265 lines
30 KiB
XML
265 lines
30 KiB
XML
<document id="A0E6307936968507A5C828B861B2C5A3" ID-DOI="10.1649/072.065.0208" ID-ISSN="1938-4394" ID-Zenodo-Dep="4893087" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1622637701652" checkinUser="carolina" docAuthor="Ratcliffe, Brett C. & Curoe, Daniel J." docDate="2011" docId="0380D251FFB37674FF7AFB4B42D30E4D" docLanguage="en" docName="TheColeopBull.65.2.139-143.pdf.imf" docOrigin="The Coleopterists Bulletin 65 (2)" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Hemiphileurus panamanius Ratcliffe and Curoe, new species" docType="treatment" docVersion="3" lastPageNumber="141" masterDocId="FFB9AA29FFB27676FFEDFA5541330A34" masterDocTitle="A New Species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Panama, with a Revised Key to the Panamanian Species" masterLastPageNumber="143" masterPageNumber="139" pageNumber="140" updateTime="1698952875464" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="29F6A332019B1D041DFA41C20109FDC0" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="45EDF7B68D546BC6E63569897AF4E029">
|
||
<mods:title id="152C1DA5372D49AE1D49769194569BEB">A New Species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Panama, with a Revised Key to the Panamanian Species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="630B0C3BD4965F850BA5A2979EA94526" type="personal">
|
||
<mods:role id="C17B2286F3AF6E8FC4B04B8D5ED6ED5B">
|
||
<mods:roleTerm id="57C4602A011434C09D8A8536316787CB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2E3F857DA637E8663FE3DF774C4398C0">Ratcliffe, Brett C.</mods:namePart>
|
||
<mods:affiliation id="EA2192427387C94232A5CAC008AEBCF5">Systematics Research Collections, W- 436 Nebraska Hall University of Nebraska State Museum, Lincoln, NE 68588 - 0514, U. S. A.</mods:affiliation>
|
||
<mods:nameIdentifier id="838C008A72AA35747D3E0E30FD81CA8D" type="email">bratcliffel@unl.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="BF8D877AAE6268C8FDCED55DF6DC4943" type="personal">
|
||
<mods:role id="1EAD27F3170A8D302596F5D585C543D4">
|
||
<mods:roleTerm id="8047C36A6BA80B8012562EC2F77600C9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F1CC8361F9111BC7667E4F53014E0518">Curoe, Daniel J.</mods:namePart>
|
||
<mods:affiliation id="008FE65611946CC5D56962C95399DF4E">Schiller 524 México D. F. 11580, MEXICO</mods:affiliation>
|
||
<mods:nameIdentifier id="6BDE5D1919275A6113FC3B4C92BB9E2D" type="email">dcuroe@hotmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="447A01D8E3189327C160F6EDEB688203">text</mods:typeOfResource>
|
||
<mods:relatedItem id="5DAE185DE7A83D25E6B7646B93E96C12" type="host">
|
||
<mods:titleInfo id="738CB538892AA2B4E0027FB111A4CBCE">
|
||
<mods:title id="91301FF0397B3006DB93981A076D4DDB">The Coleopterists Bulletin</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="10F5425CD1EE1283A0C214E8A6465D04">
|
||
<mods:date id="07FDBDC1D9081B4EF9C7369797A09992">2011</mods:date>
|
||
<mods:detail id="0D8D0C66C2A99E82EFE9A5DA5D3CC7BD" type="pubDate">
|
||
<mods:number id="16046CAF7073E8F304631B77B0405A28">2011-06-01</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="887F30F059928E90B3B8EE5A5E2BA99B" type="volume">
|
||
<mods:number id="CD6E7584CA3A43AEDC4BDF847FA21F63">65</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="53A82CF09C732BD3147F0DB5AA55141E" type="issue">
|
||
<mods:number id="70C0E800A51A88D497A994FFEC78E8CF">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="90B1BE3BD11A7D5334A4008BF582EC7B" unit="page">
|
||
<mods:start id="E0224115985225F693CC5F4FCD32F4B9">139</mods:start>
|
||
<mods:end id="FBCF72B4FCA6A4DF3DA37E86DF0F2BF1">143</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="36E34F44C532DB420682870BEDE97C29">journal article</mods:classification>
|
||
<mods:identifier id="B946961506A0752AE6FCFD01B80129AB" type="DOI">10.1649/072.065.0208</mods:identifier>
|
||
<mods:identifier id="0E32261953069BACD3E4EB69B1E5A4E0" type="ISSN">1938-4394</mods:identifier>
|
||
