber="331" pagination="131 - 156" refId="ref15464" refString="Menegazzo MC, Catuneanu O, Chang HK. 2016. The South American retroarc foreland system: the development of the Bauru Basin in the back-bulge province. Marine and Petroleum Geology 73: 131 - 156." type="journal article" year="2016">
Menegazzo
<emphasis id="B91E3E5CFFC3936BFDE53C8FFD1F92D6" box="[610,673,688,709]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
(2016)
</bibRefCitation>
suggests a Coniacian age for the Santo Anastácio Formation.
</paragraph>
<paragraph id="8BD5E24EFFC3936BFF2E3D33FED6966F" blockId="19.[145,762,197,1660]" pageId="19" pageNumber="331">
The Adamantina and Araçatuba Formations from the Bauru Group and the Locoche and Bajo de La Carpa Formations from the
<collectingRegion id="49AE2CACFFC3936BFE633D76FDEE934C" box="[484,592,841,863]" country="Argentina" name="Neuquen" pageId="19" pageNumber="331">Neuquén</collectingRegion>
Group can be dated to the Campanian–Maastrichtian ages (
<bibRefCitation id="EFFB9FBFFFC3936BFD243D57FEFA9388" author="Leanza AH &amp; Apesteguia S &amp; Novas FE &amp; Fuente MS de la" pageId="19" pageNumber="331" pagination="61 - 87" refId="ref15251" refString="Leanza AH, Apesteguia S, Novas FE, Fuente MS de la. 2004. Cretaceous terrestrial beds from the Neuquen Basin (Argentina) and their tetrapod assemblages. Cretaceous Research 25: 61 - 87." type="journal article" year="2004">Leanza &amp; Novas, 2004</bibRefCitation>
) considering assemblages of ostracods such as
<emphasis id="B91E3E5CFFC3936BFEB63D9AFDF193A8" box="[305,591,933,955]" italics="true" pageId="19" pageNumber="331">Ilyocipris argentinensis</emphasis>
,
<taxonomicName id="4C6A99CDFFC3936BFDE73D99FF0F93CA" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="riogradensis">
<emphasis id="B91E3E5CFFC3936BFDE73D99FF0F93CA" italics="true" pageId="19" pageNumber="331">I. riogradensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFB9FBFFFC3936BFF443DFBFD9193CA" author="Gobbo-Rodrigues SR &amp; Petri S &amp; Bertini RJ" box="[195,559,964,986]" pageId="19" pageNumber="331" pagination="3 - 13" refId="ref14795" refString="Gobbo-Rodrigues SR, Petri S, Bertini RJ. 1999 a. Ocorrencias de ostracodes na Formacao Adamantina do Grupo Bauru, Cretaceo Superior da Bacia do Parana e possibilidades de correlacao com depositos isocronos argentinos. Parte I-Familia Ilyocyprididae. Acta Geologica Leopoldinenese 23: 3 - 13." type="journal article" year="1999">
Gobbo-Rodrigues
<emphasis id="B91E3E5CFFC3936BFE1C3DFBFE7193CA" box="[411,463,964,985]" italics="true" pageId="19" pageNumber="331">et al</emphasis>
., 1999a
</bibRefCitation>
, b;
<bibRefCitation id="EFFB9FBFFFC3936BFDDA3DFBFE9493E4" author="Carignano AP &amp; Varela JA" pageId="19" pageNumber="331" pagination="169 - 178" refId="ref14464" refString="Carignano AP, Varela JA. 2011. Ostracodos (Crustacea) de la Formacion Allen (Cretacico Tardio), Cuenca Neuquina, Argentina. Revista Brasileira de Paleontologia 14: 169 - 178." type="journal article" year="2011">Carignano &amp; Varela, 2011</bibRefCitation>
) and
<emphasis id="B91E3E5CFFC3936BFEF63DDCFE4E93EB" box="[369,496,995,1016]" italics="true" pageId="19" pageNumber="331">Vecticipris</emphasis>
