treatments-xml/data/03/A8/B7/03A8B723553AFFA8FDBFE401FAC5C9B2.xml

<document id="F95924178624CE56455F806FCF09C1D0" ID-CLB-Dataset="304964" ID-DOI="10.5852/ejt.2024.967.2699" ID-GBIF-Dataset="0647b761-1496-4dcd-abb6-495b67abe85f" ID-ISSN="2118-9773" ID-Zenodo-Dep="13990819" ID-ZooBank="C08B8027-50CC-417E-BCD4-5183B9FF6738" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1729849508920" checkinUser="plazi" docAuthor="Zonstein, Sergei L." docDate="2024" docId="03A8B723553AFFA8FDBFE401FAC5C9B2" docLanguage="en" docName="EJT.2024.967.1-185.pdf" docOrigin="European Journal of Taxonomy 967" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola Zonstein 1987" docType="key" docVersion="7" lastPageNumber="19" masterDocId="FF91CF5B552AFFBAFFEFE72AFFB2CE5B" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia" masterLastPageNumber="185" masterPageNumber="1" pageNumber="17" updateTime="1730721133903" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EFE43D3EE528742C54FCDD7BEA271AF3">A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia</mods:title>
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<mods:namePart id="C30A022C741410EA769B1873A217D9E7">Zonstein, Sergei L.</mods:namePart>
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<subSubSection id="C31B55BE553AFFAAFDBFE401FC51CD1D" box="[592,995,811,838]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BBE0635553AFFAAFDBFE401FC51CD1D" blockId="16.[592,995,811,838]" box="[592,995,811,838]" pageId="16" pageNumber="17">
<heading id="D0F6B159553AFFAAFDBFE401FC51CD1D" box="[592,995,811,838]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
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<taxonomicName id="4C017DB6553AFFAAFDBFE401FD78CD1D" ID-CoL="77BJ" authority="Zonstein, 1987" authorityName="Zonstein" authorityYear="1987" box="[592,714,811,838]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Raveniola</taxonomicName>
concolor
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species group
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<emphasis id="B975DA27553AFFAAFF52E445FE7ECDD1" bold="true" box="[189,460,879,906]" pageId="16" pageNumber="17">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF52E48AFC9BCA9B" blockId="16.[186,1400,928,1216]" pageId="16" pageNumber="17">
Maxilla with numerous cuspules arranged in a triangular area (
<figureCitation id="133A1AB0553AFFAAFC5CE48AFBE8CDE0" box="[947,1114,928,955]" captionStart-0="Figs 202–210" captionStart-1="Figs 211–219" captionStartId-0="129.[189,241,1851,1877]" captionStartId-1="130.[189,241,1847,1873]" captionTargetBox-0="[189,1398,265,1816]" captionTargetBox-1="[189,1398,265,1816]" captionTargetId-0="figure-88@129.[189,1398,265,1816]" captionTargetId-1="figure-85@130.[189,1398,265,1816]" captionTargetPageId-0="129" captionTargetPageId-1="130" captionText-0="Figs 202–210. Raveniola spp., ♂♂, holotypes (except 204), sternum, labium and maxillae, ventral views. 202. R. caudata Zonstein, 2009 (SMNH). 203. R. inopinata sp. nov. (SMNH). 204. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 205. R. alajensis sp. nov. (SMNH). 206. R. concolor Zonstein, 2000 (MCSN AR 18). 207. R. cucullata sp. nov. (SMNH). 208. R. dolosa sp. nov. (SMNH). 209. R. hirta sp. nov. (SMNH). 210. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990878" figureDoi-1="http://doi.org/10.5281/zenodo.13990880" httpUri-0="https://zenodo.org/record/13990878/files/figure.png" httpUri-1="https://zenodo.org/record/13990880/files/figure.png" pageId="16" pageNumber="17">Figs 205–215</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFB86E48AFB62CDE1" box="[1129,1232,928,954]" captionStart-0="Figs 229–237" captionStart-1="Figs 238–246" captionStartId-0="132.[189,241,1827,1853]" captionStartId-1="133.[189,241,1820,1846]" captionTargetBox-0="[200,1385,265,1784]" captionTargetBox-1="[208,1381,265,1770]" captionTargetId-0="figure-90@132.[200,1385,265,1784]" captionTargetId-1="figure-91@133.[208,1381,265,1770]" captionTargetPageId-0="132" captionTargetPageId-1="133" captionText-0="Figs 229–237. Raveniola spp., ♀♀, paratypes (except 229–230), sternum, labium and maxillae, ventral views. 229. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 230. R. afghana sp. nov., holotype (NHMD). 231. R. alajensis sp. nov. (SMNH). 232. R. concolor Zonstein, 2000 (MCSN AR 19). 233– 234. R. cucullata sp. nov. from Shikildara (SMNH) and Shahob (SMNH), respectively. 235. R. dolosa sp. nov.(SMNH). 236. R. hirta sp. nov. (SMNH). 237. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990884" figureDoi-1="http://doi.org/10.5281/zenodo.13990887" httpUri-0="https://zenodo.org/record/13990884/files/figure.png" httpUri-1="https://zenodo.org/record/13990887/files/figure.png" pageId="16" pageNumber="17">230–242</figureCitation>
). PLS chiefly medium-sized, sometimes small (
<figureCitation id="133A1AB0553AFFAAFDA3E4EFFD44CDBB" box="[588,758,965,992]" captionStart-0="Figs 555–564" captionStart-1="Figs 565–574" captionStart-2="Figs 575–583" captionStart-3="Figs 584–592" captionStartId-0="160.[189,241,1786,1812]" captionStartId-1="161.[189,241,1825,1851]" captionStartId-2="162.[189,241,1857,1883]" captionStartId-3="163.[189,241,1821,1847]" captionTargetBox-0="[213,1375,265,1755]" captionTargetBox-1="[202,1386,265,1765]" captionTargetBox-2="[189,1398,265,1816]" captionTargetBox-3="[189,1398,265,1789]" captionTargetId-0="figure-109@160.[213,1375,265,1755]" captionTargetId-1="figure-100@161.[202,1386,265,1784]" captionTargetId-2="figure-74@162.[189,1398,265,1816]" captionTargetId-3="figure-92@163.[189,1398,265,1789]" captionTargetPageId-1="161" captionTargetPageId-2="162" captionTargetPageId-3="163" captionText-0="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 565–574. Raveniola spp., ♂♂ (565, 568, 571, 573) and ♀♀ (566–567, 569–570, 572), paratypes (except 568, 571, 573). Spinnerets, ventral (565–566, 571–574), lateral (567–568) and ventrolateral (569–570) views. 565–566. R. cucullata sp. nov., paratypes from Shahob (SMNH). 567. R. cucullata, from Sangdara (SMNH). 568. R. dolosa sp. nov. (SMNH). 569–570. R. dolosa, paratypes (SMNH). 571. R. hirta sp. nov., holotype (SMNH). 572. R. hirta, paratype (SMNH). 573. R. ignobilis sp. nov. (SMNH). 574. R. ignobilis, paratype (SMNH). Scale bars: 1.0 mm." captionText-2="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." captionText-3="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14018401" figureDoi-1="http://doi.org/10.5281/zenodo.14018403" figureDoi-2="http://doi.org/10.5281/zenodo.14018406" figureDoi-3="http://doi.org/10.5281/zenodo.14018408" httpUri-0="https://zenodo.org/record/14018401/files/figure.png" httpUri-1="https://zenodo.org/record/14018403/files/figure.png" httpUri-2="https://zenodo.org/record/14018406/files/figure.png" httpUri-3="https://zenodo.org/record/14018408/files/figure.png" pageId="16" pageNumber="17">Figs 559–588</figureCitation>
). Apical segment of PLS mostly triangular, though in some species shortly digitiform (as in 
<figureCitation id="133A1AB0553AFFAAFD7DE4C1FCC5CA5D" box="[658,887,1003,1030]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="16" pageNumber="17">Figs 577–579, 581</figureCitation>
). Males: tibiae and metatarsi I–II in many species of the group with long, thin and erect hairs (
<figureCitation id="133A1AB0553AFFAAFCCCE33AFC3CCA71" box="[803,910,1040,1066]" captionStart="Figs 256–264" captionStartId="135.[189,241,1824,1850]" captionTargetBox="[200,1388,265,1788]" captionTargetId="figure-97@135.[200,1388,265,1788]" captionTargetPageId="135" captionText="Figs 256–264. Raveniola spp., ♂♂; holotypes (except 260, 264), tibia and metatarsus I (256, 258, 260– 264) and tarsus IV (257, 259); retrolateral views. 256–257. R. caudata Zonstein, 2009 (SMNH). 258– 259. R. inopinatasp. nov. (SMNH). 260. R. redikorzevi(Spassky, 1937), ♂ fromAkar-Cheshme (SMNH). 261. R. alajensis sp. nov. (SMNH) 262. R. concolor Zonstein, 2000 (MCSN AR 18). 263. R. cucullata sp. nov. (SMNH). 264. R. cucullata, paratype from Shahob (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990893" httpUri="https://zenodo.org/record/13990893/files/figure.png" pageId="16" pageNumber="17">Figs 262</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFC74E33AFBC9CA71" box="[923,1147,1040,1067]" captionStart="Figs 265–273" captionStartId="136.[189,241,1765,1791]" captionTargetBox="[189,1398,265,1731]" captionTargetId="figure-88@136.[189,1398,265,1731]" captionTargetPageId="136" captionText="Figs 265–273. Raveniola spp., ♂♂, holotypes (except 272), tibia and metatarsus I, retrolateral views. 265. R. dolosa sp. nov. (SMNH). 266. R. hirta sp. nov. (SMNH). 267. R. ignobilis sp. nov. (SMNH). 268. R. insolita sp. nov. (ZISP). 269. R. ornata sp. nov. (SMNH). 270. R. ornatula sp. nov. (SMNH). 271. R. pamira sp. nov. (SMNH). 272. R. pamira, paratype (SMNH). 273. R. diluta sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990895" httpUri="https://zenodo.org/record/13990895/files/figure.png" pageId="16" pageNumber="17">265, 267, 269–272</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFB68E33AFB41CA71" box="[1159,1267,1040,1066]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="16" pageNumber="17">297–302</figureCitation>
); cymbium very short (as in 
<figureCitation id="133A1AB0553AFFAAFE97E31FFDA9CA0B" box="[376,539,1077,1104]" captionStart-0="Figs 349–363" captionStart-1="Figs 364–378" captionStartId-0="143.[189,241,1727,1753]" captionStartId-1="144.[189,241,1714,1740]" captionTargetBox-0="[189,1390,270,1690]" captionTargetBox-1="[189,1398,271,1690]" captionTargetId-0="figure-139@143.[189,1398,265,1690]" captionTargetId-1="figure-142@144.[189,1398,265,1690]" captionTargetPageId-0="143" captionTargetPageId-1="144" captionText-0="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990913" figureDoi-1="http://doi.org/10.5281/zenodo.13990915" httpUri-0="https://zenodo.org/record/13990913/files/figure.png" httpUri-1="https://zenodo.org/record/13990915/files/figure.png" pageId="16" pageNumber="17">Figs 352–364</figureCitation>
); tapering embolus bent and screwed subapically, with or without subapical keels (
<figureCitation id="133A1AB0553AFFAAFEEAE371FE18CA2D" box="[261,426,1115,1142]" captionStart-0="Figs 379–388" captionStart-1="Figs 389–399" captionStart-2="Figs 400–408" captionStart-3="Figs 409–417" captionStart-4="Figs 418–428" captionStartId-0="145.[189,241,1844,1870]" captionStartId-1="146.[189,241,1854,1880]" captionStartId-2="147.[189,241,1853,1879]" captionStartId-3="148.[189,241,1847,1873]" captionStartId-4="149.[189,241,1845,1871]" captionTargetBox-0="[189,1398,265,1806]" captionTargetBox-1="[189,1398,265,1814]" captionTargetBox-2="[198,1398,265,1817]" captionTargetBox-3="[189,1398,265,1801]" captionTargetBox-4="[189,1398,265,1794]" captionTargetId-0="figure-75@145.[189,1398,265,1806]" captionTargetId-1="figure-68@146.[189,1398,265,1814]" captionTargetId-2="figure-50@147.[189,1398,265,1817]" captionTargetId-3="figure-49@148.[189,1398,265,1801]" captionTargetId-4="figure-44@149.[189,1398,265,1794]" captionTargetPageId-0="145" captionTargetPageId-1="146" captionTargetPageId-2="147" captionTargetPageId-3="148" captionTargetPageId-4="149" captionText-0="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." captionText-1="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." captionText-2="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." captionText-3="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." captionText-4="Figs 418–428. Raveniola pamira sp. nov, ♂♂, holotype (418–422) and two paratypes (423–426 and 427–428, respectively), details of copulatoty bulb; retrolateral (418, 427), retroventral (419, 428), ventral (420, 423, 426), proventral (421, 425) and prolateral (422, 424) views (all SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990917" figureDoi-1="http://doi.org/10.5281/zenodo.13990919" figureDoi-2="http://doi.org/10.5281/zenodo.13990921" figureDoi-3="http://doi.org/10.5281/zenodo.13990923" figureDoi-4="http://doi.org/10.5281/zenodo.13990925" httpUri-0="https://zenodo.org/record/13990917/files/figure.png" httpUri-1="https://zenodo.org/record/13990919/files/figure.png" httpUri-2="https://zenodo.org/record/13990921/files/figure.png" httpUri-3="https://zenodo.org/record/13990923/files/figure.png" httpUri-4="https://zenodo.org/record/13990925/files/figure.png" pageId="16" pageNumber="17">Figs 385–428</figureCitation>
). Females: spermathecae mostly U- or V-shaped with wide bases and with inner and outer branches more or less widely separated apart (
<figureCitation id="133A1AB0553AFFAAFCC9E3AAFBF6CAC0" box="[806,1092,1152,1179]" captionStart-0="Figs 487–503" captionStart-1="Figs 504–521" captionStart-2="Figs 522–536" captionStartId-0="156.[189,241,1695,1721]" captionStartId-1="157.[189,241,1719,1745]" captionStartId-2="158.[189,241,1728,1754]" captionTargetBox-0="[189,1398,265,1653]" captionTargetBox-1="[189,1398,265,1675]" captionTargetBox-2="[189,1398,265,1688]" captionTargetId-0="figure-102@156.[189,1398,265,1653]" captionTargetId-1="figure-86@157.[189,1398,265,1675]" captionTargetId-2="figure-119@158.[189,1398,265,1688]" captionTargetPageId-0="156" captionTargetPageId-1="157" captionTargetPageId-2="158" captionText-0="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." captionText-1="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." captionText-2="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990941" figureDoi-1="http://doi.org/10.5281/zenodo.13990943" figureDoi-2="http://doi.org/10.5281/zenodo.13990945" httpUri-0="https://zenodo.org/record/13990941/files/figure.png" httpUri-1="https://zenodo.org/record/13990943/files/figure.png" httpUri-2="https://zenodo.org/record/13990945/files/figure.png" pageId="16" pageNumber="17">Figs 490–512, 520–525</figureCitation>
), or spermathecae narrow F- or Y-shaped in some species (
<figureCitation id="133A1AB0553AFFAAFDD5E38FFD17CA9B" box="[570,677,1189,1216]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="16" pageNumber="17">Figs 489</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFD5FE38FFCAECA9B" box="[688,796,1189,1216]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="16" pageNumber="17">513–519</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE553AFFAAFF52E3C3FB26CB24" pageId="16" pageNumber="17" type="taxon_list">