<mods:identifier id="BA956F641CEF1A3CDC1BD78243345720" type="Zenodo-Dep">4893087</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0380D251FFB37674FF7AFB4B42D30E4D" LSID="urn:lsid:plazi:treatment:0380D251FFB37674FF7AFB4B42D30E4D" httpUri="http://treatment.plazi.org/id/0380D251FFB37674FF7AFB4B42D30E4D" lastPageId="2" lastPageNumber="141" pageId="1" pageNumber="140">
|
||
<subSubSection id="C33330CCFFB37677FF7AFB4B40F60B7B" pageId="1" pageNumber="140" type="nomenclature">
|
||
<paragraph id="8B966347FFB37677FF7AFB4B40F60B7B" blockId="1.[151,638,286,362]" pageId="1" pageNumber="140">
|
||
<heading id="D0DED42BFFB37677FF7AFB4B40F60B7B" centered="true" fontSize="10" level="2" pageId="1" pageNumber="140" reason="2">
|
||
<emphasis id="B95DBF55FFB37677FF7AFB4B40F60B7B" bold="true" pageId="1" pageNumber="140">
|
||
<taxonomicName id="4C2918C4FFB37677FF7AFB4B434A0B00" authority="Ratcliffe and Curoe" authorityName="Ratcliffe and Curoe" box="[151,633,286,308]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="panamanius" status="sp. nov.">
|
||
<emphasis id="B95DBF55FFB37677FF7AFB4B409A0B00" bold="true" box="[151,425,286,308]" italics="true" pageId="1" pageNumber="140">Hemiphileurus panamanius</emphasis>
|
||
Ratcliffe and Curoe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26E022EFFB37677FEA2FB6F40F60B7B" box="[335,453,314,335]" pageId="1" pageNumber="140" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33330CCFFB37677FEAFFB0140E00B5D" box="[322,467,340,362]" pageId="1" pageNumber="140" type="description">
|
||
<paragraph id="8B966347FFB37677FEAFFB0140E00B5D" blockId="1.[151,638,286,362]" box="[322,467,340,362]" pageId="1" pageNumber="140">
|
||
(
|
||
<figureCitation id="13127FC2FFB37677FEA4FB0140940B5D" box="[329,423,340,362]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="1.[175,208,1686,1705]" captionStartId-1="1.[709,750,938,957]" captionTargetBox-0="[184,590,981,1617]" captionTargetBox-1="[715,1141,206,902]" captionTargetId-0="figure-385@1.[151,637,944,1663]" captionTargetId-1="figure-394@1.[685,1172,183,914]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="Fig. 1. Hemiphileurus panamanius, habitus of holotype." captionText-1="Figs. 2–3. Hemiphileurus panamanius, parameres." figureDoi-0="http://doi.org/10.5281/zenodo.4893093" figureDoi-1="http://doi.org/10.5281/zenodo.4893089" httpUri-0="https://zenodo.org/record/4893093/files/figure.png" httpUri-1="https://zenodo.org/record/4893089/files/figure.png" pageId="1" pageNumber="140">Figs. 1–3</figureCitation>
|
||
,
|
||
<figureCitation id="13127FC2FFB37677FE59FB0140FF0B5D" box="[436,460,340,361]" captionStart="Fig" captionStartId="3.[175,207,908,927]" captionTargetBox="[151,1171,194,885]" captionTargetId="figure-361@3.[151,1172,191,885]" captionTargetPageId="3" captionText="Fig. 14. Cloud forest on Cerro Gaital near El Valle, Panama, where the paratype was collected. Photo by Daniel Curoe." figureDoi="http://doi.org/10.5281/zenodo.4893097" httpUri="https://zenodo.org/record/4893097/files/figure.png" pageId="1" pageNumber="140">14</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33330CCFFB37674FF42FB2A425F0AC6" lastPageId="2" lastPageNumber="141" pageId="1" pageNumber="140" type="materials_examined">
|
||
<paragraph id="8B966347FFB37677FF42FB2A401008A9" blockId="1.[151,639,383,908]" pageId="1" pageNumber="140">
|
||
<emphasis id="B95DBF55FFB37677FF42FB2A40780BA0" bold="true" box="[175,331,383,404]" pageId="1" pageNumber="140">Type Material.</emphasis>
|
||
<materialsCitation id="3B41691AFFB37677FEB4FB2A401008A9" collectingDate="2000-05-29" collectingDateMax="2001-05-26" collectingDateMin="2000-05-29" collectorName="D. Curoe & B. Eya" country="Panama" elevation="2" location="Estacion Pirre" pageId="1" pageNumber="140" specimenCount="5" stateProvince="Darien" typeStatus="holotype">
|
||
H o l o t y p e m a l e, l a b e l e d “
|
||
<collectingCountry id="F33E23D7FFB37677FF4EFBCC40380B9A" box="[163,267,409,430]" name="Panama" pageId="1" pageNumber="140">PANAMA</collectingCountry>
|
||
P.N.
|
||
<collectingRegion id="49EDADA5FFB37677FEBEFBCC40800B9A" box="[339,435,409,430]" country="Panama" name="Darien" pageId="1" pageNumber="140">DARIEN</collectingRegion>
|
||
,
|
||
<location id="8EF6359CFFB37677FE2BFBCC436F0B9A" LSID="urn:lsid:plazi:treatment:0380D251FFB37674FF7AFB4B42D30E4D:8EF6359CFFB37677FE2BFBCC436F0B9A" box="[454,604,409,430]" country="Panama" name="Estacion Pirre" pageId="1" pageNumber="140" stateProvince="Darien">Estacion Pirre</location>
|
||
<quantity id="4CD1CEA2FFB37677FD86FBCC41FE0BFD" metricMagnitude="0" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="140" unit="m" value="1.75">
|
||
<elevation id="00048474FFB37677FD86FBCC41FE0BFD" metricMagnitude="0" metricUnit="m" metricValue="1.75" pageId="1" pageNumber="140" unit="m" value="1.75">1, 75 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF974587FFB37677FF35FBE140510BFD" box="[216,354,435,457]" pageId="1" pageNumber="140" value="2001-05-16" valueMax="2001-05-26" valueMin="2001-05-16">
|
||
<collectingDate id="EFD3BC6FFFB37677FF35FBE140510BFD" box="[216,354,435,457]" pageId="1" pageNumber="140" value="2001-05-16" valueMax="2001-05-26" valueMin="2001-05-16">16–26 v 2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DC0691FFB37677FE83FBE140FB0BFD" box="[366,456,436,457]" pageId="1" pageNumber="140">D. Curoe</collectorName>
|
||
” and with our red
|
||
<typeStatus id="5492DDE5FFB37677FF7AFB9B41C30BD7" box="[151,240,462,483]" pageId="1" pageNumber="140" type="holotype">holotype</typeStatus>
|
||
label. Single male
|
||
<typeStatus id="5492DDE5FFB37677FE22FB9B43140BD7" box="[463,551,462,483]" pageId="1" pageNumber="140" type="paratype">paratype</typeStatus>
|
||
labeled “
|
||
<collectingCountry id="F33E23D7FFB37677FF4FFBBC403D0BCA" box="[162,270,489,510]" name="Panama" pageId="1" pageNumber="140">PANAMA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49EDADA5FFB37677FEF3FBBC40690BCA" box="[286,346,489,510]" country="Panama" name="Cocle" pageId="1" pageNumber="140">Cocle</collectingRegion>
|
||
, El Valle: La Mesa,
|
||
<quantity id="4CD1CEA2FFB37677FDD2FBBC434A0BCA" box="[575,633,489,510]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="140" unit="m" value="850.0">
|
||
<elevation id="00048474FFB37677FDD2FBBC434A0BCA" box="[575,633,489,510]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="140" unit="m" value="850.0">850m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF974587FFB37677FF7AF8514017082D" box="[151,292,516,537]" pageId="1" pageNumber="140" value="2000-05-29" valueMax="2000-06-09" valueMin="2000-05-29">
|
||
<collectingDate id="EFD3BC6FFFB37677FF7AF8514017082D" box="[151,292,516,537]" pageId="1" pageNumber="140" value="2000-05-29" valueMax="2000-06-09" valueMin="2000-05-29">29v-9vi 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DC0691FFB37677FEDEF85140A7082D" box="[307,404,516,537]" pageId="1" pageNumber="140">D. Curoe</collectorName>
|
||
,
|
||
<collectorName id="26DC0691FFB37677FE49F85140D8082D" box="[420,491,516,537]" pageId="1" pageNumber="140">B. Eya</collectorName>
|
||
col//Taken in decaying log” and with our yellow
|
||
<typeStatus id="5492DDE5FFB37677FE1CF84B43710807" box="[497,578,542,563]" pageId="1" pageNumber="140" type="paratype">paratype</typeStatus>
|
||
label.
|
||
<typeStatus id="5492DDE5FFB37677FF7AF86C41C3087A" box="[151,240,569,590]" pageId="1" pageNumber="140" type="holotype">Holotype</typeStatus>
|
||
deposited at the University of Nebraska State Museum (Lincoln, NE, USA) and
|
||
<typeStatus id="5492DDE5FFB37677FDC7F806434E085C" box="[554,637,595,616]" pageId="1" pageNumber="140" type="paratype">paratype</typeStatus>
|
||
deposited in the Daniel Curoe collection (Mexico City, Mexico).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B966347FFB37677FF42F8F6434D09B8" blockId="1.[151,639,383,908]" pageId="1" pageNumber="140">
|
||
<emphasis id="B95DBF55FFB37677FF42F8F64020088C" bold="true" box="[175,275,675,696]" pageId="1" pageNumber="140">
|
||
<typeStatus id="5492DDE5FFB37677FF42F8F6403D088C" box="[175,270,675,696]" pageId="1" pageNumber="140" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Length 24.4 mm; width across humeri 11.2 mm. Color black.
|
||
<emphasis id="B95DBF55FFB37677FE20F8EB433708E7" bold="true" box="[461,516,702,723]" pageId="1" pageNumber="140">Head</emphasis>
|
||
: Frons with moderately dense, moderately large punctures, with longitudinal tubercle either side of slightly concave center. Clypeus triangular, apex acute and reflexed, a small carina extending from apex to base of each horn; surface laterad of each carina declivous and with small, sparse punctures. Interocular width equals 4.9 transverse eye diameters. Antenna 10-
|
||
</paragraph>
|
||
<caption id="DF5633CFFFB37677FD28F9FF45A00988" ID-DOI="http://doi.org/10.5281/zenodo.4893089" ID-Zenodo-Dep="4893089" box="[709,1171,937,957]" httpUri="https://zenodo.org/record/4893089/files/figure.png" pageId="1" pageNumber="140" startId="1.[709,750,938,957]" targetBox="[715,1141,206,902]" targetPageId="1">
|
||
<paragraph id="8B966347FFB37677FD28F9FF45A00988" blockId="1.[685,1171,937,981]" box="[709,1171,937,957]" pageId="1" pageNumber="140">
|
||
<emphasis id="B95DBF55FFB37677FD28F9FF42100989" bold="true" box="[709,803,937,957]" pageId="1" pageNumber="140">Figs. 2–3.</emphasis>
|
||
<taxonomicName id="4C2918C4FFB37677FCD4F9FC451A0988" box="[825,1065,937,956]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB37677FCD4F9FC451A0988" box="[825,1065,937,956]" italics="true" pageId="1" pageNumber="140">Hemiphileurus panamanius</emphasis>
|
||
</taxonomicName>
|
||
, parameres.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B966347FFB37677FD40F99742FF09E1" blockId="1.[685,1171,937,981]" box="[685,972,962,981]" pageId="1" pageNumber="140">
|
||
<emphasis id="B95DBF55FFB37677FD40F99743F209E1" bold="true" box="[685,705,962,981]" pageId="1" pageNumber="140">2)</emphasis>
|
||
Caudal view;
|
||
<emphasis id="B95DBF55FFB37677FCACF997426709E1" bold="true" box="[833,852,962,981]" pageId="1" pageNumber="140">3)</emphasis>
|
||
Lateral view.
