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<document id="B2CCFC0FA3426162730388FCF8B9101F" ID-CLB-Dataset="7896" ID-DOI="10.11646/zootaxa.4875.1.1" ID-GBIF-Dataset="a75d6e42-221a-4df9-9a7d-5f33dbff6a56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4578779" ID-ZooBank="278200CE-E16F-45B4-9A89-60C2052415C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610024385246" checkinUser="plazi" docAuthor="Görn, Sebastian" docDate="2020" docId="807787F6B835FFD549A1D616FBD13CDD" docLanguage="en" docName="zootaxa.4875.1.1.pdf" docOrigin="Zootaxa 4875 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heptodonta halensis, sp. nov." docType="treatment" docVersion="7" lastPageNumber="14" masterDocId="7C4EFF8EB83EFFD84936D36BFFA83A3C" masterDocTitle="Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="12" updateTime="1698923305811" updateUser="ExternalLinkService">
<mods:mods id="8C275704585F0C002C8B5A3642FC921E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5A1D8F0A0F4656235E7B415E820D10B2">Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="01EFB459341651BBD41A91070216BD03">Görn, Sebastian</mods:namePart>
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<mods:title id="46568507410A904FCC0989154C869CF6">Zootaxa</mods:title>
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<mods:date id="77D8FF6502CBB15BACC6FA2A36E0C167">2020</mods:date>
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<mods:number id="F7EF36617EE5BC412C44B1BCE782CC8F">2020-11-06</mods:number>
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<mods:number id="A1016FCA8E42E6EAB32668E7310419D9">4875</mods:number>
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<mods:classification id="5370E964290E7041DEDD6E7C6E512022">journal article</mods:classification>
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<mods:identifier id="37B4A56060DFD0F70D40A57042C71B81" type="DOI">10.11646/zootaxa.4875.1.1</mods:identifier>
<mods:identifier id="9E6DC9F91C7AAD4D0B17EDB2AD36670C" type="GBIF-Dataset">a75d6e42-221a-4df9-9a7d-5f33dbff6a56</mods:identifier>
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<treatment id="807787F6B835FFD549A1D616FBD13CDD" ID-DOI="http://doi.org/10.5281/zenodo.4579785" ID-GBIF-Taxon="179988675" ID-Zenodo-Dep="4579785" LSID="urn:lsid:plazi:treatment:807787F6B835FFD549A1D616FBD13CDD" httpUri="http://treatment.plazi.org/id/807787F6B835FFD549A1D616FBD13CDD" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<subSubSection id="40C4656BB835FFD349A1D616FE5D3FAB" box="[151,501,1405,1432]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="086136E0B835FFD349A1D616FE5D3FAB" blockId="11.[151,501,1405,1467]" box="[151,501,1405,1432]" pageId="11" pageNumber="12">
<heading id="5329818CB835FFD349A1D616FE5D3FAB" bold="true" box="[151,501,1405,1432]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="4">
<emphasis id="3AAAEAF2B835FFD349A1D616FE5D3FAB" bold="true" box="[151,501,1405,1432]" pageId="11" pageNumber="12">
<taxonomicName id="CFDE4D63B835FFD349A1D616FE273FA4" ID-CoL="8MK6B" authority="Gorn. 2020" box="[151,399,1405,1432]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
<emphasis id="3AAAEAF2B835FFD349A1D616FE273FA4" bold="true" box="[151,399,1405,1432]" italics="true" pageId="11" pageNumber="12">Heptodonta halensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21995789B835FFD348A1D616FE5D3FAB" box="[407,501,1405,1431]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB835FFD349A1D6CAFEB43F87" box="[151,284,1441,1467]" pageId="11" pageNumber="12" type="description">
<paragraph id="086136E0B835FFD349A1D6CAFEB43F87" blockId="11.[151,501,1405,1467]" box="[151,284,1441,1467]" pageId="11" pageNumber="12">
<figureCitation id="90E52A65B835FFD349A1D6CAFEB43F87" box="[151,284,1441,1467]" captionStart="FIGURES 2329" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,186,1856]" captionTargetId="figure-20@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 2329. Heptodonta halensis sp. nov.: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="11" pageNumber="12">Figs. 2329</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB835FFD349A1D68FFAAD3DB6" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="086136E0B835FFD349A1D68FFC873FC2" blockId="11.[151,1437,1508,2039]" box="[151,815,1508,1535]" pageId="11" pageNumber="12">
<materialsCitation id="B8B63CBDB835FFD349A1D68FFDB73FC3" ID-GBIF-Occurrence="3045076427" box="[151,543,1508,1535]" country="Philippines" location="Leyte" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Southern Leyte" typeStatus="holotype">
<emphasis id="3AAAEAF2B835FFD349A1D68FFEE03FC2" bold="true" box="[151,328,1508,1534]" pageId="11" pageNumber="12">
<typeStatus id="D7658842B835FFD349A1D68FFF7A3FC2" box="[151,210,1508,1534]" pageId="11" pageNumber="12">Type</typeStatus>
localities.
</emphasis>
<collectingCountry id="70C97670B835FFD34879D68FFE7A3FC2" box="[335,466,1508,1534]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
:
<collectingRegion id="CA1AF802B835FFD348EAD68EFDB73FC3" box="[476,543,1509,1535]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="12">Leyte</collectingRegion>
</materialsCitation>
; Luzon; Panaon;
<collectingRegion id="CA1AF802B835FFD34BD4D68FFC843FC2" box="[738,812,1508,1534]" country="Philippines" name="Samar" pageId="11" pageNumber="12">Samar</collectingRegion>
.
</paragraph>
<paragraph id="086136E0B835FFD349F1D563FAAD3DB6" blockId="11.[151,1437,1508,2039]" pageId="11" pageNumber="12">
<emphasis id="3AAAEAF2B835FFD349F1D563FE433C1E" bold="true" box="[199,491,1544,1570]" pageId="11" pageNumber="12">
Type material.
<typeStatus id="D7658842B835FFD34848D563FE433C1E" box="[382,491,1544,1570]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="1ED8FD69B835FFD348CDD562FDB83C1F" box="[507,528,1545,1571]" pageId="11" pageNumber="12" type="male"></specimenCount>
in JWCW, labelled: “
<collectingCountry id="70C97670B835FFD34A31D563FC233C1E" box="[775,907,1544,1570]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
, /
<date id="7C601020B835FFD34A91D563FBCF3C1F" box="[935,1127,1544,1571]" pageId="11" pageNumber="12" value="2005-05-26" valueMax="2005-05-30" valueMin="2005-05-26">26. - 30. 5. 2005</date>
, /
<collectingRegion id="CA1AF802B835FFD34DB5D562FB6B3C1F" box="[1155,1219,1545,1571]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="12">Leyte</collectingRegion>
Island, / Mt. Balo- caue,
<quantity id="CF269B05B835FFD349EFD547FE8A3C7B" box="[217,290,1580,1607]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="600.0">600 m</quantity>
, / C. Pottgens leg.” [yellow, printed] // “
<taxonomicName id="CFDE4D63B835FFD34BD5D547FCC13C7A" authorityName="Analis. Hope" authorityYear="1838" box="[739,873,1580,1606]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ analis analis / (
<bibRefCitation id="6C4F4B11B835FFD34D11D546FB713C7A" author="Fabricius, J. C." box="[1063,1241,1580,1607]" pageId="11" pageNumber="12" refId="ref38373" refString="Fabricius, J. C. (1801) Systema Eleutheratorum secundum, ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiel, xxiv + 506 pp." type="book" year="1801">FABRICIUS, 1801</bibRefCitation>
) / det. J.
<bibRefCitation id="6C4F4B11B835FFD34C76D547FF7A3C56" author="Wiesner, J." pageId="11" pageNumber="12" pagination="457 - 461" refId="ref41421" refString="Wiesner, J. (2006) New records of tiger beetles from Myanmar (Coleoptera: Cicindelidae). Lambillionea, 106, 457 - 461." type="journal article" year="2006">Wiesner 2006</bibRefCitation>
” [printed].