sp. (
<bibRefCitation id="EFFB9FBFFFC3936BFDAA3DDDFF759405" author="Dias-Brito D &amp; Musacchio EA &amp; Castro JC &amp; Maranhao MSAS &amp; Suarez JM &amp; Rodrigues R" pageId="19" pageNumber="331" pagination="245 - 304" refId="ref14621" refString="Dias-Brito D, Musacchio EA, Castro JC, Maranhao MSAS, Suarez JM, Rodrigues R. 2001. Grupo Bauru: uma unidade continental do Cretaceo no Brasil. Revue de Paleobiologie 20: 245 - 304." type="journal article" year="2001">
Dias-Brito
<emphasis id="B91E3E5CFFC3936BFD323DDCFD4C93E4" box="[693,754,994,1016]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2001
</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC3936BFF5B3A3EFDA19405" author="Carignano AP &amp; Varela JA" box="[220,543,1025,1047]" pageId="19" pageNumber="331" pagination="169 - 178" refId="ref14464" refString="Carignano AP, Varela JA. 2011. Ostracodos (Crustacea) de la Formacion Allen (Cretacico Tardio), Cuenca Neuquina, Argentina. Revista Brasileira de Paleontologia 14: 169 - 178." type="journal article" year="2011">Carignano &amp; Varela, 2011</bibRefCitation>
). There are similarities between the aeolosaurine
<taxonomicName id="4C6A99CDFFC3936BFDDC3A1FFE949440" class="Reptilia" genus="Brasilotitan" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="nemaphagus">
<emphasis id="B91E3E5CFFC3936BFDDC3A1FFE949440" italics="true" pageId="19" pageNumber="331">Brasilotitan nemaphagus</emphasis>
</taxonomicName>
from the
<taxonomicName id="4C6A99CDFFC3936BFE233A01FEF29461" authority="Formation and Antarctasaurus" authorityName="Formation and Antarctasaurus" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
Adamantina Formation and
<emphasis id="B91E3E5CFFC3936BFF163A62FEF29461" box="[145,332,1117,1138]" italics="true" pageId="19" pageNumber="331">Antarctasaurus</emphasis>
</taxonomicName>
<emphasis id="B91E3E5CFFC3936BFED23A62FD9B9461" box="[341,549,1117,1138]" italics="true" pageId="19" pageNumber="331">whichmannianus</emphasis>
and
<taxonomicName id="4C6A99CDFFC3936BFDE43A61FF4B9482" authorityName="Apesteguia" authorityYear="2004" class="Reptilia" family="Nemegtosauridae" genus="Bonitasaura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="salgadoi">
<emphasis id="B91E3E5CFFC3936BFDE43A61FF4B9482" italics="true" pageId="19" pageNumber="331">Bonitasaura salgadoi</emphasis>
</taxonomicName>
from the Santonian of
<collectingCountry id="F37DA2DEFFC3936BFD803A43FDC19481" box="[519,639,1148,1170]" name="Argentina" pageId="19" pageNumber="331">Argentina</collectingCountry>
(
<bibRefCitation id="EFFB9FBFFFC3936BFD083A43FEAF94BC" author="Machado EB &amp; Avilla LS &amp; Nava WR &amp; Campos DA &amp; Kellner AWA" pageId="19" pageNumber="331" pagination="301 - 321" refId="ref15332" refString="Machado EB., Avilla LS., Nava WR., Campos DA., Kellner AWA. 2013. A new titanosaur sauropod from the Late Cretaceous of Brazil. Zootaxa 3701: 301 - 321." type="journal article" year="2013">
Machado
<emphasis id="B91E3E5CFFC3936BFF163AA4FF7594BC" box="[145,203,1178,1200]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2013
</bibRefCitation>
).
