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<mods:title id="150081B860245F2023A6A2697AC867EF">Revision of Eugenia and Syzygium (Myrtaceae) from the Comoros archipelago</mods:title>
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<paragraph id="8BD7F709FF8F6B01FF60BF07B602F331" blockId="15.[136,849,266,293]" box="[136,849,266,293]" pageId="15" pageNumber="178">
<heading id="D09F4065FF8F6B01FF60BF07B602F331" box="[136,849,266,293]" fontSize="11" level="2" pageId="15" pageNumber="178" reason="4">
<emphasis id="B91C2B1BFF8F6B01FF60BF07B5CEF331" bold="true" box="[136,157,268,293]" pageId="15" pageNumber="178">4.</emphasis>
<taxonomicName id="4C688C8AFF8F6B01FF4CBF01B602F331" ID-CoL="54678" authority="(H. Perrier) Labat &amp; Schatz (2002: 203)" authorityName="Labat &amp; Schatz" authorityPageNumber="203" authorityYear="2002" baseAuthorityName="H. Perrier" box="[164,849,266,293]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="178" phylum="Tracheophyta" rank="species" species="humblotii">
<emphasis id="B91C2B1BFF8F6B01FF4CBF01B4D9F331" bold="true" box="[164,394,266,293]" italics="true" pageId="15" pageNumber="178">Syzygium humblotii</emphasis>
(H. Perrier)
<bibRefCitation id="EFF98AF8FF8F6B01FDF1BF07B602F331" author="Labat, J. - N. &amp; Schatz, G. E." box="[537,849,268,293]" pageId="15" pageNumber="178" pagination="201 - 205" refId="ref15422" refString="Labat, J. - N. &amp; Schatz, G. E. (2002) New combinations and a new name in Syzygium (Myrtaceae) from Madagascar and the Comoro Islands. Novon 12: 201 - 205. http: // dx. doi. org / 10.2307 / 3392957" type="journal article" year="2002">Labat &amp; Schatz (2002: 203)</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C372A482FF8F6B01FF60BF5DB4D3F109" pageId="15" pageNumber="178" type="reference_group">
<paragraph id="8BD7F709FF8F6B01FF60BF5DB194F39B" blockId="15.[136,1451,342,399]" pageId="15" pageNumber="178">
<treatmentCitationGroup id="AB78D027FF8F6B01FF60BF5DB194F39B" pageId="15" pageNumber="178">
<treatmentCitation id="0AC9D118FF8F6B01FF60BF5DB77BF37F" author="Perrier de la Bathie, H." box="[136,552,342,363]" page=")" pageId="15" pageNumber="178" year="1953">
<taxonomicName id="4C688C8AFF8F6B01FF60BF5DB77BF37F" ID-CoL="3C4YF" authority="H. Perrier (1953: 193)" authorityName="H. Perrier" authorityPageNumber="193" authorityYear="1953" box="[136,552,342,363]" class="Magnoliopsida" family="Myrtaceae" genus="Eugenia" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="178" phylum="Tracheophyta" rank="species" species="humblotii">
<emphasis id="B91C2B1BFF8F6B01FF60BF5DB46EF37F" box="[136,317,342,363]" italics="true" pageId="15" pageNumber="178">Eugenia humblotii</emphasis>
<materialsCitation id="3B00FD54FF8F6B01FEAEBF5DB77BF37F" box="[326,552,342,363]" collectingDate="1884-10-19" collectorName="de Combani" country="Mayotte" location="Foret de Combani" pageId="15" pageNumber="178" specimenCount="2" typeStatus="holotype">
H.
