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<document id="9E5F59271E8947A3E4869E8565900E5C" ID-CLB-Dataset="3079" ID-DOI="10.1649/072.068.0mo4.1" ID-GBIF-Dataset="62322993-ca80-4b6c-ac95-861cbb236abd" ID-ISSN="1938-4394" ID-Zenodo-Dep="4907735" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623093043626" checkinUser="carolina" docAuthor="Park, J. - S. &amp; Carlton, C. E." docDate="2014" docId="03D82A3EBDE611C33CD2FB9DFB43FA19" docLanguage="en" docName="TheColeopBull.68.mo13.1-156.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 13) 68" docStyle="DocumentStyle:4CAA047AD53AEBA766D4E826C2986B94.5:TheColeopBull.2010-2016.monograph" docStyleId="4CAA047AD53AEBA766D4E826C2986B94" docStyleName="TheColeopBull.2010-2016.monograph" docStyleVersion="5" docTitle="Sagola gyudongi Park and Carlton, new species" docType="treatment" docVersion="23" lastPageNumber="144" masterDocId="FFE15246BD69114C3C45FFF3FFD6FF9A" masterDocTitle="A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)" masterLastPageNumber="156" masterPageNumber="1" pageNumber="144" updateTime="1699001444256" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EC51A0C36FDA9C3D0F2606C07FB779D7">A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae)</mods:title>
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<mods:namePart id="722CA499606550FD2854FA954510CF20">Park, J. - S.</mods:namePart>
<mods:affiliation id="8F23CDEA91D1D99DEB292718D9CE4823">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:namePart id="5C8F57A065BC2672CEE8F3E639D2791C">Carlton, C. E.</mods:namePart>
<mods:affiliation id="7BF43540E71E3438F3BF93407DA06D95">Louisiana State Arthropod Museum, Department of Entomology, LSB 404 Louisiana State University Agric</mods:affiliation>
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<mods:title id="F2DAFF2B644BB8811FE409E5D099DACD">The Coleopterists Bulletin</mods:title>
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<mods:date id="63712D7757D22AB42A564017A09574BB">2014</mods:date>
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<mods:title id="8E598D47564971586C570E706064E328">mo 13</mods:title>
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<treatment id="03D82A3EBDE611C33CD2FB9DFB43FA19" ID-DOI="http://doi.org/10.5281/zenodo.5765397" ID-GBIF-Taxon="191244978" ID-Zenodo-Dep="5765397" LSID="urn:lsid:zoobank.org:act:900C3440-B99A-4B29-A031-5B27583DB86F" httpUri="http://treatment.plazi.org/id/03D82A3EBDE611C33CD2FB9DFB43FA19" lastPageNumber="144" pageId="143" pageNumber="144">
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<heading id="D0862C44BDE611C33CD2FB9DFDABFB1E" box="[151,637,1134,1156]" centered="true" fontSize="10" level="2" pageId="143" pageNumber="144" reason="2">
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<taxonomicName id="4C71E0ABBDE611C33CD2FB9DFE2BFB1E" ID-CoL="4TYY4" authority="Park and Carlton" authorityName="Park and Carlton" box="[151,509,1134,1156]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="gyudongi" status="sp. nov.">
<emphasis id="B905473ABDE611C33CD2FB9DFEEBFB19" bold="true" box="[151,317,1134,1155]" italics="true" pageId="143" pageNumber="144">Sagola gyudongi</emphasis>
Park and Carlton
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,
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<paragraph id="8BCE9B28BDE611C33D5DFB7AFE2BFB04" blockId="143.[151,637,1134,1182]" box="[280,509,1161,1182]" pageId="143" pageNumber="144">
(
<figureCitation id="134A87ADBDE611C33D5AFB7AFEAEFB04" box="[287,376,1161,1182]" captionStart="Fig" captionStartId="124.[197,229,1292,1311]" captionTargetBox="[176,1192,186,1268]" captionTargetId="figure-148@124.[173,1193,183,1268]" captionTargetPageId="124" captionText="Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907920" httpUri="https://zenodo.org/record/4907920/files/figure.png" pageId="143" pageNumber="144">
Figs.