<taxonomicName id="4C6A99CDFFC3936BFEAF3AA5FE6D94BC" authorityName="Carvalho &amp; Bertini" authorityYear="1999" box="[296,467,1178,1199]" class="Reptilia" family="Candidodontidae" genus="Mariliasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFEAF3AA5FE6D94BC" box="[296,467,1178,1199]" italics="true" pageId="19" pageNumber="331">Mariliasuchus</emphasis>
</taxonomicName>
material has been found in the lower layers of the
<taxonomicName id="4C6A99CDFFC3936BFE553A86FD4D94DC" authority="Formation" authorityName="Formation" box="[466,755,1209,1231]" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">Adamantina Formation</taxonomicName>
, in contact with the Araçatuba Formation, which is dated as Coniacian–early Santonian (
<bibRefCitation id="EFFB9FBFFFC3936BFDF73AC9FE949539" author="Andrade MB &amp; Bertini RJ" pageId="19" pageNumber="331" pagination="3 - 62" refId="ref13736" refString="Andrade MB, Bertini RJ. 2008. Morphological and anatomical observations about Mariliasuchus amarali and Notosuchus terrestris (Mesoeucrocodylia) and their relationships with other South American notosuchians. Arquivos do Museu Nacional 66: 3 - 62." type="journal article" year="2008">Andrade &amp; Bertini, 2008</bibRefCitation>
;
<bibRefCitation id="EFFB9FBFFFC3936BFEB03B2AFD839538" author="Menegazzo MC &amp; Catuneanu O &amp; Chang HK" box="[311,573,1301,1323]" pageId="19" pageNumber="331" pagination="131 - 156" refId="ref15464" refString="Menegazzo MC, Catuneanu O, Chang HK. 2016. The South American retroarc foreland system: the development of the Bauru Basin in the back-bulge province. Marine and Petroleum Geology 73: 131 - 156." type="journal article" year="2016">
Menegazzo
<emphasis id="B91E3E5CFFC3936BFE383B29FE469539" box="[447,504,1301,1323]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
, 2016
</bibRefCitation>
). These correlations and the distribution of some morphotypes along the same time span in these geological units point to at least a Campanian age for the
<taxonomicName id="4C6A99CDFFC3936BFDD83B4EFEA995B5" authority="Formation" authorityName="Formation" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">Adamantina Formation</taxonomicName>
, possibly Campanian–Maastrichtian. Supporting this hypothesis, there are morphological similarities between crocodyliform taxa from those geological units, as proposed by
<bibRefCitation id="EFFB9FBFFFC3936BFD8C3BD3FD4C9611" author="Bertini RJ &amp; Santucci RM &amp; Arruda-Campos A" box="[523,754,1516,1538]" pageId="19" pageNumber="331" pagination="93 - 103" refId="ref13867" refString="Bertini RJ, Santucci RM, Arruda-Campos A. 2001. Titanossauros (Sauropoda: Saurischia) no Cretaceo Superior continental (Formacao Marilia, Membro Echapora) de Monte Alto, Estado de Sao Paulo, e correlacao com formas associadas do Triangulo Mineiro. UNESP, Geociencias 20: 93 - 103." type="journal article" year="2001">
Bertini
<emphasis id="B91E3E5CFFC3936BFDE03BD3FD1F9612" box="[615,673,1516,1537]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
(2001)
</bibRefCitation>
,
<bibRefCitation id="EFFB9FBFFFC3936BFF163835FDA7960C" author="Gobbo-Rodrigues SR &amp; Petri S &amp; Bertini RJ" box="[145,537,1546,1568]" pageId="19" pageNumber="331" pagination="3 - 13" refId="ref14795" refString="Gobbo-Rodrigues SR, Petri S, Bertini RJ. 1999 a. Ocorrencias de ostracodes na Formacao Adamantina do Grupo Bauru, Cretaceo Superior da Bacia do Parana e possibilidades de correlacao com depositos isocronos argentinos. Parte I-Familia Ilyocyprididae. Acta Geologica Leopoldinenese 23: 3 - 13." type="journal article" year="1999">
Gobbo-Rodrigues
<emphasis id="B91E3E5CFFC3936BFEF13834FE06960C" box="[374,440,1546,1568]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
(1999a
</bibRefCitation>
, b) and
<bibRefCitation id="EFFB9FBFFFC3936BFD0C3835FEE2962C" author="Santucci RM &amp; Bertini RJ" pageId="19" pageNumber="331" pagination="307 - 314" refId="ref15906" refString="Santucci RM, Bertini RJ. 2001. Distribuicao paleogeografica e biocronologica dos titanossauros (Saurischia, Sauropoda) do Grupo Bauru, Cretaceo Superior do Sudeste brasileiro. Revista Brasileira de Geociencias 31: 307 - 314." type="journal article" year="2001">Santucci &amp; Bertini (2001)</bibRefCitation>
, in opposition to the Turonian to Santonian proposal of
<bibRefCitation id="EFFB9FBFFFC3936BFE1B3877FDC0964D" author="Castro JC &amp; Dias-Brito D &amp; Musacchio EA &amp; Suarez J &amp; Maranhao MSAS &amp; Rodrigues R" box="[412,638,1608,1630]" pageId="19" pageNumber="331" pagination="509 - 515" refId="ref14500" refString="Castro JC, Dias-Brito D, Musacchio EA, Suarez J, Maranhao MSAS, Rodrigues R. 1999. Arcabouco estratigrafico do Grupo Bauru no Oeste Paulista. Boletim do 5 o Simposio sobre o Cretaceo do Brasil. Serra Negra: UNESP-Rio Claro, 509 - 515." type="book chapter" year="1999">
Castro
<emphasis id="B91E3E5CFFC3936BFE753877FD93964E" box="[498,557,1608,1629]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
(1999)
</bibRefCitation>
and Dias- Brito
<emphasis id="B91E3E5CFFC3936BFF523858FEB19668" box="[213,271,1638,1660]" italics="true" pageId="19" pageNumber="331">et al.</emphasis>
(2001).
</paragraph>
<paragraph id="8BD5E24EFFC3936BFEF33886FDAB96C3" blockId="19.[372,533,1721,1746]" box="[372,533,1721,1746]" pageId="19" pageNumber="331">CONCLUSIONS</paragraph>
<paragraph id="8BD5E24EFFC3936BFF1638DEFBAC9167" blockId="19.[145,761,1761,1875]" lastBlockId="19.[809,1425,197,1078]" pageId="19" pageNumber="331">
The new data provided by
<taxonomicName id="4C6A99CDFFC3936BFE4938DEFD6696E5" authority="MCT 1723" authorityName="MCT" authorityYear="1723" box="[462,728,1761,1782]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFE4938DEFDED96E5" box="[462,595,1761,1782]" italics="true" pageId="19" pageNumber="331">Pepesuchus</emphasis>
MCT 1723
</taxonomicName>
-R, regarding taxa closely related to
<taxonomicName id="4C6A99CDFFC3936BFD8C393EFDC29706" authorityName="Price" authorityYear="1955" box="[523,636,1792,1814]" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="jesuinoi">
<emphasis id="B91E3E5CFFC3936BFD8C393EFDC29706" box="[523,636,1792,1814]" italics="true" pageId="19" pageNumber="331">I. jesuinoi</emphasis>
</taxonomicName>
, generates phylogenetic stability and is a significant addition to the fossil record from the Bauru Group. MCT 1723-R shares synapomorphies with
<taxonomicName id="4C6A99CDFFC3936BFB133EF9FAB790C9" box="[1172,1289,197,219]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC3936BFB133EF9FAB790C9" box="[1172,1289,197,219]" italics="true" pageId="19" pageNumber="331">P. deiseae</emphasis>
</taxonomicName>
that allow the specimen to be inserted in the
<taxonomicName id="4C6A99CDFFC3936BFB3A3EDBFAFE90EA" authorityName="MCT" authorityYear="1723" box="[1213,1344,228,249]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFB3A3EDBFAFE90EA" box="[1213,1344,228,249]" italics="true" pageId="19" pageNumber="331">Pepesuchus</emphasis>
</taxonomicName>
genus. These synapomorphies include the dorsal edge of the rostrum starting from the anterior verge of the concave orbits and the L-shaped lacrimal inferior process contacting the jugal.