<bibRefCitation id="EFF98AF8FF8F6B01FE8FBF5DB77BF37F" author="Perrier de la Bathie, H." box="[359,552,342,363]" pageId="15" pageNumber="178" pagination="1 - 80" refId="ref16109" refString="Perrier de la Bathie, H. (1953) Myrtaceae. In: Humbert, H. (Ed.) Flore de Madagascar et des Comores 152: 1 - 80." type="journal article" year="1953">Perrier (1953: 193)</bibRefCitation>
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. Type:
<emphasis id="B91C2B1BFF8F6B01FD82BF5DB7D1F37F" bold="true" box="[618,642,342,363]" pageId="15" pageNumber="178"></emphasis>
<collectingCountry id="F37FB799FF8F6B01FD6ABF5DB7A6F37F" box="[642,757,342,363]" name="Mayotte" pageId="15" pageNumber="178">MAYOTTE</collectingCountry>
[
<collectingCountry id="F37FB799FF8F6B01FCECBF5DB638F37F" box="[772,875,342,363]" name="France" pageId="15" pageNumber="178">FRANCE</collectingCountry>
]. Grande Terre,
<location id="8EB7A1D2FF8F6B01FBE5BF5DB190F37F" LSID="urn:lsid:plazi:treatment:03C1461FFF8F6B01FF60BF07B7D9FA1A:8EB7A1D2FF8F6B01FBE5BF5DB190F37F" box="[1037,1219,342,363]" country="Mayotte" name="Foret de Combani" pageId="15" pageNumber="178">
Foret
<collectorName id="269D92DFFF8F6B01FBA1BF5DB190F37F" box="[1097,1219,342,363]" pageId="15" pageNumber="178">de Combani</collectorName>
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,
<date id="FFD6D1C9FF8F6B01FB38BF5DB02BF37F" box="[1232,1400,342,363]" pageId="15" pageNumber="178" value="1884-10-19">
<collectingDate id="EF922821FF8F6B01FB38BF5DB02BF37F" box="[1232,1400,342,363]" pageId="15" pageNumber="178" value="1884-10-19">19 October 1884</collectingDate>
</date>
(fl.),
<emphasis id="B91C2B1BFF8F6B01FF55BF71B414F39B" box="[189,327,378,399]" italics="true" pageId="15" pageNumber="178">Humblot 1329</emphasis>
(
<typeStatus id="54D349ABFF8F6B01FEBCBF71B4F9F39B" box="[340,426,378,399]" pageId="15" pageNumber="178" type="holotype">holotype</typeStatus>
P! [barcode P00118029];
<typeStatus id="54D349ABFF8F6B01FD4EBF71B7B8F39B" box="[678,747,378,399]" pageId="15" pageNumber="178" type="isotype">isotype</typeStatus>
P! [barcodes P00118028, P00118030]).
<figureCitation id="1353EB8CFF8F6B01FB99BF71B190F39B" box="[1137,1219,378,399]" captionStart="FIGURE 8" captionStartId="15.[136,229,1679,1700]" captionTargetBox="[164,1423,831,1642]" captionTargetId="figure-281@15.[151,1436,821,1654]" captionTargetPageId="15" captionText="FIGURE 8. Holotype of Syzygium humblotii (H. Perrier) Labat &amp; Schatz. Humblot 1329. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.13673324" httpUri="https://zenodo.org/record/13673324/files/figure.png" pageId="15" pageNumber="178">Figure 8</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD7F709FF8F6B01FF60BFCBB4D3F109" blockId="15.[136,1452,448,797]" pageId="15" pageNumber="178">
Trees or shrubs, height unknown; bark grey to reddish-brown. Branchlets angular, grey to reddish-brown. Petioles ca. 23 ×
<quantity id="4C905AECFF8F6B01FF3CBFEFB44BF3E9" box="[212,280,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="178" unit="mm" value="1.0">1 mm</quantity>
, slender. Leaves coriaceous; drying dark to light brown above, paler below, dull on both sides. Blades 24.4 ×
<quantity id="4C905AECFF8F6B01FF0BBC03B41DF035" box="[227,334,520,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.2" metricValueMin="1.0" pageId="15" pageNumber="178" unit="cm" value="1.6" valueMax="2.2" valueMin="1.0">12.2 cm</quantity>
, elliptic or oblong, base cuneate, apex acute or acuminate, to
<quantity id="4C905AECFF8F6B01FC1FBC03B100F035" box="[1015,1107,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="15" pageNumber="178" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long; margin slightly revolute; secondary vein pairs 815,
<quantity id="4C905AECFF8F6B01FE28BC27B74DF051" box="[448,542,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="15" pageNumber="178" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">13 mm</quantity>
apart, more or less prominent on both sides, angle of divergence from mid vein 5570°; tertiary veins few, faint; intramarginal vein ca.
<quantity id="4C905AECFF8F6B01FCE3BC5BB637F07D" box="[779,868,592,617]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="178" unit="mm" value="0.5">0.5 mm</quantity>
from leaf margin. Bracts and bracteoles ca.
<quantity id="4C905AECFF8F6B01FA80BC5BB0FFF07D" box="[1384,1452,592,617]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="178" unit="mm" value="1.0">1 mm</quantity>
long, deltoid, caducous. Inflorescences terminal (or in subterminal leaf axils),
<quantity id="4C905AECFF8F6B01FBE9BC7FB139F099" box="[1025,1130,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="15" pageNumber="178" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 cm</quantity>
long, congested, axes terete to subangular, flowers up to 30, in clusters of 89. Flowers ca.