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,
<figureCitation id="134A87ADBDE611C33DC1FB7AFE73FB04" box="[388,421,1161,1182]" captionStart="Fig" captionStartId="125.[175,208,1532,1551]" captionTargetBox="[153,1171,197,1508]" captionTargetId="figure-0@125.[151,1171,183,1509]" captionTargetPageId="125" captionText="Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907922" httpUri="https://zenodo.org/record/4907922/files/figure.png" pageId="143" pageNumber="144">
<quantity id="4C8936CDBDE611C33DC1FB7AFE73FB04" box="[388,421,1161,1182]" metricMagnitude="4" metricUnit="kg" metricValue="6.9" pageId="143" pageNumber="144" unit="t" value="69.0">69t</quantity>
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,
<figureCitation id="134A87ADBDE611C33DF4FB7AFE07FB04" box="[433,465,1161,1182]" captionStart="Fig" captionStartId="126.[197,230,1302,1321]" captionTargetBox="[177,1188,193,1268]" captionTargetId="figure-135@126.[173,1193,183,1278]" captionTargetPageId="126" captionText="Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907928" httpUri="https://zenodo.org/record/4907928/files/figure.png" pageId="143" pageNumber="144">
<quantity id="4C8936CDBDE611C33DF4FB7AFE07FB04" box="[433,465,1161,1182]" metricMagnitude="4" metricUnit="kg" metricValue="7.0" pageId="143" pageNumber="144" unit="t" value="70.0">70t</quantity>
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,
<figureCitation id="134A87ADBDE611C33D98FB7AFE20FB04" box="[477,502,1161,1182]" captionStart="Fig" captionStartId="143.[175,207,907,926]" captionTargetBox="[154,636,188,881]" captionTargetId="figure-621@143.[151,637,183,884]" captionTargetPageId="143" captionText="Fig. 74. Known collection localities of New Zealand Sagola species. S. hanae: black circles; S. gyudongi: triangles; S. egmontensis: black squares; S. boonei: stars; S. stewartensis: white circles; S. jeongnamae: white squares." figureDoi="http://doi.org/10.5281/zenodo.4907942" httpUri="https://zenodo.org/record/4907942/files/figure.png" pageId="143" pageNumber="144">74</figureCitation>
)
</paragraph>
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<emphasis id="B905473ABDE611C33CEAFB47FE90FB53" bold="true" box="[175,326,1204,1225]" pageId="143" pageNumber="144">Type Material.</emphasis>
<materialsCitation id="3B199175BDE611C33D11FB47FDE5FAD4" ID-GBIF-Occurrence="3415153797" collectingDate="1967-07-25" collectionCode="NZAC, MB" collectorName="G. Kuschel" country="New Zealand" location="Marlborough" pageId="143" pageNumber="144" specimenCount="1" specimenCount-male="1" stateProvince="Marlborough" typeStatus="holotype">
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<typeStatus id="54CA258ABDE611C33D11FB47FE65FB53" box="[340,435,1204,1225]" pageId="143" pageNumber="144" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F366DBB8BDE611C33DFBFB47FDAFFB53" box="[446,633,1204,1225]" name="New Zealand" pageId="143" pageNumber="144">NEW ZEALAND</collectingCountry>
:
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:
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(
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), aedeagus dissected and mounted in balsam on a clear plastic card, “
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<collectingCountry id="F366DBB8BDE611C33CE6FAF0FEBDFA82" box="[163,363,1283,1304]" name="New Zealand" pageId="143" pageNumber="144">NEW ZEALAND</collectingCountry>
:
<collectionCode id="ED6003EDBDE611C33D3FFAF0FE73FA82" box="[378,421,1283,1304]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="143" pageNumber="144">MB</collectionCode>
:
</emphasis>
Pelorus Bridge,
<date id="FFCFBDE8BDE611C33E20FAF0FF38FAA9" pageId="143" pageNumber="144" value="1967-07-25">
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<collectorName id="2684FEFEBDE611C33CB1FAEDFEB7FAA9" box="[244,353,1310,1331]" pageId="143" pageNumber="144">G. Kuschel</collectorName>
, litter 67/220”, “
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<taxonomicName id="4C71E0ABBDE611C33CD2FACBFDFFFAD7" authority="Park and Carlton 2013" authorityName="Park and Carlton" authorityYear="2013" box="[151,553,1336,1357]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="gyudongi">
<emphasis id="B905473ABDE611C33CD2FACBFE96FAD7" bold="true" box="[151,320,1336,1357]" italics="true" pageId="143" pageNumber="144">Sagola gyudongi</emphasis>
Park and Carlton 2013
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.
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<typeStatus id="54CA258ABDE611C33E07FACBFF07FAF2" pageId="143" pageNumber="144" type="paratype">Paratypes</typeStatus>
(
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</emphasis>
=
<emphasis id="B905473ABDE611C33D5FFAA0FEA8FA19" bold="true" pageId="143" pageNumber="144">
10:
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;
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).