<emphasis id="3AAAEAF2B835FFD34867D53BFE613C56" bold="true" box="[337,457,1616,1642]" pageId="11" pageNumber="12">
<typeStatus id="D7658842B835FFD34867D53BFE613C56" box="[337,457,1616,1642]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
</emphasis>
(18):
<specimenCount id="1ED8FD69B835FFD34B3AD53BFDE23C57" box="[524,586,1616,1643]" pageId="11" pageNumber="12" type="male">3 ♂♂</specimenCount>
<specimenCount id="1ED8FD69B835FFD34B66D53BFD273C57" box="[592,655,1616,1643]" pageId="11" pageNumber="12" type="female">4 ♀♀</specimenCount>
in JWCW,
<specimenCount id="1ED8FD69B835FFD34A38D53BFC9F3C57" box="[782,823,1616,1643]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
in SGCH,
<specimenCount id="1ED8FD69B835FFD34A87D53BFC733C57" box="[945,987,1616,1643]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
in SMNS: “
<collectingCountry id="70C97670B835FFD34D57D53BFB4D3C56" box="[1121,1253,1616,1642]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
,
<collectingRegion id="CA1AF802B835FFD34DD9D53AFA9A3C57" box="[1263,1330,1617,1643]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="12">Leyte</collectingRegion>
, /
<date id="7C601020B835FFD34C7CD53BFA303C56" box="[1354,1432,1616,1642]" pageId="11" pageNumber="12" value="2014-05">v.2014</date>
, / Noel Mohagan leg.” [yellow, printed] // “
<taxonomicName id="CFDE4D63B835FFD34BB7D51FFCAF3CB2" authorityName="Analis. Hope" authorityYear="1838" box="[641,775,1652,1678]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ analis analis / (
<bibRefCitation id="6C4F4B11B835FFD34AF9D51EFB2B3CB2" author="Fabricius, J. C." box="[975,1155,1652,1679]" pageId="11" pageNumber="12" refId="ref38373" refString="Fabricius, J. C. (1801) Systema Eleutheratorum secundum, ordines, genera, species, adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiel, xxiv + 506 pp." type="book" year="1801">FABRICIUS, 1801</bibRefCitation>
) / det. J. Wiesner 2014” [printed].
<specimenCount id="1ED8FD69B835FFD3483FD5F3FE9D3C8F" box="[265,309,1688,1715]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
in JWCW: “
<collectingCountry id="70C97670B835FFD348FED5F3FDE43C8E" box="[456,588,1688,1714]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
,
<collectingRegion id="CA1AF802B835FFD34B6FD5F2FD753C8F" box="[601,733,1689,1715]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="12">Leyte Prov.</collectingRegion>
/ Reg. VIII, Marplag, Barangay / Mt. Bulocaue,
<quantity id="CF269B05B835FFD34C27D5F3FAF23C8F" box="[1297,1370,1688,1715]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="12" unit="m" value="600.0">600 m</quantity>
/ leg. F. Mohagan jr.” [yellow margin, printed] // “
<taxonomicName id="CFDE4D63B835FFD34BABD5D7FC8B3CEA" authorityName="Analis. Hope" authorityYear="1838" box="[669,803,1724,1750]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ melanopyga /
<bibRefCitation id="6C4F4B11B835FFD34AD2D5D7FB2E3CEA" author="Schaum, H." box="[996,1158,1724,1751]" pageId="11" pageNumber="12" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">SCHAUM, 1862</bibRefCitation>
/ det. J. Wiesner 2005” [printed].
<specimenCount id="1ED8FD69B835FFD34831D58BFE983CC7" box="[263,304,1760,1787]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
<specimenCount id="1ED8FD69B835FFD34800D58BFEC83CC7" box="[310,352,1760,1787]" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
in PSCH: “
<collectingCountry id="70C97670B835FFD348D6D58BFD233CC6" box="[480,651,1760,1786]" name="Philippines" pageId="11" pageNumber="12">PHILIPPINES</collectingCountry>
, North Luzon /
<collectingRegion id="CA1AF802B835FFD34A0CD58BFC2E3CC6" box="[826,902,1760,1786]" country="Philippines" name="Ifugao" pageId="11" pageNumber="12">Ifugao</collectingRegion>
, Banaue, /
<date id="7C601020B835FFD34D36D58BFBC53CC6" box="[1024,1133,1760,1786]" pageId="11" pageNumber="12" value="2016-07">July 2016</date>
/ local collector” [printed].
<specimenCount id="1ED8FD69B835FFD349A1D46FFF693D23" box="[151,193,1796,1823]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
in PSCH: “
<collectingCountry id="70C97670B835FFD34872D46FFE473D22" box="[324,495,1796,1822]" name="Philippines" pageId="11" pageNumber="12">PHILIPPINES</collectingCountry>
,
<collectingRegion id="CA1AF802B835FFD348CCD46FFDEB3D22" box="[506,579,1796,1822]" country="Philippines" name="Samar" pageId="11" pageNumber="12">Samar</collectingRegion>
, / Central Visayas, Lope de Vega /
<date id="7C601020B835FFD34AFDD46FFB913D22" box="[971,1081,1796,1822]" pageId="11" pageNumber="12" value="2016-07">July 2016</date>
/ local collector” [printed].
<specimenCount id="1ED8FD69B835FFD34C44D46FFA343D23" box="[1394,1436,1796,1823]" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
in ZSM: “Panaon / Philippin” [printed].
<specimenCount id="1ED8FD69B835FFD34B6FD443FD303D7F" box="[601,664,1832,1859]" pageId="11" pageNumber="12" type="male">2 ♂♂</specimenCount>
<specimenCount id="1ED8FD69B835FFD34BA9D443FD763D7F" box="[671,734,1832,1859]" pageId="11" pageNumber="12" type="female">2 ♀♀</specimenCount>
in MNB: “
<collectingCountry id="70C97670B835FFD34A6FD443FC753D7E" box="[857,989,1832,1858]" name="Philippines" pageId="11" pageNumber="12">Philippinen</collectingCountry>
/ Panaon” [handwritten] // “
<taxonomicName id="CFDE4D63B835FFD34C21D443FA353D7E" authorityName="Analis. Hope" authorityYear="1838" box="[1303,1437,1832,1858]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/
<taxonomicName id="CFDE4D63B835FFD34993D427FE093D5B" authority="(Schaum)" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[165,417,1868,1895]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melanopyga">melanopyga (Schaum)</taxonomicName>
/ Museum f. Naturkunde / Berlin” [printed]. All type specimens labelled: “
<typeStatus id="D7658842B835FFD34DE8D427FADD3D5A" box="[1246,1397,1868,1894]" pageId="11" pageNumber="12" type="holotype">HOLOTYPE</typeStatus>
(or
<typeStatus id="D7658842B835FFD349A1D41BFE8A3DB6" box="[151,290,1904,1930]" pageId="11" pageNumber="12" type="paratype">PARATYPE</typeStatus>
respectively) /
<taxonomicName id="CFDE4D63B835FFD348E6D41BFDFB3DB6" authorityName="Analis. Hope" authorityYear="1838" box="[464,595,1904,1930]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ halensis
<taxonomicNameLabel id="21995789B835FFD34BFFD41AFC883DB7" box="[713,800,1905,1931]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
/ ded. Sebastian Görn 2020” [red, printed].