|
||
</paragraph>
|
||
<paragraph id="8B966347FFB37677FD40F9AC42AE0CF5" blockId="1.[685,1173,1017,1730]" pageId="1" pageNumber="140">
|
||
segmented, club subequal in length to segments 2–7. Mandibles arcuate on lateral edge, apex acute.
|
||
<emphasis id="B95DBF55FFB37677FD40FE7A42220E70" bold="true" box="[685,785,1071,1092]" pageId="1" pageNumber="140">Pronotum</emphasis>
|
||
: Surface punctate; punctures moderately dense, large, deep, umbilicate, becoming very large on disc. Center with shallow, longitudinal furrow extending from just before base to just beyond middle, furrow slightly broader anteriorly. All margins with strong, marginal bead.
|
||
<emphasis id="B95DBF55FFB37677FC17FEE645090EFC" bold="true" box="[1018,1082,1203,1224]" pageId="1" pageNumber="140">Elytra</emphasis>
|
||
: Surface with punctate striae in furrows; punctures large, deep, ocellate. Intervals strongly convex, all of equal height.
|
||
<emphasis id="B95DBF55FFB37677FCDEFF5642A00F2C" bold="true" box="[819,915,1283,1304]" pageId="1" pageNumber="140">Pygidium</emphasis>
|
||
: Surface densely punctate along base and in angles, becoming moderately punctate in apical half; punctures small to moderate at base and in angles, moderately large to large on apical half, setigerous; setae minute, tawny. In lateral view, surface strongly convex.
|
||
<emphasis id="B95DBF55FFB37677FBE6FFDD45090FA9" bold="true" box="[1035,1082,1416,1437]" pageId="1" pageNumber="140">Legs</emphasis>
|
||
: Protibia tridentate, teeth long, subequally spaced. Apex of metatibia with upper angle prolonged into sharp tooth with several stout spinules below tooth. Apex of first tarsomere of metatarsus prolonged into long spine.
|
||
<emphasis id="B95DBF55FFB37677FD00FC58421E0C16" bold="true" box="[749,813,1549,1570]" pageId="1" pageNumber="140">Venter</emphasis>
|
||
: Prosternal process tall, broadly subrectangular, apex broadly subtriangular, posterior face with small projection near base. Last sternite with small, dense punctures except at center apex where punctures sparse.
|
||
<emphasis id="B95DBF55FFB37677FC7FFC2242CF0CB8" bold="true" box="[914,1020,1655,1676]" pageId="1" pageNumber="140">Parameres</emphasis>
|
||
: Each paramere with large, deep, suboval depression centered at about middle (
|
||
<figureCitation id="13127FC2FFB37677FCDAFCF942A30CF5" box="[823,912,1708,1730]" captionStart="Figs" captionStartId="1.[709,750,938,957]" captionTargetBox="[715,1141,206,902]" captionTargetId="figure-394@1.[685,1172,183,914]" captionTargetPageId="1" captionText="Figs. 2–3. Hemiphileurus panamanius, parameres." figureDoi="http://doi.org/10.5281/zenodo.4893089" httpUri="https://zenodo.org/record/4893089/files/figure.png" pageId="1" pageNumber="140">Figs. 2–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5633CFFFB37677FF42FCC341DB0CF5" ID-DOI="http://doi.org/10.5281/zenodo.4893093" ID-Zenodo-Dep="4893093" httpUri="https://zenodo.org/record/4893093/files/figure.png" pageId="1" pageNumber="140" startId="1.[175,208,1686,1705]" targetBox="[184,590,981,1617]" targetPageId="1">
|
||
<paragraph id="8B966347FFB37677FF42FCC341DB0CF5" blockId="1.[151,638,1686,1729]" pageId="1" pageNumber="140">
|
||
<emphasis id="B95DBF55FFB37677FF42FCC341C00C9D" bold="true" box="[175,243,1686,1705]" pageId="1" pageNumber="140">Fig. 1.</emphasis>
|
||
<taxonomicName id="4C2918C4FFB37677FEE5FCC3433B0C9D" box="[264,520,1686,1705]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB37677FEE5FCC3433B0C9D" box="[264,520,1686,1705]" italics="true" pageId="1" pageNumber="140">Hemiphileurus panamanius</emphasis>
|
||
</taxonomicName>
|
||
, habitus of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B966347FFB07674FD2EFA97425F0AC6" blockId="2.[707,1193,194,1145]" pageId="2" pageNumber="141">
|
||
<collectingCountry id="F33E23D7FFB07674FD2EFA97423D0AE3" box="[707,782,194,215]" name="Panama" pageId="2" pageNumber="141">Panama</collectingCountry>
|
||
” as in the more standard use of the suffix, panamaensis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33330CCFFB07674FD36FAA2450A0BA5" pageId="2" pageNumber="141" type="distribution">
|
||
<paragraph id="8B966347FFB07674FD36FAA242B90B75" blockId="2.[707,1193,194,1145]" pageId="2" pageNumber="141">
|
||
<emphasis id="B95DBF55FFB07674FD36FAA2426E0B38" bold="true" box="[731,861,247,268]" pageId="2" pageNumber="141">Distribution.</emphasis>
|
||
<taxonomicName id="4C2918C4FFB07674FC86FAA245BE0B38" box="[875,1165,247,268]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB07674FC86FAA245BE0B38" box="[875,1165,247,268]" italics="true" pageId="2" pageNumber="141">Hemiphileurus panamanius</emphasis>
|
||
</taxonomicName>
|
||
is known only from rain forests (
|
||
<figureCitation id="13127FC2FFB07674FC18FB47450E0B13" box="[1013,1085,274,295]" captionStart="Fig" captionStartId="3.[175,207,908,927]" captionTargetBox="[151,1171,194,885]" captionTargetId="figure-361@3.[151,1172,191,885]" captionTargetPageId="3" captionText="Fig. 14. Cloud forest on Cerro Gaital near El Valle, Panama, where the paratype was collected. Photo by Daniel Curoe." figureDoi="http://doi.org/10.5281/zenodo.4893097" httpUri="https://zenodo.org/record/4893097/files/figure.png" pageId="2" pageNumber="141">Fig. 14</figureCitation>
|
||
) in central and eastern
|
||
<collectingCountry id="F33E23D7FFB07674FCD5FB7942B50B75" box="[824,902,300,321]" name="Panama" pageId="2" pageNumber="141">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B966347FFB07674FD36FB1245310B43" blockId="2.[707,1193,194,1145]" pageId="2" pageNumber="141">
|
||
<emphasis id="B95DBF55FFB07674FD36FB12425E0B68" bold="true" box="[731,877,327,348]" pageId="2" pageNumber="141">Locality Data.</emphasis>
|
||
<materialsCitation id="3B41691AFFB07674FC96FB1243C40B43" collectorName="El Valle" country="Panama" location="Cocle" pageId="2" pageNumber="141" specimenCount="1" stateProvince="Cocle">
|
||
<emphasis id="B95DBF55FFB07674FC96FB1242DA0B68" bold="true" box="[891,1001,327,348]" pageId="2" pageNumber="141">
|
||
<collectingCountry id="F33E23D7FFB07674FC96FB1242DA0B68" box="[891,1001,327,348]" name="Panama" pageId="2" pageNumber="141">PANAMA</collectingCountry>
|
||
</emphasis>
|
||
(2):
|
||
<collectingRegion id="49EDADA5FFB07674FBCCFB1245680B68" box="[1057,1115,327,348]" country="Panama" name="Cocle" pageId="2" pageNumber="141">Coclé</collectingRegion>
|
||
(1):
|
||
<collectorName id="26DC0691FFB07674FB7EFB1243C40B43" pageId="2" pageNumber="141">El Valle</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<collectingRegion id="49EDADA5FFB07674FCEFFB3742710B43" box="[770,834,354,375]" country="Panama" name="Darien" pageId="2" pageNumber="141">Darién</collectingRegion>
|
||
(1): Estacíon Pirre.
|
||
</paragraph>
|
||
<paragraph id="8B966347FFB07674FD36FB29450A0BA5" blockId="2.[707,1193,194,1145]" box="[731,1081,380,401]" pageId="2" pageNumber="141">
|
||
<emphasis id="B95DBF55FFB07674FD36FB2942490BA5" bold="true" box="[731,890,380,401]" pageId="2" pageNumber="141">Temporal Data.</emphasis>
|
||
May (1), June (1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33330CCFFB07674FD36FBC242D30E4D" pageId="2" pageNumber="141" type="diagnosis">
|
||
<paragraph id="8B966347FFB07674FD36FBC242C00822" blockId="2.[707,1193,194,1145]" pageId="2" pageNumber="141">
|
||
<emphasis id="B95DBF55FFB07674FD36FBC242770B98" bold="true" box="[731,836,407,428]" pageId="2" pageNumber="141">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C2918C4FFB07674FCBCFBC345670B9F" box="[849,1108,406,427]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB07674FCBCFBC345670B9F" box="[849,1108,406,427]" italics="true" pageId="2" pageNumber="141">Hemiphileurus panamanius</emphasis>
|
||
</taxonomicName>
|
||
will key to couplet
|
||
<quantity id="4CD1CEA2FFB07674FCC1FBE4426C0BF2" box="[812,863,433,454]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="2" pageNumber="141" unit="in" value="41.0">41 in</quantity>
|
||
<bibRefCitation id="EFB81EB6FFB07674FC8BFBE442C80BF2" author="Endrodi" box="[870,1019,433,454]" pageId="2" pageNumber="141" refId="ref2518" refString="Endrodi, S. 1985. The Dynastinae of the World. Dr. W. Junk Publishers, Dordrecht, The Netherlands." type="book" year="1985">Endrödi (1985)</bibRefCitation>
|
||
where, after that, the remaining two choices do not fit. The most diagnostic feature of this species is the form of the male parameres (
|
||
<figureCitation id="13127FC2FFB07674FC66F85442D50822" box="[907,998,512,534]" captionStart="Figs" captionStartId="1.[709,750,938,957]" captionTargetBox="[715,1141,206,902]" captionTargetId="figure-394@1.[685,1172,183,914]" captionTargetPageId="1" captionText="Figs. 2–3. Hemiphileurus panamanius, parameres." figureDoi="http://doi.org/10.5281/zenodo.4893089" httpUri="https://zenodo.org/record/4893089/files/figure.png" pageId="2" pageNumber="141">Figs. 2–3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B966347FFB07674FD36F84E42D30E4D" blockId="2.[707,1193,194,1145]" pageId="2" pageNumber="141">
|
||
The parameres of
|
||
<taxonomicName id="4C2918C4FFB07674FC52F84E45430804" box="[959,1136,539,560]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB07674FC52F84E45430804" box="[959,1136,539,560]" italics="true" pageId="2" pageNumber="141">H. panamanius</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2918C4FFB07674FD2EF8604537087F" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2003" box="[707,1028,565,587]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="curoei">
|
||
<emphasis id="B95DBF55FFB07674FD2EF8604291087E" box="[707,930,565,586]" italics="true" pageId="2" pageNumber="141">Hemiphileurus curoei</emphasis>
|
||
Ratcliffe
|
||
</taxonomicName>
|
||
are remarkably similar in that they both have a large, deep, suboval depression at about the middle of each paramere (
|
||
<figureCitation id="13127FC2FFB07674FCDCF8D342A608AF" box="[817,917,645,667]" captionStart="Figs" captionStartId="1.[709,750,938,957]" captionTargetBox="[715,1141,206,902]" captionTargetId="figure-394@1.[685,1172,183,914]" captionTargetPageId="1" captionText="Figs. 2–3. Hemiphileurus panamanius, parameres." figureDoi="http://doi.org/10.5281/zenodo.4893089" httpUri="https://zenodo.org/record/4893089/files/figure.png" pageId="2" pageNumber="141">Figs. 2–3</figureCitation>
|
||
,
|
||
<figureCitation id="13127FC2FFB07674FC4BF8D342FE08AF" box="[934,973,645,667]" captionStart="Figs" captionStartId="2.[197,237,1415,1434]" captionTargetBox="[173,660,192,1390]" captionTargetId="figure-439@2.[173,660,191,1391]" captionTargetPageId="2" captionText="Figs. 4–13. Caudal and lateral views of the parameres of 4–5) Hemiphileurus curoei; 6–7) Hemiphileurus laevicauda; 8–9) Hemiphileurus pygidiopunctissimus; 10–11) Hemiphileurus variolosus; 12–13) Hemiphileurus vicarius." figureDoi="http://doi.org/10.5281/zenodo.4893095" httpUri="https://zenodo.org/record/4893095/files/figure.png" pageId="2" pageNumber="141">4–5</figureCitation>
|
||
, respectively). These two species differ in that the last abdominal sternite in
|
||
<taxonomicName id="4C2918C4FFB07674FCE8F8EF42A408FB" box="[773,919,698,719]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB07674FCE8F8EF42A408FB" box="[773,919,698,719]" italics="true" pageId="2" pageNumber="141">H. panamanius</emphasis>
|
||
</taxonomicName>
|
||
is moderately densely punctate all over (more so along base of segment), while in
|
||
<taxonomicName id="4C2918C4FFB07674FD30F8A5420B0931" authorityName="Ratcliffe" authorityYear="2003" box="[733,824,752,773]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="curoei">
|
||
<emphasis id="B95DBF55FFB07674FD30F8A5420B0931" box="[733,824,752,773]" italics="true" pageId="2" pageNumber="141">H. curoei</emphasis>
|
||
</taxonomicName>
|
||
this segment is punctate in a narrow, transverse band along the base only and sparsely punctate elsewhere. This is usually difficult to ascertain without comparative material. In addition, the elytral intervals in
|
||
<taxonomicName id="4C2918C4FFB07674FC41F90F4576095B" box="[940,1093,858,879]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB07674FC41F90F4576095B" box="[940,1093,858,879]" italics="true" pageId="2" pageNumber="141">H. panamanius</emphasis>
|
||
</taxonomicName>
|
||
are regularly rounded ridges, whereas the elytral intervals in
|
||
<taxonomicName id="4C2918C4FFB07674FD31F9DA42060990" authorityName="Ratcliffe" authorityYear="2003" box="[732,821,911,932]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="curoei">
|
||
<emphasis id="B95DBF55FFB07674FD31F9DA42060990" box="[732,821,911,932]" italics="true" pageId="2" pageNumber="141">H. curoei</emphasis>
|
||
</taxonomicName>
|
||
are crenulated because the large punctures in the striae impinge on the sides of the raised intervals. Lastly, the parameres, while similar, do have differences. In
|
||
<taxonomicName id="4C2918C4FFB07674FC0CF98A454809C0" box="[993,1147,991,1012]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="panamanius">
|
||
<emphasis id="B95DBF55FFB07674FC0CF98A454809C0" box="[993,1147,991,1012]" italics="true" pageId="2" pageNumber="141">H. panamanius</emphasis>
|
||
</taxonomicName>
|
||
, the lateral, cariniform margin of the suboval depression is strongly, evenly arcuate in lateral view, and in
|
||
<taxonomicName id="4C2918C4FFB07674FCE6FE7B425B0E77" authorityName="Ratcliffe" authorityYear="2003" box="[779,872,1070,1091]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="curoei">
|
||
<emphasis id="B95DBF55FFB07674FCE6FE7B425B0E77" box="[779,872,1070,1091]" italics="true" pageId="2" pageNumber="141">H. curoei</emphasis>
|
||
</taxonomicName>
|
||
the cariniform margin is nearly straight and with a small, heavily sclerotized bump or “tubercle” (
|
||
<figureCitation id="13127FC2FFB07674FCBDFE3142BF0E4D" box="[848,908,1124,1145]" captionStart="Figs" captionStartId="2.[197,237,1415,1434]" captionTargetBox="[173,660,192,1390]" captionTargetId="figure-439@2.[173,660,191,1391]" captionTargetPageId="2" captionText="Figs. 4–13. Caudal and lateral views of the parameres of 4–5) Hemiphileurus curoei; 6–7) Hemiphileurus laevicauda; 8–9) Hemiphileurus pygidiopunctissimus; 10–11) Hemiphileurus variolosus; 12–13) Hemiphileurus vicarius." figureDoi="http://doi.org/10.5281/zenodo.4893095" httpUri="https://zenodo.org/record/4893095/files/figure.png" pageId="2" pageNumber="141">Fig. 5</figureCitation>
|
||
, arrow).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |