253 lines
27 KiB
XML
253 lines
27 KiB
XML
<document id="F2339455686E0AD029B6CB36246AA7F4" ID-DOI="10.11646/zootaxa.4007.3.10" ID-GBIF-Dataset="e659252e-1ea0-4783-af2f-6398fa8b6aa4" ID-ISSN="1175-5326" ID-Zenodo-Dep="240333" ID-ZooBank="A48D1205-6480-4727-90DC-55C880E5F52C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461219968982" checkinUser="plazi" docAuthor="Frolov, Andrey V. & Vaz-De-Mello, Fernando Z." docDate="2015" docId="03A51F7C0C7CFFF57BDDFF07FCB8FA72" docLanguage="en" docName="zt04007p436.pdf" docOrigin="Zootaxa 4007 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onorius inexpectatus Frolov" docType="treatment" docVersion="8" lastPageNumber="435" masterDocId="FF9C67040C7DFFF77B4AFF9DFFA5FF85" masterDocTitle="A new genus and species of Orphninae (Coleoptera: Scarabaeidae) associated with epiphytes in an Andean cloud forest in Ecuador" masterLastPageNumber="436" masterPageNumber="433" pageNumber="434" updateTime="1698603874954" updateUser="plazi">
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<mods:mods id="386452DA0C09F4D38644A3219D57C36C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DA6BF7A2DCEBA35B4F1E51BB897A7EAC">
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<mods:title id="AFA4A0E4369821F50CAC96829BC68C48">A new genus and species of Orphninae (Coleoptera: Scarabaeidae) associated with epiphytes in an Andean cloud forest in Ecuador</mods:title>
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</mods:titleInfo>
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<mods:name id="B0059AF5116CF5E7E9BF48E16064A856" type="personal">
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<mods:role id="6770F52185454A5A19AC7706ECB412B1">
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<mods:roleTerm id="7797F801E81EF172035DC99D2A36B200">Author</mods:roleTerm>
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<mods:namePart id="D0707F1556F92AFDEBFC134C84121672">Frolov, Andrey V.</mods:namePart>
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</mods:name>
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<mods:name id="951C2635AA0937034BC624C0CDD917C2" type="personal">
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<mods:role id="B675EDD63A61570CFECD9447B358EA55">
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<mods:roleTerm id="87A3B330AC94C4FE32AC463B62996F80">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F46B9BAB290A52C1498629BC6A6D354F">Vaz-De-Mello, Fernando Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8C185D3C0806DA28E9CECB8AF9131255">text</mods:typeOfResource>
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<mods:titleInfo id="C9848C1936A3D7A893E3DAD2C1F9313C">
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<mods:title id="E4A38591CAC8AB1D7BED70383D791063">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4BC1E93CE954EB0E352E2B95AFAAAE02">
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<mods:date id="BB10AF2D164DB08469A31B57AE17B243">2015</mods:date>
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<mods:detail id="51A1B86A9EEA89603ACCF213EFA2517D" type="volume">
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<mods:number id="7F4E9C72FFAF1F682F24E21D69EE916B">4007</mods:number>
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</mods:detail>
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<mods:detail id="B389D1BBCAEE0428ADB69E4BD8B93800" type="issue">
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<mods:number id="ED499F13E5128752C09CA9D45E0AB2AA">3</mods:number>
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</mods:detail>
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<mods:start id="E0B1B8651CB489E1F1F21F8B4C351EEF">433</mods:start>
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<mods:end id="67B1B6779A0BE05AD6779E2FBFBDEF9E">436</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="2A99A334B7987440C6EA6A930025F0D0">journal article</mods:classification>
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<mods:identifier id="806A0929E76351353C5BDAA0020516D4" type="DOI">10.11646/zootaxa.4007.3.10</mods:identifier>
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<mods:identifier id="49094A88793E6A25507EB61E770507D0" type="GBIF-Dataset">e659252e-1ea0-4783-af2f-6398fa8b6aa4</mods:identifier>