<quantity id="4C905AECFF8F6B01FC81BC93B6BAF0A5" box="[873,1001,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="15" pageNumber="178" unit="cm" value="0.75" valueMax="0.8" valueMin="0.7">0.70.8 cm</quantity>
in diameter at anthesis, buds not seen, pseudostalk
<quantity id="4C905AECFF8F6B01FEFDBCB7B427F0C1" box="[277,372,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="15" pageNumber="178" unit="mm" value="1.0" valueMax="2.0" valueMin="0.0">02 mm</quantity>
long. Hypanthium 47 ×
<quantity id="4C905AECFF8F6B01FD76BCB7B7ACF0C1" box="[670,767,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="15" pageNumber="178" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
, pyriform. Calyx lobes ca. 0.51 ×
<quantity id="4C905AECFF8F6B01FB70BCB7B18EF0C1" box="[1176,1245,700,725]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="178" unit="mm" value="1.0">1 mm</quantity>
, broadly rounded. Petals ca. 2 ×
<quantity id="4C905AECFF8F6B01FEC2BCEBB4D0F0ED" box="[298,387,736,761]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="178" unit="mm" value="1.5">1.5 mm</quantity>
, calyptrate, orbicular. Stamens
<quantity id="4C905AECFF8F6B01FD00BCEBB615F0ED" box="[744,838,736,761]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="15" pageNumber="178" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long,?white; anthers ca.
<quantity id="4C905AECFF8F6B01FB83BCEBB190F0ED" box="[1131,1219,736,761]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="178" unit="mm" value="0.3">0.3 mm</quantity>
long. Style
<quantity id="4C905AECFF8F6B01FAA6BCEBB0FFF0ED" box="[1358,1452,736,761]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="15" pageNumber="178" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">47 mm</quantity>
long. Fruits unknown.
</paragraph>
</subSubSection>
<caption id="DF17A781FF8F6B01FF60B884B1C7F4B0" ID-DOI="http://doi.org/10.5281/zenodo.13673324" ID-Zenodo-Dep="13673324" box="[136,1172,1679,1700]" httpUri="https://zenodo.org/record/13673324/files/figure.png" pageId="15" pageNumber="178" startId="15.[136,229,1679,1700]" targetBox="[164,1423,831,1642]" targetPageId="15" targetType="figure">
<paragraph id="8BD7F709FF8F6B01FF60B884B1C7F4B0" blockId="15.[136,1172,1679,1700]" box="[136,1172,1679,1700]" pageId="15" pageNumber="178">
<emphasis id="B91C2B1BFF8F6B01FF60B884B5AEF4B0" bold="true" box="[136,253,1679,1700]" pageId="15" pageNumber="178">FIGURE 8.</emphasis>
Holotype of
<taxonomicName id="4C688C8AFF8F6B01FE95B884B68CF4B0" authority="(H. Perrier) Labat &amp; Schatz. Humblot 1329" authorityName="Labat &amp; Schatz. Humblot" authorityYear="1329" baseAuthorityName="H. Perrier" box="[381,991,1679,1700]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="15" pageNumber="178" phylum="Tracheophyta" rank="species" species="humblotii">
<emphasis id="B91C2B1BFF8F6B01FE95B884B76AF4B0" box="[381,569,1679,1700]" italics="true" pageId="15" pageNumber="178">Syzygium humblotii</emphasis>
(H. Perrier) Labat &amp; Schatz.
<emphasis id="B91C2B1BFF8F6B01FCBDB884B68CF4B0" box="[853,991,1679,1700]" italics="true" pageId="15" pageNumber="178">Humblot 1329</emphasis>
</taxonomicName>
. Scale bar = 2 cm.
</paragraph>
</caption>
<subSubSection id="C372A482FF8F6B01FF55B8DEB7D9FA1A" pageId="15" pageNumber="178" type="distribution">
<paragraph id="8BD7F709FF8F6B01FF55B8DEB796F4FA" blockId="15.[136,1452,1749,2062]" box="[189,709,1749,1774]" pageId="15" pageNumber="178">
<emphasis id="B91C2B1BFF8F6B01FF55B8DEB41CF4FA" bold="true" box="[189,335,1749,1774]" pageId="15" pageNumber="178">Distribution</emphasis>
:
<emphasis id="B91C2B1BFF8F6B01FEBFB8DEB420F4FA" bold="true" box="[343,371,1749,1774]" pageId="15" pageNumber="178"></emphasis>
Endemic to
<collectingCountry id="F37FB799FF8F6B01FE13B8DEB70BF4FA" box="[507,600,1749,1774]" name="Mayotte" pageId="15" pageNumber="178">Mayotte</collectingCountry>
[
<collectingCountry id="F37FB799FF8F6B01FD8EB8DEB7E8F4FA" box="[614,699,1749,1774]" name="France" pageId="15" pageNumber="178">France</collectingCountry>
].