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB835FFD549F1D4FFFDE33BB5" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="086136E0B835FFD549F1D4FFFDE33BB5" blockId="11.[151,1437,1508,2039]" lastBlockId="13.[151,1437,151,1761]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<emphasis id="3AAAEAF2B835FFD349F1D4FFFEEA3D92" bold="true" box="[199,322,1940,1966]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
This species has been confused with
<taxonomicName id="CFDE4D63B835FFD34BE8D4FEFCEB3D92" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[734,835,1940,1966]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="3AAAEAF2B835FFD34BE8D4FEFCEB3D92" box="[734,835,1940,1966]" italics="true" pageId="11" pageNumber="12">H. analis</emphasis>
</taxonomicName>
and
<taxonomicName id="CFDE4D63B835FFD34A4ED4FEFB8A3D92" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[888,1058,1940,1966]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B835FFD34A4ED4FEFB8A3D92" box="[888,1058,1940,1966]" italics="true" pageId="11" pageNumber="12">H. melanopyga</emphasis>
</taxonomicName>
, but it is immediately distinguishable from
<taxonomicName id="CFDE4D63B835FFD3483ED4D2FEC63DEE" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[264,366,1976,2002]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="3AAAEAF2B835FFD3483ED4D2FEC63DEE" box="[264,366,1976,2002]" italics="true" pageId="11" pageNumber="12">H. analis</emphasis>
</taxonomicName>
by the smooth, silkyshining pronotum, and from
<taxonomicName id="CFDE4D63B835FFD34AA4D4D2FB943DEE" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[914,1084,1976,2002]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B835FFD34AA4D4D2FB943DEE" box="[914,1084,1976,2002]" italics="true" pageId="11" pageNumber="12">H. melanopyga</emphasis>
</taxonomicName>
by the mediolateral and anteroapical elytral impressions (from dorsal view appearing dark and iridescent, respectively). From
<taxonomicName id="CFDE4D63B835FFD34DE3D4B6FA2D3DCA" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[1237,1413,2013,2038]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B835FFD34DE3D4B6FA2D3DCA" box="[1237,1413,2013,2038]" italics="true" pageId="11" pageNumber="12">H. nigrosericea</emphasis>
</taxonomicName>
it differs by the iridescent green dorsal colouration. There might be some confusion between bluish-green
<taxonomicName id="CFDE4D63B833FFD54C29D3FCFA343A8D" box="[1311,1436,151,177]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
<emphasis id="3AAAEAF2B833FFD54C29D3FCFA343A8D" box="[1311,1436,151,177]" italics="true" pageId="13" pageNumber="14">H. halensis</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B833FFD549A1D3D0FF583AE9" bold="true" box="[151,240,187,213]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="21995789B833FFD549A1D3D0FF583AE9" box="[151,240,187,213]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens and rather pale bluish-yellow
<taxonomicName id="CFDE4D63B833FFD54B8AD3D0FCC33AE8" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[700,875,187,212]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B833FFD54B8AD3D0FCC33AE8" box="[700,875,187,212]" italics="true" pageId="13" pageNumber="14">H. nigrosericea</emphasis>
</taxonomicName>
specimens, but females of
<taxonomicName id="CFDE4D63B833FFD54DAED3D0FABD3AE9" box="[1176,1301,187,213]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
<emphasis id="3AAAEAF2B833FFD54DAED3D0FABD3AE9" box="[1176,1301,187,213]" italics="true" pageId="13" pageNumber="14">H. halensis</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B833FFD54C2DD3D0FADC3AE9" bold="true" box="[1307,1396,187,213]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="21995789B833FFD54C2DD3D0FADC3AE9" box="[1307,1396,187,213]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are clearly characterized by a testaceous labrum, in contrast to the black to dark-brown female labrum of
<taxonomicName id="CFDE4D63B833FFD54C3DD3B4FF163B20" baseAuthorityName="W. Horn" baseAuthorityYear="1930" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B833FFD54C3DD3B4FF163B20" italics="true" pageId="13" pageNumber="14">H. nigrosericea</emphasis>
</taxonomicName>
. It is also similar to
<taxonomicName id="CFDE4D63B833FFD54897D268FD823B21" box="[417,554,259,285]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="abasileia" status="sp. nov.">