<paragraph id="8BBE0635553AFFAAFF52E3C3FE3ECB58" blockId="16.[189,396,1257,1283]" box="[189,396,1257,1283]" pageId="16" pageNumber="17">
<heading id="D0F6B159553AFFAAFF52E3C3FE3ECB58" bold="true" box="[189,396,1257,1283]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B975DA27553AFFAAFF52E3C3FE3ECB58" bold="true" box="[189,396,1257,1283]" pageId="16" pageNumber="17">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF52E230FB26CB24" blockId="16.[189,1400,1306,1407]" pageId="16" pageNumber="17">
<taxonomicName id="4C017DB6553AFFAAFF52E230FE2CCB6F" authorityName="Zonstein" authorityYear="2024" box="[189,414,1306,1332]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="afghana" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFF52E230FE2CCB6F" box="[189,414,1306,1332]" italics="true" pageId="16" pageNumber="17">Raveniola afghana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFE46E230FDBBCB6F" box="[425,521,1306,1332]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFDF8E231FD11CB6F" authorityName="Zonstein" authorityYear="2024" box="[535,675,1306,1332]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alajensis" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFDF8E231FD11CB6F" box="[535,675,1306,1332]" italics="true" pageId="16" pageNumber="17">R. alajensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFD41E230FCBCCB6F" box="[686,782,1306,1332]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFCF3E231FBDBCB6F" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[796,1129,1306,1332]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B975DA27553AFFAAFCF3E231FC1ACB6F" box="[796,936,1306,1332]" italics="true" pageId="16" pageNumber="17">R. concolor</emphasis>
<bibRefCitation id="EF907BC4553AFFAAFC5DE230FBDBCB6F" author="Zonstein S. L." box="[946,1129,1306,1332]" journalOrPublisher="Tethys Entomological Research" pageId="16" pageNumber="17" pagination="49 - 52" part="2" refId="ref83098" refString="Zonstein S. L. 2000 a. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C017DB6553AFFAAFB98E231FABACB6F" authorityName="Zonstein" authorityYear="2024" box="[1143,1288,1306,1332]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cucullata" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFB98E231FABACB6F" box="[1143,1288,1306,1332]" italics="true" pageId="16" pageNumber="17">R. cucullata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFAFDE230FAC0CB6F" box="[1298,1394,1306,1332]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFF52E26AFE9FCB02" authorityName="Zonstein" authorityYear="2024" box="[189,301,1343,1369]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dolosa" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFF52E26AFE9FCB02" box="[189,301,1343,1369]" italics="true" pageId="16" pageNumber="17">R. dolosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFED9E26AFE26CB01" box="[310,404,1344,1370]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFE4EE26AFE48CB02" authorityName="Zonstein" authorityYear="2024" box="[417,506,1343,1369]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hirta" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFE4EE26AFE48CB02" box="[417,506,1343,1369]" italics="true" pageId="16" pageNumber="17">R. hirta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFDEBE26AFDD0CB01" box="[516,610,1344,1370]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFD80E26AFD44CB02" authorityName="Zonstein" authorityYear="2024" box="[623,758,1343,1369]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ignobilis" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFD80E26AFD44CB02" box="[623,758,1343,1369]" italics="true" pageId="16" pageNumber="17">R. ignobilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFD10E26AFCEFCB01" box="[767,861,1344,1370]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFC85E26AFC51CB02" authorityName="Zonstein" authorityYear="2024" box="[874,995,1343,1369]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="insolita" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFC85E26AFC51CB02" box="[874,995,1343,1369]" italics="true" pageId="16" pageNumber="17">R. insolita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFC03E26AFBF8CB01" box="[1004,1098,1344,1370]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFBB8E26AFAB9CB02" authorityName="Zonstein" authorityYear="2024" box="[1111,1291,1343,1369]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="karategensis" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFBB8E26AFAB9CB02" box="[1111,1291,1343,1369]" italics="true" pageId="16" pageNumber="17">R. karategensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFAFAE26AFAC1CB01" box="[1301,1395,1344,1370]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFF51E24FFE9FCB25" authorityName="Zonstein" authorityYear="2024" box="[190,301,1381,1406]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFF51E24FFE9FCB25" box="[190,301,1381,1406]" italics="true" pageId="16" pageNumber="17">R. ornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFEDCE24FFE22CB24" box="[307,400,1381,1407]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFE73E24FFD90CB25" authorityName="Zonstein" authorityYear="2024" box="[412,546,1380,1406]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ornatula" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFE73E24FFD90CB25" box="[412,546,1380,1406]" italics="true" pageId="16" pageNumber="17">R. ornatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFDC6E24FFD34CB24" box="[553,646,1381,1407]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFD7DE24FFCB5CB25" authorityName="Zonstein" authorityYear="2024" box="[658,775,1381,1406]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pamira" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFD7DE24FFCB5CB25" box="[658,775,1381,1406]" italics="true" pageId="16" pageNumber="17">R. pamira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFCE1E24FFCDBCB24" box="[782,873,1381,1407]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C017DB6553AFFAAFC4DE24FFB80CB25" authorityName="Zonstein" authorityYear="2024" box="[930,1074,1380,1406]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sororcula" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFC4DE24FFB80CB25" box="[930,1074,1380,1406]" italics="true" pageId="16" pageNumber="17">R. sororcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFBD6E24FFB26CB24" box="[1081,1172,1381,1407]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31B55BE553AFFA8FF52E282FAC5C9B2" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="key">
<paragraph id="8BBE0635553AFFAAFF52E282FD1ECB99" blockId="16.[189,684,1448,1475]" box="[189,684,1448,1475]" pageId="16" pageNumber="17">
<heading id="D0F6B159553AFFAAFF52E282FD1ECB99" bold="true" box="[189,684,1448,1475]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B975DA27553AFFAAFF52E282FD1ECB99" bold="true" box="[189,684,1448,1475]" pageId="16" pageNumber="17">
Key to the species of the 
<taxonomicName id="4C017DB6553AFFAAFE1EE282FDE9CB98" ID-CoL="4RMTP" authorityName="Zonstein" authorityYear="2000" box="[497,603,1448,1475]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B975DA27553AFFAAFE1EE282FDE9CB98" bold="true" box="[497,603,1448,1475]" italics="true" pageId="16" pageNumber="17">concolor</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF05E2F3FE87CBA8" blockId="16.[189,1379,1497,1561]" box="[234,309,1497,1523]" pageId="16" pageNumber="17">
<emphasis id="B975DA27553AFFAAFF05E2F3FE87CBA8" bold="true" box="[234,309,1497,1523]" pageId="16" pageNumber="17">Males</emphasis>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF52E2D4FAD1C843" blockId="16.[189,1379,1497,1561]" box="[189,1379,1534,1561]" pageId="16" pageNumber="17">
Males of 
<taxonomicName id="4C017DB6553AFFAAFEC3E2D4FDB8C843" authorityName="Zonstein" authorityYear="2024" box="[300,522,1534,1560]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="afghana" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFEC3E2D4FDB8C843" box="[300,522,1534,1560]" italics="true" pageId="16" pageNumber="17">Raveniola afghana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFDFDE2D5FDDDC842" box="[530,623,1535,1561]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C017DB6553AFFAAFD95E2D5FC9DC843" authorityName="Zonstein" authorityYear="2024" box="[634,815,1534,1561]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="karategensis" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFD95E2D5FD3EC843" box="[634,652,1535,1560]" italics="true" pageId="16" pageNumber="17">R</emphasis>
. 
<emphasis id="B975DA27553AFFAAFD74E2D4FC9DC843" box="[667,815,1534,1560]" italics="true" pageId="16" pageNumber="17">karategensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFCD8E2D5FC23C842" box="[823,913,1535,1561]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C017DB6553AFFAAFC25E2D5FBE8C843" authorityName="Zonstein" authorityYear="2024" box="[970,1114,1534,1561]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sororcula" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFC25E2D5FC6EC843" box="[970,988,1535,1560]" italics="true" pageId="16" pageNumber="17">R</emphasis>
. 
<emphasis id="B975DA27553AFFAAFC05E2D4FBE8C843" box="[1002,1114,1534,1560]" italics="true" pageId="16" pageNumber="17">sororcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFB8DE2D5FB0EC842" box="[1122,1212,1535,1561]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
are unknown.
</paragraph>
<key id="FF905104553AFFABFF52E168FAC5CB18" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<keyStep id="30F51E90553AFFAAFF52E168FAC5C897" pageId="16" pageNumber="17">
<paragraph id="8BBE0635553AFFAAFF52E168FAC5C8DA" blockId="16.[189,1399,1602,1740]" pageId="16" pageNumber="17">
<keyLead id="30F0A500553AFFAAFF52E168FAC5C8DA" pageId="16" pageNumber="17">
1. Hairs on tibia and metatarsus IV unmodified. Embolus either subequal to tegulum in length, or considerably longer than the latter (
<figureCitation id="133A1AB0553AFFAAFD64E14DFC82C8D9" box="[651,816,1639,1666]" captionStart-0="Figs 379–388" captionStart-1="Figs 389–399" captionStartId-0="145.[189,241,1844,1870]" captionStartId-1="146.[189,241,1854,1880]" captionTargetBox-0="[189,1398,265,1806]" captionTargetBox-1="[189,1398,265,1814]" captionTargetId-0="figure-75@145.[189,1398,265,1806]" captionTargetId-1="figure-68@146.[189,1398,265,1814]" captionTargetPageId-0="145" captionTargetPageId-1="146" captionText-0="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." captionText-1="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990917" figureDoi-1="http://doi.org/10.5281/zenodo.13990919" httpUri-0="https://zenodo.org/record/13990917/files/figure.png" httpUri-1="https://zenodo.org/record/13990919/files/figure.png" pageId="16" pageNumber="17">Figs 385–399</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFCD4E14DFC16C8DA" box="[827,932,1639,1665]" captionStart-0="Figs 400–408" captionStart-1="Figs 409–417" captionStart-2="Figs 418–428" captionStartId-0="147.[189,241,1853,1879]" captionStartId-1="148.[189,241,1847,1873]" captionStartId-2="149.[189,241,1845,1871]" captionTargetBox-0="[198,1398,265,1817]" captionTargetBox-1="[189,1398,265,1801]" captionTargetBox-2="[189,1398,265,1794]" captionTargetId-0="figure-50@147.[189,1398,265,1817]" captionTargetId-1="figure-49@148.[189,1398,265,1801]" captionTargetId-2="figure-44@149.[189,1398,265,1794]" captionTargetPageId-0="147" captionTargetPageId-1="148" captionTargetPageId-2="149" captionText-0="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." captionText-1="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." captionText-2="Figs 418–428. Raveniola pamira sp. nov, ♂♂, holotype (418–422) and two paratypes (423–426 and 427–428, respectively), details of copulatoty bulb; retrolateral (418, 427), retroventral (419, 428), ventral (420, 423, 426), proventral (421, 425) and prolateral (422, 424) views (all SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990921" figureDoi-1="http://doi.org/10.5281/zenodo.13990923" figureDoi-2="http://doi.org/10.5281/zenodo.13990925" httpUri-0="https://zenodo.org/record/13990921/files/figure.png" httpUri-1="https://zenodo.org/record/13990923/files/figure.png" httpUri-2="https://zenodo.org/record/13990925/files/figure.png" pageId="16" pageNumber="17">403–428</figureCitation>
) ........................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF2EE1A7FAC5C897" blockId="16.[189,1399,1602,1740]" pageId="16" pageNumber="17">
<keyLead id="30F0A500553AFFAAFF2EE1A7FAC5C897" pageId="16" pageNumber="17">
– Tibia and metatarsus IV with dense thin and long modified hairs (
<figureCitation id="133A1AB0553AFFAAFC14E1A6FBD1C8FC" box="[1019,1123,1676,1703]" captionStart="Figs 310–318" captionStartId="140.[189,241,1786,1812]" captionTargetBox="[189,1398,265,1765]" captionTargetId="figure-96@140.[189,1398,265,1765]" captionTargetPageId="140" captionText="Figs 310–318. Raveniola spp., ♀♀ (310–312, 314–318) and ♂ (313), tibia and metatarsus IV, retrolateral views. 310. R. afghana sp. nov., holotype (NHMD). 311. R. alajensis sp. nov., paratype from Berksu. 312. R. cucullata sp. nov., paratype from Mullokoni (SMNH). 313–314. R. hirta sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 315. R. ignobilis sp. nov., paratype from Sangdara (SMNH). 316–317. R. karategensis sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 318. R. pamira sp. nov., paratype from Ubagandara (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990905" httpUri="https://zenodo.org/record/13990905/files/figure.png" pageId="16" pageNumber="17">Fig. 313</figureCitation>
). Embolus shorter than tegulum (
<figureCitation id="133A1AB0553AFFAAFEB5E198FE4CC897" box="[346,510,1714,1740]" captionStart="Figs 400–408" captionStartId="147.[189,241,1853,1879]" captionTargetBox="[198,1398,265,1817]" captionTargetId="figure-50@147.[189,1398,265,1817]" captionTargetPageId="147" captionText="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990921" httpUri="https://zenodo.org/record/13990921/files/figure.png" pageId="16" pageNumber="17">Figs 400–402</figureCitation>
)............................................................................................. 