</paragraph>
<paragraph id="8BD7F709FF8F6B01FF55B8F2B665F506" blockId="15.[136,1452,1749,2062]" box="[189,822,1785,1810]" pageId="15" pageNumber="178">
<emphasis id="B91C2B1BFF8F6B01FF55B8F2B44AF506" bold="true" box="[189,281,1785,1810]" pageId="15" pageNumber="178">Habitat</emphasis>
:
<emphasis id="B91C2B1BFF8F6B01FEC9B8F2B46EF506" bold="true" box="[289,317,1785,1810]" pageId="15" pageNumber="178"></emphasis>
Montane forests of Combani; ca.
<quantity id="4C905AECFF8F6B01FD5CB8F2B7A8F506" box="[692,763,1785,1810]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="178" unit="m" value="500.0">500 m</quantity>
elev.
</paragraph>
<paragraph id="8BD7F709FF8F6B01FF55B916B692F5B6" blockId="15.[136,1452,1749,2062]" pageId="15" pageNumber="178">
<emphasis id="B91C2B1BFF8F6B01FF55B916B4FEF522" bold="true" box="[189,429,1821,1846]" pageId="15" pageNumber="178">Conservation status</emphasis>
:
<emphasis id="B91C2B1BFF8F6B01FE5CB916B483F522" bold="true" box="[436,464,1821,1846]" pageId="15" pageNumber="178"></emphasis>
Extinct (EX) according to the IUCN Red List Criteria (
<bibRefCitation id="EFF98AF8FF8F6B01FBA3B916B184F522" author="IUCN &amp; Species Survival Commission, IUCN" box="[1099,1239,1821,1846]" pageId="15" pageNumber="178" refId="ref15129" refString="IUCN (2014) IUCN Red List Categories and Criteria: Version 3.1. 2 nd Edition. IUCN Species Survival Commission, IUCN Gland &amp; Cambridge. Available from: http: // jr. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 1 January 2015)" type="book" year="2014">IUCN 2014</bibRefCitation>
). The species only was known from the
<typeStatus id="54D349ABFF8F6B01FE9EB94AB4F5F54E" box="[374,422,1857,1882]" pageId="15" pageNumber="178">type</typeStatus>
locality and after thorough surveys by Pascal in the 1990s, the second author in the 2000s and a visit by the first author in
<date id="FFD6D1C9FF8F6B01FE07B96EB7F4F56A" box="[495,679,1893,1918]" pageId="15" pageNumber="178" value="2011-11">November 2011</date>
, it has not been recollected. Because the forest has been degraded by surrounding farms and non-native species it should be considered extinct.
</paragraph>
<paragraph id="8BD7F709FF8F6B01FF55B9A6B7D9FA1A" blockId="15.[136,1452,1749,2062]" pageId="15" pageNumber="178">
<emphasis id="B91C2B1BFF8F6B01FF55B9A6B453F5D2" bold="true" box="[189,256,1965,1990]" pageId="15" pageNumber="178">Notes</emphasis>
:
<emphasis id="B91C2B1BFF8F6B01FEE0B9A6B477F5D2" bold="true" box="[264,292,1965,1990]" pageId="15" pageNumber="178"></emphasis>
The
<typeStatus id="54D349ABFF8F6B01FEB0B9A6B4DBF5D2" box="[344,392,1965,1990]" pageId="15" pageNumber="178">type</typeStatus>
locality and collection number were described erroneously in the protologue, which
<bibRefCitation id="EFF98AF8FF8F6B01FAA6B9A6B475F5FE" author="Labat, J. - N. &amp; Schatz, G. E." pageId="15" pageNumber="178" pagination="201 - 205" refId="ref15422" refString="Labat, J. - N. &amp; Schatz, G. E. (2002) New combinations and a new name in Syzygium (Myrtaceae) from Madagascar and the Comoro Islands. Novon 12: 201 - 205. http: // dx. doi. org / 10.2307 / 3392957" type="journal article" year="2002">Labat &amp; Schatz (2002)</bibRefCitation>
discuss in more detail with the correct details cited above. Prior to this study it was the only endemic
<taxonomicName id="4C688C8AFF8F6B01FF60B9FEB419FA1A" box="[136,330,2037,2062]" pageId="15" pageNumber="178">
<emphasis id="B91C2B1BFF8F6B01FF60B9FEB5A1FA1A" box="[136,242,2037,2062]" italics="true" pageId="15" pageNumber="178">Syzygium</emphasis>
species
</taxonomicName>
recorded in the archipelago.
</paragraph>
</subSubSection>
</treatment>
</document>