<emphasis id="3AAAEAF2B833FFD54897D268FD823B21" box="[417,554,259,285]" italics="true" pageId="13" pageNumber="14">H. abasileia</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B833FFD54B07D268FD223B21" bold="true" box="[561,650,259,285]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="21995789B833FFD54B07D268FD223B21" box="[561,650,259,285]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but black apical colouration expands over less than one quarter of the profemora (best seen from lateral view). The aedeagus dorsoapically has a considerably protruding crown-shaped sclerite (best seen in right lateral aspect), aedeagal apex a little less bent ventrally and aedeagal base more steeply sloping than in to
<taxonomicName id="CFDE4D63B833FFD54857D204FE433BB5" box="[353,491,367,393]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="abasileia" status="sp. nov.">
<emphasis id="3AAAEAF2B833FFD54857D204FE433BB5" box="[353,491,367,393]" italics="true" pageId="13" pageNumber="14">H. abasileia</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B833FFD548C7D204FDE33BB5" bold="true" box="[497,587,367,393]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="21995789B833FFD548C7D204FDE33BB5" box="[497,587,367,393]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<caption id="5CA16668B832FFD449A1D41AFB113DED" ID-DOI="http://doi.org/10.5281/zenodo.4578791" ID-Zenodo-Dep="4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,264,1905,1929]" targetBox="[151,1435,186,1856]" targetPageId="12">
<paragraph id="086136E0B832FFD449A1D41AFB113DED" blockId="12.[151,1437,1905,2002]" pageId="12" pageNumber="13">
<emphasis id="3AAAEAF2B832FFD449A1D41AFEFF3DB5" bold="true" box="[151,343,1905,1929]" pageId="12" pageNumber="13">FIGURES 2329.</emphasis>
<taxonomicName id="CFDE4D63B832FFD44856D41AFD9D3DB5" box="[352,565,1905,1929]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
<emphasis id="3AAAEAF2B832FFD44856D41AFD9D3DB5" box="[352,565,1905,1929]" italics="true" pageId="12" pageNumber="13">Heptodonta halensis</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B832FFD44B08D419FD3A3DB6" bold="true" box="[574,658,1906,1930]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="21995789B832FFD44B08D419FD3A3DB6" box="[574,658,1906,1930]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="40C4656BB833FFD549F1D2F8FB273B92" box="[199,1167,403,430]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="086136E0B833FFD549F1D2F8FB273B92" blockId="13.[151,1437,151,1761]" box="[199,1167,403,430]" pageId="13" pageNumber="14">
<emphasis id="3AAAEAF2B833FFD549F1D2F8FEE63B91" bold="true" box="[199,334,403,429]" pageId="13" pageNumber="14">Etymology.</emphasis>
This beautiful species is dedicated to my lovely hometown, Halle (Saale).
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB833FFD549F1D2DDFC7F3C81" pageId="13" pageNumber="14" type="description">
<paragraph id="086136E0B833FFD549F1D2DDFAB43BED" blockId="13.[151,1437,151,1761]" box="[199,1308,438,466]" pageId="13" pageNumber="14">
<emphasis id="3AAAEAF2B833FFD549F1D2DDFEF13BEC" bold="true" box="[199,345,438,464]" pageId="13" pageNumber="14">Description.</emphasis>
Body size: Length
<quantity id="CF269B05B833FFD54B02D2DCFD7E3BED" box="[564,726,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.235" metricValueMax="1.29" metricValueMin="1.18" pageId="13" pageNumber="14" unit="mm" value="12.35" valueMax="12.9" valueMin="11.8">11.812.9 mm</quantity>
(without labrum), width 3.44.0 mm (
<figureCitation id="90E52A65B833FFD54DB0D2DCFAB83BED" box="[1158,1296,439,465]" captionStart="FIGURES 2329" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,186,1856]" captionTargetId="figure-20@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 2329. Heptodonta halensis sp. nov.: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="13" pageNumber="14">Figs. 2324</figureCitation>
).
</paragraph>
<paragraph id="086136E0B833FFD549F1D2B0FBAC3999" blockId="13.[151,1437,151,1761]" pageId="13" pageNumber="14">
Head with dorsal surface greenish, with more or less expanded bronze colouration on frons and occiput, temples blue. Frons transversely to irregularly grooved, crumpled wrinkles in transition to vertex. Vertex median with parallel longitudinal striation that converge in transition to the orbital plates. Orbital plates with fine, convex to parallel converging striae, and with two setae on each side. Occiput with transverse grooves, anteromedian irregularly rugose. Genae glabrous and shallowly grooved, violet-blue with iridescent green anterior apex. Clypeus glabrous, iridescent blue-green with an anteromedian bronze spot and two anterior blue spots. Labrum testaceous, with four setae, five apical teeth, and one lateral tooth on each side, three median teeth acute, third apical teeth reduced, shape variable, male labrum rectangular and short (