<collectingCountry id="F366DBB8BDE611C33E07FAA0FEC9FA19" name="New Zealand" pageId="143" pageNumber="144">NEW ZEALAND</collectingCountry>
:
<collectingRegion id="49B555CABDE611C33D74FA9DFEA8FA19" box="[305,382,1390,1411]" pageId="143" pageNumber="144">Buller</collectingRegion>
</emphasis>
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<emphasis id="B905473ABDE611C33D3BFA9DFE55FA19" bold="true" box="[382,387,1390,1411]" pageId="143" pageNumber="144">:</emphasis>
<materialsCitation id="3B199175BDE611C33DCAFA9DFDAFFA19" ID-GBIF-Occurrence="3415154165" box="[399,633,1389,1411]" collectionCode="DSC, FMNH" country="New Zealand" pageId="143" pageNumber="144" specimenCount="2" specimenCount-male="2" stateProvince="Buller" typeStatus="paratype">
<specimenCount id="9D7750A1BDE611C33DCAFA9DFE15FA19" box="[399,451,1389,1411]" pageId="143" pageNumber="144" type="male">2♂♂</specimenCount>
(
<collectionCode id="ED6003EDBDE611C33D92FA9DFDDBFA19" box="[471,525,1390,1411]" country="USA" httpUri="http://grbio.org/cool/s9x3-awht" name="Dicty Stock Center" pageId="143" pageNumber="144">DSC</collectionCode>
&amp;
<collectionCode id="ED6003EDBDE611C33E64FA9DFDA5FA19" box="[545,627,1390,1411]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="143" pageNumber="144" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B199175BDE611C33CD2FA7BFF17FA77" ID-GBIF-Occurrence="3415153998" collectingDate="1984-12-14" collectingDateMax="1985-01-06" collectingDateMin="1984-12-14" collectionCode="FIT" collectorName="A. Newton &amp; M. Thayer" country="New Zealand" county="Lakes" elevation="670" location="St. Arnaud" municipality="Lake Rotoiti" pageId="143" pageNumber="144" specimenCount="1" stateProvince="Nelson" typeStatus="paratype">
<collectingRegion id="49B555CABDE611C33CD2FA7BFF34FA07" box="[151,226,1416,1437]" country="New Zealand" name="Nelson" pageId="143" pageNumber="144">Nelson</collectingRegion>
<collectingCounty id="62AFE3A4BDE611C33CABFA7BFEFAFA07" box="[238,300,1416,1437]" pageId="143" pageNumber="144">Lakes</collectingCounty>
NP,
<collectingMunicipality id="6BAA0152BDE611C33D2CFA7BFE20FA07" box="[361,502,1416,1437]" pageId="143" pageNumber="144">Lake Rotoiti</collectingMunicipality>
,
<location id="8EAECDF3BDE611C33E43FA7BFDA8FA07" LSID="urn:lsid:plazi:treatment:03D82A3EBDE611C33CD2FB9DFB43FA19:8EAECDF3BDE611C33E43FA7BFDA8FA07" box="[518,638,1416,1437]" country="New Zealand" county="Lakes" municipality="Lake Rotoiti" name="St. Arnaud" pageId="143" pageNumber="144" stateProvince="Nelson">St. Arnaud</location>
tr,
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,
<date id="FFCFBDE8BDE611C33D40FA50FE5DFA22" box="[261,395,1443,1464]" pageId="143" pageNumber="144" value="1984-12-14">
<collectingDate id="EF8B4400BDE611C33D40FA50FE5DFA22" box="[261,395,1443,1464]" pageId="143" pageNumber="144" value="1984-12-14">14 XII 1984</collectingDate>
</date>
<date id="FFCFBDE8BDE611C33DDDFA50FE2FFA22" box="[408,505,1443,1464]" pageId="143" pageNumber="144" value="1985-01-06">
<collectingDate id="EF8B4400BDE611C33DDDFA50FE2FFA22" box="[408,505,1443,1464]" pageId="143" pageNumber="144" value="1985-01-06">6 I 1985</collectingDate>
</date>
,
<taxonomicName id="4C71E0ABBDE611C33E4FFA50FDABFA22" box="[522,637,1443,1464]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="143" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="B905473ABDE611C33E4FFA50FDABFA22" box="[522,637,1443,1464]" italics="true" pageId="143" pageNumber="144">Nothofagus</emphasis>
</taxonomicName>
forest,
<collectorName id="2684FEFEBDE611C33C99FA4EFE9DFA48" box="[220,331,1469,1490]" pageId="143" pageNumber="144">A. Newton</collectorName>
,
<collectorName id="2684FEFEBDE611C33D12FA4EFE69FA48" box="[343,447,1469,1490]" pageId="143" pageNumber="144">M. Thayer</collectorName>
706,
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&amp;
<collectingMethod id="5230E33FBDE611C33E74FA4EFF17FA77" pageId="143" pageNumber="144">window trap</collectingMethod>
</materialsCitation>
;
<materialsCitation id="3B199175BDE611C33C96FA2BFE7AFA77" ID-GBIF-Occurrence="3415154102" box="[211,428,1495,1517]" collectionCode="FMNH" country="New Zealand" pageId="143" pageNumber="144" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Buller" typeStatus="paratype">
<specimenCount id="9D7750A1BDE611C33C96FA2BFED3FA77" box="[211,261,1495,1517]" pageId="143" pageNumber="144" type="male">2♂♂</specimenCount>
<specimenCount id="9D7750A1BDE611C33D57FA2BFE95FA77" box="[274,323,1495,1517]" pageId="143" pageNumber="144" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED6003EDBDE611C33D12FA2BFE72FA77" box="[343,420,1496,1517]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="143" pageNumber="144" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B199175BDE611C33DF8FA2BFE38F9A6" ID-GBIF-Occurrence="3415154588" collectingDate="1984-12-14" collectingDateMax="1985-01-06" collectingDateMin="1984-12-14" collectorName="A. Newton &amp; M. Thayer" country="New Zealand" county="Lakes" elevation="645" location="St. Arnaud" municipality="Lake Rotoiti" pageId="143" pageNumber="144" specimenCount="1" stateProvince="Nelson" typeStatus="paratype">
<collectingRegion id="49B555CABDE611C33DF8FA2BFDD2FA77" box="[445,516,1496,1517]" country="New Zealand" name="Nelson" pageId="143" pageNumber="144">Nelson</collectingRegion>
<collectingCounty id="62AFE3A4BDE611C33E54FA2BFD9BFA77" box="[529,589,1496,1517]" pageId="143" pageNumber="144">Lakes</collectingCounty>
NP,
<collectingMunicipality id="6BAA0152BDE611C33CD2FA01FEF0F99D" box="[151,294,1522,1543]" pageId="143" pageNumber="144">Lake Rotoiti</collectingMunicipality>
,
<location id="8EAECDF3BDE611C33D7DFA01FE67F99D" LSID="urn:lsid:plazi:treatment:03D82A3EBDE611C33CD2FB9DFB43FA19:8EAECDF3BDE611C33D7DFA01FE67F99D" box="[312,433,1522,1543]" country="New Zealand" county="Lakes" municipality="Lake Rotoiti" name="St. Arnaud" pageId="143" pageNumber="144" stateProvince="Nelson">St. Arnaud</location>
tr,
<quantity id="4C8936CDBDE611C33DA6FA01FDF7F99D" box="[483,545,1522,1543]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="143" pageNumber="144" unit="m" value="645.0">
<elevation id="005C7C1BBDE611C33DA6FA01FDF7F99D" box="[483,545,1522,1543]" metricMagnitude="2" metricUnit="m" metricValue="6.45" pageId="143" pageNumber="144" unit="m" value="645.0">645m</elevation>
</quantity>
,
<date id="FFCFBDE8BDE611C33E76FA01FF11F9B8" pageId="143" pageNumber="144" value="1984-12-14">
<collectingDate id="EF8B4400BDE611C33E76FA01FF11F9B8" pageId="143" pageNumber="144" value="1984-12-14">14 XII 1984</collectingDate>
</date>
<date id="FFCFBDE8BDE611C33C96F9FEFEFEF9B8" box="[211,296,1549,1570]" pageId="143" pageNumber="144" value="1985-01-06">
<collectingDate id="EF8B4400BDE611C33C96F9FEFEFEF9B8" box="[211,296,1549,1570]" pageId="143" pageNumber="144" value="1985-01-06">6 I 1985</collectingDate>
</date>
,
<taxonomicName id="4C71E0ABBDE611C33D76F9FEFE77F9B8" box="[307,417,1549,1570]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="143" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="B905473ABDE611C33D76F9FEFE77F9B8" box="[307,417,1549,1570]" italics="true" pageId="143" pageNumber="144">Nothofagus</emphasis>
</taxonomicName>
forest,
<collectorName id="2684FEFEBDE611C33DAEF9FEFD8EF9B8" box="[491,600,1549,1570]" pageId="143" pageNumber="144">A. Newton</collectorName>
,
<collectorName id="2684FEFEBDE611C33E27F9FEFF0AF9A6" pageId="143" pageNumber="144">M. Thayer</collectorName>
705,
<collectingMethod id="5230E33FBDE611C33D5DF9D4FE4EF9A6" box="[280,408,1575,1596]" pageId="143" pageNumber="144">window trap</collectingMethod>
;
<collectingRegion id="49B555CABDE611C33DE1F9D4FE38F9A6" box="[420,494,1575,1596]" country="New Zealand" name="Nelson" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33DE1F9D4FE38F9A6" bold="true" box="[420,494,1575,1596]" pageId="143" pageNumber="144">Nelson</emphasis>
</collectingRegion>
</materialsCitation>
<emphasis id="B905473ABDE611C33DABF9D4FE24F9A6" bold="true" box="[494,498,1575,1596]" pageId="143" pageNumber="144">:</emphasis>
<materialsCitation id="3B199175BDE611C33DBEF9D4FDACF9A6" ID-GBIF-Occurrence="3415153717" box="[507,634,1575,1597]" collectionCode="FMNH" country="New Zealand" pageId="143" pageNumber="144" specimenCount="1" specimenCount-male="1" stateProvince="Buller" typeStatus="paratype">
<specimenCount id="9D7750A1BDE611C33DBEF9D4FDCFF9A7" box="[507,537,1575,1597]" pageId="143" pageNumber="144" type="male">1♂</specimenCount>
(
<collectionCode id="ED6003EDBDE611C33E6DF9D4FDA5F9A6" box="[552,627,1575,1596]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="143" pageNumber="144" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B199175BDE611C33CD2F9B1FEFDF916" ID-GBIF-Occurrence="3415153513" collectingDate="1982-05-21" collectionCode="V" collectorName="S. Peck" country="New Zealand" location="Takaka" pageId="143" pageNumber="144" specimenCode="FMHD#2005-591" specimenCount="1" stateProvince="Tasman" typeStatus="paratype">
<quantity id="4C8936CDBDE611C33CD2F9B1FF02F9CD" box="[151,212,1602,1623]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="143" pageNumber="144" unit="km" value="20.0">20km</quantity>
ne
<location id="8EAECDF3BDE611C33D40F9B1FE82F9CD" LSID="urn:lsid:plazi:treatment:03D82A3EBDE611C33CD2FB9DFB43FA19:8EAECDF3BDE611C33D40F9B1FE82F9CD" box="[261,340,1602,1623]" country="New Zealand" name="Takaka" pageId="143" pageNumber="144" stateProvince="Tasman">Takaka</location>
,
<collectingRegion id="49B555CABDE611C33D2CF9B1FE6BF9CD" box="[361,445,1602,1623]" country="New Zealand" name="Tasman" pageId="143" pageNumber="144">Tasman</collectingRegion>
NP,
<date id="FFCFBDE8BDE611C33DBEF9B1FDA0F9CD" box="[507,630,1602,1623]" pageId="143" pageNumber="144" value="1982-05-21">
<collectingDate id="EF8B4400BDE611C33DBEF9B1FDA0F9CD" box="[507,630,1602,1623]" pageId="143" pageNumber="144" value="1982-05-21">
21
<collectionCode id="ED6003EDBDE611C33E67F9B1FDE5F9CD" box="[546,563,1602,1623]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="143" pageNumber="144" type="Museum">V</collectionCode>
1982
</collectingDate>
</date>
,
<specimenCode id="DBD73353BDE611C33CD2F9AEFE86F9E8" box="[151,336,1629,1650]" collectionCode="FMHD" pageId="143" pageNumber="144">FMHD#2005-591</specimenCode>
, mixed forest litter,
<collectorName id="2684FEFEBDE611C33E6DF9AEFDACF9E8" box="[552,634,1629,1650]" pageId="143" pageNumber="144">S. Peck</collectorName>
;
<collectingRegion id="49B555CABDE611C33CD2F984FEFDF916" box="[151,299,1655,1676]" country="New Zealand" name="Marlborough" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33CD2F984FEFDF916" bold="true" box="[151,299,1655,1676]" pageId="143" pageNumber="144">Marlborough</emphasis>
</collectingRegion>
</materialsCitation>
<materialsCitation id="3B199175BDE611C33D7DF984FDE5F916" ID-GBIF-Occurrence="3415153394" box="[312,563,1655,1677]" collectionCode="LUNZ" country="New Zealand" location="Sounds" pageId="143" pageNumber="144" specimenCount="3" specimenCount-male="3" stateProvince="Sounds" typeStatus="paratype">
<emphasis id="B905473ABDE611C33D7DF984FE47F916" bold="true" box="[312,401,1655,1676]" pageId="143" pageNumber="144">
<collectingRegion id="49B555CABDE611C33D7DF984FE5AF916" box="[312,396,1655,1676]" pageId="143" pageNumber="144">Sounds</collectingRegion>
:
</emphasis>
<specimenCount id="9D7750A1BDE611C33DD8F984FE19F917" box="[413,463,1655,1677]" pageId="143" pageNumber="144" type="male">3♂♂</specimenCount>
(
<collectionCode id="ED6003EDBDE611C33DA7F984FDFAF916" box="[482,556,1655,1676]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="143" pageNumber="144">LUNZ</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B199175BDE611C33E06F984FDBEF95B" ID-GBIF-Occurrence="3415154151" collectingDate="1988-02-11" collectingDateMax="1988-02-16" collectingDateMin="1988-02-11" collectorName="Outer Chetwode I &amp; M. H. Bowie" country="New Zealand" location="Te Kakaho" pageId="143" pageNumber="144" specimenCount="1" stateProvince="Sounds" typeStatus="paratype">
<collectorName id="2684FEFEBDE611C33E06F984FEDBF93D" pageId="143" pageNumber="144">Outer Chetwode I</collectorName>
, (
<location id="8EAECDF3BDE611C33D67F961FE43F93D" LSID="urn:lsid:plazi:treatment:03D82A3EBDE611C33CD2FB9DFB43FA19:8EAECDF3BDE611C33D67F961FE43F93D" box="[290,405,1682,1703]" country="New Zealand" name="Te Kakaho" pageId="143" pageNumber="144" stateProvince="Sounds">Te Kakaho</location>
),
<date id="FFCFBDE8BDE611C33DECF961FDEBF93D" box="[425,573,1681,1703]" pageId="143" pageNumber="144" value="1988-02-11" valueMax="1988-02-16" valueMin="1988-02-11">
<collectingDate id="EF8B4400BDE611C33DECF961FDEBF93D" box="[425,573,1681,1703]" pageId="143" pageNumber="144" value="1988-02-11" valueMax="1988-02-16" valueMin="1988-02-11">1116 II 1988</collectingDate>
</date>
,
<collectorName id="2684FEFEBDE611C33E0EF961FF01F95B" pageId="143" pageNumber="144">M.H. Bowie</collectorName>
,
<collectingMethod id="5230E33FBDE611C33CA0F95FFEA9F95B" box="[229,383,1708,1729]" pageId="143" pageNumber="144">yellow pan trap</collectingMethod>
in shoreline vegetation
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BDE611C33E80FF31FC56FEDB" pageId="143" pageNumber="144" type="etymology">
<paragraph id="8BCE9B28BDE611C33E80FF31FC56FEDB" blockId="143.[685,1173,194,1411]" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33E80FF31FCEEFF4D" bold="true" box="[709,824,194,215]" pageId="143" pageNumber="144">Etymology.</emphasis>
This species is named for one of the enthusiastic supporters of this study, Gyu-Dong Han. He is one of the first author s colleagues and assisted with molecular components of the phylogenetic analysis.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BDE611C33E80FEB4FBF3FDD1" pageId="143" pageNumber="144" type="diagnosis">
<paragraph id="8BCE9B28BDE611C33E80FEB4FBF3FDD1" blockId="143.[685,1173,194,1411]" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33E80FEB4FCF8FEC6" bold="true" box="[709,814,327,348]" pageId="143" pageNumber="144">Diagnosis.</emphasis>
This species can be distinguished from the other species of the
<taxonomicName id="4C71E0ABBDE611C33FAFFE91FBEEFEED" authorityName="Broun, 1893 b:" authorityYear="1049" box="[1002,1080,354,375]" class="Insecta" family="Staphylinidae" genus="Sagola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B905473ABDE611C33FAFFE91FBEEFEED" box="[1002,1080,354,375]" italics="true" pageId="143" pageNumber="144">insignis</emphasis>
</taxonomicName>
speciesgroup by the following combination of characters: body length 2.63.0 mm; temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; elytra subquadrate; shape of antennomeres and genitalia unique to species.
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BDE611C33E80FDA2FBACFB64" pageId="143" pageNumber="144" type="description">
<paragraph id="8BCE9B28BDE611C33E80FDA2FBACFB64" blockId="143.[685,1173,194,1411]" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33E80FDA2FC97FDFC" bold="true" box="[709,833,593,614]" pageId="143" pageNumber="144">Description.</emphasis>
Length 2.63.0 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler (
<figureCitation id="134A87ADBDE611C33EF0FD75FCDDFD01" box="[693,779,646,667]" captionStart="Fig" captionStartId="124.[197,229,1292,1311]" captionTargetBox="[176,1192,186,1268]" captionTargetId="figure-148@124.[173,1193,183,1268]" captionTargetPageId="124" captionText="Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907920" httpUri="https://zenodo.org/record/4907920/files/figure.png" pageId="143" pageNumber="144">
Fig.
<quantity id="4C8936CDBDE611C33EA2FD75FCDDFD01" box="[743,779,646,667]" metricMagnitude="4" metricUnit="kg" metricValue="6.8" pageId="143" pageNumber="144" unit="t" value="68.0">68t</quantity>
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).
<emphasis id="B905473ABDE611C33F64FD75FCB7FD01" bold="true" box="[801,865,646,667]" pageId="143" pageNumber="144">Head:</emphasis>
Male head triangular, widest across temples (
<figureCitation id="134A87ADBDE611C33F10FD53FC7CFD2F" box="[853,938,672,693]" captionStart="Fig" captionStartId="125.[175,208,1532,1551]" captionTargetBox="[153,1171,197,1508]" captionTargetId="figure-0@125.[151,1171,183,1509]" captionTargetPageId="125" captionText="Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907922" httpUri="https://zenodo.org/record/4907922/files/figure.png" pageId="143" pageNumber="144">
Fig.
<quantity id="4C8936CDBDE611C33FC2FD53FC7CFD2F" box="[903,938,672,693]" metricMagnitude="4" metricUnit="kg" metricValue="6.9" pageId="143" pageNumber="144" unit="t" value="69.0">69t</quantity>
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). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 28 longer than wide, 910 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with flattened ventral surface (
<figureCitation id="134A87ADBDE611C33EF1FC59FCD2FC25" box="[692,772,938,959]" captionStart="Fig" captionStartId="125.[175,208,1532,1551]" captionTargetBox="[153,1171,197,1508]" captionTargetId="figure-0@125.[151,1171,183,1509]" captionTargetPageId="125" captionText="Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907922" httpUri="https://zenodo.org/record/4907922/files/figure.png" pageId="143" pageNumber="144">
Fig.
<quantity id="4C8936CDBDE611C33EA7FC59FCD2FC25" box="[738,772,938,959]" metricMagnitude="4" metricUnit="kg" metricValue="6.9" pageId="143" pageNumber="144" unit="t" value="69.0">69t</quantity>
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). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae (
<figureCitation id="134A87ADBDE611C33873FC2CFB5FFC6E" box="[1078,1161,991,1012]" captionStart="Fig" captionStartId="125.[175,208,1532,1551]" captionTargetBox="[153,1171,197,1508]" captionTargetId="figure-0@125.[151,1171,183,1509]" captionTargetPageId="125" captionText="Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907922" httpUri="https://zenodo.org/record/4907922/files/figure.png" pageId="143" pageNumber="144">
Fig.
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).
<emphasis id="B905473ABDE611C33EE8FC09FCD2FB95" bold="true" box="[685,772,1018,1039]" pageId="143" pageNumber="144">Thorax:</emphasis>
Prosternum as long as wide, widest at one-third length. Elytra subquadrate (
<figureCitation id="134A87ADBDE611C33876FBE7FB5EFBB3" box="[1075,1160,1044,1065]" captionStart="Fig" captionStartId="124.[197,229,1292,1311]" captionTargetBox="[176,1192,186,1268]" captionTargetId="figure-148@124.[173,1193,183,1268]" captionTargetPageId="124" captionText="Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907920" httpUri="https://zenodo.org/record/4907920/files/figure.png" pageId="143" pageNumber="144">
Fig.
<quantity id="4C8936CDBDE611C33820FBE7FB5EFBB3" box="[1125,1160,1044,1065]" metricMagnitude="4" metricUnit="kg" metricValue="6.8" pageId="143" pageNumber="144" unit="t" value="68.0">68t</quantity>
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). Meso- and metathorax trapezoidal, as long as wide.
<emphasis id="B905473ABDE611C33EE8FBBAFCCEFBC4" bold="true" box="[685,792,1097,1118]" pageId="143" pageNumber="144">Abdomen:</emphasis>
Male tergite IV with or without pair of small, round patches of microtrichia, patches absent in female.
<emphasis id="B905473ABDE611C33F1DFB8CFC17FB0E" bold="true" box="[856,961,1151,1172]" pageId="143" pageNumber="144">Aedeagus:</emphasis>
Median lobe of genitalia broad, with triangular apical lobe (
<figureCitation id="134A87ADBDE611C3387DFB6AFB5FFB34" box="[1080,1161,1177,1198]" captionStart="Fig" captionStartId="126.[197,230,1302,1321]" captionTargetBox="[177,1188,193,1268]" captionTargetId="figure-135@126.[173,1193,183,1278]" captionTargetPageId="126" captionText="Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907928" httpUri="https://zenodo.org/record/4907928/files/figure.png" pageId="143" pageNumber="144">
Fig.
<quantity id="4C8936CDBDE611C3382DFB6AFB5FFB34" box="[1128,1161,1177,1198]" metricMagnitude="4" metricUnit="kg" metricValue="7.0" pageId="143" pageNumber="144" unit="t" value="70.0">70t</quantity>
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). Phallobase of median lobe symmetrical and rounded (
<figureCitation id="134A87ADBDE611C33F43FB3DFC87FB79" box="[774,849,1230,1251]" captionStart="Fig" captionStartId="126.[197,230,1302,1321]" captionTargetBox="[177,1188,193,1268]" captionTargetId="figure-135@126.[173,1193,183,1278]" captionTargetPageId="126" captionText="Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907928" httpUri="https://zenodo.org/record/4907928/files/figure.png" pageId="143" pageNumber="144">
Fig.
<quantity id="4C8936CDBDE611C33F75FB3DFC87FB79" box="[816,849,1230,1251]" metricMagnitude="4" metricUnit="kg" metricValue="7.0" pageId="143" pageNumber="144" unit="t" value="70.0">70t</quantity>
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). Parameres symmetrical and triangular, setose from midpoints to apices (
<figureCitation id="134A87ADBDE611C33864FB1AFBB9FB64" box="[1057,1135,1257,1278]" captionStart="Fig" captionStartId="126.[197,230,1302,1321]" captionTargetBox="[177,1188,193,1268]" captionTargetId="figure-135@126.[173,1193,183,1278]" captionTargetPageId="126" captionText="Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4907928" httpUri="https://zenodo.org/record/4907928/files/figure.png" pageId="143" pageNumber="144">
Fig.
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).
</paragraph>
</subSubSection>
<subSubSection id="C36BC8A3BDE611C33E80FAF0FB43FA19" pageId="143" pageNumber="144" type="distribution">
<paragraph id="8BCE9B28BDE611C33E80FAF0FBEAFAA9" blockId="143.[685,1173,194,1411]" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33E80FAF0FC9EFA82" bold="true" box="[709,840,1283,1304]" pageId="143" pageNumber="144">Distribution.</emphasis>
Buller,
<collectingRegion id="49B555CABDE611C33FEDFAF0FBE3FA82" box="[936,1077,1283,1304]" country="New Zealand" name="Marlborough" pageId="143" pageNumber="144">Marlborough</collectingRegion>
,
<collectingRegion id="49B555CABDE611C33806FAF0FB59FA82" box="[1091,1167,1283,1304]" country="New Zealand" name="Nelson" pageId="143" pageNumber="144">Nelson</collectingRegion>
,
<collectingRegion id="49B555CABDE611C33EE8FAEDFCFCFAA9" box="[685,810,1310,1331]" country="New Zealand" name="Marlborough" pageId="143" pageNumber="144">Marlborough</collectingRegion>
Sounds (
<figureCitation id="134A87ADBDE611C33FCDFAEDFC07FAA9" box="[904,977,1310,1331]" captionStart="Fig" captionStartId="143.[175,207,907,926]" captionTargetBox="[154,636,188,881]" captionTargetId="figure-621@143.[151,637,183,884]" captionTargetPageId="143" captionText="Fig. 74. Known collection localities of New Zealand Sagola species. S. hanae: black circles; S. gyudongi: triangles; S. egmontensis: black squares; S. boonei: stars; S. stewartensis: white circles; S. jeongnamae: white squares." figureDoi="http://doi.org/10.5281/zenodo.4907942" httpUri="https://zenodo.org/record/4907942/files/figure.png" pageId="143" pageNumber="144">Fig. 74</figureCitation>
: triangles).
</paragraph>
<paragraph id="8BCE9B28BDE611C33E80FACBFB43FA19" blockId="143.[685,1173,194,1411]" pageId="143" pageNumber="144">
<emphasis id="B905473ABDE611C33E80FACBFCCCFAD7" bold="true" box="[709,794,1336,1357]" pageId="143" pageNumber="144">Habitat.</emphasis>
Specimens of this species were collected using window and yellow pan traps or by sifting leaf litter in
<taxonomicName id="4C71E0ABBDE611C33F06FA9DFC79FA19" box="[835,943,1390,1411]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="143" pageNumber="144" phylum="Tracheophyta" rank="genus">
<emphasis id="B905473ABDE611C33F06FA9DFC79FA19" box="[835,943,1390,1411]" italics="true" pageId="143" pageNumber="144">Nothofagus</emphasis>
</taxonomicName>
forests or on shorelines.
</paragraph>
</subSubSection>
</treatment>
</document>