</paragraph>
<paragraph id="8BD5E24EFFC3936BFCC63F42FB9B921E" blockId="19.[809,1425,197,1078]" pageId="19" pageNumber="331">
The differences between
<taxonomicName id="4C6A99CDFFC3936BFBE33F41FB6C9181" box="[1124,1234,381,403]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="species" species="deiseae">
<emphasis id="B91E3E5CFFC3936BFBE33F41FB6C9181" box="[1124,1234,381,403]" italics="true" pageId="19" pageNumber="331">P. deiseae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC3936BFA8A3F42FC1A91A2" authority="MCT 1723" authorityName="MCT" authorityYear="1723" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFA8A3F42FA2E9181" box="[1293,1424,381,402]" italics="true" pageId="19" pageNumber="331">Pepesuchus</emphasis>
MCT 1723
</taxonomicName>
-R cannot be determined as taxonomic or intraspecific variation, because there are few specimens of
<taxonomicName id="4C6A99CDFFC3936BFC0D3FE6FB3391FD" authority="MCT 1723" authorityName="MCT" authorityYear="1723" box="[906,1165,473,494]" class="Reptilia" family="Peirosauridae" genus="Pepesuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFC0D3FE6FBB291FD" box="[906,1036,473,494]" italics="true" pageId="19" pageNumber="331">Pepesuchus</emphasis>
MCT 1723
</taxonomicName>
-R, and the material is in a fragmented state.
</paragraph>
<paragraph id="8BD5E24EFFC3936BFCC63C29FB789425" blockId="19.[809,1425,197,1078]" pageId="19" pageNumber="331">
The description of MCT 1723-R indicates characteristics that solved internal relationships among
<taxonomicName id="4C6A99CDFFC3936BFCAE3C6BFBF0927A" authority=", Pepesuchusxi" authorityName="Pepesuchusxi" box="[809,1102,596,617]" class="Reptilia" family="Itasuchidae" genus="Itasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFCAE3C6BFC21927A" box="[809,927,596,617]" italics="true" pageId="19" pageNumber="331">Itasuchus</emphasis>
,
<emphasis id="B91E3E5CFFC3936BFC283C6BFBF0927A" box="[943,1102,596,617]" italics="true" pageId="19" pageNumber="331">Pepesuchusxi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6A99CDFFC3936BFB173C6BFAF4927A" authorityName="Iori &amp; Garcia" authorityYear="2012" box="[1168,1354,596,617]" class="Reptilia" family="Trematochampsidae" genus="Barreirosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">
<emphasis id="B91E3E5CFFC3936BFB173C6BFAF4927A" box="[1168,1354,596,617]" italics="true" pageId="19" pageNumber="331">Barreirosuchus</emphasis>
</taxonomicName>
, with polytomies prior to the addition of the MCT 1723-R material. Two characteristic suture patterns occur in
<taxonomicName id="4C6A99CDFFC3936BFCAE3C8FFC7192D5" authority="Geroto &amp; Bertini, 2019" authorityName="Geroto &amp; Bertini" authorityYear="2019" box="[809,975,688,710]" class="Reptilia" family="Peirosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="331" phylum="Chordata" rank="subFamily" status="subfam. nov." subFamily="Pepesuchinae">Pepesuchinae</taxonomicName>
: the insertion of the frontals into the anterior region between the nasals, and a posterolateral process inserting between the prefrontal and lacrimal. The frontal inserted between the prefrontal is a diagnostic characteristic that appears as a synapomorphy from a more inclusive group, containing taxa from both the Adamantina and Marília formations. The fact that distinct geological units show not only species with morphological similarities, but also the occurrence of the same genus in both geological units, points to the correlation of their ages and the placement of the
<taxonomicName id="4C6A99CDFFC3936BFBAF3A3EFAFF9404" authority="Formation" authorityName="Formation" box="[1064,1345,1025,1047]" class="Chondrichthyes" genus="Adamantina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="331" phylum="Chordata" rank="genus">Adamantina Formation</taxonomicName>
in the Campanian–Maastrichtian interval.
</paragraph>
</subSubSection>
</treatment>
</document>
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