<quantity id="CF269B05B833FFD54BB9D1BDFC9B38CD" box="[655,819,726,753]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="6.1" metricValueMin="5.2" pageId="13" pageNumber="14" unit="mm" value="0.565" valueMax="0.61" valueMin="0.52">0.520.61 mm</quantity>
long,
<quantity id="CF269B05B833FFD54A4CD1BDFBB538CD" box="[890,1053,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" metricValueMax="1.55" metricValueMin="1.43" pageId="13" pageNumber="14" unit="mm" value="1.49" valueMax="1.55" valueMin="1.43">1.431.55 mm</quantity>
wide,
<figureCitation id="90E52A65B833FFD54D5ED1BCFB1238CD" box="[1128,1210,726,753]" captionStart="FIGURES 2329" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,186,1856]" captionTargetId="figure-20@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 2329. Heptodonta halensis sp. nov.: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="13" pageNumber="14">Fig. 25</figureCitation>
), female labrum antero-triangular or three median teeth protruding (
<quantity id="CF269B05B833FFD54B8ED190FCF53929" box="[696,861,763,789]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="10.3" metricValueMin="9.0" pageId="13" pageNumber="14" unit="mm" value="0.965" valueMax="1.03" valueMin="0.9">0.901.03 mm</quantity>
long,
<quantity id="CF269B05B833FFD54A95D191FBEE3929" box="[931,1094,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" metricValueMax="1.65" metricValueMin="1.47" pageId="13" pageNumber="14" unit="mm" value="1.56" valueMax="1.65" valueMin="1.47">1.471.65 mm</quantity>
wide,
<figureCitation id="90E52A65B833FFD54DA6D190FB4B392A" box="[1168,1251,763,790]" captionStart="FIGURES 2329" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,186,1856]" captionTargetId="figure-20@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 2329. Heptodonta halensis sp. nov.: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="13" pageNumber="14">Fig. 26</figureCitation>
). Mandibles testaceous, teeth dark rufous-brown to black. Labial and maxillary palpi testaceous with dark-brown to black terminal palpomeres, in female pre-terminal maxillary palpomere dark-testaceous. Antennae slender, extending back over the first quarter of the elytra, metallic black, pedicel testaceous at base, scape with a single apical seta, antennomeres 34 with few scattered setae, antennomeres 511 finely and evenly pubescent.
</paragraph>
<paragraph id="086136E0B833FFD549F1D0C4FE3E3E41" blockId="13.[151,1437,151,1761]" pageId="13" pageNumber="14">
Thorax entirely glabrous. Pronotum iridescent green with more or less expanded bronze areas, blue lateral margin, median line moderately pronounced with blue margin, proportions variable, pronotal length (
<quantity id="CF269B05B833FFD54DDBD0B8FA3C39D1" box="[1261,1428,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="2.475" metricValueMax="2.66" metricValueMin="2.29" pageId="13" pageNumber="14" unit="mm" value="2.475" valueMax="2.66" valueMin="2.29">2.292.66 mm</quantity>
) and width (
<quantity id="CF269B05B833FFD5482FD09CFE693E2D" box="[281,449,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.39" metricValueMax="2.6" metricValueMin="2.18" pageId="13" pageNumber="14" unit="mm" value="2.39" valueMax="2.6" valueMin="2.18">2.182.60 mm</quantity>
) nearly identical, shallow transverse grooves, pronounced anterior and posterior sulci, anterior lobe wider than posterior, lateral margins of the median lobe moderately converging to the base. Episterna violet-blue, metepisternum and anterior proepisternum with bluish-green lustre. Sterna and epimera violet-blue with iridescent green lustre.
</paragraph>
<paragraph id="086136E0B833FFD549F1D7ECFED33F69" blockId="13.[151,1437,151,1761]" pageId="13" pageNumber="14">
Elytra elongate, length
<quantity id="CF269B05B833FFD548FDD7ECFDFC3E9D" box="[459,596,1159,1185]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.1" metricValueMin="7.2" pageId="13" pageNumber="14" unit="mm" value="7.65" valueMax="8.1" valueMin="7.2">7.28.1 mm</quantity>
, slightly dilated laterally to almost parallel, maximal width posterior, green with yellow-bronze lustre, lateral margin violet, apex blue to bluish-green; juxtahumeral, discal, mediolateral, anteroapical, and apical impressions considerably pronounced, impression on posterior declivity moderate or lacking, posterior gibbosity significant, basodiscal convexity moderate, blue punctures slightly decreasing in size from base to apex, punctures fused to transverse lines, converging to apex on anteroapical impression; irregularly punctured basally and apically.
</paragraph>
<paragraph id="086136E0B833FFD549F1D634FF673FD9" blockId="13.[151,1437,151,1761]" pageId="13" pageNumber="14">Coxae dark-testaceous with iridescent blue-green lustre, pro- and mesocoxae anteriorly, metacoxae laterally densely covered with long white setae, mesocoxae posterior with few white setae. Trochanters and femora testaceous. Femoral distal apices, tibiae, and tarsi metallic-black. Claws black proximally and rufous-testaceous distally.</paragraph>
<paragraph id="086136E0B833FFD549F1D684FC093C11" blockId="13.[151,1437,151,1761]" pageId="13" pageNumber="14">
Abdominal sternites glabrous (apart from few long setae at posterior margins), dark-brown with violet-blue reflections, anal sternite with dark-testaceous apical margin (
<figureCitation id="90E52A65B833FFD54A76D578FC3D3C11" box="[832,917,1555,1581]" captionStart="FIGURES 2329" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,186,1856]" captionTargetId="figure-20@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 2329. Heptodonta halensis sp. nov.: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="13" pageNumber="14">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="086136E0B833FFD549F1D55CFC7F3C81" blockId="13.[151,1437,151,1761]" pageId="13" pageNumber="14">
Aedeagus arched, length
<quantity id="CF269B05B833FFD548D5D55CFD213C6D" box="[483,649,1591,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" metricValueMax="2.66" metricValueMin="2.62" pageId="13" pageNumber="14" unit="mm" value="2.64" valueMax="2.66" valueMin="2.62">2.622.66 mm</quantity>
, dorsal and medioventral regions convex, apex slightly bent ventrally, with round tip and convex dorsal aspect, aedeagal base comparatively steeply sloping. Inner sac in left lateral aspect medioventral with elongate vertical sclerite and medially with short horizontal sclerite; in right lateral aspect with a spiny dorsoapical sclerite protruding from the apical orifice (
<figureCitation id="90E52A65B833FFD54A77D5C8FC633C81" box="[833,971,1699,1725]" captionStart="FIGURES 2329" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,186,1856]" captionTargetId="figure-20@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 2329. Heptodonta halensis sp. nov.: 23, 25, 2729—♂, holotype, Philippines, Leyte, Mt. Balocaue (JWCW): 23—habitus, 25—head, 27—lateral body, 28—aedeagus in left lateral aspect, 29—aedeagus in right lateral aspect; 24, 26—♀, paratype, Philippines, Leyte (JWCW): 24—habitus, 26—head. Scales: 2327 = 1 mm, 2829 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578791" httpUri="https://zenodo.org/record/4578791/files/figure.png" pageId="13" pageNumber="14">Figs. 2829</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB833FFD549F1D5ADFBD13CDD" box="[199,1145,1734,1761]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="086136E0B833FFD549F1D5ADFBD13CDD" blockId="13.[151,1437,151,1761]" box="[199,1145,1734,1761]" pageId="13" pageNumber="14">
<emphasis id="3AAAEAF2B833FFD549F1D5ADFEC93CDC" bold="true" box="[199,353,1734,1760]" pageId="13" pageNumber="14">Distribution.</emphasis>
Northern and eastern
<emphasis id="3AAAEAF2B833FFD54B6FD5ADFCA43CDC" bold="true" box="[601,780,1734,1760]" pageId="13" pageNumber="14">
<collectingCountry id="70C97670B833FFD54B6FD5ADFCA43CDC" box="[601,780,1734,1760]" name="Philippines" pageId="13" pageNumber="14">PHILIPPINES</collectingCountry>
</emphasis>
(
<collectingRegion id="CA1AF802B833FFD54A2DD5ACFCF73CDD" box="[795,863,1735,1761]" country="Philippines" name="Southern Leyte" pageId="13" pageNumber="14">Leyte</collectingRegion>
, Luzon, Panaon,
<collectingRegion id="CA1AF802B833FFD54D16D5ACFBC53CDD" box="[1056,1133,1735,1761]" country="Philippines" name="Samar" pageId="13" pageNumber="14">Samar</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>