<taxonomicName id="4C017DB6553AFFAAFB57E198FAA7C897" authorityName="Zonstein" authorityYear="2024" box="[1208,1301,1713,1740]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hirta" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFB57E198FAA7C897" bold="true" box="[1208,1301,1713,1740]" italics="true" pageId="16" pageNumber="17">R. hirta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFAF3E198FAC5C897" box="[1308,1399,1714,1740]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553AFFAAFF52E1DFFAC5C96E" pageId="16" pageNumber="17">
<paragraph id="8BBE0635553AFFAAFF52E1DFFAC5C954" blockId="16.[189,1399,1781,1845]" box="[189,1399,1781,1808]" pageId="16" pageNumber="17">
<keyLead id="30F0A500553AFFAAFF52E1DFFAC5C954" box="[189,1399,1781,1808]" pageId="16" pageNumber="17">
2. Embolus approximately as long as tegulum (
<figureCitation id="133A1AB0553AFFAAFD1AE1DFFC28C954" box="[757,922,1781,1807]" captionStart="Figs 389–399" captionStartId="146.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1814]" captionTargetId="figure-68@146.[189,1398,265,1814]" captionTargetPageId="146" captionText="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990919" httpUri="https://zenodo.org/record/13990919/files/figure.png" pageId="16" pageNumber="17">Figs 397–399</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFC49E1DFFBBDC954" box="[934,1039,1781,1807]" captionStart="Figs 400–408" captionStartId="147.[189,241,1853,1879]" captionTargetBox="[198,1398,265,1817]" captionTargetId="figure-50@147.[189,1398,265,1817]" captionTargetPageId="147" captionText="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990921" httpUri="https://zenodo.org/record/13990921/files/figure.png" pageId="16" pageNumber="17">403–408</figureCitation>
)............................................. 3
</keyLead>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF2EE031FAC5C96E" blockId="16.[189,1399,1781,1845]" box="[193,1399,1818,1845]" pageId="16" pageNumber="17">
<keyLead id="30F0A500553AFFAAFF2EE031FAC5C96E" box="[193,1399,1818,1845]" pageId="16" pageNumber="17">
– Embolus considerably longer than tegulum (
<figureCitation id="133A1AB0553AFFAAFD1CE031FC2AC96E" box="[755,920,1818,1845]" captionStart-0="Figs 379–388" captionStart-1="Figs 389–399" captionStartId-0="145.[189,241,1844,1870]" captionStartId-1="146.[189,241,1854,1880]" captionTargetBox-0="[189,1398,265,1806]" captionTargetBox-1="[189,1398,265,1814]" captionTargetId-0="figure-75@145.[189,1398,265,1806]" captionTargetId-1="figure-68@146.[189,1398,265,1814]" captionTargetPageId-0="145" captionTargetPageId-1="146" captionText-0="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." captionText-1="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990917" figureDoi-1="http://doi.org/10.5281/zenodo.13990919" httpUri-0="https://zenodo.org/record/13990917/files/figure.png" httpUri-1="https://zenodo.org/record/13990919/files/figure.png" pageId="16" pageNumber="17">Figs 385–396</figureCitation>
, 
<figureCitation id="133A1AB0553AFFAAFC4BE031FBBEC96E" box="[932,1036,1819,1845]" captionStart-0="Figs 409–417" captionStart-1="Figs 418–428" captionStartId-0="148.[189,241,1847,1873]" captionStartId-1="149.[189,241,1845,1871]" captionTargetBox-0="[189,1398,265,1801]" captionTargetBox-1="[189,1398,265,1794]" captionTargetId-0="figure-49@148.[189,1398,265,1801]" captionTargetId-1="figure-44@149.[189,1398,265,1794]" captionTargetPageId-0="148" captionTargetPageId-1="149" captionText-0="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." captionText-1="Figs 418–428. Raveniola pamira sp. nov, ♂♂, holotype (418–422) and two paratypes (423–426 and 427–428, respectively), details of copulatoty bulb; retrolateral (418, 427), retroventral (419, 428), ventral (420, 423, 426), proventral (421, 425) and prolateral (422, 424) views (all SMNH). Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990923" figureDoi-1="http://doi.org/10.5281/zenodo.13990925" httpUri-0="https://zenodo.org/record/13990923/files/figure.png" httpUri-1="https://zenodo.org/record/13990925/files/figure.png" pageId="16" pageNumber="17">409–428</figureCitation>
)............................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553AFFAAFF52E074FAC5C9B2" pageId="16" pageNumber="17">
<paragraph id="8BBE0635553AFFAAFF52E074FAC5C9C5" blockId="16.[189,1399,1886,2025]" pageId="16" pageNumber="17">
<keyLead id="30F0A500553AFFAAFF52E074FAC5C9C5" pageId="16" pageNumber="17">
3. Proximal part of embolus (viewed from lateral aspect) with two raised opposite keels (
<figureCitation id="133A1AB0553AFFAAFAEAE074FEAAC9C6" captionStart="Figs 400–408" captionStartId="147.[189,241,1853,1879]" captionTargetBox="[198,1398,265,1817]" captionTargetId="figure-50@147.[189,1398,265,1817]" captionTargetPageId="147" captionText="Figs 400–408. Raveniola spp., ♂♂, holotypes (except 406–408), details of copulatoty bulb; retrolateral (400, 403, 406–407), retroventral (401, 404) and ventral (402, 405, 408) views. 400–402. R. hirta sp. nov. (SMNH). 403–405. R. ignobilis sp. nov. (SMNH). 406–408. R. ignobilis, paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990921" httpUri="https://zenodo.org/record/13990921/files/figure.png" pageId="16" pageNumber="17">Figs 403, 406</figureCitation>
) ...................................................................................................................... 
<taxonomicName id="4C017DB6553AFFAAFB65E0AEFAA7C9C5" authorityName="Zonstein" authorityYear="2024" box="[1162,1301,1923,1950]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ignobilis" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFB65E0AEFAA7C9C5" bold="true" box="[1162,1301,1923,1950]" italics="true" pageId="16" pageNumber="17">R. ignobilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFAF3E0AEFAC5C9C5" box="[1308,1399,1924,1950]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE0635553AFFAAFF2EE080FAC5C9B2" blockId="16.[189,1399,1886,2025]" pageId="16" pageNumber="17">
<keyLead id="30F0A500553AFFAAFF2EE080FAC5C9B2" pageId="16" pageNumber="17">
– Proximal part of embolus with two poorly discernible vestigial keels (
<figureCitation id="133A1AB0553AFFAAFB03E083FEA5C9B3" captionStart="Figs 389–399" captionStartId="146.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1814]" captionTargetId="figure-68@146.[189,1398,265,1814]" captionTargetPageId="146" captionText="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990919" httpUri="https://zenodo.org/record/13990919/files/figure.png" pageId="16" pageNumber="17">Figs 397– 398</figureCitation>
) ........................................................................................................................ 
<taxonomicName id="4C017DB6553AFFAAFB4BE0E5FAA7C9B2" authorityName="Zonstein" authorityYear="2024" box="[1188,1301,1998,2025]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="dolosa" status="sp. nov.">
<emphasis id="B975DA27553AFFAAFB4BE0E5FAA7C9B2" bold="true" box="[1188,1301,1998,2025]" italics="true" pageId="16" pageNumber="17">R. dolosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553AFFAAFAF3E0E5FAC5C9B2" box="[1308,1399,1999,2025]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E623FAC5CFD0" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E623FAC5CF1C" blockId="17.[189,1399,265,396]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E623FAC5CF1C" pageId="17" pageNumber="18">
4. Sternum broadly oval (
<figureCitation id="133A1AB0553BFFABFDEEE623FC91CF7F" box="[513,803,265,292]" captionStart-0="Figs 202–210" captionStart-1="Figs 211–219" captionStartId-0="129.[189,241,1851,1877]" captionStartId-1="130.[189,241,1847,1873]" captionTargetBox-0="[189,1398,265,1816]" captionTargetBox-1="[189,1398,265,1816]" captionTargetId-0="figure-88@129.[189,1398,265,1816]" captionTargetId-1="figure-85@130.[189,1398,265,1816]" captionTargetPageId-0="129" captionTargetPageId-1="130" captionText-0="Figs 202–210. Raveniola spp., ♂♂, holotypes (except 204), sternum, labium and maxillae, ventral views. 202. R. caudata Zonstein, 2009 (SMNH). 203. R. inopinata sp. nov. (SMNH). 204. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 205. R. alajensis sp. nov. (SMNH). 206. R. concolor Zonstein, 2000 (MCSN AR 18). 207. R. cucullata sp. nov. (SMNH). 208. R. dolosa sp. nov. (SMNH). 209. R. hirta sp. nov. (SMNH). 210. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990878" figureDoi-1="http://doi.org/10.5281/zenodo.13990880" httpUri-0="https://zenodo.org/record/13990878/files/figure.png" httpUri-1="https://zenodo.org/record/13990880/files/figure.png" pageId="17" pageNumber="18">Figs 205–211, 214–215</figureCitation>
). PMS medium-sized to moderately small; apical segment of PLS triangular (
<figureCitation id="133A1AB0553BFFABFDC0E606FD66CF1C" box="[559,724,300,327]" captionStart="Figs 555–564" captionStartId="160.[189,241,1786,1812]" captionTargetBox="[213,1375,265,1755]" captionTargetId="figure-109@160.[213,1375,265,1755]" captionText="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018401" httpUri="https://zenodo.org/record/14018401/files/figure.png" pageId="17" pageNumber="18">Figs 560, 563</figureCitation>
, 565, 568, 571, 573, 575, 584)........................................ 5
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE665FAC5CFD0" blockId="17.[189,1399,265,396]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE665FAC5CFD0" pageId="17" pageNumber="18">
– Sternum subcircular (
<figureCitation id="133A1AB0553BFFABFE09E664FD3FCF32" box="[486,653,334,361]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="17" pageNumber="18">Figs 212–213</figureCitation>
). PMS small and thin; apical segment of PLS shortly digitiform (
<figureCitation id="133A1AB0553BFFABFF1DE65BFEEFCFD7" box="[242,349,369,396]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="17" pageNumber="18">Figs 577</figureCitation>
, 781)................................................................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E698FAC5CFB4" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E698FAC5CF97" blockId="17.[189,1399,434,496]" box="[189,1399,434,461]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E698FAC5CF97" box="[189,1399,434,461]" pageId="17" pageNumber="18">
5. Abdomen with darker chevron-like pattern (
<figureCitation id="133A1AB0553BFFABFD1FE698FC4BCF97" box="[752,1017,434,460]" captionStart-0="Figs 1–9" captionStart-1="Figs 10–18" captionStartId-0="107.[189,241,1848,1874]" captionStartId-1="108.[189,241,1826,1852]" captionTargetBox-0="[189,1398,268,1805]" captionTargetBox-1="[189,1398,265,1785]" captionTargetId-0="figure-84@107.[189,1398,265,1805]" captionTargetId-1="figure-83@108.[189,1398,265,1785]" captionTargetPageId-0="107" captionTargetPageId-1="108" captionText-0="Figs 1–9.Raveniola spp., ♂♂, holotypes (except 3, 6), habitus, dorsal views. 1. R. caudataZonstein, 2009 (SMNH). 2. R. inopinata sp. nov. (SMNH). 3. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 4. R. alajensis sp. nov. (SMNH). 5. R. cucullata sp. nov. (SMNH). 6. R. cucullata, paratype from Shahob (SMNH). 7. R. dolosa sp. nov. (SMNH). 8. R. hirta sp. nov. (SMNH). 9. R. ignobilis sp. nov. (SMNH). Scale bars: 5.0 mm." captionText-1="Figs 10–18. Raveniola spp., ♂♂, holotypes (except 14, 16), habitus, dorsal views. 10. R. insolita sp. nov. (ZISP). 11. R. ornata sp. nov. (SMNH). 12. R. ornatula sp. nov. (SMNH). 13. R. pamira sp. nov. (SMNH). 14. R. pamira sp. nov., paratype (SMNH). 15. R. diluta sp. nov. (SMNH). 16. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 17. R. pallens sp. nov. (SMNH). 18. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990824" figureDoi-1="http://doi.org/10.5281/zenodo.13990826" httpUri-0="https://zenodo.org/record/13990824/files/figure.png" httpUri-1="https://zenodo.org/record/13990826/files/figure.png" pageId="17" pageNumber="18">Figs 4–7, 9–10, 13–14</figureCitation>
) ............................................... 6
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE6FCFAC5CFB4" blockId="17.[189,1399,434,496]" box="[193,1399,469,496]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE6FCFAC5CFB4" box="[193,1399,469,496]" pageId="17" pageNumber="18">
– Abdomen uniformly dark coloured ......................................................... 
<taxonomicName id="4C017DB6553BFFABFBDCE6FFFAC5CFB4" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[1075,1399,469,496]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B975DA27553BFFABFBDCE6FFFB0DCFAB" bold="true" box="[1075,1215,469,496]" italics="true" pageId="17" pageNumber="18">R. concolor</emphasis>
<bibRefCitation id="EF907BC4553BFFABFB29E6FFFAC5CFB4" author="Zonstein S. L." box="[1222,1399,469,495]" journalOrPublisher="Tethys Entomological Research" pageId="17" pageNumber="18" pagination="49 - 52" part="2" refId="ref83098" refString="Zonstein S. L. 2000 a. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E53CFAC5CCC2" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E53CFAC5CC08" blockId="17.[189,1399,534,665]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E53CFAC5CC08" pageId="17" pageNumber="18">
6. Palpal tibia very long, 4.7–5.1 times as long as wide (
<figureCitation id="133A1AB0553BFFABFC89E53CFBBDCC6B" box="[870,1039,534,561]" captionStart="Figs 349–363" captionStartId="143.[189,241,1727,1753]" captionTargetBox="[189,1390,270,1690]" captionTargetId="figure-139@143.[189,1398,265,1690]" captionTargetPageId="143" captionText="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990913" httpUri="https://zenodo.org/record/13990913/files/figure.png" pageId="17" pageNumber="18">Figs 352, 360</figureCitation>
). Embolus with long proximal and short distal sections, as in 
<figureCitation id="133A1AB0553BFFABFDBEE513FD46CC0F" box="[593,756,569,596]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="17" pageNumber="18">Figs 385–387</figureCitation>
, 
<figureCitation id="133A1AB0553BFFABFCEFE513FCD7CC08" box="[768,869,569,595]" captionStart="Figs 409–417" captionStartId="148.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1801]" captionTargetId="figure-49@148.[189,1398,265,1801]" captionTargetPageId="148" captionText="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990923" httpUri="https://zenodo.org/record/13990923/files/figure.png" pageId="17" pageNumber="18">409–411</figureCitation>
..................................................................... 7
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE576FAC5CCC2" blockId="17.[189,1399,534,665]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE576FAC5CCC2" pageId="17" pageNumber="18">
– Palpal tibia shorter, 4–4.5 times as long as wide (
<figureCitation id="133A1AB0553BFFABFCFBE576FB99CC2D" box="[788,1067,603,630]" captionStart-0="Figs 349–363" captionStart-1="Figs 364–378" captionStartId-0="143.[189,241,1727,1753]" captionStartId-1="144.[189,241,1714,1740]" captionTargetBox-0="[189,1390,270,1690]" captionTargetBox-1="[189,1398,271,1690]" captionTargetId-0="figure-139@143.[189,1398,265,1690]" captionTargetId-1="figure-142@144.[189,1398,265,1690]" captionTargetPageId-0="143" captionTargetPageId-1="144" captionText-0="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." captionText-1="Figs 364–378. Raveniola spp., ♂♂, holotypes (except 366, 369, 376, 378), palpal tibia, cymbium and copulatory bulb, retrolateral (364–377) and retroventral (378) views. 364. R. pamira sp. nov. (SMNH). 365. R. diluta sp. nov. (SMNH). 366. R. fedotovi (Charitonov, 1946), lectotype (ZMMU). 367. R. pallens sp. nov. (SMNH). 368. R. zyuzini sp. nov. (SMNH). 369. R. zyuzini, paratype (SMNH). 370. R. ferghanensis (Zonstein, 1984) (ZISP). 371. R. kirgizica sp. nov. (SMNH).372. R. kopetdaghensis (Fet, 1984) (ZISP). 373. R. mikhailovi Zonstein, 2021 (SMNH). 374. R. nenilini sp. nov. (SMNH). 375. R. ovchinnikovi sp. nov. (SMNH). 376. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 377. R. vulpina sp. nov. (SMNH). 378. R. vulpina, paratype from Tereksai (SMNH). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990913" figureDoi-1="http://doi.org/10.5281/zenodo.13990915" httpUri-0="https://zenodo.org/record/13990913/files/figure.png" httpUri-1="https://zenodo.org/record/13990915/files/figure.png" pageId="17" pageNumber="18">Figs 354–355, 363–364</figureCitation>
). Embolus differs (
<figureCitation id="133A1AB0553BFFABFAECE576FEABCCC3" captionStart="Figs 389–399" captionStartId="146.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1814]" captionTargetId="figure-68@146.[189,1398,265,1814]" captionTargetPageId="146" captionText="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990919" httpUri="https://zenodo.org/record/13990919/files/figure.png" pageId="17" pageNumber="18">Figs 389– 396</figureCitation>
, 
<figureCitation id="133A1AB0553BFFABFECAE555FE3FCCC2" box="[293,397,639,665]" captionStart="Figs 418–428" captionStartId="149.[189,241,1845,1871]" captionTargetBox="[189,1398,265,1794]" captionTargetId="figure-44@149.[189,1398,265,1794]" captionTargetPageId="149" captionText="Figs 418–428. Raveniola pamira sp. nov, ♂♂, holotype (418–422) and two paratypes (423–426 and 427–428, respectively), details of copulatoty bulb; retrolateral (418, 427), retroventral (419, 428), ventral (420, 423, 426), proventral (421, 425) and prolateral (422, 424) views (all SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990925" httpUri="https://zenodo.org/record/13990925/files/figure.png" pageId="17" pageNumber="18">418–428</figureCitation>
) .................................................................................................................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E5EAFAC5CD19" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E5EAFAC5CCA6" blockId="17.[189,1399,703,834]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E5EAFAC5CCA6" pageId="17" pageNumber="18">
7. Palpal tibia longer, 5–5.1 times as long as wide (
<figureCitation id="133A1AB0553BFFABFCD7E595FC10CC81" box="[824,930,703,730]" captionStart="Figs 349–363" captionStartId="143.[189,241,1727,1753]" captionTargetBox="[189,1390,270,1690]" captionTargetId="figure-139@143.[189,1398,265,1690]" captionTargetPageId="143" captionText="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990913" httpUri="https://zenodo.org/record/13990913/files/figure.png" pageId="17" pageNumber="18">Fig. 352</figureCitation>
). Proximal part of embolus stouter and clearly keeled (
<figureCitation id="133A1AB0553BFFABFE4FE5C8FDF1CCA6" box="[416,579,738,765]" captionStart="Figs 379–388" captionStartId="145.[189,241,1844,1870]" captionTargetBox="[189,1398,265,1806]" captionTargetId="figure-75@145.[189,1398,265,1806]" captionTargetPageId="145" captionText="Figs 379–388. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb, retrolateral (379, 381, 383, 385, 388), ventral (380, 382, 384, 387) and retroventral (386) views. 379–380. R. caudata Zonstein, 2009 (SMNH). 381–382. R. inopinata sp. nov. (SMNH). 383–384. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH).385–387. R. alajensis sp. nov. (SMNH). 388. R. concolor Zonstein, 2000 (MCSN AR 18). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990917" httpUri="https://zenodo.org/record/13990917/files/figure.png" pageId="17" pageNumber="18">Figs 385–387</figureCitation>
) ............................................................................. 
<taxonomicName id="4C017DB6553BFFABFB66E5C9FAA7CCA6" authorityName="Zonstein" authorityYear="2024" box="[1161,1301,738,765]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alajensis" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB66E5C9FAA7CCA6" bold="true" box="[1161,1301,738,765]" italics="true" pageId="17" pageNumber="18">R. alajensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E5C9FAC5CCA6" box="[1308,1399,739,765]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE42CFAC5CD19" blockId="17.[189,1399,703,834]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE42CFAC5CD19" pageId="17" pageNumber="18">
– Palpal tibia relatively shorter, 4.7 times as long as wide (
<figureCitation id="133A1AB0553BFFABFC61E42FFC45CD44" box="[910,1015,773,799]" captionStart="Figs 349–363" captionStartId="143.[189,241,1727,1753]" captionTargetBox="[189,1390,270,1690]" captionTargetId="figure-139@143.[189,1398,265,1690]" captionTargetPageId="143" captionText="Figs 349–363. Raveniola spp., ♂♂, holotypes (except 351, 355, 363), palpal tibia, cymbium and copulatory bulb, retrolateral (349–358, 360–361, 363) and retroventral (359, 362) views.349. R. caudata Zonstein, 2009 (SMNH). 350. R. inopinata sp. nov. (SMNH). 351. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 352. R. alajensis sp. nov. (SMNH). 353. R. concolor Zonstein, 2000 (MCSN AR 18). 354. R. cucullata sp. nov. (SMNH). 355. R. cucullata, paratype from Shahob (SMNH). 356. R. dolosa sp. nov. (SMNH). 357. R. hirta sp. nov. (SMNH). 358–359. R. ignobilis sp. nov. (SMNH). 360. R. insolita sp. nov. (ZISP). 361. R. ornata sp. nov. (SMNH). 362. R. ornatula sp. nov. (SMNH). 363. R. pamira sp. nov., paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990913" httpUri="https://zenodo.org/record/13990913/files/figure.png" pageId="17" pageNumber="18">Fig. 360</figureCitation>
). Proximal part of embolus with keel vestige and thinner (
<figureCitation id="133A1AB0553BFFABFDFDE402FD07CD19" box="[530,693,808,834]" captionStart="Figs 409–417" captionStartId="148.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1801]" captionTargetId="figure-49@148.[189,1398,265,1801]" captionTargetPageId="148" captionText="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990923" httpUri="https://zenodo.org/record/13990923/files/figure.png" pageId="17" pageNumber="18">Figs 409–411</figureCitation>
)................................................................ 
<taxonomicName id="4C017DB6553BFFABFB76E402FAA7CD19" authorityName="Zonstein" authorityYear="2024" box="[1177,1301,807,834]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="insolita" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB76E402FAA7CD19" bold="true" box="[1177,1301,807,834]" italics="true" pageId="17" pageNumber="18">R. insolita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E402FAC5CD19" box="[1308,1399,808,834]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E443FAC5CA0F" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E443FAC5CDB7" blockId="17.[189,1399,873,1108]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E443FAC5CDB7" pageId="17" pageNumber="18">
8. Tibia and metatarsus I elongate, slender, and covered with very long thin setae (
<figureCitation id="133A1AB0553BFFABFB17E443FEA9CDFD" captionStart="Figs 265–273" captionStartId="136.[189,241,1765,1791]" captionTargetBox="[189,1398,265,1731]" captionTargetId="figure-88@136.[189,1398,265,1731]" captionTargetPageId="136" captionText="Figs 265–273. Raveniola spp., ♂♂, holotypes (except 272), tibia and metatarsus I, retrolateral views. 265. R. dolosa sp. nov. (SMNH). 266. R. hirta sp. nov. (SMNH). 267. R. ignobilis sp. nov. (SMNH). 268. R. insolita sp. nov. (ZISP). 269. R. ornata sp. nov. (SMNH). 270. R. ornatula sp. nov. (SMNH). 271. R. pamira sp. nov. (SMNH). 272. R. pamira, paratype (SMNH). 273. R. diluta sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990895" httpUri="https://zenodo.org/record/13990895/files/figure.png" pageId="17" pageNumber="18">Figs 271– 272</figureCitation>
). Metatarsus I, viewed from ventral aspect, with narrowed arcuate proximal section (
<figureCitation id="133A1AB0553BFFABFF1CE484FE2DCD93" box="[243,415,942,968]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="17" pageNumber="18">Figs 301, 302</figureCitation>
). Embolus with almost equally short and low dorsal and ventral keels (
<figureCitation id="133A1AB0553BFFABFB11E484FEAACDB0" captionStart="Figs 418–428" captionStartId="149.[189,241,1845,1871]" captionTargetBox="[189,1398,265,1794]" captionTargetId="figure-44@149.[189,1398,265,1794]" captionTargetPageId="149" captionText="Figs 418–428. Raveniola pamira sp. nov, ♂♂, holotype (418–422) and two paratypes (423–426 and 427–428, respectively), details of copulatoty bulb; retrolateral (418, 427), retroventral (419, 428), ventral (420, 423, 426), proventral (421, 425) and prolateral (422, 424) views (all SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990925" httpUri="https://zenodo.org/record/13990925/files/figure.png" pageId="17" pageNumber="18">Figs 418– 428</figureCitation>
) ....................................................................................................................... 
<taxonomicName id="4C017DB6553BFFABFB73E4FBFAA7CDB0" authorityName="Zonstein" authorityYear="2024" box="[1180,1301,977,1003]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="pamira" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB73E4FBFAA7CDB0" bold="true" box="[1180,1301,977,1003]" italics="true" pageId="17" pageNumber="18">R. pamira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E4F8FAC5CDB7" box="[1308,1399,978,1004]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE4DEFAC5CA0F" blockId="17.[189,1399,873,1108]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE4DEFAC5CA0F" pageId="17" pageNumber="18">
– Tibia and metatarsus I stouter and covered with less modified shorter setae (
<figureCitation id="133A1AB0553BFFABFBAEE4DEFB54CA55" box="[1089,1254,1012,1038]" captionStart="Figs 256–264" captionStartId="135.[189,241,1824,1850]" captionTargetBox="[200,1388,265,1788]" captionTargetId="figure-97@135.[200,1388,265,1788]" captionTargetPageId="135" captionText="Figs 256–264. Raveniola spp., ♂♂; holotypes (except 260, 264), tibia and metatarsus I (256, 258, 260– 264) and tarsus IV (257, 259); retrolateral views. 256–257. R. caudata Zonstein, 2009 (SMNH). 258– 259. R. inopinatasp. nov. (SMNH). 260. R. redikorzevi(Spassky, 1937), ♂ fromAkar-Cheshme (SMNH). 261. R. alajensis sp. nov. (SMNH) 262. R. concolor Zonstein, 2000 (MCSN AR 18). 263. R. cucullata sp. nov. (SMNH). 264. R. cucullata, paratype from Shahob (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990893" httpUri="https://zenodo.org/record/13990893/files/figure.png" pageId="17" pageNumber="18">Figs 263–264</figureCitation>
). Metatarsus I, viewed from ventral aspect, without certain basal constriction (
<figureCitation id="133A1AB0553BFFABFC30E33DFB35CA6A" box="[991,1159,1047,1073]" captionStart="Figs 290–309" captionStartId="139.[189,241,1678,1704]" captionTargetBox="[189,1398,265,1651]" captionTargetId="figure-180@139.[189,1398,265,1651]" captionTargetPageId="139" captionText="Figs 290–309. Raveniola spp., ♂♂, holotypes (except 290, 294, 300, 302, 308), metatarsus I, ventral views. 290. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme (SMNH). 291. R. alajensis sp. nov. (SMNH). 292. R. concolor Zonstein, 2000 (MCSN AR 18). 293. R. cucullata sp. nov. (SMNH). 294. R. cucullata, paratype from Shahob (SMNH).295. R. dolosa sp. nov.(SMNH). 296. R. hirtasp. nov. (SMNH). 297. R. ignobilis sp. nov. (SMNH). 298. R. ornata sp. nov. (SMNH). 299. R. ornatula sp. nov. (SMNH). 300. R. ornatula, paratype (SMNH).301. R. pamirasp. nov. (SMNH). 302. R. pamira, paratype (SMNH). 303. R. diluta sp. nov. (SMNH). 304. R. kirgizica sp. nov. (SMNH). 305. R. kopetdaghensis (Fet, 1984) (ZISP). 306. R. mikhailovi Zonstein, 2021 (SMNH). 307. R. nenilini sp. nov. (SMNH). 308. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 309. R. vulpina sp. nov. (SMNH). Modified hairs are indicated with arrows. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990903" httpUri="https://zenodo.org/record/13990903/files/figure.png" pageId="17" pageNumber="18">Figs 293–294</figureCitation>
). Embolus with long and raised ventral keel; dorsal keel absent (
<figureCitation id="133A1AB0553BFFABFD0BE313FC3BCA08" box="[740,905,1081,1107]" captionStart="Figs 389–399" captionStartId="146.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1814]" captionTargetId="figure-68@146.[189,1398,265,1814]" captionTargetPageId="146" captionText="Figs 389–399. Raveniola spp., ♂♂, holotypes (except 392–396), details of copulatoty bulb, retrolateral (389, 392, 395, 397), retroventral (390, 393, 398) and ventral (391, 394, 396, 399) views. 389– 391. R. cucullata sp. nov. (SMNH). 392–394. R. cucullata sp. nov., paratype from Shahob (SMNH). 395–396. R. cucullata, paratype from Sangdara (SMNH). 397–399. R. dolosa sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990919" httpUri="https://zenodo.org/record/13990919/files/figure.png" pageId="17" pageNumber="18">Figs 389–396</figureCitation>
, 
<figureCitation id="133A1AB0553BFFABFC7BE313FC4FCA08" box="[916,1021,1081,1107]" captionStart="Figs 466–474" captionStartId="154.[189,241,1854,1880]" captionTargetBox="[189,1398,265,1817]" captionTargetId="figure-57@154.[189,1398,265,1817]" captionTargetPageId="154" captionText="Figs 466–474. Raveniola spp., ♂♂, details of copulatoty bulb (466–467, 469, 471, 473) and tip of embolus (468, 470, 472, 474), ventral (466, 473–474), retrolateral (467–468) and retoventral (469–472) views. 466–468. R. redikorzevi (Spassky, 1937) (SMNH). 469–470. R. cucullata sp. nov. (SMNH). 471–474. R. diluta sp. nov. (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.13990937" httpUri="https://zenodo.org/record/13990937/files/figure.png" pageId="17" pageNumber="18">469–470</figureCitation>
)................ 
<taxonomicName id="4C017DB6553BFFABFB6DE310FAA6CA0F" authorityName="Zonstein" authorityYear="2024" box="[1154,1300,1081,1108]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cucullata" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB6DE310FAA6CA0F" bold="true" box="[1154,1300,1081,1108]" italics="true" pageId="17" pageNumber="18">R. cucullata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E310FAC5CA0F" box="[1308,1399,1082,1108]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E351FAC5CB18" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E351FAC5CA80" blockId="17.[189,1399,1146,1347]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E351FAC5CA80" pageId="17" pageNumber="18">
9. Labium longer and narrower, maxillae with relatively smaller but more numerous cuspules (
<figureCitation id="133A1AB0553BFFABFF1CE3B7FEE9CAEC" box="[243,347,1181,1207]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="17" pageNumber="18">Fig. 212</figureCitation>
). Tarsi I–IV apically with relatively dense laterodistal tufts of long setae. Embolus longer and less tapering, with relatively shorter distal part (
<figureCitation id="133A1AB0553BFFABFCA2E3EAFC43CA81" box="[845,1009,1216,1242]" captionStart="Figs 409–417" captionStartId="148.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1801]" captionTargetId="figure-49@148.[189,1398,265,1801]" captionTargetPageId="148" captionText="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990923" httpUri="https://zenodo.org/record/13990923/files/figure.png" pageId="17" pageNumber="18">Figs 412–414</figureCitation>
)....................... 
<taxonomicName id="4C017DB6553BFFABFB4CE3EAFAA7CA81" authorityName="Zonstein" authorityYear="2024" box="[1187,1301,1216,1242]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB4CE3EAFAA7CA81" bold="true" box="[1187,1301,1216,1242]" italics="true" pageId="17" pageNumber="18">R. ornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E3EBFAC5CA80" box="[1308,1399,1217,1243]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE3C9FAC5CB18" blockId="17.[189,1399,1146,1347]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE3C9FAC5CB18" pageId="17" pageNumber="18">
– Labium shorter and broader, maxillae with relatively larger but less numerous cuspules (
<figureCitation id="133A1AB0553BFFABFAEDE3C9FADBCAA6" box="[1282,1385,1251,1277]" captionStart="Figs 211–219" captionStartId="130.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-85@130.[189,1398,265,1816]" captionTargetPageId="130" captionText="Figs 211–219. Raveniola spp., ♂♂, holotypes (except 215, 217), sternum, labium and maxillae, ventral views. 211. R. insolita sp. nov. (ZISP). 212. R. ornata sp. nov. (SMNH). 213. R. ornatula sp. nov. (SMNH). 214. R. pamira sp. nov. (SMNH). 215. R. pamira, paratype (SMNH). 216. R. diluta sp. nov. (SMNH). 217. R. fedotovi (Charitonov, 1946), ♂ from Hojakurgan (SMNH). 218. R. pallens sp. nov. (SMNH). 219. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990880" httpUri="https://zenodo.org/record/13990880/files/figure.png" pageId="17" pageNumber="18">Fig. 213</figureCitation>
). Tarsi I–IV apically with sparse laterodistal tufts of short setae. Embolus shorter and more tapering, with relatively longer distal part (
<figureCitation id="133A1AB0553BFFABFD9CE202FCA5CB18" box="[627,791,1320,1347]" captionStart="Figs 409–417" captionStartId="148.[189,241,1847,1873]" captionTargetBox="[189,1398,265,1801]" captionTargetId="figure-49@148.[189,1398,265,1801]" captionTargetPageId="148" captionText="Figs 409–417. Raveniola spp., ♂♂, holotypes, details of copulatoty bulb; retroventral (409, 413, 416), retrolateral (410, 412, 415) and ventral (411, 414, 417) views. 409–411. R. insolita sp. nov. (ZISP). 412–414. R. ornata sp. nov. (SMNH). 415–417. R. ornatula sp. nov. (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990923" httpUri="https://zenodo.org/record/13990923/files/figure.png" pageId="17" pageNumber="18">Figs 415–417</figureCitation>
) ................................................ 
<taxonomicName id="4C017DB6553BFFABFB65E203FAA7CB18" authorityName="Zonstein" authorityYear="2024" box="[1162,1301,1320,1347]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ornatula" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB65E203FAA7CB18" bold="true" box="[1162,1301,1320,1347]" italics="true" pageId="17" pageNumber="18">R. ornatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E203FAC5CB18" box="[1308,1399,1321,1347]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
<paragraph id="8BBE0635553BFFABFF05E240FEE3CBDF" blockId="17.[189,805,1386,1447]" box="[234,337,1386,1412]" pageId="17" pageNumber="18">
<emphasis id="B975DA27553BFFABFF05E240FEE3CBDF" bold="true" box="[234,337,1386,1412]" pageId="17" pageNumber="18">Females</emphasis>
</paragraph>
<paragraph id="8BBE0635553BFFABFF52E2A6FC97CBFD" blockId="17.[189,805,1386,1447]" box="[189,805,1420,1447]" pageId="17" pageNumber="18">
Females of 
<taxonomicName id="4C017DB6553BFFABFEA9E2A6FDAECBFD" authorityName="Zonstein" authorityYear="2024" box="[326,540,1420,1446]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="insolita" status="sp. nov.">
<emphasis id="B975DA27553BFFABFEA9E2A6FDAECBFD" box="[326,540,1420,1446]" italics="true" pageId="17" pageNumber="18">Raveniola insolita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFDCCE2A7FDCCCBFC" box="[547,638,1421,1447]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
are unknown.
</paragraph>
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<keyStep id="30F51E90553BFFABFF52E2E7FAC5C828" pageId="17" pageNumber="18">
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1. Anterodistal edge of chelicera with dense brush of stout setae in front of fang base. Tibia and metatarsus IV with dense, evenly thin, long, and more or less uniformly arranged modified hairs (
<figureCitation id="133A1AB0553BFFABFF1CE139FDF7C876" box="[243,581,1555,1581]" captionStart="Figs 310–318" captionStartId="140.[189,241,1786,1812]" captionTargetBox="[189,1398,265,1765]" captionTargetId="figure-96@140.[189,1398,265,1765]" captionTargetPageId="140" captionText="Figs 310–318. Raveniola spp., ♀♀ (310–312, 314–318) and ♂ (313), tibia and metatarsus IV, retrolateral views. 310. R. afghana sp. nov., holotype (NHMD). 311. R. alajensis sp. nov., paratype from Berksu. 312. R. cucullata sp. nov., paratype from Mullokoni (SMNH). 313–314. R. hirta sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 315. R. ignobilis sp. nov., paratype from Sangdara (SMNH). 316–317. R. karategensis sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 318. R. pamira sp. nov., paratype from Ubagandara (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990905" httpUri="https://zenodo.org/record/13990905/files/figure.png" pageId="17" pageNumber="18">Figs 310–311, 314, 316–317</figureCitation>
)........................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE11CFAC5C828" blockId="17.[189,1399,1485,1651]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE11CFAC5C828" pageId="17" pageNumber="18">
– Anterodistal edge of chelicera without dense brush of stout setae. Hairs on tibia and metatarsus IV heterogeneous and unmodified (as in 
<figureCitation id="133A1AB0553BFFABFD4DE172FCCCC829" box="[674,894,1624,1651]" captionStart="Figs 310–318" captionStartId="140.[189,241,1786,1812]" captionTargetBox="[189,1398,265,1765]" captionTargetId="figure-96@140.[189,1398,265,1765]" captionTargetPageId="140" captionText="Figs 310–318. Raveniola spp., ♀♀ (310–312, 314–318) and ♂ (313), tibia and metatarsus IV, retrolateral views. 310. R. afghana sp. nov., holotype (NHMD). 311. R. alajensis sp. nov., paratype from Berksu. 312. R. cucullata sp. nov., paratype from Mullokoni (SMNH). 313–314. R. hirta sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 315. R. ignobilis sp. nov., paratype from Sangdara (SMNH). 316–317. R. karategensis sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 318. R. pamira sp. nov., paratype from Ubagandara (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990905" httpUri="https://zenodo.org/record/13990905/files/figure.png" pageId="17" pageNumber="18">Figs 312, 315, 318</figureCitation>
) ................................................................ 5
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</paragraph>
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<keyStep id="30F51E90553BFFABFF52E1B3FAC5C947" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E1B3FAC5C88D" blockId="17.[189,1399,1689,1820]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E1B3FAC5C88D" pageId="17" pageNumber="18">
2. Clypeus shorter then diameter of ALE; anterior eye row nearly straight (
<figureCitation id="133A1AB0553BFFABFBBDE1B3FB09C8E8" box="[1106,1211,1689,1715]" captionStart="Figs 160–171" captionStartId="125.[189,241,1732,1758]" captionTargetBox="[189,1398,265,1691]" captionTargetId="figure-124@125.[189,1398,265,1691]" captionTargetPageId="125" captionText="Figs 160–171. Raveniola spp., ♂♂ (160–165) and ♀♀ (166–171), clypeus and eye tubercle, dorsal views. 160. R. nenilini sp. nov., holotype (SMNH). 161. R. ovchinnikovi sp. nov., holotype (SMNH). 162. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 163. R. virgata, ♂ from Gava (SMF). 164. R. vulpina sp. nov., holotype (SMNH). 165. R. vulpina, paratype from Tereksai (SMNH). 166. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 167. R. afghana sp. nov., holotype (NHMD). 168. R. alajensis sp. nov. (SMNH). 169. R. concolor Zonstein, 2000 (MCSN AR 19). 170– 171. R. cucullata sp. nov., paratypes from Shikildara and Shahob (SMNH), respectively. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990870" httpUri="https://zenodo.org/record/13990870/files/figure.png" pageId="17" pageNumber="18">Fig. 167</figureCitation>
). Spermathecae with short and straight branches (
<figureCitation id="133A1AB0553BFFABFD9CE196FCA5C88D" box="[627,791,1724,1750]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="17" pageNumber="18">Figs 491–493</figureCitation>
, 
<figureCitation id="133A1AB0553BFFABFCCBE196FBB2C88C" box="[804,1024,1724,1751]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="17" pageNumber="18">504–508, 510–512</figureCitation>
)............................................... 3
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE1F5FAC5C947" blockId="17.[189,1399,1689,1820]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE1F5FAC5C947" pageId="17" pageNumber="18">
– Clypeus noticeably wider than diameter of ALE; anterior eye row strictly procurved (
<figureCitation id="133A1AB0553BFFABFB10E1F5FADBC8A2" box="[1279,1385,1759,1785]" captionStart="Figs 160–171" captionStartId="125.[189,241,1732,1758]" captionTargetBox="[189,1398,265,1691]" captionTargetId="figure-124@125.[189,1398,265,1691]" captionTargetPageId="125" captionText="Figs 160–171. Raveniola spp., ♂♂ (160–165) and ♀♀ (166–171), clypeus and eye tubercle, dorsal views. 160. R. nenilini sp. nov., holotype (SMNH). 161. R. ovchinnikovi sp. nov., holotype (SMNH). 162. R. virgata (Simon, 1891), ♂ from vicinity of Arslanbob (SMNH). 163. R. virgata, ♂ from Gava (SMF). 164. R. vulpina sp. nov., holotype (SMNH). 165. R. vulpina, paratype from Tereksai (SMNH). 166. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 167. R. afghana sp. nov., holotype (NHMD). 168. R. alajensis sp. nov. (SMNH). 169. R. concolor Zonstein, 2000 (MCSN AR 19). 170– 171. R. cucullata sp. nov., paratypes from Shikildara and Shahob (SMNH), respectively. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990870" httpUri="https://zenodo.org/record/13990870/files/figure.png" pageId="17" pageNumber="18">Fig. 167</figureCitation>
). Spermathecae with long twisted branches (
<figureCitation id="133A1AB0553BFFABFD0EE028FC37C947" box="[737,901,1794,1820]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="17" pageNumber="18">Figs 489–490</figureCitation>
) .................................. 
<taxonomicName id="4C017DB6553BFFABFB61E028FAA7C947" authorityName="Zonstein" authorityYear="2024" box="[1166,1301,1793,1820]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="afghana" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB61E028FAA7C947" bold="true" box="[1166,1301,1793,1820]" italics="true" pageId="17" pageNumber="18">R. afghana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E028FAC5C947" box="[1308,1399,1794,1820]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
</keyStep>
<keyStep id="30F51E90553BFFABFF52E069FAC5C9B2" pageId="17" pageNumber="18">
<paragraph id="8BBE0635553BFFABFF52E069FAC5C9F9" blockId="17.[189,1399,1859,2025]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF52E069FAC5C9F9" pageId="17" pageNumber="18">
3. Abdomen dorsally uniformly dark or with poorly discernible dark chevron-like pattern (
<figureCitation id="133A1AB0553BFFABFB0CE069FAD8C906" box="[1251,1386,1859,1885]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="17" pageNumber="18">Figs 42, 44</figureCitation>
). Spermathecae with unevenly wide pair of branches: outer branch clearly slenderer than the massive and often multilobate inner branch (
<figureCitation id="133A1AB0553BFFABFD60E0A2FC1BC9F8" box="[655,937,1928,1955]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="17" pageNumber="18">Figs 504–508, 510–512</figureCitation>
) .......................................................... 4
</keyLead>
</paragraph>
<paragraph id="8BBE0635553BFFABFF2EE086FAC5C9B2" blockId="17.[189,1399,1859,2025]" pageId="17" pageNumber="18">
<keyLead id="30F0A500553BFFABFF2EE086FAC5C9B2" pageId="17" pageNumber="18">
– Abdomen dorsally with distinct, dark chevron-like pattern (
<figureCitation id="133A1AB0553BFFABFC4BE081FC4EC99E" box="[932,1020,1963,1989]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="17" pageNumber="18">Fig. 38</figureCitation>
). Each spermatheca with almost uniformly slender inner and outer branches (
<figureCitation id="133A1AB0553BFFABFD1BE0E4FC2AC9B3" box="[756,920,1998,2024]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="17" pageNumber="18">Figs 491–493</figureCitation>
)............................... 
<taxonomicName id="4C017DB6553BFFABFB66E0E5FAA7C9B2" authorityName="Zonstein" authorityYear="2024" box="[1161,1301,1998,2025]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="alajensis" status="sp. nov.">
<emphasis id="B975DA27553BFFABFB66E0E5FAA7C9B2" bold="true" box="[1161,1301,1998,2025]" italics="true" pageId="17" pageNumber="18">R. alajensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C553BFFABFAF3E0E5FAC5C9B2" box="[1308,1399,1999,2025]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E623FAC5CF86" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E623FAC5CF35" blockId="18.[189,1399,265,477]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E623FAC5CF35" pageId="18" pageNumber="19">
4. Abdomen dorsally uniformly dark brown or with almost indistinct paler chevrons (
<figureCitation id="133A1AB05538FFA8FB25E623FA97CF78" box="[1226,1317,265,291]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="18" pageNumber="19">Fig. 42</figureCitation>
). Tibia and metatarsus IV dorsally with extremely dense modified hairs (
<figureCitation id="133A1AB05538FFA8FBE6E604FBC7CF13" box="[1033,1141,302,328]" captionStart="Figs 310–318" captionStartId="140.[189,241,1786,1812]" captionTargetBox="[189,1398,265,1765]" captionTargetId="figure-96@140.[189,1398,265,1765]" captionTargetPageId="140" captionText="Figs 310–318. Raveniola spp., ♀♀ (310–312, 314–318) and ♂ (313), tibia and metatarsus IV, retrolateral views. 310. R. afghana sp. nov., holotype (NHMD). 311. R. alajensis sp. nov., paratype from Berksu. 312. R. cucullata sp. nov., paratype from Mullokoni (SMNH). 313–314. R. hirta sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 315. R. ignobilis sp. nov., paratype from Sangdara (SMNH). 316–317. R. karategensis sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 318. R. pamira sp. nov., paratype from Ubagandara (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990905" httpUri="https://zenodo.org/record/13990905/files/figure.png" pageId="18" pageNumber="19">Fig. 314</figureCitation>
). Spermathecae as in 
<figureCitation id="133A1AB05538FFA8FF05E679FE39CF35" box="[234,395,339,366]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">Figs 504–508</figureCitation>
............................................................................................................. 
<taxonomicName id="4C017DB65538FFA8FB57E67EFAA7CF35" authorityName="Zonstein" authorityYear="2024" box="[1208,1301,339,366]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="hirta" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB57E67EFAA7CF35" bold="true" box="[1208,1301,339,366]" italics="true" pageId="18" pageNumber="19">R. hirta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E67EFAC5CF35" box="[1308,1399,340,366]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE653FAC5CF86" blockId="18.[189,1399,265,477]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE653FAC5CF86" pageId="18" pageNumber="19">
– Abdomen dorsally medium brown with poorly discernible dark brown chevron-like pattern (
<figureCitation id="133A1AB05538FFA8FF1CE6B7FEFFCFE3" box="[243,333,413,440]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="18" pageNumber="19">Fig. 44</figureCitation>
). Tibia and metatarsus IV dorsally with moderately dense modified hairs (
<figureCitation id="133A1AB05538FFA8FB2DE6B7FAD9CFEC" box="[1218,1387,413,439]" captionStart="Figs 310–318" captionStartId="140.[189,241,1786,1812]" captionTargetBox="[189,1398,265,1765]" captionTargetId="figure-96@140.[189,1398,265,1765]" captionTargetPageId="140" captionText="Figs 310–318. Raveniola spp., ♀♀ (310–312, 314–318) and ♂ (313), tibia and metatarsus IV, retrolateral views. 310. R. afghana sp. nov., holotype (NHMD). 311. R. alajensis sp. nov., paratype from Berksu. 312. R. cucullata sp. nov., paratype from Mullokoni (SMNH). 313–314. R. hirta sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 315. R. ignobilis sp. nov., paratype from Sangdara (SMNH). 316–317. R. karategensis sp. nov., holotype (SMNH) and paratype (SMNH), respectively. 318. R. pamira sp. nov., paratype from Ubagandara (SMNH). Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990905" httpUri="https://zenodo.org/record/13990905/files/figure.png" pageId="18" pageNumber="19">Figs 316–317</figureCitation>
). Spermathecae as in 
<figureCitation id="133A1AB05538FFA8FE3BE6E8FDC7CF86" box="[468,629,450,477]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">Figs 510–512</figureCitation>
.................................................................. 
<taxonomicName id="4C017DB65538FFA8FBB4E6E9FAA7CF86" authorityName="Zonstein" authorityYear="2024" box="[1115,1301,450,477]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="karategensis" status="sp. nov.">
<emphasis id="B975DA275538FFA8FBB4E6E9FAA7CF86" bold="true" box="[1115,1301,450,477]" italics="true" pageId="18" pageNumber="19">R. karategensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E6E9FAC5CF86" box="[1308,1399,451,477]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E52CFAC5CCEF" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E52CFAC5CC31" blockId="18.[189,1399,518,693]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E52CFAC5CC31" pageId="18" pageNumber="19">
5. Abdomen with darker chevron-like or reticulate pattern (
<figureCitation id="133A1AB05538FFA8FC94E52CFB9ECC7B" box="[891,1068,518,544]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="18" pageNumber="19">Figs 39–41, 43</figureCitation>
, 
<figureCitation id="133A1AB05538FFA8FBD8E52CFB37CC7A" box="[1079,1157,518,545]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="18" pageNumber="19">45–49</figureCitation>
). Spermathecae with dissimilar inner and outer branches on low common base (
<figureCitation id="133A1AB05538FFA8FC43E501FBE1CC1D" box="[940,1107,555,582]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="18" pageNumber="19">Figs 495–503</figureCitation>
, 
<figureCitation id="133A1AB05538FFA8FB8DE501FAB8CC1D" box="[1122,1290,555,582]" captionStart-0="Figs 504–521" captionStart-1="Figs 522–536" captionStartId-0="157.[189,241,1719,1745]" captionStartId-1="158.[189,241,1728,1754]" captionTargetBox-0="[189,1398,265,1675]" captionTargetBox-1="[189,1398,265,1688]" captionTargetId-0="figure-86@157.[189,1398,265,1675]" captionTargetId-1="figure-119@158.[189,1398,265,1688]" captionTargetPageId-0="157" captionTargetPageId-1="158" captionText-0="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." captionText-1="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990943" figureDoi-1="http://doi.org/10.5281/zenodo.13990945" httpUri-0="https://zenodo.org/record/13990943/files/figure.png" httpUri-1="https://zenodo.org/record/13990945/files/figure.png" pageId="18" pageNumber="19">509, 520–525</figureCitation>
), or with main trunk and lateral diverticulum (
<figureCitation id="133A1AB05538FFA8FD73E57AFCF2CC30" box="[668,832,592,619]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">Figs 513–519</figureCitation>
)......................................................................... 6
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE55CFAC5CCEF" blockId="18.[189,1399,518,693]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE55CFAC5CCEF" pageId="18" pageNumber="19">
– Abdomen uniformly dark coloured. Spermathecae with similar inner and outer branches on high common base (
<figureCitation id="133A1AB05538FFA8FE72E5B0FDB3CCEF" box="[413,513,666,692]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="18" pageNumber="19">Fig. 494</figureCitation>
) .......................................................................... 
<taxonomicName id="4C017DB65538FFA8FBDCE5B1FAC5CCEF" ID-CoL="4RMTP" authority="Zonstein, 2000" authorityName="Zonstein" authorityYear="2000" box="[1075,1399,666,693]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="concolor">
<emphasis id="B975DA275538FFA8FBDCE5B1FB0DCCEE" bold="true" box="[1075,1215,666,693]" italics="true" pageId="18" pageNumber="19">R. concolor</emphasis>
<bibRefCitation id="EF907BC45538FFA8FB29E5B0FAC5CCEF" author="Zonstein S. L." box="[1222,1399,666,692]" journalOrPublisher="Tethys Entomological Research" pageId="18" pageNumber="19" pagination="49 - 52" part="2" refId="ref83098" refString="Zonstein S. L. 2000 a. Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia. Tethys Entomological Research 2: 49 - 52." title="Two new species of the mygalomorph spider genus Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from southwestern Asia" type="journal article" year="2000">Zonstein, 2000</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E5F4FAC4CDEA" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E5F4FAC5CD19" blockId="18.[189,1400,733,946]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E5F4FAC5CD19" pageId="18" pageNumber="19">
6. Sternum broadly oval (
<figureCitation id="133A1AB05538FFA8FE10E5F4FD56CCA3" box="[511,740,733,760]" captionStart="Figs 229–237" captionStartId="132.[189,241,1827,1853]" captionTargetBox="[200,1385,265,1784]" captionTargetId="figure-90@132.[200,1385,265,1784]" captionTargetPageId="132" captionText="Figs 229–237. Raveniola spp., ♀♀, paratypes (except 229–230), sternum, labium and maxillae, ventral views. 229. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 230. R. afghana sp. nov., holotype (NHMD). 231. R. alajensis sp. nov. (SMNH). 232. R. concolor Zonstein, 2000 (MCSN AR 19). 233– 234. R. cucullata sp. nov. from Shikildara (SMNH) and Shahob (SMNH), respectively. 235. R. dolosa sp. nov.(SMNH). 236. R. hirta sp. nov. (SMNH). 237. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990884" httpUri="https://zenodo.org/record/13990884/files/figure.png" pageId="18" pageNumber="19">Figs 233–235, 237</figureCitation>
, 
<figureCitation id="133A1AB05538FFA8FD1DE5F4FCECCCA3" box="[754,862,734,760]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="18" pageNumber="19">241–242</figureCitation>
). Abdomen without darker ventral pattern or with a few dark marks on ventral side. PMS mostly medium-sized; apical segment of PLS shorter and triangular (as in 
<figureCitation id="133A1AB05538FFA8FE31E402FD0FCD18" box="[478,701,808,835]" captionStart="Figs 565–574" captionStartId="161.[189,241,1825,1851]" captionTargetBox="[202,1386,265,1765]" captionTargetId="figure-100@161.[202,1386,265,1784]" captionText="Figs 565–574. Raveniola spp., ♂♂ (565, 568, 571, 573) and ♀♀ (566–567, 569–570, 572), paratypes (except 568, 571, 573). Spinnerets, ventral (565–566, 571–574), lateral (567–568) and ventrolateral (569–570) views. 565–566. R. cucullata sp. nov., paratypes from Shahob (SMNH). 567. R. cucullata, from Sangdara (SMNH). 568. R. dolosa sp. nov. (SMNH). 569–570. R. dolosa, paratypes (SMNH). 571. R. hirta sp. nov., holotype (SMNH). 572. R. hirta, paratype (SMNH). 573. R. ignobilis sp. nov. (SMNH). 574. R. ignobilis, paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018403" httpUri="https://zenodo.org/record/14018403/files/figure.png" pageId="18" pageNumber="19">Figs 566–567, 574</figureCitation>
, 585–588)........................................................................... 7
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE464FAC4CDEA" blockId="18.[189,1400,733,946]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE464FAC4CDEA" pageId="18" pageNumber="19">
– Sternum subcircular (
<figureCitation id="133A1AB05538FFA8FE08E467FD3CCD3C" box="[487,654,845,871]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="18" pageNumber="19">Figs 239–240</figureCitation>
). Abdomen with more or less dense maculate pattern on ventral side (
<figureCitation id="133A1AB05538FFA8FEC2E458FE68CDD6" box="[301,474,882,909]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="18" pageNumber="19">Figs 580, 583</figureCitation>
). PMS small and thin; apical segment of PLS longer and shortly digitiform (
<figureCitation id="133A1AB05538FFA8FF1CE4BDFE63CDE9" box="[243,465,919,946]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="18" pageNumber="19">Figs 578–579, 582</figureCitation>
)..........................................................................................................................11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E4F1FAC4CA3F" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E4F1FAC5CA41" blockId="18.[189,1399,987,1125]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E4F1FAC5CA41" pageId="18" pageNumber="19">
7. Habitus as in 
<figureCitation id="133A1AB05538FFA8FE66E4F1FDBDCDAE" box="[393,527,987,1013]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="18" pageNumber="19">Figs 39–40</figureCitation>
, 
<figureCitation id="133A1AB05538FFA8FDF5E4F1FDDACDAE" box="[538,616,987,1013]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="18" pageNumber="19">48–49</figureCitation>
. Each paired spermatheca U-shaped, with outer and inner branches distinctly separated from each other; inner branch often multilobate (
<figureCitation id="133A1AB05538FFA8FBFEE32AFB04CA41" box="[1041,1206,1024,1050]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="18" pageNumber="19">Figs 494–499</figureCitation>
, 
<figureCitation id="133A1AB05538FFA8FB2DE32AFA99CA40" box="[1218,1323,1024,1051]" captionStart-0="Figs 504–521" captionStart-1="Figs 522–536" captionStartId-0="157.[189,241,1719,1745]" captionStartId-1="158.[189,241,1728,1754]" captionTargetBox-0="[189,1398,265,1675]" captionTargetBox-1="[189,1398,265,1688]" captionTargetId-0="figure-86@157.[189,1398,265,1675]" captionTargetId-1="figure-119@158.[189,1398,265,1688]" captionTargetPageId-0="157" captionTargetPageId-1="158" captionText-0="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." captionText-1="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990943" figureDoi-1="http://doi.org/10.5281/zenodo.13990945" httpUri-0="https://zenodo.org/record/13990943/files/figure.png" httpUri-1="https://zenodo.org/record/13990945/files/figure.png" pageId="18" pageNumber="19">520–525</figureCitation>
) ..... 8
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE30FFAC4CA3F" blockId="18.[189,1399,987,1125]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE30FFAC4CA3F" pageId="18" pageNumber="19">
– Habitus as in 
<figureCitation id="133A1AB05538FFA8FE65E30FFDA3CA64" box="[394,529,1061,1087]" captionStart="Figs 36–44" captionStartId="111.[189,241,1837,1863]" captionTargetBox="[201,1398,270,1792]" captionTargetId="figure-83@111.[189,1398,265,1792]" captionTargetPageId="111" captionText="Figs 36–44. Raveniola spp., ♀♀, paratypes (except 36–37, 44), habitus, dorsal views. 36. R. redikorzevi (Spassky, 1937), from Kepele (SMNH). 37. R. afghana sp. nov., holotype (NHMD). 38. R. alajensis sp. nov. (SMNH). 39–40. R. cucullata sp. nov. from Shikildara (SMNH) and Childara (SMNH), respectively. 41. R. dolosa sp. nov. (SMNH). 42. R. hirta sp. nov. (SMNH). 43. R. ignobilis sp. nov. (SMNH). 44. R. karategensis sp. nov., holotype (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990834" httpUri="https://zenodo.org/record/13990834/files/figure.png" pageId="18" pageNumber="19">Figs 41, 43</figureCitation>
. Each paired spermatheca V-shaped, with outer and inner branches arising very close to each other; inner branch entire (
<figureCitation id="133A1AB05538FFA8FD12E360FC10CA3E" box="[765,930,1098,1125]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="18" pageNumber="19">Figs 500–503</figureCitation>
, 
<figureCitation id="133A1AB05538FFA8FC41E360FC6ECA3E" box="[942,988,1098,1125]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">509</figureCitation>
) ................................................. 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E3A7FAC5CBDC" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E3A7FAC5CAA9" blockId="18.[189,1400,1165,1415]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E3A7FAC5CAA9" pageId="18" pageNumber="19">
8. Maxillae with 70–90 cuspules (
<figureCitation id="133A1AB05538FFA8FDB3E3A7FCB6CAFC" box="[604,772,1165,1191]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="18" pageNumber="19">Figs 241–242</figureCitation>
). Tarsus II with entire scopula. Spermathecae usually shorter and broader spaced from each other (
<figureCitation id="133A1AB05538FFA8FCEDE398FC19CA96" box="[770,939,1202,1229]" captionStart-0="Figs 504–521" captionStart-1="Figs 522–536" captionStartId-0="157.[189,241,1719,1745]" captionStartId-1="158.[189,241,1728,1754]" captionTargetBox-0="[189,1398,265,1675]" captionTargetBox-1="[189,1398,265,1688]" captionTargetId-0="figure-86@157.[189,1398,265,1675]" captionTargetId-1="figure-119@158.[189,1398,265,1688]" captionTargetPageId-0="157" captionTargetPageId-1="158" captionText-0="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." captionText-1="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990943" figureDoi-1="http://doi.org/10.5281/zenodo.13990945" httpUri-0="https://zenodo.org/record/13990943/files/figure.png" httpUri-1="https://zenodo.org/record/13990945/files/figure.png" pageId="18" pageNumber="19">Figs 520–525</figureCitation>
). Basal segment of PLS ventrally with denser spigots confined to about posterior 0.35–0.40 of its length (
<figureCitation id="133A1AB05538FFA8FC1AE3FDFB2BCAA9" box="[1013,1177,1239,1266]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="18" pageNumber="19">Figs 585, 587</figureCitation>
).......................... 9
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE3D7FAC5CBDC" blockId="18.[189,1400,1165,1415]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE3D7FAC5CBDC" pageId="18" pageNumber="19">
– Maxillae with 35–60 cuspules (
<figureCitation id="133A1AB05538FFA8FDB0E3D7FCBACB4C" box="[607,776,1277,1303]" captionStart="Figs 229–237" captionStartId="132.[189,241,1827,1853]" captionTargetBox="[200,1385,265,1784]" captionTargetId="figure-90@132.[200,1385,265,1784]" captionTargetPageId="132" captionText="Figs 229–237. Raveniola spp., ♀♀, paratypes (except 229–230), sternum, labium and maxillae, ventral views. 229. R. redikorzevi (Spassky, 1937), ♀ from Kepele (SMNH). 230. R. afghana sp. nov., holotype (NHMD). 231. R. alajensis sp. nov. (SMNH). 232. R. concolor Zonstein, 2000 (MCSN AR 19). 233– 234. R. cucullata sp. nov. from Shikildara (SMNH) and Shahob (SMNH), respectively. 235. R. dolosa sp. nov.(SMNH). 236. R. hirta sp. nov. (SMNH). 237. R. ignobilis sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990884" httpUri="https://zenodo.org/record/13990884/files/figure.png" pageId="18" pageNumber="19">Figs 233–234</figureCitation>
). Scopula on tarsus II widely divided. Spermathecae usually longer and closer spaced from each other (
<figureCitation id="133A1AB05538FFA8FC67E208FB8BCB66" box="[904,1081,1314,1341]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="18" pageNumber="19">Figs 495–499</figureCitation>
). Basal segment of PLS ventrally with sparser spigots confined to about posterior 0.20–0.25 of its length (
<figureCitation id="133A1AB05538FFA8FAEFE26DFAC0CB39" box="[1280,1394,1351,1378]" captionStart="Figs 555–564" captionStartId="160.[189,241,1786,1812]" captionTargetBox="[213,1375,265,1755]" captionTargetId="figure-109@160.[213,1375,265,1755]" captionText="Figs 555–564. Raveniola spp., ♂♂ (555–557, 560, 563) and ♀♀ (558–559, 561–562, 564), holotypes (except 557, 558, 561, 562, 564). Spinnerets, ventral (555–558, 560–561, 563–564) and lateral (559, 562) views. 555. R. caudata Zonstein, 2009, holotype (SMNH). 556. R. inopinata sp. nov. (SMNH). 557–558. R. redikorzevi (Spassky, 1937), ♂ from Akar-Cheshme and ♀ from Kepele, respectively (both SMNH). 559. R. afghana sp. nov. (NHMD). 560. R. alajensis sp. nov. (SMNH). 561–562. R. alajensis, paratype (SMNH). 563. R. cucullata sp. nov. (SMNH). 564. R. cucullata, paratype from Shikildara (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018401" httpUri="https://zenodo.org/record/14018401/files/figure.png" pageId="18" pageNumber="19">Figs 564</figureCitation>
, 566) .................................................................................................................... 
<taxonomicName id="4C017DB65538FFA8FB6DE246FAA6CBDD" authorityName="Zonstein" authorityYear="2024" box="[1154,1300,1387,1414]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cucullata" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB6DE246FAA6CBDD" bold="true" box="[1154,1300,1387,1414]" italics="true" pageId="18" pageNumber="19">R. cucullata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E247FAC5CBDC" box="[1308,1399,1389,1415]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E285FAC5C862" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E285FAC5CBB4" blockId="18.[189,1399,1455,1593]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E285FAC5CBB4" pageId="18" pageNumber="19">
9. Spermathecae with stouter multilobate inner branches (
<figureCitation id="133A1AB05538FFA8FC7DE285FB8CCB91" box="[914,1086,1455,1482]" captionStart-0="Figs 504–521" captionStart-1="Figs 522–536" captionStartId-0="157.[189,241,1719,1745]" captionStartId-1="158.[189,241,1728,1754]" captionTargetBox-0="[189,1398,265,1675]" captionTargetBox-1="[189,1398,265,1688]" captionTargetId-0="figure-86@157.[189,1398,265,1675]" captionTargetId-1="figure-119@158.[189,1398,265,1688]" captionTargetPageId-0="157" captionTargetPageId-1="158" captionText-0="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." captionText-1="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13990943" figureDoi-1="http://doi.org/10.5281/zenodo.13990945" httpUri-0="https://zenodo.org/record/13990943/files/figure.png" httpUri-1="https://zenodo.org/record/13990945/files/figure.png" pageId="18" pageNumber="19">Figs 520–523</figureCitation>
). Apical segment of PLS longer and about 1.5 times as long as wide (
<figureCitation id="133A1AB05538FFA8FD02E2FEFC23CBB4" box="[749,913,1492,1519]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="18" pageNumber="19">Figs 585–586</figureCitation>
)................................... 
<taxonomicName id="4C017DB65538FFA8FB73E2FFFAA7CBB5" authorityName="Zonstein" authorityYear="2024" box="[1180,1301,1492,1519]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="pamira" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB73E2FFFAA7CBB5" bold="true" box="[1180,1301,1492,1519]" italics="true" pageId="18" pageNumber="19">R. pamira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E2FFFAC5CBB4" box="[1308,1399,1493,1519]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE2D0FAC5C862" blockId="18.[189,1399,1455,1593]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE2D0FAC5C862" pageId="18" pageNumber="19">
– Spermathecae with thinner and smaller inner branches (
<figureCitation id="133A1AB05538FFA8FC79E2D3FBF0C84F" box="[918,1090,1529,1556]" captionStart="Figs 522–536" captionStartId="158.[189,241,1728,1754]" captionTargetBox="[189,1398,265,1688]" captionTargetId="figure-119@158.[189,1398,265,1688]" captionTargetPageId="158" captionText="Figs 522–536. Raveniola spp., ♀♀, spermathecae, dorsal views. 522–523. R. pamira sp. nov., paratype and female from Shahob, respectively (both SMNH). 524. R. sororcula sp. nov., holotype (SMNH). 525. R. sororcula, paratype (SMNH). 526–527. R. diluta sp. nov., paratype from Kondara (SMNH). 528–529. R. fedotovi (Charitonov, 1946), from Hojakurgan (SMNH). 530–531. R. pallens sp. nov., paratype (SMNH). 532–533. R. zyuzini sp. nov., paratype (SMNH). 534–536. R. ferghanensis (Zonstein, 1984), from Andijan (SMNH), non-type specimen from environs of Jalal-Abad (SMNH), and paratype from Jalal-Abad (SMNH), respectively. Abbreviations: ld = lateral diverticulum; sb = spermathecal base; st = spermathecal trunk. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990945" httpUri="https://zenodo.org/record/13990945/files/figure.png" pageId="18" pageNumber="19">Figs 524–525</figureCitation>
). Apical segment of PLS shorter and about as long as wide (
<figureCitation id="133A1AB05538FFA8FD6EE135FC96C862" box="[641,804,1566,1593]" captionStart="Figs 584–592" captionStartId="163.[189,241,1821,1847]" captionTargetBox="[189,1398,265,1789]" captionTargetId="figure-92@163.[189,1398,265,1789]" captionText="Figs 584–592. Raveniola spp., ♂♂ (584, 589, 591) and ♀♀ (585–588, 590, 592), holotypes (except 585–586, 590, 592). Spinnerets, ventral (584–585, 587, 589, 591–592) and lateral (586, 588, 590) views. 584. R. pamira sp. nov. (SMNH). 585–586. R. pamira, paratype (SMNH). 587–588. R. sororcula sp. nov. (SMNH). 589. R. diluta sp. nov. (SMNH). 590. R. diluta, paratype (SMNH). 591. R. fedotovi (Charitonov, 1946), paralectotype (PSU). 592. R. fedotovi, ♀ from Hojakurgan (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018408" httpUri="https://zenodo.org/record/14018408/files/figure.png" pageId="18" pageNumber="19">Figs 587–588</figureCitation>
) ............................................. 
<taxonomicName id="4C017DB65538FFA8FB90E135FAA7C862" authorityName="Zonstein" authorityYear="2024" box="[1151,1301,1566,1593]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sororcula" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB90E135FAA7C862" bold="true" box="[1151,1301,1566,1593]" italics="true" pageId="18" pageNumber="19">R. sororcula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E135FAC5C862" box="[1308,1399,1567,1593]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E148FAC5C8B7" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E148FAC5C8F9" blockId="18.[189,1399,1634,1772]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E148FAC5C8F9" pageId="18" pageNumber="19">
10. PMS smaller; apical segment of PLS longer (
<figureCitation id="133A1AB05538FFA8FD15E148FC12C826" box="[762,928,1634,1661]" captionStart="Figs 565–574" captionStartId="161.[189,241,1825,1851]" captionTargetBox="[202,1386,265,1765]" captionTargetId="figure-100@161.[202,1386,265,1784]" captionText="Figs 565–574. Raveniola spp., ♂♂ (565, 568, 571, 573) and ♀♀ (566–567, 569–570, 572), paratypes (except 568, 571, 573). Spinnerets, ventral (565–566, 571–574), lateral (567–568) and ventrolateral (569–570) views. 565–566. R. cucullata sp. nov., paratypes from Shahob (SMNH). 567. R. cucullata, from Sangdara (SMNH). 568. R. dolosa sp. nov. (SMNH). 569–570. R. dolosa, paratypes (SMNH). 571. R. hirta sp. nov., holotype (SMNH). 572. R. hirta, paratype (SMNH). 573. R. ignobilis sp. nov. (SMNH). 574. R. ignobilis, paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018403" httpUri="https://zenodo.org/record/14018403/files/figure.png" pageId="18" pageNumber="19">Figs 569–570</figureCitation>
). Inner and outer spermathecal branches shorter (
<figureCitation id="133A1AB05538FFA8FEA4E1ADFE5DC8F9" box="[331,495,1671,1698]" captionStart="Figs 487–503" captionStartId="156.[189,241,1695,1721]" captionTargetBox="[189,1398,265,1653]" captionTargetId="figure-102@156.[189,1398,265,1653]" captionTargetPageId="156" captionText="Figs 487–503. Raveniola spp., ♀♀, paratypes (except 487–489), spermathecae, dorsal views. 487–488. R. redikorzevi (Spassky, 1937), Kepele (SMNH). 489. R. afghana sp. nov. (NHMD). 490. Same, holotype. 491–493. R. alajensis sp. nov., Berksu. 494. R. concolor Zonstein, 2000 (MCSN AR 19). 495–496. R. cucullata sp. nov., Shikildara (SMNH). 497. R. cucullata, Shahob (SMNH). 498– 499. R. cucullata, Sangdara (SMNH). 500–503. R. dolosa sp. nov. (SMNH). Abbreviations: isb = inner spermathecal branch; osb = outer spermathecal branch; sb = spermathecal base. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990941" httpUri="https://zenodo.org/record/13990941/files/figure.png" pageId="18" pageNumber="19">Figs 500–503</figureCitation>
)............................................................................................. 
<taxonomicName id="4C017DB65538FFA8FB4BE1ADFAA7C8F9" authorityName="Zonstein" authorityYear="2024" box="[1188,1301,1671,1698]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="dolosa" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB4BE1ADFAA7C8F9" bold="true" box="[1188,1301,1671,1698]" italics="true" pageId="18" pageNumber="19">R. dolosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E1A2FAC5C8F9" box="[1308,1399,1672,1698]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE187FAC5C8B7" blockId="18.[189,1399,1634,1772]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE187FAC5C8B7" pageId="18" pageNumber="19">
– PMS larger; apical segment of PLS shorter (
<figureCitation id="133A1AB05538FFA8FD04E186FCE0C89C" box="[747,850,1708,1735]" captionStart="Figs 565–574" captionStartId="161.[189,241,1825,1851]" captionTargetBox="[202,1386,265,1765]" captionTargetId="figure-100@161.[202,1386,265,1784]" captionText="Figs 565–574. Raveniola spp., ♂♂ (565, 568, 571, 573) and ♀♀ (566–567, 569–570, 572), paratypes (except 568, 571, 573). Spinnerets, ventral (565–566, 571–574), lateral (567–568) and ventrolateral (569–570) views. 565–566. R. cucullata sp. nov., paratypes from Shahob (SMNH). 567. R. cucullata, from Sangdara (SMNH). 568. R. dolosa sp. nov. (SMNH). 569–570. R. dolosa, paratypes (SMNH). 571. R. hirta sp. nov., holotype (SMNH). 572. R. hirta, paratype (SMNH). 573. R. ignobilis sp. nov. (SMNH). 574. R. ignobilis, paratype (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018403" httpUri="https://zenodo.org/record/14018403/files/figure.png" pageId="18" pageNumber="19">Fig. 574</figureCitation>
). Inner and outer spermathecal branches longer (
<figureCitation id="133A1AB05538FFA8FF1DE1FBFEE5C8B7" box="[242,343,1745,1772]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">Fig. 509</figureCitation>
) ............................................................................................................. 
<taxonomicName id="4C017DB65538FFA8FB65E1F8FAA7C8B7" authorityName="Zonstein" authorityYear="2024" box="[1162,1301,1745,1772]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ignobilis" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB65E1F8FAA7C8B7" bold="true" box="[1162,1301,1745,1772]" italics="true" pageId="18" pageNumber="19">R. ignobilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E1F8FAC5C8B7" box="[1308,1399,1746,1772]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F51E905538FFA8FF52E03FFAC5C9B2" pageId="18" pageNumber="19">
<paragraph id="8BBE06355538FFA8FF52E03FFAC5C921" blockId="18.[189,1400,1813,2025]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF52E03FFAC5C921" pageId="18" pageNumber="19">
11. Habitus as in 
<figureCitation id="133A1AB05538FFA8FE7FE03FFDAAC96B" box="[400,536,1813,1840]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="18" pageNumber="19">Figs 45–46</figureCitation>
. Labium longer and narrower, maxillae with relatively smaller but more numerous cuspules (
<figureCitation id="133A1AB05538FFA8FE32E010FDF7C90F" box="[477,581,1850,1876]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="18" pageNumber="19">Fig. 239</figureCitation>
). Spermathecae with entire trunks (
<figureCitation id="133A1AB05538FFA8FC0AE010FB3EC90E" box="[997,1164,1850,1877]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">Figs 513–519</figureCitation>
). Ventral abdominal pattern sparser (
<figureCitation id="133A1AB05538FFA8FE48E075FDBEC921" box="[423,524,1887,1914]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="18" pageNumber="19">Fig. 580</figureCitation>
)......................................................................................... 
<taxonomicName id="4C017DB65538FFA8FB4CE075FAA7C922" authorityName="Zonstein" authorityYear="2024" box="[1187,1301,1887,1913]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ornata" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB4CE075FAA7C922" bold="true" box="[1187,1301,1887,1913]" italics="true" pageId="18" pageNumber="19">R. ornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E04AFAC5C921" box="[1308,1399,1888,1914]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BBE06355538FFA8FF2EE0AFFAC5C9B2" blockId="18.[189,1400,1813,2025]" pageId="18" pageNumber="19">
<keyLead id="30F0A5005538FFA8FF2EE0AFFAC5C9B2" pageId="18" pageNumber="19">
– Habitus as in 
<figureCitation id="133A1AB05538FFA8FE66E0AEFE52C9C5" box="[393,480,1924,1950]" captionStart="Figs 45–53" captionStartId="112.[189,241,1842,1868]" captionTargetBox="[190,1398,265,1803]" captionTargetId="figure-85@112.[189,1398,265,1803]" captionTargetPageId="112" captionText="Figs 45–53. Raveniola spp., ♀♀, paratypes (except 49, 51), habitus, dorsal views. 45–46. R. ornata sp. nov., from Ganjina and Sanglok (both SMNH), respectively. 47. R. ornatula sp. nov. (SMNH). 48. R. pamira sp. nov. (SMNH). 49. R. sororcula sp. nov., holotype (SMNH). 50. R. diluta sp. nov. (SMNH). 51. R. fedotovi(Charitonov, 1946), from Hojakurgan (SMNH). 52. R. pallenssp. nov. (SMNH). 53. R. zyuzini sp. nov. (SMNH). Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990836" httpUri="https://zenodo.org/record/13990836/files/figure.png" pageId="18" pageNumber="19">Fig. 47</figureCitation>
. Labium shorter and broader, maxillae with relatively larger but less numerous cuspules (
<figureCitation id="133A1AB05538FFA8FEB0E083FE7AC998" box="[351,456,1961,1987]" captionStart="Figs 238–246" captionStartId="133.[189,241,1820,1846]" captionTargetBox="[208,1381,265,1770]" captionTargetId="figure-91@133.[208,1381,265,1770]" captionTargetPageId="133" captionText="Figs 238–246. Raveniola spp., ♀♀, paratypes (except 238, 242, 244), sternum, labium and maxillae, ventral views. 238. R. karategensis sp. nov., holotype (SMNH). 239. R. ornata sp. nov., from Sanglok (SMNH). 240. R. ornatula sp. nov. (SMNH). 241. R. pamira sp. nov. (SMNH). 242. R. sororcula sp. nov., holotype (SMNH). 243. R. diluta sp. nov. (SMNH). 244. R. fedotovi (Charitonov, 1946), ♀ from Hojakurgan (SMNH). 245. R. pallens sp. nov. (SMNH). 246. R. zyuzini sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13990887" httpUri="https://zenodo.org/record/13990887/files/figure.png" pageId="18" pageNumber="19">Fig. 240</figureCitation>
). Spermathecae with mostly two- or three-lobate trunks (
<figureCitation id="133A1AB05538FFA8FB89E083FABCC99F" box="[1126,1294,1961,1988]" captionStart="Figs 504–521" captionStartId="157.[189,241,1719,1745]" captionTargetBox="[189,1398,265,1675]" captionTargetId="figure-86@157.[189,1398,265,1675]" captionTargetPageId="157" captionText="Figs 504–521. Raveniola spp., ♀♀, spermathecae, dorsal views. 504–508. R. hirta sp. nov., paratypes (SMNH). 509. R. ignobilis sp. nov., paratype (SMNH). 510. R. karategensis sp. nov., holotype (SMNH). 511–512. R. karategensis, paratypes (ZISP). 513–514. R. ornata sp. nov., paratypes from Ganjina (SMNH). 515–516. R. ornata, paratypes from Sanglok (SMNH). 517–519. R. ornatula sp. nov., paratypes (SMNH). 520–521. R. pamira sp. nov., paratypes (SMNH). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13990943" httpUri="https://zenodo.org/record/13990943/files/figure.png" pageId="18" pageNumber="19">Figs 517–519</figureCitation>
). Ventral abdominal pattern denser (
<figureCitation id="133A1AB05538FFA8FDCCE0E4FD35C9B2" box="[547,647,1998,2025]" captionStart="Figs 575–583" captionStartId="162.[189,241,1857,1883]" captionTargetBox="[189,1398,265,1816]" captionTargetId="figure-74@162.[189,1398,265,1816]" captionText="Figs 575–583. Raveniola spp., ♂♂ (575, 577, 581) and ♀♀ (576, 578–580, 582–583); holotypes (575– 577, 581) and paratypes (578–580, 582–583). Spinnerets (575–579, 581–582) and entire abdomen (580, 583), ventral (575–576, 579–580, 582–583) and lateral (577–578, 581) views. 575. R. insolita sp. nov. (ZISP). 576. R. karategensis sp. nov. (SMNH). 577–580. R. ornata sp. nov. (SMNH). 581– 583. R. ornatula sp. nov. (SMNH). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14018406" httpUri="https://zenodo.org/record/14018406/files/figure.png" pageId="18" pageNumber="19">Fig. 583</figureCitation>
).................................................................... 
<taxonomicName id="4C017DB65538FFA8FB65E0E5FAA7C9B2" authorityName="Zonstein" authorityYear="2024" box="[1162,1301,1998,2025]" class="Arachnida" family="Nemesiidae" genus="Raveniola" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ornatula" status="sp. nov.">
<emphasis id="B975DA275538FFA8FB65E0E5FAA7C9B2" bold="true" box="[1162,1301,1998,2025]" italics="true" pageId="18" pageNumber="19">R. ornatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A246675C5538FFA8FAF3E0E5FAC5C9B2" box="[1308,1399,1999,2025]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>