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<mods:identifier id="CA7AA2498A5CADA5578EA5D9D46D9214" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6582AE42716F8B53CC68A308364F5A6C" type="Zenodo-Dep">240333</mods:identifier>
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</mods:mods>
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<treatment id="03A51F7C0C7CFFF57BDDFF07FCB8FA72" ID-DOI="http://doi.org/10.5281/zenodo.6107961" ID-GBIF-Taxon="119631542" ID-Zenodo-Dep="6107961" LSID="urn:lsid:plazi:treatment:03A51F7C0C7CFFF57BDDFF07FCB8FA72" httpUri="http://treatment.plazi.org/id/03A51F7C0C7CFFF57BDDFF07FCB8FA72" lastPageId="2" lastPageNumber="435" pageId="1" pageNumber="434">
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<subSubSection id="C316FDE10C7CFFF67BDDFF07FCFEFF30" box="[151,859,154,181]" pageId="1" pageNumber="434" type="nomenclature">
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<paragraph id="8BB3AE6A0C7CFFF67BDDFF07FCFEFF30" blockId="1.[151,859,154,181]" box="[151,859,154,181]" pageId="1" pageNumber="434">
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<heading id="D0FB19060C7CFFF67BDDFF07FCFEFF30" bold="true" box="[151,859,154,181]" fontSize="11" level="1" pageId="1" pageNumber="434" reason="1">
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<emphasis id="B97872780C7CFFF67BDDFF07FCFEFF30" bold="true" box="[151,859,154,181]" pageId="1" pageNumber="434">
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<taxonomicName id="4C0CD5E90C7CFFF67BDDFF07FE56FF30" authority="Frolov" authorityName="Frolov" box="[151,499,154,181]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B97872780C7CFFF67BDDFF07FE3DFF31" bold="true" box="[151,408,154,180]" italics="true" pageId="1" pageNumber="434">Onorius inexpectatus</emphasis>
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Frolov
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</taxonomicName>
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& Vaz-de-Mello, new species
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316FDE10C7CFFF57BDDFF7DFA86FB3A" lastPageId="2" lastPageNumber="435" pageId="1" pageNumber="434" type="materials_examined">
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<paragraph id="8BB3AE6A0C7CFFF67BDDFF7DFD01FE07" blockId="1.[151,1436,224,455]" pageId="1" pageNumber="434">
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<emphasis id="B97872780C7CFFF67BDDFF7DFE93FF72" bold="true" box="[151,310,224,247]" pageId="1" pageNumber="434">
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<typeStatus id="54B710C80C7CFFF67BDDFF7DFF6AFF72" box="[151,207,224,247]" pageId="1" pageNumber="434">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="54B710C80C7CFFF67A0FFF7DFE0EFF72" box="[325,427,224,247]" pageId="1" pageNumber="434" type="holotype">Holotype</typeStatus>
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, male: “
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<collectingCountry id="F31BEEFA0C7CFFF67949FF7DFD23FF72" box="[515,646,224,247]" name="Ecuador" pageId="1" pageNumber="434">ECUADOR</collectingCountry>
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COTOPAXI OTONGA
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<quantity id="4CF4038F0C7CFFF678D8FF7DFC41FF72" box="[914,996,224,247]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="434" unit="m" value="1900.0">1900 m</quantity>
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<date id="FFB288AA0C7CFFF678A6FF7DFBFAFF72" box="[1004,1119,224,247]" pageId="1" pageNumber="434" value="1998-02-03">3FEB1998</date>
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GOnore // Ex docel
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<quantity id="4CF4038F0C7CFFF67E09FF7DFADEFF72" box="[1347,1403,224,247]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="434" unit="m" value="16.0">16 m</quantity>
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de alture” (
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<collectionCode id="ED1D36AF0C7CFFF67BA7FE9EFE99FE9F" box="[237,316,259,282]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="434">QCAZ</collectionCode>
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||
).
