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<document id="FEA11EF6E0142B85FCB43C151B8C0CB6" ID-DOI="10.3897/zookeys.1202.121633" ID-ZooBank="F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7" ID-publisher-id="121633" URI-arpha="CB9CB628-9D1E-53A4-84B5-3B9C013566B4" URI-zoobank="http://zoobank.org/F2DE6CAC-842E-4275-B3C1-C84D7CEDF5B7" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1716826741008" checkinUser="pensoft" docAuthor="Zhang, Ludan, Wu, Zheng, Li, Shuqiang &amp; Yao, Zhiyuan" docDate="2024" docId="D72D17B6CC225D3CBB6C874820714C8B" docLanguage="en" docName="ZooKeys 1202: 255-286" docOrigin="ZooKeys 1202" docSource="https://zookeys.pensoft.net/article/121633/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Belisana lincang Zhang, Li &amp; Yao 2024, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="286" masterDocId="CB9CB6289D1E53A484B53B9C013566B4" masterDocTitle="Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China" masterLastPageNumber="286" masterPageNumber="255" pageNumber="255" updateTime="1716918634710" updateUser="admin">
<mods:mods id="4E8963CEEED28F9F2C8A05ECA6B6D475" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5DA70D10FAFBE677EB3F2FD7568EA15E">Eight new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae), with an updated overview of Belisana species from Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name id="1D611B055B9F476ABBFE9079F9D55289" type="personal">
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<mods:namePart id="4E352B3DBF650BA6D1FB32C7277AA16E">Zhang, Ludan</mods:namePart>
<mods:nameIdentifier id="74E7F3CA6AE1064177648D947D8886FA" type="ORCID">0000-0001-5657-751X</mods:nameIdentifier>
<mods:affiliation id="3CB42D0BE3D7B790883F377C650B200D">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
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<mods:namePart id="86E1F305D3AA030222623079B1818BDF">Wu, Zheng</mods:namePart>
<mods:affiliation id="681DD45D91E6FE53C76FFA7E82725CEA">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:namePart id="550ECA840405E7F76255C79F19CEC232">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="6A75ECF53A2A8613511CBA5C4299C641" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="3ECF087F676E073095D9D80DF7C1DFFA">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name id="2E5DA101AC8F5E7FB6F3C9A8792E7C61" type="personal">
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<mods:namePart id="C12CD5BEBB70A96198442C09DEB3A2C9">Yao, Zhiyuan</mods:namePart>
<mods:nameIdentifier id="D891B955E98CFA195E7C2753D40C601E" type="ORCID">0000-0002-1631-0949</mods:nameIdentifier>
<mods:affiliation id="06B24BA15CA3F0611ED03FBCD95FDDAE">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:typeOfResource id="BAF008484A8305DFAEC35BEF24110356">text</mods:typeOfResource>
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<mods:title id="C08DADAD098973426A2FCC0C83A9F19C">ZooKeys</mods:title>
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<mods:part id="A36C524A0A36A3FAA3CF375CC5F795C9">
<mods:date id="37122202CB0A4F0FDFA9B498A757E900">2024</mods:date>
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<mods:number id="4D742F72D0FAC9A499F57E9031855ED6">2024-05-27</mods:number>
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<mods:number id="5A43D69AC778DE525BC92BE6AC024F93">1202</mods:number>
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<mods:identifier id="E05E7BCC0D285ED85F34FD2D39577A1B" type="DOI">10.3897/zookeys.1202.121633</mods:identifier>
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<treatment id="D72D17B6CC225D3CBB6C874820714C8B" ID-DOI="http://doi.org/10.5281/zenodo.11356046" ID-Zenodo-Dep="11356046" ID-arpha="D72D17B6-CC22-5D3C-BB6C-874820714C8B" ID-zoobank="https://zoobank.org/7401890D-953B-478C-AFBC-3E8E477689D1" LSID="urn:lsid:plazi:treatment:D72D17B6CC225D3CBB6C874820714C8B" LSID-ZBK="urn:lsid:zoobank.org:act:7401890D-953B-478C-AFBC-3E8E477689D1" httpUri="http://treatment.plazi.org/id/D72D17B6CC225D3CBB6C874820714C8B">
<subSubSection id="C2CB90EA5D983EAF1C462269750FE875" type="nomenclature">
<paragraph id="AE516C1BD73A8EBFBBDE986D955D2F30">
<taxonomicName id="6CB2FA2E5466C73B647FA8CC3F43C9CF" authority="Zhang, Li &amp; Yao" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lincang" status="sp. nov.">
<emphasis id="E47FCFDFFDF68DAFBCF7E09D7502503B" italics="true">Belisana lincang</emphasis>
Zhang, Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="1B4C24C69ECD0428765276E64963646B" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="68C5B182C408A2A38F7913EC290113D2" type="description">
<paragraph id="90C1CABDAFF847B4E9714D262F700050">
<figureCitation id="A7AD350D193E9706FC78E1DA44BF9735" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">Figs 6</figureCitation>
,
<figureCitation id="671807DF31AE2142B7F72C58613C1672" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7</figureCitation>
,
<figureCitation id="D07C8BF47F681C16041E92F6D1ADEB8C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E, F</figureCitation>
,
<figureCitation id="F5FE63B6B6D669BD9D5CD530CE500045" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E1TAG" type="type material">
<paragraph id="4E4BDCEAE3D3CBC628EB6C6C84735B3D">
<heading id="B221E6D7F001859D7383642F8B965346" reason="title">Type material.</heading>
</paragraph>
<paragraph id="098B24D8C547A65F8E394088BBDFFE53">
<materialsCitation id="264C0830617BC0553A68B12717254469" collectingDate="2010-08-05" collectionCode="IZCAS" collectorName="C Wang &amp; Q Zhao &amp; L Lin" country="China" county="Yun County" elevation="1619" latitude="24.331034" location="Bianfu Cave" longLatPrecision="1" longitude="100.23335" municipality="Lincang" specimenCode="Ar 44962, Ar 44963" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="E5FA25968183B1D91D5642A4E5483BA8" bold="true" italics="true">
<typeStatus id="A797886CE2B32DCBF19F1316BE59A005" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="3DACCDA687E71E848B9BE73343475859" count="1" type="male">♂</specimenCount>
(
<named-content id="19FC0BDFC7468A7B321E62D97549267F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="A129B8D42E791136263349D255A7BDC4" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="3AC2A53BA7F1BF5B97199F3259C032E5">Ar 44962</specimenCode>
) and
<emphasis id="3A30207A1423D2A9033480B78A6D1C55" bold="true" italics="true">
<typeStatus id="B2CB09DEEACE8EDFC97623BC4BBCE888" type="paratype">paratype</typeStatus>
</emphasis>
<specimenCount id="1561F4DF6D83B8253C595825EAAD5948" count="1" type="female">1 ♀</specimenCount>
(
<named-content id="DCF7F327DE1BC55AB37FC31AD3AD48C0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="312DD2AD47C172D2C58726A22EE1D7E6" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="2615C7E5DF79179E04DEBA992ED5B595">Ar 44963</specimenCode>
),
<location id="1B696EE84F7F2BEEF148B80488F92DE8" LSID="urn:lsid:plazi:treatment:D72D17B6CC225D3CBB6C874820714C8B:1B696EE84F7F2BEEF148B80488F92DE8" country="China" county="Yun County" latitude="24.331034" longLatPrecision="1" longitude="100.23335" municipality="Lincang" name="Bianfu Cave" stateProvince="Yunnan">Bianfu Cave</location>
(
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ETUAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[100.233350,24.331033]}">
<geoCoordinate id="CC7E40B8D4D2382BC077AAB93135DD6B" degrees="24" direction="north" minutes="19.862" orientation="latitude" precision="1" value="24.331034">24 ° 19.862 ' N</geoCoordinate>
,
<geoCoordinate id="BE649FD2DCDABFDDEFDAA29719416132" degrees="100" direction="east" minutes="14.001" orientation="longitude" precision="1" value="100.23335">100 ° 14.001 ' E</geoCoordinate>
</named-content>
,
<quantity id="0BB9C049DECBCED85CC3546CA24D55DE" metricMagnitude="3" metricUnit="m" metricValue="1.619" unit="m" value="1619.0">
<elevation id="8D55F4B75F51CA66F0D50C80B4900B39" metricMagnitude="3" metricUnit="m" metricValue="1.619" unit="m" value="1619.0">1619 m</elevation>
</quantity>
),
<location id="9A42D5DA7412D1B09D73B61FBB0A38D2" LSID="urn:lsid:plazi:treatment:D72D17B6CC225D3CBB6C874820714C8B:9A42D5DA7412D1B09D73B61FBB0A38D2" country="China" county="Yun County" latitude="24.331034" longLatPrecision="1" longitude="100.23335" municipality="Lincang" name="Yankou Village" stateProvince="Yunnan">Yankou Village</location>
,
<location id="4843B183CFBAB17DAF0FC4D062162ED1" LSID="urn:lsid:plazi:treatment:D72D17B6CC225D3CBB6C874820714C8B:4843B183CFBAB17DAF0FC4D062162ED1" country="China" county="Yun County" latitude="24.331034" longLatPrecision="1" longitude="100.23335" municipality="Lincang" name="Laoxu Town" stateProvince="Yunnan">Laoxu Town</location>
,
<collectingCounty id="A078D781E31A118CFC84252431E8D3EE">Yun County</collectingCounty>
,
<collectingMunicipality id="7710BB649B430FEBA6210421131AFDE2">Lincang</collectingMunicipality>
,
<emphasis id="9731AD2B5909CCD28BB39C0E3CE43995" bold="true">
<collectingRegion id="9536CEF2431EC2D41C5C0A7BC9F4F1B4" country="China" name="Yunnan">Yunnan</collectingRegion>
</emphasis>
,
<emphasis id="D16C9C1B17B3B6ACCC693D7371D32018" bold="true">
<collectingCountry id="F8C3769589F27F1C32B879A49D339687" name="China">China</collectingCountry>
</emphasis>
,
<date id="5C6D3B4296830ACDB9E115F273992E6C" value="2010-08-05">
<collectingDate id="A3C0D97647AC8EA058DB74D17AC7D16A" value="2010-08-05">05 / 08 / 2010</collectingDate>
</date>
,
<collectorName id="AF87691E59E81BBDBC8FF06DBDA4507F">C Wang</collectorName>
,
<collectorName id="ED462585299DBE4AD438B3CD49046398">Q Zhao</collectorName>
&amp;
<collectorName id="5ED3E451EC1E2B50EB21035D79D2C39D">L Lin</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3UAG" type="etymology">
<paragraph id="83F1FAE3720198C97D8832E00B315F9C">
<heading id="3198C1364585350C3BCAE75A3FA03918" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="0FBD24BA39104CB9499FF53F56E94A59">
The specific name refers to the
<typeStatus id="2FD161D26C7379B3496E7D92342A07BE">type</typeStatus>
locality, which is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBVAG" type="diagnosis">
<paragraph id="379AD808F5CFE984358734669E398AA1">
<heading id="FBB90807FD94B61C0429F3B22FB408BE" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="61B56B3C9038B3F16A4B1AF998DBCA40">
The new species resembles
<taxonomicName id="F3743FFD6B1D8ADA82A9DEF008829E55" authority="Yao, Pham &amp; Li, 2015" authorityName="Yao, Pham &amp; Li" authorityYear="2015" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="phungae">
<emphasis id="9194CD809243E137C31889BDD61EBEB2" italics="true">B. phungae</emphasis>
<bibRefCitation id="5BDD21AEB41A476FF54A3EFF43F1ED67" DOI="10.11646/zootaxa.3909.1.1" author="Yao Z &amp; Pham DS &amp; Li S" issue="1" journalOrPublisher="Zootaxa" pagination="1-82" refId="B7" refString="Yao Z, Pham DS, Li S (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa 3909 (1): 1 - 82. https: // doi. org / 10.11646 / zootaxa. 3909.1. 1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species." volume="3909" year="2015">Yao, Pham &amp; Li, 2015</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="0A2BC513E4A0B25914197047EAC93260" DOI="10.11646/zootaxa.3909.1.1" author="Yao Z &amp; Pham DS &amp; Li S" etAl="et al." firstAuthor="Yao" issue="1" journalOrPublisher="Zootaxa" page="9" pagination="1-82" refId="B7" refString="Yao Z, Pham DS, Li S (2015) Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species. Zootaxa 3909 (1): 1–82. https://doi.org/10.11646/zootaxa.3909.1.1" title="Pholcid spiders (Araneae: Pholcidae) from northern Vietnam, with descriptions of nineteen new species." volume="3909" year="2015">Yao et al. 2015: 9</bibRefCitation>
, figs 19 A – D, 20 A – G, 21 A – E) by having similar male chelicerae, bulbal apophyses, and epigyne (Figs
<figureCitation id="6961145B99DB7B3526546103A6732C5C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 A, C, D</figureCitation>
,
<figureCitation id="33AE15B62EDD68DC09CD181168969A4B" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 E</figureCitation>
), but can be distinguished by differences in males: procursus with dorso-distal membranous lamella (arrow
<quantity id="C09960C1000DE831A3233DB0268FCFEA" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="916B1F02927BBBBC08C08B58A67CE0AE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="A6FC719D5CB01994600DBE358032DDC1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
vs absent), distally pointed ventro-subdistal membranous lamella (arrow
<quantity id="94FBC11DD79DC70D1EF1CAFF8A178C20" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
Figs
<figureCitation id="6825ED97A83EAB93DD8330DB6A7EB44B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="FFA47D4740E587080F5A0B7A9471C0BA" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
vs nearly half-round) and without dorso-subdistal sclerite (Figs
<figureCitation id="AA704960E1450547B4D78708BA6E47A9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="1DDB274A7B58BECCC11BD8F80CD912AB" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
vs present); differences in females: epigynal pockets closer to each other (Figs
<figureCitation id="995BFD3A5922B3C49918545350371897" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 A, B</figureCitation>
,
<figureCitation id="29232CE1334D0AF0D8E2E2A09F07C5E5" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 E, F</figureCitation>
vs widely separated), pore plates curved and narrow (Figs
<figureCitation id="969579EE67F268E5E1CB1079B40DB063" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 B</figureCitation>
,
<figureCitation id="F20D98994E843C12338F26A4209A8776" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 F</figureCitation>
vs nearly round).
</paragraph>
<caption id="E313204CFFC10F49403853AEEA7DB120" ID-DOI="10.3897/zookeys.1202.121633.figure6" ID-arpha="005FDAF5-F3F7-509D-A0B9-653F3FEE1F07" httpUri="https://binary.pensoft.net/fig/1055871" startId="F6">
<paragraph id="8458FBBF58ADD55EA2E016A31CD5A5DE">
<label id="86C2871C20924D6F608C1833F8F57C7B">Figure 6.</label>
</paragraph>
<paragraph id="72BCF8569B955112B62AF28C645CDF3D">
<taxonomicName id="E00E771F3B769548F34172D114E089AC" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lincang" status="sp. nov.">
<emphasis id="466F53A450534AE4FEFA1D7C04183DD5" italics="true">Belisana lincang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F90D02ECE39AAC121AEB128A7BCDE78D" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="1147DB51AED4F8ADA47D157D70F59FD8" bold="true">A, B</emphasis>
palp (
<emphasis id="9FAC074A0354BF78E1F150ADE3316FFF" bold="true">A</emphasis>
prolateral view
<emphasis id="072BAD2FF63855A079E7ABC39A6C69CF" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="85084DD0078A556FF8C71023BB377410" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="5FD0A237ECD789BB7284B57335C81B23" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella
<emphasis id="E08599AF48542CCBD5CD6FC052AC73AA" bold="true">D</emphasis>
retrolateral view). Abbreviations: b = bulb,
<abbrev id="ABBRID0EFYAG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus,
<abbrev id="ABBRID0EJYAG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 (
<emphasis id="C0C533357620334C6386F41A80BA82E2" bold="true">A, B</emphasis>
); 0.02 (
<emphasis id="B2D1C92051E21CC6DA9AF7E8E46B7E91" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="B5DB9E83A2D13B107EC5ECC59AD40B64" ID-DOI="10.3897/zookeys.1202.121633.figure7" ID-arpha="D025EE55-EC16-5A0F-A0C4-08EC07263B76" httpUri="https://binary.pensoft.net/fig/1055872" startId="F7">
<paragraph id="74C97DF4C9D2A049F5084F558ACF0A92">
<label id="DD9E896E3C523E0E828565811D7A5004">Figure 7.</label>
</paragraph>
<paragraph id="6ED87B650C44B415C3A74BE4D4D2A04C">
<taxonomicName id="89762BDEB31252E07FAAD3F28CF2E5D5" authority="Zhang &amp; Wu &amp; Li &amp; Yao, 2024" authorityName="Zhang &amp; Wu &amp; Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lincang" status="sp. nov.">
<emphasis id="53F0C85436AF2BE01EAEC719058487EE" italics="true">Belisana lincang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2D80A75A3833BB6CC64E22DEC079674" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="A75513FB563D97D19C7042804E279915" bold="true">C – F</emphasis>
) and paratype female (
<emphasis id="2D88B48FDAC99819B3A3A69E6B49B4DA" bold="true">A, B, G, H</emphasis>
)
<emphasis id="9A3CE9C24FB6CD5AB7BEF9BB22640C19" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="6D090529F551385D14229D35B4380783" bold="true">B</emphasis>
vulva, dorsal view, arrow points at sac-like structure
<emphasis id="778648A2AC99753E0FCB67B30B667F0A" bold="true">C</emphasis>
right bulb, prolateral view
<emphasis id="FB12DE66CEDC99E01D13C0813EE80BC9" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="B363F86B510733BA86F2294890973B30" bold="true">E – H</emphasis>
habitus (
<emphasis id="E56494EB418C19773D5FBB0D700FA776" bold="true">E, G</emphasis>
dorsal view
<emphasis id="E5BA2F07BBE7D480989011ACBE8BDBFA" bold="true">F</emphasis>
lateral view
<emphasis id="0FA46CA97813A8EBEA0363A14E6FCDE5" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0E4ZAG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0EB1AG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EF1AG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EJ1AG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EN1AG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0ER1AG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 (
<emphasis id="D552B4038AD9BFA6E2022D015FA5E2F1" bold="true">A – D</emphasis>
); 0.50 (
<emphasis id="39C0CEDC7388B5EF0B15DA0A917F7385" bold="true">E – H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E51AG" type="description">
<paragraph id="BAF679EB5E0993DD47B1A8DF59A0D185">
<heading id="A21934673AB0689502B49A65B10F9346" reason="title">Description.</heading>
</paragraph>
<paragraph id="26B0DBA459139888C6CDE1FA543638FB">
<emphasis id="8C45B6C040CCFF647AA227F0D143B81A" bold="true">Male</emphasis>
(
<emphasis id="11082C59D0D5D237B77FB8A9B5EB08E7" bold="true" italics="true">
<typeStatus id="6D564AC71EC299ABDA60363C16597AEF" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.78 (1.90 with clypeus), carapace 0.85 long, 0.88 wide, opisthosoma 0.93 long, 0.85 wide. Leg I missing, leg II: 6.69 (1.78, 0.31, 1.62, 2.18, 0.80), leg III: 5.18 (1.42, 0.26, 1.19, 1.68, 0.63), leg IV: 6.70 (1.90, 0.28, 1.70, 2.13, 0.69). Eye inter-distances and diameters:
<abbrev id="ABBRID0EJ2AG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EN2AG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0ER2AG" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EV2AG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EZ2AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.03,
<abbrev id="ABBRID0E42AG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.61 / 0.56. Habitus as in Fig.
<figureCitation id="0994397B362F81C9C88E4E12A25E76B2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 E, F</figureCitation>
. Carapace yellowish, with brown median band; sternum yellowish. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae (Fig.
<figureCitation id="BC0A8E91A54E7A2C722D467D94915537" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 D</figureCitation>
) with a pair of proximo-lateral apophyses and a pair of curved distal apophyses (distance between tips: 0.15). Palp as in Fig.
<figureCitation id="6A913F2C2948EFF9E89836E73CBF753B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 A, B</figureCitation>
; trochanter with ventral apophysis (2 × longer than wide, arrow
<quantity id="ECCF7E3F0D8FDA79E3E3DB408772F63E" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="459F00792BFFB57E5AC017D33270A266" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 B</figureCitation>
); femur with tiny retrolatero-proximal protrusion (arrow
<quantity id="D61618E79510A3C915318BB3CDE57910" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="37E0497765D3D896057E2B25D338AA8C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-distal membranous process (arrow
<quantity id="51494C49D47B9E91CD1242F4ED862F23" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="1F5CB045D1276EC93298C3DD72BAB9E3" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="2F0A78E2E2F04D4CE7E55141ED186483" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
) bearing narrow lateral sclerite, sclerotized prolatero-distal apophysis (arrow
<quantity id="57435108163E1244362E082992E889E8" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="7C221704703ED337C87A39A7D27B2C47" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="8D3F2FFF88A93BCB4C51B5CB0F782F8C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
), dorso-distal membranous lamella (arrow
<quantity id="C368758F75070DE8BFC31AA6E9BC1C95" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
Figs
<figureCitation id="C309B9F83C267318C20821E7B644DF3D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="C0891F967B8917842F47DBA5A73D631C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
), and ventro-subdistal membranous lamella (arrow
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Figs
<figureCitation id="65A8EF3B4395C3470465E9D054664192" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana lincang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-distal membranous process, arrow 2 points at sclerotized prolatero-distal apophysis, arrow 3 points at dorso-distal membranous lamella, arrow 4 points at ventro-subdistal membranous lamella D retrolateral view). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (A, B); 0.02 (C, D)." figureDoi="10.3897/zookeys.1202.121633.figure6" httpUri="https://binary.pensoft.net/fig/1055871">6 C</figureCitation>
,
<figureCitation id="569543AFC3887ABC9045D559D66DB510" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov. E, F B. lincang sp. nov. G, H B. luxi sp. nov. I, J B. tengchong sp. nov. K, L B. tongi sp. nov. Abbreviation: f = flap. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure18" httpUri="https://binary.pensoft.net/fig/1055888">18 E</figureCitation>
); bulb (Fig.
<figureCitation id="267D3B7EDAC9FC2C2ADE62F573A7F10C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 C</figureCitation>
) with distally angular apophysis and simple embolus.
</paragraph>
<paragraph id="7BDCD7352AA0B2790589B8ED1ADB4B00">
<emphasis id="D3076282C38D3DD303C19FD0DB6E398F" bold="true">Female</emphasis>
(
<emphasis id="5013CCD7761535AF5C3840513D806729" bold="true" italics="true">
<typeStatus id="FA9ED780E365ECFB6512379E24C0BAA9" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="44316F27AD136C066D04E081ECCFEB9B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="508B9CD5AC1457C1461184CBE2C3E28F" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="1C4E3F6A183C912EB12C2C45F7283515">Ar 44963</specimenCode>
): Similar to male, habitus as in Fig.
<figureCitation id="91782DFEB2A416CD5C9CC8DC69FA12DB" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 G, H</figureCitation>
. Total length 2.48 (2.62 with clypeus), carapace 0.86 long, 0.86 wide, opisthosoma 1.62 long, 1.31 wide. Leg I: 9.08 (2.33, 0.33, 2.30, 2.97, 1.15); tibia I
<abbrev id="ABBRID0EI5AG" xlink_title="length/diameter">L / d</abbrev>
: 26. Eye inter-distances and diameters:
<abbrev id="ABBRID0EM5AG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EQ5AG" xlink_title="posterior median eye">PME</abbrev>
0.09,
<abbrev id="ABBRID0EU5AG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EY5AG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0E35AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EA6AG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum approximately as wide as long (0.60). Epigyne (Figs
<figureCitation id="9A1236824692B65731D9CAA473C1E238" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 A</figureCitation>
,
<figureCitation id="A9751FBE8B500DC816443DDB0102B6CD" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 E</figureCitation>
) simple and flat, with dark internal shade and a pair of postero-median pockets 0.05 apart. Vulva (Figs
<figureCitation id="6C1DC26314B6196A6DDC6FA2DD39E335" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 B</figureCitation>
,
<figureCitation id="1B0FE10A5A489D947E3BCC2B04A625BB" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 F</figureCitation>
) with sac-like structure (arrow in Figs
<figureCitation id="214E0D204E9859D3607ABEADED6D13F5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana lincang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view, arrow points at sac-like structure C right bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A – D); 0.50 (E – H)." figureDoi="10.3897/zookeys.1202.121633.figure7" httpUri="https://binary.pensoft.net/fig/1055872">7 B</figureCitation>
,
<figureCitation id="E01DF131BE4EAB81EC6C3912DE996E40" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Female genitalia in ventral and dorsal views A, B Belisana honghe sp. nov. C, D B. jiuxiang sp. nov., arrow points at lateral sclerite E, F B. lincang sp. nov., arrow points at sac-like structure G, H B. luxi sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="10.3897/zookeys.1202.121633.figure20" httpUri="https://binary.pensoft.net/fig/1055890">20 F</figureCitation>
), sclerotized anterior arch, and a pair of curved, long elliptic pore plates (5 × longer than wide). Retrolateral trichobothria on tibia I at 9 % proximally; legs with short vertical setae on metatarsi; tarsus I with 20 distinct pseudosegments.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E36AG" type="habitat">
<paragraph id="8B6B68B17102DD9A7C2196D3D404F040">
<heading id="8C7B4900EC8A81E84404CD30E20CE433" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="E0A6FDE29539BD0017934A82CB82B4FA">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EBABG" type="distribution">
<paragraph id="B3E73CC89D40444641DD4278A2658BAA">
<heading id="631F998ECF2E50B29D8539AB5186315F" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="4B0A1E6E6A9F679CC96449D93BEA9645">
<collectingCountry id="478E5F064E17C3850035EBE05D78546E" name="China">China</collectingCountry>
(
<collectingRegion id="38D08389869C9DA997AD43B06B7B4461" country="China" name="Yunnan">Yunnan</collectingRegion>
,
<typeStatus id="CB5F0E74DA7DDE90D401B773CEF1ECB9">type</typeStatus>
locality; Fig.
<figureCitation id="389E2E12B339CB64F81A490842275282" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the new Belisana species from Yunnan, China 1 Belisana honghe sp. nov. 2 B. jiuxiang sp. nov. 3 B. lincang sp. nov. 4 B. luxi sp. nov. 5 B. tengchong sp. nov. 6 B. tongi sp. nov. 7 B. yongsheng sp. nov. 8 B. yunnan sp. nov." figureDoi="10.3897/zookeys.1202.121633.figure1" httpUri="https://binary.pensoft.net/fig/1055866">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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