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<typeStatus id="54B710C80C7CFFF67A07FE9EFE1CFE9F" box="[333,441,259,282]" pageId="1" pageNumber="434" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D0A65E30C7CFFF67A88FE9EFD8DFE9F" box="[450,552,259,282]" pageId="1" pageNumber="434" type="female">2 females</specimenCount>
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with the same data as the
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<typeStatus id="54B710C80C7CFFF67877FE9EFC3FFE9F" box="[829,922,259,282]" pageId="1" pageNumber="434" type="holotype">holotype</typeStatus>
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but the label “Ex docel
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<quantity id="4CF4038F0C7CFFF67FD3FE9EFB6BFE9F" box="[1177,1230,259,282]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="434" unit="m" value="16.0">16 m</quantity>
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de alture” (
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<collectionCode id="ED1D36AF0C7CFFF67E01FE9EFA3DFE9F" box="[1355,1432,259,282]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="434">QCAZ</collectionCode>
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,
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||
<collectionCode id="ED1D36AF0C7CFFF67BDDFEB8FF43FEB9" box="[151,230,293,316]" pageId="1" pageNumber="434">CEMT</collectionCode>
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||
);
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<specimenCount id="9D0A65E30C7CFFF67BB7FEB8FECEFEB9" box="[253,363,293,316]" pageId="1" pageNumber="434" type="female">2 females</specimenCount>
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, “
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<collectingCountry id="F31BEEFA0C7CFFF67ACEFEB8FDA2FEB9" box="[388,519,293,316]" name="Ecuador" pageId="1" pageNumber="434">ECUADOR</collectingCountry>
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COTOPAXI OTONGA
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<date id="FFB288AA0C7CFFF67854FEB8FC28FEB9" box="[798,909,293,316]" pageId="1" pageNumber="434" value="1996-08">AUG1996</date>
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FRios” (
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<collectionCode id="ED1D36AF0C7CFFF678BDFEB8FBE1FEB9" box="[1015,1092,293,316]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="434">QCAZ</collectionCode>
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,
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<collectionCode id="ED1D36AF0C7CFFF67F19FEB8FB21FEB9" box="[1107,1156,293,316]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="434">ZIN</collectionCode>
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);
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<specimenCount id="9D0A65E30C7CFFF67FD1FEB8FB5AFEB9" box="[1179,1279,293,316]" pageId="1" pageNumber="434" type="female">1 female</specimenCount>
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, “
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<collectingCountry id="F31BEEFA0C7CFFF67E52FEB8FA3EFEB9" box="[1304,1435,293,316]" name="Ecuador" pageId="1" pageNumber="434">ECUADOR</collectingCountry>
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COTOPAXI LAS PALMAS
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<quantity id="4CF4038F0C7CFFF67A93FED5FD87FEDA" box="[473,546,328,351]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="434" unit="m" value="1800.0">1800m</quantity>
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<date id="FFB288AA0C7CFFF67964FED5FD16FEDA" box="[558,691,328,351]" pageId="1" pageNumber="434" value="1994-12-04">04DEC1994</date>
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ETapia” (
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<collectionCode id="ED1D36AF0C7CFFF67862FED5FCDDFEDA" box="[808,888,328,351]" pageId="1" pageNumber="434">CEMT</collectionCode>
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||
);
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||
<specimenCount id="9D0A65E30C7CFFF678C5FED5FC56FEDA" box="[911,1011,328,351]" pageId="1" pageNumber="434" type="female">1 female</specimenCount>
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||
, “
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<collectingCountry id="F31BEEFA0C7CFFF67F46FED5FB2AFEDA" box="[1036,1167,328,351]" name="Ecuador" pageId="1" pageNumber="434">ECUADOR</collectingCountry>
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||
COTOPAXI SIGCHOS ASACHE
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||
<quantity id="4CF4038F0C7CFFF67A42FEF6FEF7FE07" box="[264,338,363,386]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="434" unit="m" value="1800.0">1800m</quantity>
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<date id="FFB288AA0C7CFFF67A13FEF6FE7BFE07" box="[345,478,363,386]" pageId="1" pageNumber="434" value="1995-12-31">31DEC1995</date>
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||
ETapia” (
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||
<collectionCode id="ED1D36AF0C7CFFF67900FEF6FD3CFE07" box="[586,665,363,386]" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="1" pageNumber="434">QCAZ</collectionCode>
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||
).
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||
</paragraph>
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||
<paragraph id="8BB3AE6A0C7CFFF67B8DFE10FEDDFE42" blockId="1.[151,1436,224,455]" pageId="1" pageNumber="434">
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||
<emphasis id="B97872780C7CFFF67B8DFE10FE1BFE21" bold="true" box="[199,446,397,420]" pageId="1" pageNumber="434">
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||
<typeStatus id="54B710C80C7CFFF67B8DFE10FE8AFE21" box="[199,303,397,420]" pageId="1" pageNumber="434" type="holotype">Holotype</typeStatus>
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description.
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</emphasis>
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Male (Fig. 1). Body length
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<quantity id="4CF4038F0C7CFFF679B2FE10FCEBFE21" box="[760,846,397,420]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="434" unit="mm" value="9.6">9.6 mm</quantity>
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, width of elytra
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<quantity id="4CF4038F0C7CFFF67F4DFE10FBF8FE21" box="[1031,1117,397,420]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="434" unit="mm" value="4.5">4.5 mm</quantity>
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, width of pronotum
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||
<quantity id="4CF4038F0C7CFFF67E0BFE10FA3DFE21" box="[1345,1432,397,420]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="434" unit="mm" value="3.8">3.8 mm</quantity>
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. Color uniform black.
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</paragraph>
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||
<paragraph id="8BB3AE6A0C7CFFF67BDDF82AFE16F86F" blockId="1.[151,1436,1975,2026]" pageId="1" pageNumber="434">
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<emphasis id="B97872780C7CFFF67BDDF82AFEA2F848" bold="true" box="[151,263,1975,1997]" pageId="1" pageNumber="434">FIGURES.</emphasis>
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<taxonomicName id="4C0CD5E90C7CFFF67A58F825FE44F848" box="[274,481,1976,1997]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="434" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B97872780C7CFFF67A58F825FE44F848" box="[274,481,1976,1997]" italics="true" pageId="1" pageNumber="434">Onorius inexpectatus</emphasis>
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</taxonomicName>
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. 1–2—habitus (1—
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<typeStatus id="54B710C80C7CFFF679ECF82AFCA5F848" box="[678,768,1975,1997]" pageId="1" pageNumber="434" type="holotype">holotype</typeStatus>
|
||
, male, 2—
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||
<typeStatus id="54B710C80C7CFFF6783AF82AFC61F848" box="[880,964,1975,1997]" pageId="1" pageNumber="434" type="paratype">paratype</typeStatus>
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||
, female), 3—
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||
<typeStatus id="54B710C80C7CFFF67F1AF82AFB02F848" box="[1104,1191,1975,1997]" pageId="1" pageNumber="434" type="holotype">holotype</typeStatus>
|
||
, aedeagus in lateral and dorsal view, 4—locality map.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFF05FC12FF71" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">Clypeus rounded anteriorly, with weak sinuation on the right side making clypeus somewhat asymmetrical. Head densely punctate with round punctures separated by 2–3 puncture diameters. Head without traces of medial horn or tubercle. Mandibles and labrum not protruding past clypeus in dorsal view.</paragraph>
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||
<paragraph id="8BB3AE6A0C7FFFF57B8DFE9DFC5FFED9" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFE9DFE94FE92" box="[199,305,256,279]" italics="true" pageId="2" pageNumber="435">Pronotum</emphasis>
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1.5 times wider than long, widest at basal 0.4 of its length. Anterior margin without border, basal and lateral margins bordered. Disc of pronotum with shallow fossa anteriorly and small tubercle apicad of this fossa. Surface of pronotum punctate with round punctures separated by 2–4 puncture diameters.
|
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</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFEF5FAE7FEFA" blockId="2.[151,1436,152,1527]" box="[199,1346,360,383]" pageId="2" pageNumber="435">
|
||
<taxonomicName id="4C0CD5E90C7FFFF57B8DFEF5FE95FEFA" box="[199,304,360,383]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="435" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97872780C7FFFF57B8DFEF5FE95FEFA" box="[199,304,360,383]" italics="true" pageId="2" pageNumber="435">Scutellum</emphasis>
|
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</taxonomicName>
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triangular, narrowly rounded apically, about 1/15 length of elytra, with somewhat rugose surface.
|
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</paragraph>
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||
<paragraph id="8BB3AE6A0C7FFFF57B8DFE17FAB6FE62" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFE17FEAFFE24" box="[199,266,394,417]" italics="true" pageId="2" pageNumber="435">Elytra</emphasis>
|
||
convex, with marked humeral and apical humps, without ridges or other elevated areas. Maximum width approximately at the middle. Striae indistinct. Elytra punctate with round, moderately dense punctures separated by 2–4 puncture diameters; elytral punctures somewhat larger than those on pronotum. Base of elytra not bordered.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFE6FFE18FD8C" blockId="2.[151,1436,152,1527]" box="[199,445,498,521]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFE6FFEADFD8C" box="[199,264,498,521]" italics="true" pageId="2" pageNumber="435">Wings</emphasis>
|
||
fully developed.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFD88FB96FD11" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFD88FE8EFDA9" box="[199,299,533,556]" italics="true" pageId="2" pageNumber="435">Protibiae</emphasis>
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||
with 3 outer teeth and a smaller inner tooth. Lateral margin basad of outer teeth not crenulate. Apex and internal margin of tibia with a few slender setae. Protarsi well developed, about 2/3 length of protibiae. Claws 1/3 length of tarsomere 5. Tarsomere 5 as long as tarsomeres 2–4 combined, somewhat thicker than other tarsomeres. Tarsomere 1 as long as tarsomeres 2–4. Ventral surface of femora punctate with elongate punctures.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFD3DFECBFCC4" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFD3DFE66FD32" box="[199,451,672,695]" italics="true" pageId="2" pageNumber="435">Mesolegs and metalegs</emphasis>
|
||
similar in shape; metafemora and metatibiae about 1.2 times longer than mesofemora and mesotibiae. Femora sparsely punctate with elongate punctures. Tibiae somewhat triangular, with two apical spurs. Upper tibial spur as long as tarsomeres 1–3; lower spur as long as or a bit longer than tarsomere 1. Claws 1/3 length of tarsomere 5. Tarsomere 5 relatively robust, as wide as the other tarsomeres, as long as tarsomeres 2–4 combined, and as long as tarsomere 1.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFCD0FCD2FC02" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFCD0FE39FCE1" box="[199,412,845,868]" italics="true" pageId="2" pageNumber="435">Abdominal sternites</emphasis>
|
||
punctate with elongate punctures. Sternite 8 about 2 times wider than sternites 4–7 medially.
|
||
<taxonomicName id="4C0CD5E90C7FFFF57BDDFCEDFF58FC02" box="[151,253,880,903]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="435" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97872780C7FFFF57BDDFCEDFF58FC02" box="[151,253,880,903]" italics="true" pageId="2" pageNumber="435">Pygidium</emphasis>
|
||
</taxonomicName>
|
||
transverse, irregularly punctate, partly hidden under elytra.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFC0FFC65FBD2" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFC0FFE93FC2C" box="[199,310,914,937]" italics="true" pageId="2" pageNumber="435">Aedeagus.</emphasis>
|
||
Phallobase tube-shaped, 1.3 times longer than parameres. Parameres are of complex shape, with 3 processes apically but without setation (Fig. 3). The shape of the parameres of
|
||
<taxonomicName id="4C0CD5E90C7FFFF578AAFC28FB20FC49" box="[992,1157,949,972]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B97872780C7FFFF578AAFC28FB20FC49" box="[992,1157,949,972]" italics="true" pageId="2" pageNumber="435">O. inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
is similar to that of a new species of
|
||
<taxonomicName id="4C0CD5E90C7FFFF57A40FC45FE4CFC6A" authority="Westwood" authorityName="Westwood" box="[266,489,984,1007]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97872780C7FFFF57A40FC45FED5FC6A" box="[266,368,984,1007]" italics="true" pageId="2" pageNumber="435">Aegidium</emphasis>
|
||
Westwood
|
||
</taxonomicName>
|
||
recently found in the Atlantic Forest ecoregion (
|
||
<bibRefCitation id="EF9DD39B0C7FFFF578B6FC45FA34FC6A" author="Frolov" box="[1020,1425,984,1007]" pageId="2" pageNumber="435" refString="Frolov, A. V., Grossi, P. C. & Vaz-de-Mello, F. Z. (2015) A new species of Aegidium Arrow (Coleoptera: Scarabaeidae) from the Atlantic Forest ecoregion in South America. Zootaxa, 4007 (3), 437 - 439. http: // dx. doi. org / 10.11646 / zootaxa. 4007.3.11" type="journal article" year="2015">Frolov, Grossi & Vaz-de-Mello 2015</bibRefCitation>
|
||
). Except for the unusual shape of the parameres, the latter, however, has all the diagnostic characters of
|
||
<taxonomicName id="4C0CD5E90C7FFFF57E4CFC67FAC9FB94" box="[1286,1388,1018,1041]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97872780C7FFFF57E4CFC67FAC9FB94" box="[1286,1388,1018,1041]" italics="true" pageId="2" pageNumber="435">Aegidium</emphasis>
|
||
</taxonomicName>
|
||
and externally is dissimilar to
|
||
<taxonomicName id="4C0CD5E90C7FFFF57AF2FB80FDC4FBB1" box="[440,609,1053,1076]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B97872780C7FFFF57AF2FB80FE6EFBB1" box="[440,459,1053,1076]" italics="true" pageId="2" pageNumber="435">O</emphasis>
|
||
.
|
||
<emphasis id="B97872780C7FFFF57A96FB80FDC4FBB1" box="[476,609,1053,1076]" italics="true" pageId="2" pageNumber="435">inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
. It is possible that the complex shape of the parameres is a plesiomorphic character state and thus does not suggest close relationships of the two taxa.
|
||
</paragraph>
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFBFFFA86FB3A" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<typeStatus id="54B710C80C7FFFF57B8DFBFFFE9CFBFC" box="[199,313,1122,1145]" pageId="2" pageNumber="435" type="paratype">
|
||
<emphasis id="B97872780C7FFFF57B8DFBFFFE9CFBFC" bold="true" box="[199,313,1122,1145]" pageId="2" pageNumber="435">Paratypes</emphasis>
|
||
</typeStatus>
|
||
. All the
|
||
<typeStatus id="54B710C80C7FFFF57AD0FBFFFDA5FBFC" box="[410,512,1122,1145]" pageId="2" pageNumber="435" type="paratype">paratypes</typeStatus>
|
||
are females differing from the male in having a fully developed apical protibial spur, no inner apical tooth, convex pronotum without fossa and tubercle anteromedially, relatively narrower and more tapering apically pronotum (Fig 2), and longer abdominal sternites 8 and 7. Body length of the
|
||
<typeStatus id="54B710C80C7FFFF57F65FB35FB30FB3A" box="[1071,1173,1192,1215]" pageId="2" pageNumber="435" type="paratype">paratypes</typeStatus>
|
||
9.0–
|
||
<quantity id="4CF4038F0C7FFFF57F81FB35FABAFB3A" box="[1227,1311,1192,1215]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="435" unit="mm" value="9.5">9.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316FDE10C7FFFF57B8DFB57FCB0FA51" pageId="2" pageNumber="435" type="distribution">
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFB57FCB0FA51" blockId="2.[151,1436,152,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFB57FE49FB64" bold="true" box="[199,492,1226,1249]" pageId="2" pageNumber="435">Distribution and habitat.</emphasis>
|
||
The
|
||
<typeStatus id="54B710C80C7FFFF57961FB57FDFCFB64" box="[555,601,1226,1249]" pageId="2" pageNumber="435">type</typeStatus>
|
||
series was collected in a small area in the Toachi River valley (Cotopaxi Province, Central
|
||
<collectingCountry id="F31BEEFA0C7FFFF57A17FB70FE1DFA81" box="[349,440,1261,1284]" name="Ecuador" pageId="2" pageNumber="435">Ecuador</collectingCountry>
|
||
) (Fig. 4). The area is occupied by native cloud forest, characterized by a thick cloud layer covering the forest even in the dry season (
|
||
<bibRefCitation id="EF9DD39B0C7FFFF5793EFA8DFD50FAA2" author="Jarrin" box="[628,757,1296,1319]" pageId="2" pageNumber="435" refString="Jarrin, V. P. (2001) Mamiferos en la niebla: Otonga, un bosque nublado del Ecuador. Publicaciones especiales 5. Museo de Zoologia, Centro de Biodiversidad y Ambiente, Pontificia Universidad Catolica del Ecuador, Quito, Ecuador." type="book chapter" year="2001">Jarrín 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9DD39B0C7FFFF57848FA8DFC37FAA2" author="Zunino" box="[770,914,1296,1319]" pageId="2" pageNumber="435" refString="Zunino, F. (2007) Conservation and new forms of action: Otonga, Ecuador. International Journal of Environment and Health, 1, 243 - 267." type="journal article" year="2007">Zunino 2007</bibRefCitation>
|
||
). The specimens were collected by sifting dead plant material accumulated between epiphytes and tree bark up to
|
||
<quantity id="4CF4038F0C7FFFF5783CFAAFFC0AFACC" box="[886,943,1330,1353]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="435" unit="m" value="16.0">16 m</quantity>
|
||
high (Giovanni Onore, Franz Ríos, personal communication). This is the first record of the
|
||
<taxonomicName id="4C0CD5E90C7FFFF579F1FAC8FC89FAE9" box="[699,812,1365,1388]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="subFamily" subFamily="Orphninae">Orphninae</taxonomicName>
|
||
members collected form such a special habitat. Other orphnines, known from both New World and Old World, are presumably litter dwellers or specialist geobionts (
|
||
<bibRefCitation id="EF9DD39B0C7FFFF57E00FAE5FF6AFA34" author="Paulian" pageId="2" pageNumber="435" refString="Paulian, R. (1948) Revision des Orphnus Africains (Coleoptera, Scarabaeidae). Annales de la Societe entomologique de France, 117, 1 - 75." type="journal article" year="1948">Paulian 1948</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9DD39B0C7FFFF57B95FA07FED8FA34" author="Gourves" box="[223,381,1434,1457]" pageId="2" pageNumber="435" refString="Gourves, J. (1988) Observations sur Hybalus rotroui Peyerimhoff au Maroc (col. Scarabaeidae). L'Entomologiste, 44, 50." type="journal article" year="1988">Gourvès 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9DD39B0C7FFFF57AC4FA07FDBCFA34" author="Frolov" box="[398,537,1434,1457]" pageId="2" pageNumber="435" refString="Frolov, A. V. (2012) Diagnosis, classification, and phylogenetic relationships of the orphnine scarab beetles (Coleoptera, Scarabaeidae: Orphninae). Entomological Review, 92, 782 - 797. http: // dx. doi. org / 10.1134 / S 0013873812070056" type="journal article" year="2012">Frolov 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9DD39B0C7FFFF57963FA07FCC3FA34" author="Gradinarov" box="[553,870,1434,1457]" pageId="2" pageNumber="435" refString="Gradinarov, D. & Petrova, Y. (2012) Development of Steinernema feltiae (Rhabditida: Steinernematidae) in larvae of Chaetonyx robustus (Coleoptera: Orphnidae). ZooNotes, 36, 1 - 4." type="journal article" year="2012">Gradinarov & Petrova 2012</bibRefCitation>
|
||
). Further investigations of scarabs associated with epiphytes in the Andes may reveal other, yet unknown taxa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316FDE10C7FFFF57B8DFA7DFCB8FA72" box="[199,797,1504,1527]" pageId="2" pageNumber="435" type="etymology">
|
||
<paragraph id="8BB3AE6A0C7FFFF57B8DFA7DFCB8FA72" blockId="2.[151,1436,152,1527]" box="[199,797,1504,1527]" pageId="2" pageNumber="435">
|
||
<emphasis id="B97872780C7FFFF57B8DFA7DFEE5FA72" bold="true" box="[199,320,1504,1527]" pageId="2" pageNumber="435">Etymology</emphasis>
|
||
. From Latin, “
|
||
<taxonomicName id="4C0CD5E90C7FFFF57A95FA7DFDC1FA72" box="[479,612,1504,1527]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="435" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B97872780C7FFFF57A95FA7DFDC1FA72" box="[479,612,1504,1527]" italics="true" pageId="2" pageNumber="435">inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
” for unexpected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |