treatments-xml/data/80/77/87/807787F6B82CFFCB49A1D3FCFCD639D1.xml
2024-06-21 12:41:55 +02:00

478 lines
57 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="54ABBAE7B699AADD7C740E4B790EE44C" ID-CLB-Dataset="7896" ID-DOI="10.11646/zootaxa.4875.1.1" ID-GBIF-Dataset="a75d6e42-221a-4df9-9a7d-5f33dbff6a56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4578779" ID-ZooBank="278200CE-E16F-45B4-9A89-60C2052415C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610024385246" checkinUser="plazi" docAuthor="Görn, Sebastian" docDate="2020" docId="807787F6B82CFFCB49A1D3FCFCD639D1" docLanguage="en" docName="zootaxa.4875.1.1.pdf" docOrigin="Zootaxa 4875 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heptodonta nigrosericea, stat. nov." docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="7C4EFF8EB83EFFD84936D36BFFA83A3C" masterDocTitle="Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="19" updateTime="1698923305811" updateUser="ExternalLinkService">
<mods:mods id="8B27164320B44C93A6FCC88FCE301266" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B6C51538E021C90E85A8FBD0E4197E64">
<mods:title id="BCA612534874FD7ED0C7748DF031C698">Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)</mods:title>
</mods:titleInfo>
<mods:name id="D84B0DC27B2E2BBE67104970CE4B7F55" type="personal">
<mods:role id="099DD169C82B7E7B5A2D19928ED2891F">
<mods:roleTerm id="9314FA9C086CBD2FB8C78F07BC825D84">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B94575AFAE9C89238386D9A00E8A235">Görn, Sebastian</mods:namePart>
</mods:name>
<mods:typeOfResource id="2EB73AF3443F384C3B5DCFDCBC1FD66D">text</mods:typeOfResource>
<mods:relatedItem id="259D2C6F334FFAF5371596DF798CBAC9" type="host">
<mods:titleInfo id="F989972CD63229BE20727B44E88D2804">
<mods:title id="E0CAF47C3C0F595263C0A0872553654D">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5DCF3B37F9BB91BBDEB80596AB69B326">
<mods:date id="575A40A02833BF687BABB0304097AC00">2020</mods:date>
<mods:detail id="E8271BF81D50DF9D566EFA92842D5005" type="pubDate">
<mods:number id="33988CA8104462E0C74DFD6993D50DC2">2020-11-06</mods:number>
</mods:detail>
<mods:detail id="F41ED8E1B4D1403C4B53AA236481DDA0" type="volume">
<mods:number id="877599139938FC6504B0CC01031E9266">4875</mods:number>
</mods:detail>
<mods:detail id="F4016AA6BC4C780E7907D08AEE0ED532" type="issue">
<mods:number id="C1C892BF6240B10B46EBEA491D9EDF91">1</mods:number>
</mods:detail>
<mods:extent id="88E0301AA807C209C6E1DAEA1EF4EAB9" unit="page">
<mods:start id="C652B931B66254F0EF96742EDA357710">1</mods:start>
<mods:end id="DFC22A32BEF16C48603DA93939CC036C">62</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3D6874901EECE025272DDC3FEE2587C3">journal article</mods:classification>
<mods:identifier id="091278251C532B7AB63BEE95F63E0B43" type="CLB-Dataset">7896</mods:identifier>
<mods:identifier id="705B0C33B3B62EDB796C7B956AD6F9FA" type="DOI">10.11646/zootaxa.4875.1.1</mods:identifier>
<mods:identifier id="3F3A64B6D5E85E0AAFF190FE47307280" type="GBIF-Dataset">a75d6e42-221a-4df9-9a7d-5f33dbff6a56</mods:identifier>
<mods:identifier id="47916EDE463E65D37A98299B8FA6A062" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5178959F0337B8178E12A97243825C34" type="Zenodo-Dep">4578779</mods:identifier>
<mods:identifier id="3F4FBDA3C49394129AC59CBEFCDAB676" type="ZooBank">278200CE-E16F-45B4-9A89-60C2052415C7</mods:identifier>
</mods:mods>
<treatment id="807787F6B82CFFCB49A1D3FCFCD639D1" ID-DOI="http://doi.org/10.5281/zenodo.4579789" ID-GBIF-Taxon="179988679" ID-Zenodo-Dep="4579789" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1" httpUri="http://treatment.plazi.org/id/807787F6B82CFFCB49A1D3FCFCD639D1" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="40C4656BB82CFFCA49A1D3FCFCA13A8D" box="[151,777,151,178]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="086136E0B82CFFCA49A1D3FCFCA13A8D" blockId="18.[151,777,151,214]" box="[151,777,151,178]" pageId="18" pageNumber="19">
<heading id="5329818CB82CFFCA49A1D3FCFCA13A8D" bold="true" box="[151,777,151,178]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="3AAAEAF2B82CFFCA49A1D3FCFCA13A8D" bold="true" box="[151,777,151,178]" pageId="18" pageNumber="19">
<taxonomicName id="CFDE4D63B82CFFCA49A1D3FCFD3A3A8D" ID-CoL="8MGTB" authority="(W. Horn, 1930)" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[151,658,151,178]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nigrosericea" status="stat. nov.">
<emphasis id="3AAAEAF2B82CFFCA49A1D3FCFE6A3A8D" bold="true" box="[151,450,151,178]" italics="true" pageId="18" pageNumber="19">Heptodonta nigrosericea</emphasis>
(W.
<bibRefCitation id="6C4F4B11B82CFFCA48CBD3FCFD223A8D" author="Horn, W." box="[509,650,151,177]" pageId="18" pageNumber="19" pagination="1 - 9" refId="ref39412" refString="Horn, W. (1930) Beitrage zur Kenntnis neuer und alter Cicindelinen des Indo-papuanischen Faunen-Gebietes. Wiener Entomologische Zeitung, 47 (1), 1 - 9." type="journal article" year="1930">Horn, 1930</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="21995789B82CFFCA4BACD3FCFCA13A8D" box="[666,777,151,177]" pageId="18" pageNumber="19">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82CFFCA49A1D3D0FEB43AEA" box="[151,284,187,214]" pageId="18" pageNumber="19" type="description">
<paragraph id="086136E0B82CFFCA49A1D3D0FEB43AEA" blockId="18.[151,777,151,214]" box="[151,284,187,214]" pageId="18" pageNumber="19">
<figureCitation id="90E52A65B82CFFCA49A1D3D0FEB43AEA" box="[151,284,187,214]" captionStart-0="FIGURES 3743" captionStart-1="FIGURES 4447" captionStartId-0="16.[151,264,1931,1955]" captionStartId-1="17.[151,264,1908,1932]" captionTargetBox-0="[151,1435,184,1873]" captionTargetBox-1="[205,1333,189,1867]" captionTargetId-0="figure-20@16.[151,1435,181,1908]" captionTargetId-1="figure-15@17.[197,1389,181,1887]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 3743. Heptodonta nigrosericea (W. Horn, 1930): 37, 39, 4143—♂, Philippines, Mindanao, Mt. Apo (JWCW): 37—habitus, 39—head, 41—lateral body, 42—aedeagus in left lateral aspect, 43—aedeagus in right lateral aspect; 38, 40—♀, Philippines, Mindanao, Mt. Kintanlad (JWCW): 38—habitus, 40—head. Scales: 3741 = 1 mm, 4243 = 0.5 mm." captionText-1="FIGURES 4447. 4447—habitus: 4445—Heptodonta nigrosericea (W. Horn, 1930): 44—♀, lectotype, Philippines, [Min- danao], Columbogan (SDEI); 45—♂, holotype of syn. H. lumawigi Wiesner, 1980, Philippines, Canuawua (JWCW). 46—H. melanopyga (Schaum, 1862), ♂, lectotype, Philippines, Luzon (MNB). 47—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). Scales: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4578797" figureDoi-1="http://doi.org/10.5281/zenodo.4578799" httpUri-0="https://zenodo.org/record/4578797/files/figure.png" httpUri-1="https://zenodo.org/record/4578799/files/figure.png" pageId="18" pageNumber="19">Figs. 3745</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82CFFCA49A1D391FCA23B6C" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="086136E0B82CFFCA49A1D391FC9B3B2E" blockId="18.[151,819,250,336]" box="[151,819,250,274]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="28CE11CEB82CFFCA49A1D391FC9B3B2E" box="[151,819,250,274]" pageId="18" pageNumber="19">
<treatmentCitation id="897F10F1B82CFFCA49A1D391FC873B2E" author="Horn, W." box="[151,815,250,274]" page="9" pageId="18" pageNumber="19" year="1930">
<taxonomicName id="CFDE4D63B82CFFCA49A1D391FC873B2E" authority="W. Horn, 1930: 9" authorityName="W. Horn" authorityPageNumber="9" authorityYear="1930" box="[151,815,250,274]" class="Insecta" family="Carabidae" genus="Odontochila" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="analis" subGenus="Heptodonta" subSpecies="nigro-sericea">
<emphasis id="3AAAEAF2B82CFFCA49A1D391FDDF3B2E" box="[151,631,250,274]" italics="true" pageId="18" pageNumber="19">Odontochila (Heptodonta) analis nigro-sericea</emphasis>
W.
<bibRefCitation id="6C4F4B11B82CFFCA4BA9D390FC873B2E" author="Horn, W." box="[671,815,250,274]" pageId="18" pageNumber="19" pagination="1 - 9" refId="ref39412" refString="Horn, W. (1930) Beitrage zur Kenntnis neuer und alter Cicindelinen des Indo-papuanischen Faunen-Gebietes. Wiener Entomologische Zeitung, 47 (1), 1 - 9." type="journal article" year="1930">Horn, 1930: 9</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="086136E0B82CFFCA49A1D272FD0C3B0D" blockId="18.[151,819,250,336]" box="[151,676,281,305]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="28CE11CEB82CFFCA49A1D272FD0C3B0D" box="[151,676,281,305]" pageId="18" pageNumber="19">
<taxonomicName id="CFDE4D63B82CFFCA49A1D272FE703B0D" ID-CoL="8MGTB" baseAuthorityName="W.Horn" baseAuthorityYear="1930" box="[151,472,281,305]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="analis" subSpecies="nigrosericea">
<emphasis id="3AAAEAF2B82CFFCA49A1D272FE703B0D" box="[151,472,281,305]" italics="true" pageId="18" pageNumber="19">Heptodonta analis nigrosericea</emphasis>
</taxonomicName>
.
<treatmentCitation id="897F10F1B82CFFCA48D3D272FD083B0D" author="Wiesner, J." box="[485,672,281,305]" page="69" pageId="18" pageNumber="19" year="1992">
<bibRefCitation id="6C4F4B11B82CFFCA48D3D272FD083B0D" author="Wiesner, J." box="[485,672,281,305]" pageId="18" pageNumber="19" refId="ref41340" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt, Checklist of the Tiger Beetles of the World, 27. Beitrag zur Kenntnis der Cicindelidae. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner, 1992: 69</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="086136E0B82CFFCA49A1D253FCA23B6C" blockId="18.[151,819,250,336]" box="[151,778,312,336]" pageId="18" pageNumber="19">
Syn.
<taxonomicName id="CFDE4D63B82CFFCA49FDD253FD0A3B6C" ID-CoL="8LGD9" authority="Wiesner 1980: 121-123" authorityName="Wiesner" authorityPageNumber="121 - 123" authorityYear="1980" box="[203,674,312,336]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lumawigi">
<emphasis id="3AAAEAF2B82CFFCA49FDD253FE0F3B6C" box="[203,423,312,336]" italics="true" pageId="18" pageNumber="19">Heptodonta lumawigi</emphasis>
<bibRefCitation id="6C4F4B11B82CFFCA489BD253FD0A3B6C" author="Wiesner, J." box="[429,674,312,336]" pageId="18" pageNumber="19" pagination="119 - 127" refId="ref41195" refString="Wiesner, J. (1980) Beitrage zur Kenntnis der philippinischen Cicindelidae. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 119 - 127." type="journal article" year="1980">Wiesner 1980: 121123</bibRefCitation>
</taxonomicName>
,
<emphasis id="3AAAEAF2B82CFFCA4B9AD253FCA23B6C" bold="true" box="[684,778,312,336]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="21995789B82CFFCA4B9AD253FCA23B6C" box="[684,778,312,336]" pageId="18" pageNumber="19">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82CFFCA49A1D213FC343943" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="086136E0B82CFFCA49A1D213FD453BAE" blockId="18.[151,1437,375,2022]" box="[151,749,375,402]" pageId="18" pageNumber="19">
<materialsCitation id="B8B63CBDB82CFFCA49A1D213FD413BAE" ID-GBIF-Occurrence="3045076466" box="[151,745,375,402]" country="Philippines" location="Columbogan" pageId="18" pageNumber="19" specimenCount="1" typeStatus="holotype">
<emphasis id="3AAAEAF2B82CFFCA49A1D213FE9E3BAD" bold="true" box="[151,310,375,402]" pageId="18" pageNumber="19">
<typeStatus id="D7658842B82CFFCA49A1D213FF7A3BAE" box="[151,210,376,402]" pageId="18" pageNumber="19">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="70C97670B82CFFCA480AD213FE173BAE" box="[316,447,376,402]" name="Philippines" pageId="18" pageNumber="19">Philippines</collectingCountry>
:
<location id="0D01603BB82CFFCA48FFD213FDF23BAE" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82CFFCA48FFD213FDF23BAE" box="[457,602,376,402]" country="Philippines" name="Columbogan" pageId="18" pageNumber="19">Columbogan</location>
[Mindanao]
</materialsCitation>
.
</paragraph>
<paragraph id="086136E0B82CFFCA49F1D2F0FB1F38B4" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">
<emphasis id="3AAAEAF2B82CFFCA49F1D2F0FD673B89" bold="true" box="[199,719,411,437]" pageId="18" pageNumber="19">
Type material of
<taxonomicName id="CFDE4D63B82CFFCA48A0D2F0FDE23B89" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[406,586,411,437]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B82CFFCA48A0D2F0FDE23B89" bold="true" box="[406,586,411,437]" italics="true" pageId="18" pageNumber="19">H. nigrosericea</emphasis>
</taxonomicName>
.
<typeStatus id="D7658842B82CFFCA4B6FD2F0FD673B89" box="[601,719,411,437]" pageId="18" pageNumber="19" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated here):
<specimenCount id="1ED8FD69B82CFFCA4A9CD2F0FC173B89" box="[938,959,411,437]" pageId="18" pageNumber="19" type="female"></specimenCount>
in SDEI, labelled: “Columbogan / Philip- pines / VII-VIII-1917 / G. Haslam” [handwritten] // “Carn. Mus. / Acc. 5950” [printed/handwritten] // “Carnegie Mus.” [handwritten] // “Type / W. Horn” [printed] // “
<typeStatus id="D7658842B82CFFCA4BDCD28AFCFF3BC7" box="[746,855,481,507]" pageId="18" pageNumber="19" type="syntype">Syntypus</typeStatus>
” [red, printed] // “Coll. DEI / Eberswalde” [printed] // “
<taxonomicName id="CFDE4D63B82CFFCA4981D16FFE953822" authorityName="Analis. Hope" authorityYear="1838" box="[183,317,516,542]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ analis / nigrosericea / Horn” [handwritten] // “Coll W. Horn / DEI Eberswalde” [printed] // “ssp. / nigro-seri- / cea / m.” [yellow with black frame, handwritten] // “SDEI
<taxonomicName id="CFDE4D63B82CFFCA4AF6D143FB93387E" authorityName="Linnaeus" authorityYear="1758" box="[960,1083,552,578]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
/ # 301568” [printed] // “LEC- TOTYPE /
<taxonomicName id="CFDE4D63B82CFFCA482CD120FDEA385A" authority="" authorityName="Eugenia. W. Horn" authorityYear="1926" box="[282,578,587,614]" class="Insecta" family="Carabidae" genus="Odontochila" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subGenus" subGenus="Heptodonta">Odontochila (Heptodonta)</taxonomicName>
/ analis nigro-sericea / W.
<bibRefCitation id="6C4F4B11B82CFFCA4A43D120FC5E3859" author="Horn, W." box="[885,1014,587,613]" pageId="18" pageNumber="19" pagination="1 - 9" refId="ref39412" refString="Horn, W. (1930) Beitrage zur Kenntnis neuer und alter Cicindelinen des Indo-papuanischen Faunen-Gebietes. Wiener Entomologische Zeitung, 47 (1), 1 - 9." type="journal article" year="1930">Horn, 1930</bibRefCitation>
/ design. Sebastian Görn 2020” [red, printed] // “
<taxonomicName id="CFDE4D63B82CFFCA4821D105FE3538B4" authorityName="Analis. Hope" authorityYear="1838" box="[279,413,622,648]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ nigrosericea / (W.
<bibRefCitation id="6C4F4B11B82CFFCA4BB6D104FCA938B4" author="Horn, W." box="[640,769,622,649]" pageId="18" pageNumber="19" pagination="1 - 9" refId="ref39412" refString="Horn, W. (1930) Beitrage zur Kenntnis neuer und alter Cicindelinen des Indo-papuanischen Faunen-Gebietes. Wiener Entomologische Zeitung, 47 (1), 1 - 9." type="journal article" year="1930">Horn, 1930</bibRefCitation>
) / det. Sebastian Görn 2020” [printed].
</paragraph>
<paragraph id="086136E0B82CFFCA49F1D1FAFC343943" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">
<emphasis id="3AAAEAF2B82CFFCA49F1D1FAFD353897" bold="true" box="[199,669,657,684]" pageId="18" pageNumber="19">
Type material of
<taxonomicName id="CFDE4D63B82CFFCA48AFD1F9FD823890" authorityName="Wiesner" authorityYear="1980" box="[409,554,657,684]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lumawigi">
<emphasis id="3AAAEAF2B82CFFCA48AFD1F9FD823890" bold="true" box="[409,554,657,684]" italics="true" pageId="18" pageNumber="19">H. lumawigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21995789B82CFFCA4B02D1FAFD353897" box="[564,669,657,683]" pageId="18" pageNumber="19">syn. nov.</taxonomicNameLabel>
</emphasis>
<materialsCitation id="B8B63CBDB82CFFCA4B91D1FAFF5B38CE" ID-GBIF-Occurrence="3045076583" collectorName="R. Lumawig" country="Philippines" determinerName="J. Wiesner" location="Iloilo" pageId="18" pageNumber="19" specimenCount="1" specimenCount-male="1" stateProvince="Iloilo" typeStatus="holotype">
<emphasis id="3AAAEAF2B82CFFCA4B91D1FAFCBC3897" bold="true" box="[679,788,657,683]" pageId="18" pageNumber="19">
<typeStatus id="D7658842B82CFFCA4B91D1FAFCBC3897" box="[679,788,657,683]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="1ED8FD69B82CFFCA4A12D1F9FC913890" box="[804,825,658,684]" pageId="18" pageNumber="19" type="male"></specimenCount>
in JWCW, labelled: “
<collectingCountry id="70C97670B82CFFCA4D02D1FAFB133897" box="[1076,1211,657,683]" name="Philippines" pageId="18" pageNumber="19">Philippinen</collectingCountry>
,
<collectingRegion id="CA1AF802B82CFFCA4DFED1FAFAAD3897" box="[1224,1285,657,683]" country="Philippines" name="Iloilo" pageId="18" pageNumber="19">Iloilo</collectingRegion>
/ Canuawua, 5.78 / leg.
<collectorName id="A52B5336B82CFFCA4825D1DEFE0B38F2" box="[275,419,692,719]" pageId="18" pageNumber="19">R. Lumawig</collectorName>
” [yellow margin, handwritten] // “
<taxonomicName id="CFDE4D63B82CFFCA4A1DD1DFFC1938F2" authorityName="Analis. Hope" authorityYear="1838" box="[811,945,692,718]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ lumawigi m. / det.
<determinerName id="FDBDDA24B82CFFCA4D95D1DEFAB138F2" box="[1187,1305,692,719]" pageId="18" pageNumber="19">J. Wiesner</determinerName>
78” [hand- written]
</materialsCitation>
.
<emphasis id="3AAAEAF2B82CFFCA4836D1BCFECD38CD" bold="true" box="[256,357,727,753]" pageId="18" pageNumber="19">
<typeStatus id="D7658842B82CFFCA4836D1BCFECD38CD" box="[256,357,727,753]" pageId="18" pageNumber="19" type="allotype">Allotype</typeStatus>
</emphasis>
:
<specimenCount id="1ED8FD69B82CFFCA4840D1B3FE0938CE" box="[374,417,728,754]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
in JWCW with same labels as
<typeStatus id="D7658842B82CFFCA4A3DD1B3FCD938CE" box="[779,881,728,754]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
.
<emphasis id="3AAAEAF2B82CFFCA4A4BD1B3FC5D38CE" bold="true" box="[893,1013,728,754]" pageId="18" pageNumber="19">
<typeStatus id="D7658842B82CFFCA4A4BD1B3FC5D38CE" box="[893,1013,728,754]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="1ED8FD69B82CFFCA4D30D1B3FB9A38CE" box="[1030,1074,728,754]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
<specimenCount id="1ED8FD69B82CFFCA4D0CD1B3FBCE38CE" box="[1082,1126,728,754]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
in JWCW,
<specimenCount id="1ED8FD69B82CFFCA4DD8D1B3FAB238CE" box="[1262,1306,728,754]" pageId="18" pageNumber="19" type="male">1 ♂</specimenCount>
in BMNH, with same labels as
<typeStatus id="D7658842B82CFFCA484DD190FE493929" box="[379,481,763,789]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
.
<materialsCitation id="B8B63CBDB82CFFCA48DBD190FE2B3967" ID-GBIF-Occurrence="3045076399" country="Philippines" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Iloilo" typeStatus="holotype">
<specimenCount id="1ED8FD69B82CFFCA48DBD190FDB03929" box="[493,536,763,789]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
in SDEI with same labels as
<typeStatus id="D7658842B82CFFCA4A5ED190FC623929" box="[872,970,763,789]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
plus: “Coll. DEI / Eberswalde” [printed] // “lumawigi / Wiesn.” [yellow with black frame, handwritten] // “SDEI
<taxonomicName id="CFDE4D63B82CFFCA4AF2D075FB973904" authorityName="Linnaeus" authorityYear="1758" box="[964,1087,798,824]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
/ # 301574” [printed]. All type specimens labelled: “
</materialsCitation>
<materialsCitation id="B8B63CBDB82CFFCA48B5D02AFC303943" ID-GBIF-Occurrence="3045076449" country="Philippines" determinerName="Sebastian Gorn" pageId="18" pageNumber="19" specimenCount="3" stateProvince="Iloilo" typeStatus="holotype">
<typeStatus id="D7658842B82CFFCA48B5D02AFD853967" box="[387,557,833,859]" pageId="18" pageNumber="19" type="holotype">HOLOTYPUS</typeStatus>
(
<typeStatus id="D7658842B82CFFCA4B0DD02AFD483967" box="[571,736,833,859]" pageId="18" pageNumber="19" type="allotype">ALLOTYPUS</typeStatus>
or
<typeStatus id="D7658842B82CFFCA4A33D02AFC0B3967" box="[773,931,833,859]" pageId="18" pageNumber="19" type="paratype">PARATYPUS</typeStatus>
respectively)” [red, printed] // “
<taxonomicName id="CFDE4D63B82CFFCA4C3ED02AFA263967" authorityName="Analis. Hope" authorityYear="1838" box="[1288,1422,833,859]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ nigrosericea / (W.
<bibRefCitation id="6C4F4B11B82CFFCA4853D00EFE4D3943" author="Horn, W." box="[357,485,869,895]" pageId="18" pageNumber="19" pagination="1 - 9" refId="ref39412" refString="Horn, W. (1930) Beitrage zur Kenntnis neuer und alter Cicindelinen des Indo-papuanischen Faunen-Gebietes. Wiener Entomologische Zeitung, 47 (1), 1 - 9." type="journal article" year="1930">Horn, 1930</bibRefCitation>
) / det.
<determinerName id="FDBDDA24B82CFFCA4B02D00FFD483943" box="[564,736,868,895]" pageId="18" pageNumber="19">Sebastian Görn</determinerName>
2020” [printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82CFFCA49F1D0ECFC3F3E37" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="086136E0B82CFFCA49F1D0ECFC3F3E37" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">
<emphasis id="3AAAEAF2B82CFFCA49F1D0ECFEEA399D" bold="true" box="[199,322,903,929]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
Distinguished by the combination of mediolateral and anteroapical elytral impressions, and blackish colouration with blue-yellow lustre. Resembling
<taxonomicName id="CFDE4D63B82CFFCA4BF6D0C7FCE239F9" box="[704,842,939,965]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="abasileia" status="sp. nov.">
<emphasis id="3AAAEAF2B82CFFCA4BF6D0C7FCE239F9" box="[704,842,939,965]" italics="true" pageId="18" pageNumber="19">H. abasileia</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B82CFFCA4A64D0C0FC0439F9" bold="true" box="[850,940,939,965]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="21995789B82CFFCA4A64D0C0FC0439F9" box="[850,940,939,965]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and particularly
<taxonomicName id="CFDE4D63B82CFFCA4D5BD0C7FB4439F9" box="[1133,1260,939,965]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
<emphasis id="3AAAEAF2B82CFFCA4D5BD0C7FB4439F9" box="[1133,1260,939,965]" italics="true" pageId="18" pageNumber="19">H. halensis</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B82CFFCA4DC2D0C0FAE639F9" bold="true" box="[1268,1358,939,965]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="21995789B82CFFCA4DC2D0C0FAE639F9" box="[1268,1358,939,965]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
due to its impression pattern and silky shining surface, but clearly distinguished from both by dark-brown female labrum, and the usually much darker colouration lacking any greenish touch.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82CFFCB49F1D77FFA393BED" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="086136E0B82CFFCA49F1D77FFA243E13" blockId="18.[151,1437,375,2022]" box="[199,1420,1044,1071]" pageId="18" pageNumber="19">
<emphasis id="3AAAEAF2B82CFFCA49F1D77FFEDD3E12" bold="true" box="[199,373,1044,1070]" pageId="18" pageNumber="19">Redescription.</emphasis>
Body size: Length
<quantity id="CF269B05B82CFFCA4B66D77EFD5A3E13" box="[592,754,1045,1071]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.49" metricValueMin="1.17" pageId="18" pageNumber="19" unit="mm" value="13.3" valueMax="14.9" valueMin="11.7">11.714.9 mm</quantity>
(without labrum), width
<quantity id="CF269B05B82CFFCA4D3AD77EFB3B3E13" box="[1036,1171,1045,1071]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.4" metricValueMin="3.4" pageId="18" pageNumber="19" unit="mm" value="3.9" valueMax="4.4" valueMin="3.4">3.44.4 mm</quantity>
(
<figureCitation id="90E52A65B82CFFCA4D94D77FFA813E13" box="[1186,1321,1044,1071]" captionStart="FIGURES 3743" captionStartId="16.[151,264,1931,1955]" captionTargetBox="[151,1435,184,1873]" captionTargetId="figure-20@16.[151,1435,181,1908]" captionTargetPageId="16" captionText="FIGURES 3743. Heptodonta nigrosericea (W. Horn, 1930): 37, 39, 4143—♂, Philippines, Mindanao, Mt. Apo (JWCW): 37—habitus, 39—head, 41—lateral body, 42—aedeagus in left lateral aspect, 43—aedeagus in right lateral aspect; 38, 40—♀, Philippines, Mindanao, Mt. Kintanlad (JWCW): 38—habitus, 40—head. Scales: 3741 = 1 mm, 4243 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578797" httpUri="https://zenodo.org/record/4578797/files/figure.png" pageId="18" pageNumber="19">Figs. 3738</figureCitation>
,
<figureCitation id="90E52A65B82CFFCA4C03D77FFA283E13" box="[1333,1408,1044,1071]" captionStart="FIGURES 4447" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[205,1333,189,1867]" captionTargetId="figure-15@17.[197,1389,181,1887]" captionTargetPageId="17" captionText="FIGURES 4447. 4447—habitus: 4445—Heptodonta nigrosericea (W. Horn, 1930): 44—♀, lectotype, Philippines, [Min- danao], Columbogan (SDEI); 45—♂, holotype of syn. H. lumawigi Wiesner, 1980, Philippines, Canuawua (JWCW). 46—H. melanopyga (Schaum, 1862), ♂, lectotype, Philippines, Luzon (MNB). 47—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578799" httpUri="https://zenodo.org/record/4578799/files/figure.png" pageId="18" pageNumber="19">4445</figureCitation>
).
</paragraph>
<paragraph id="086136E0B82CFFCA49F1D753FC7B3FE9" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">
Dorsal surface of head blackish with a slightly blue, yellow, or bronze lustre. Frons transversely grooved. Vertex with concave striation, straight converging in transition to orbital plates. Orbital plates with fine convex striation, and with two setae on each side. Occiput with transverse grooves and a smooth anteromedian spot. Genae violetblue, glabrous and shallowly grooved. Clypeus glabrous, blue-green with some median bronze. Labrum with four setae, five apical teeth, and one lateral tooth on each side, three median teeth acute, third apical teeth often reduced, shape variable, male labrum rectangular and short (
<quantity id="CF269B05B82CFFCA4BEDD78CFC293F3E" box="[731,897,1255,1282]" metricMagnitude="-4" metricUnit="m" metricValue="7.05" metricValueMax="7.8" metricValueMin="6.3" pageId="18" pageNumber="19" unit="mm" value="0.705" valueMax="0.78" valueMin="0.63">0.630.78 mm</quantity>
long,
<quantity id="CF269B05B82CFFCA4AFFD783FBC53F3E" box="[969,1133,1256,1282]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.8" metricValueMin="1.48" pageId="18" pageNumber="19" unit="mm" value="1.64" valueMax="1.8" valueMin="1.48">1.481.80 mm</quantity>
wide,
<figureCitation id="90E52A65B82CFFCA4D8CD783FAA53F3E" box="[1210,1293,1256,1282]" captionStart="FIGURES 3743" captionStartId="16.[151,264,1931,1955]" captionTargetBox="[151,1435,184,1873]" captionTargetId="figure-20@16.[151,1435,181,1908]" captionTargetPageId="16" captionText="FIGURES 3743. Heptodonta nigrosericea (W. Horn, 1930): 37, 39, 4143—♂, Philippines, Mindanao, Mt. Apo (JWCW): 37—habitus, 39—head, 41—lateral body, 42—aedeagus in left lateral aspect, 43—aedeagus in right lateral aspect; 38, 40—♀, Philippines, Mindanao, Mt. Kintanlad (JWCW): 38—habitus, 40—head. Scales: 3741 = 1 mm, 4243 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578797" httpUri="https://zenodo.org/record/4578797/files/figure.png" pageId="18" pageNumber="19">Fig. 39</figureCitation>
), testaceous, often with a dark margin, female labrum antero-triangular or three median teeth protruding (
<quantity id="CF269B05B82CFFCA4D83D660FAF33F19" box="[1205,1371,1291,1317]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" metricValueMax="1.13" metricValueMin="0.93" pageId="18" pageNumber="19" unit="mm" value="1.03" valueMax="1.13" valueMin="0.93">0.931.13 mm</quantity>
long,
<quantity id="CF269B05B82CFFCA49A1D645FE953F75" box="[151,317,1326,1353]" metricMagnitude="-3" metricUnit="m" metricValue="1.665" metricValueMax="1.88" metricValueMin="1.45" pageId="18" pageNumber="19" unit="mm" value="1.665" valueMax="1.88" valueMin="1.45">1.451.88 mm</quantity>
wide,
<figureCitation id="90E52A65B82CFFCA48B9D645FE4D3F74" box="[399,485,1326,1352]" captionStart="FIGURES 3743" captionStartId="16.[151,264,1931,1955]" captionTargetBox="[151,1435,184,1873]" captionTargetId="figure-20@16.[151,1435,181,1908]" captionTargetPageId="16" captionText="FIGURES 3743. Heptodonta nigrosericea (W. Horn, 1930): 37, 39, 4143—♂, Philippines, Mindanao, Mt. Apo (JWCW): 37—habitus, 39—head, 41—lateral body, 42—aedeagus in left lateral aspect, 43—aedeagus in right lateral aspect; 38, 40—♀, Philippines, Mindanao, Mt. Kintanlad (JWCW): 38—habitus, 40—head. Scales: 3741 = 1 mm, 4243 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578797" httpUri="https://zenodo.org/record/4578797/files/figure.png" pageId="18" pageNumber="19">Fig. 40</figureCitation>
), dark-brown. Mandibles dark-brown to black with a dorsolateral testaceous strip. Labial and maxillary palpi testaceous with dark-brown to black terminal palpomeres, pre-terminal maxillary palpomere in males sometimes in females always dark-brown. Antennae slender, extending back over the first quarter of the elytra, metallic black, male pedicel testaceous at basal tip, scape with a single apical seta, antennomeres 34 with few scattered setae, antennomeres 511 finely and evenly pubescent.
</paragraph>
<paragraph id="086136E0B82CFFCA49F1D6B5FB623CB9" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">
Thorax entirely glabrous. Pronotum silky black with a blue-yellow and/or bronze lustre, median line moderately pronounced, proportions variable, pronotal length (
<quantity id="CF269B05B82CFFCA4BFBD56AFCD93C20" box="[717,881,1537,1564]" metricMagnitude="-3" metricUnit="m" metricValue="2.605" metricValueMax="2.93" metricValueMin="2.28" pageId="18" pageNumber="19" unit="mm" value="2.605" valueMax="2.93" valueMin="2.28">2.282.93 mm</quantity>
) and width (
<quantity id="CF269B05B82CFFCA4ACCD56AFB373C20" box="[1018,1183,1537,1564]" metricMagnitude="-3" metricUnit="m" metricValue="2.505" metricValueMax="2.83" metricValueMin="2.18" pageId="18" pageNumber="19" unit="mm" value="2.505" valueMax="2.83" valueMin="2.18">2.182.83 mm</quantity>
) nearly identical, shallow transverse grooves, pronounced anterior and posterior sulci, anterior lobe wider than posterior, lateral margins of the median lobe moderately converging to the base. Episterna, sterna, and epimera violet-blue. Prosternum, posterior mesosternum, metasternum, and proepimeron with iridescent green to bluish-green lustre.
</paragraph>
<paragraph id="086136E0B82CFFCA49F1D5E5FDAD3D64" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">
Elytra elongate, length
<quantity id="CF269B05B82CFFCA48FBD5E5FDFE3C95" box="[461,598,1678,1705]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.4" metricValueMin="7.2" pageId="18" pageNumber="19" unit="mm" value="8.3" valueMax="9.4" valueMin="7.2">7.29.4 mm</quantity>
, slightly dilated laterally to almost parallel, maximal width posterior, silky black with a blue-yellow to bronze lustre, lateral margin and apex violet to bluish-green, juxtahumeral, discal, mediolateral, anteroapical, and apical impression considerably pronounced, impression on posterior declivity moderate or lacking, posterior gibbosity significant, basodiscal convexity moderate, blue punctures slightly decreasing in size from base to apex, punctures gently fused to transverse lines, converging to apex on anteroapical impression, basal and apical irregularly punctured.
</paragraph>
<paragraph id="086136E0B82CFFCA49F1D40AFD8A3DDA" blockId="18.[151,1437,375,2022]" pageId="18" pageNumber="19">Coxae dark-testaceous with iridescent green to bluish-green lustre, pro- and mesocoxae anteriorly, metacoxae laterally densely covered with long white setae, mesocoxae posterior with few white setae. Trochanters and femora testaceous. Femoral distal apices, tibiae, and tarsi metallic-black often with some gently violet lustre. Claws rufoustestaceous, basal slightly darkened.</paragraph>
<paragraph id="086136E0B82DFFCB49F1D3FCFDFD3AC5" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
Abdominal sternites glabrous (apart from few long setae at posterior margins, and some minute setae at posterolateral margin of sternite 5), dark-brown with faintly violet-blue to blue-green reflections, anal sternite with dark-testaceous apical margin (
<figureCitation id="90E52A65B82DFFCB48C2D3B4FDE03AC5" box="[500,584,223,249]" captionStart="FIGURES 3743" captionStartId="16.[151,264,1931,1955]" captionTargetBox="[151,1435,184,1873]" captionTargetId="figure-20@16.[151,1435,181,1908]" captionTargetPageId="16" captionText="FIGURES 3743. Heptodonta nigrosericea (W. Horn, 1930): 37, 39, 4143—♂, Philippines, Mindanao, Mt. Apo (JWCW): 37—habitus, 39—head, 41—lateral body, 42—aedeagus in left lateral aspect, 43—aedeagus in right lateral aspect; 38, 40—♀, Philippines, Mindanao, Mt. Kintanlad (JWCW): 38—habitus, 40—head. Scales: 3741 = 1 mm, 4243 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578797" httpUri="https://zenodo.org/record/4578797/files/figure.png" pageId="19" pageNumber="20">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="086136E0B82DFFCB49F1D268FE783B91" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
Aedeagus arched, length
<quantity id="CF269B05B82DFFCB48C2D269FD373B21" box="[500,671,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.615" metricValueMax="2.73" metricValueMin="2.5" pageId="19" pageNumber="20" unit="mm" value="2.615" valueMax="2.73" valueMin="2.5">2.502.73 mm</quantity>
, medioventrally convex, dorsally curved, but dorsomedian often straight, apex slightly bent ventrally, with round tip and convex dorsal aspect, aedeagal base rather short and comparatively moderately sloping. Inner sac in left lateral aspect medioventral with elongate vertical sclerite and median with short horizontal sclerite, and in right lateral aspect with a spiny dorsoapical sclerite protruding from the apical orifice (
<figureCitation id="90E52A65B82DFFCB480FD2F8FE6C3B91" box="[313,452,403,429]" captionStart="FIGURES 3743" captionStartId="16.[151,264,1931,1955]" captionTargetBox="[151,1435,184,1873]" captionTargetId="figure-20@16.[151,1435,181,1908]" captionTargetPageId="16" captionText="FIGURES 3743. Heptodonta nigrosericea (W. Horn, 1930): 37, 39, 4143—♂, Philippines, Mindanao, Mt. Apo (JWCW): 37—habitus, 39—head, 41—lateral body, 42—aedeagus in left lateral aspect, 43—aedeagus in right lateral aspect; 38, 40—♀, Philippines, Mindanao, Mt. Kintanlad (JWCW): 38—habitus, 40—head. Scales: 3741 = 1 mm, 4243 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578797" httpUri="https://zenodo.org/record/4578797/files/figure.png" pageId="19" pageNumber="20">Figs. 4243</figureCitation>
).
</paragraph>
<paragraph id="086136E0B82DFFCB49F1D2DDFA393BED" blockId="19.[151,1437,151,1005]" box="[199,1425,438,465]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D2DDFEE53BEC" bold="true" box="[199,333,438,464]" pageId="19" pageNumber="20">Variability.</emphasis>
Pronotal and aedeagal shape show some variation, without losing the species-specific characters.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82DFFCB49F1D2B1FBAC38B9" pageId="19" pageNumber="20" type="distribution">
<paragraph id="086136E0B82DFFCB49F1D2B1FCB13BCA" blockId="19.[151,1437,151,1005]" box="[199,793,474,502]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D2B1FEC93BC8" bold="true" box="[199,353,474,500]" pageId="19" pageNumber="20">Distribution.</emphasis>
Southern
<emphasis id="3AAAEAF2B82DFFCB48E2D2B1FD2F3BC8" bold="true" box="[468,647,474,500]" pageId="19" pageNumber="20">
<collectingCountry id="70C97670B82DFFCB48E2D2B1FD2F3BC8" box="[468,647,474,500]" name="Philippines" pageId="19" pageNumber="20">PHILIPPINES</collectingCountry>
</emphasis>
(Mindanao).
</paragraph>
<paragraph id="086136E0B82DFFCB49F1D295FBAC38B9" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D295FD873824" bold="true" box="[199,559,510,536]" pageId="19" pageNumber="20">
<collectingRegion id="CA1AF802B82DFFCB49F1D295FEEF3824" box="[199,327,510,536]" pageId="19" pageNumber="20">Additional</collectingRegion>
material examined.
</emphasis>
<materialsCitation id="B8B63CBDB82DFFCB4B02D294FA303825" ID-GBIF-Occurrence="3045076384" box="[564,1432,510,537]" collectingDate="X. 2010, I. 2011" collectionCode="JWCW" country="Philippines" location="Mt. Apo" pageId="19" pageNumber="20" specimenCount="69" stateProvince="Mindanao">
<specimenCount id="1ED8FD69B82DFFCB4B02D294FD643825" box="[564,716,511,537]" pageId="19" pageNumber="20" type="generic">69 specimens</specimenCount>
.
<emphasis id="3AAAEAF2B82DFFCB4BE3D295FBB23824" bold="true" box="[725,1050,510,537]" pageId="19" pageNumber="20">
<collectingCountry id="70C97670B82DFFCB4BE3D295FC203824" box="[725,904,510,536]" name="Philippines" pageId="19" pageNumber="20">PHILIPPINES</collectingCountry>
:
<collectingRegion id="CA1AF802B82DFFCB4AA3D295FBBC3824" box="[917,1044,510,536]" pageId="19" pageNumber="20">Mindanao</collectingRegion>
:
</emphasis>
<location id="0D01603BB82DFFCB4D28D294FBD43825" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB4D28D294FBD43825" box="[1054,1148,511,537]" country="Philippines" name="Mt. Apo" pageId="19" pageNumber="20" stateProvince="Mindanao">Mt.Apo</location>
,
<collectingDate id="6C24E9C8B82DFFCB4DB3D294FA823825" box="[1157,1322,511,537]" isEnumeration="true" pageId="19" pageNumber="20">
<date id="7C601020B82DFFCB4DB3D294FB733825" box="[1157,1243,511,537]" pageId="19" pageNumber="20" value="2010-10">X.2010</date>
,
<date id="7C601020B82DFFCB4DD2D294FA823825" box="[1252,1322,511,537]" pageId="19" pageNumber="20" value="2011-01">I.2011</date>
</collectingDate>
(
<collectionCode id="6ECFAE25B82DFFCB4C01D294FA383825" box="[1335,1424,511,537]" pageId="19" pageNumber="20">JWCW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB49A1D148FD383801" ID-GBIF-Occurrence="3045076485" box="[151,656,546,573]" collectingDate="1984-04" collectionCode="JWCW" country="Philippines" latitude="8.5" location="Gingoog" longLatPrecision="7813" longitude="125.0" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Mindanao">
<location id="0D01603BB82DFFCB49A1D148FF543801" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB49A1D148FF543801" box="[151,252,547,573]" country="Philippines" latitude="8.5" longLatPrecision="7813" longitude="125.0" name="Gingoog" pageId="19" pageNumber="20" stateProvince="Mindanao">Gingoog</location>
,
<geoCoordinate id="6DEA5027B82DFFCB483FD149FEE63801" box="[265,334,546,573]" degrees="8.5" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="5555" value="8.5">8.5°N</geoCoordinate>
,
<geoCoordinate id="6DEA5027B82DFFCB486DD149FE103801" box="[347,440,546,573]" degrees="125.0" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="5555" value="125.0">125.0°E</geoCoordinate>
,
<date id="7C601020B82DFFCB48F0D148FDB73801" box="[454,543,547,573]" pageId="19" pageNumber="20" value="1984-04">
<collectingDate id="6C24E9C8B82DFFCB48F0D148FDB73801" box="[454,543,547,573]" pageId="19" pageNumber="20" value="1984-04">IV.1984</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82DFFCB4B19D148FD203801" box="[559,648,547,573]" pageId="19" pageNumber="20">JWCW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB4BABD148FBB83801" ID-GBIF-Occurrence="3045076362" box="[669,1040,547,573]" collectingDate="2014-10" collectionCode="JWCW" country="Philippines" location="Mt. Kintaniad" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Mindanao">
<location id="0D01603BB82DFFCB4BABD148FC963801" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB4BABD148FC963801" box="[669,830,547,573]" country="Philippines" name="Mt. Kintaniad" pageId="19" pageNumber="20" stateProvince="Mindanao">Mt. Kintaniad</location>
,
<date id="7C601020B82DFFCB4A7DD148FC373801" box="[843,927,547,573]" pageId="19" pageNumber="20" value="2014-10">
<collectingDate id="6C24E9C8B82DFFCB4A7DD148FC373801" box="[843,927,547,573]" pageId="19" pageNumber="20" value="2014-10">X.2014</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82DFFCB4A99D148FBA03801" box="[943,1032,547,573]" pageId="19" pageNumber="20">JWCW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB4D2BD148FF5B385D" ID-GBIF-Occurrence="3045076422" collectingDate="2015-10" collectionCode="PSCH" country="Philippines" location="Kabanglasan" municipality="Bukidonon" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Mindanao">
<collectingMunicipality id="E805AC9AB82DFFCB4D2BD148FB343801" box="[1053,1180,547,573]" pageId="19" pageNumber="20">Bukidonon</collectingMunicipality>
,
<location id="0D01603BB82DFFCB4D9FD148FA953801" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB4D9FD148FA953801" box="[1193,1341,547,573]" country="Philippines" municipality="Bukidonon" name="Kabanglasan" pageId="19" pageNumber="20" stateProvince="Mindanao">Kabanglasan</location>
,
<date id="7C601020B82DFFCB4C7FD149FA343801" box="[1353,1436,546,573]" pageId="19" pageNumber="20" value="2015-10">
<collectingDate id="6C24E9C8B82DFFCB4C7FD149FA343801" box="[1353,1436,546,573]" pageId="19" pageNumber="20" value="2015-10">X.2015</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82DFFCB49A9D12CFF43385D" box="[159,235,583,609]" pageId="19" pageNumber="20">PSCH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB49C8D12CFCB5385D" ID-GBIF-Occurrence="3045076505" box="[254,797,583,609]" collectingDate="IV. 2017, VII. 2017" collectionCode="PSCH" country="Philippines" location="Wao" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Lanao del Sur">
<collectingRegion id="CA1AF802B82DFFCB49C8D12CFE0B385D" box="[254,419,583,609]" country="Philippines" name="Lanao del Sur" pageId="19" pageNumber="20">Lanao Del Sur</collectingRegion>
,
<location id="0D01603BB82DFFCB489BD12CFE4B385D" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB489BD12CFE4B385D" box="[429,483,583,609]" country="Philippines" name="Wao" pageId="19" pageNumber="20" stateProvince="Lanao del Sur">Wao</location>
,
<collectingDate id="6C24E9C8B82DFFCB48D8D12CFD12385D" box="[494,698,583,609]" isEnumeration="true" pageId="19" pageNumber="20">
<date id="7C601020B82DFFCB48D8D12CFDE1385D" box="[494,585,583,609]" pageId="19" pageNumber="20" value="2017-04">IV.2017</date>
,
<date id="7C601020B82DFFCB4B62D12CFD12385D" box="[596,698,583,609]" pageId="19" pageNumber="20" value="2017-07">VII.2017</date>
</collectingDate>
(
<collectionCode id="6ECFAE25B82DFFCB4BFFD12CFCBD385D" box="[713,789,583,609]" pageId="19" pageNumber="20">PSCH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB4A1ED12CFB71385D" ID-GBIF-Occurrence="3045076303" box="[808,1241,583,609]" collectingDate="2014-08" collectionCode="SGCH" country="Philippines" location="Intavas" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Bukidnon">
<collectingRegion id="CA1AF802B82DFFCB4A1ED12CFC31385D" box="[808,921,583,609]" country="Philippines" name="Bukidnon" pageId="19" pageNumber="20">Bukidnon</collectingRegion>
,
<location id="0D01603BB82DFFCB4A92D12CFC50385D" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB4A92D12CFC50385D" box="[932,1016,583,609]" country="Philippines" name="Intavas" pageId="19" pageNumber="20" stateProvince="Bukidnon">Intavas</location>
,
<date id="7C601020B82DFFCB4D34D12CFBD9385D" box="[1026,1137,583,609]" pageId="19" pageNumber="20" value="2014-08">
<collectingDate id="6C24E9C8B82DFFCB4D34D12CFBD9385D" box="[1026,1137,583,609]" pageId="19" pageNumber="20" value="2014-08">VIII.2014</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82DFFCB4DB6D12CFB79385D" box="[1152,1233,583,609]" pageId="19" pageNumber="20">SGCH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB4DD2D12CFA30385D" ID-GBIF-Occurrence="3045076511" box="[1252,1432,583,609]" collectionCode="MNB" country="Philippines" location="Surigao" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Mindanao">
<location id="0D01603BB82DFFCB4DD2D12CFA93385D" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB4DD2D12CFA93385D" box="[1252,1339,583,609]" country="Philippines" name="Surigao" pageId="19" pageNumber="20" stateProvince="Mindanao">Surigao</location>
(
<collectionCode id="6ECFAE25B82DFFCB4C7CD12CFA27385D" box="[1354,1423,583,609]" pageId="19" pageNumber="20">MNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB49A1D100FE1538B9" ID-GBIF-Occurrence="3045076429" box="[151,445,619,645]" collectingDate="1984-10" collectionCode="ZSM" country="Philippines" location="Bukidnon" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Bukidnon">
<collectingRegion id="CA1AF802B82DFFCB49A1D100FEA038B9" box="[151,264,619,645]" country="Philippines" name="Bukidnon" pageId="19" pageNumber="20">Bukidnon</collectingRegion>
,
<date id="7C601020B82DFFCB4822D100FECF38B9" box="[276,359,619,645]" pageId="19" pageNumber="20" value="1984-10">
<collectingDate id="6C24E9C8B82DFFCB4822D100FECF38B9" box="[276,359,619,645]" pageId="19" pageNumber="20" value="1984-10">X.1984</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82DFFCB4840D100FE1D38B9" box="[374,437,619,645]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="19" pageNumber="20" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B8B63CBDB82DFFCB48FFD100FBA838B9" ID-GBIF-Occurrence="3045076454" box="[457,1024,619,645]" collectingDate="26. V. 1932, 24. VI. 1932" collectionCode="SDEI" country="Philippines" location="Silipon" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Bukidnon">
<location id="0D01603BB82DFFCB48FFD100FDB538B9" LSID="urn:lsid:plazi:treatment:807787F6B82CFFCB49A1D3FCFCD639D1:0D01603BB82DFFCB48FFD100FDB538B9" box="[457,541,619,645]" country="Philippines" name="Silipon" pageId="19" pageNumber="20" stateProvince="Bukidnon">Silipon</location>
,
<collectingRegion id="CA1AF802B82DFFCB4B1ED100FD3138B9" box="[552,665,619,645]" country="Philippines" name="Bukidnon" pageId="19" pageNumber="20">Bukidnon</collectingRegion>
,
<collectingDate id="6C24E9C8B82DFFCB4B92D100FC0C38B9" box="[676,932,619,645]" isEnumeration="true" pageId="19" pageNumber="20">
<date id="7C601020B82DFFCB4B92D100FCB138B9" box="[676,793,619,645]" pageId="19" pageNumber="20" value="1932-05-26">26.V.1932</date>
,
<date id="7C601020B82DFFCB4A13D100FC0C38B9" box="[805,932,619,645]" pageId="19" pageNumber="20" value="1932-06-24">24.VI.1932</date>
</collectingDate>
(
<collectionCode id="6ECFAE25B82DFFCB4A82D100FC5038B9" box="[948,1016,619,645]" pageId="19" pageNumber="20">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82DFFCB49F1D1E5FE2238CD" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="086136E0B82DFFCB49F1D1E5FE2238CD" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D1E5FE8F3894" bold="true" box="[199,295,654,680]" pageId="19" pageNumber="20">Biology.</emphasis>
<bibRefCitation id="6C4F4B11B82DFFCB4819D1E4FDFB3896" author="Cabras, A. A. &amp; Wiesner, J." box="[303,595,655,682]" pageId="19" pageNumber="20" pagination="1 - 6" refId="ref37814" refString="Cabras, A. A. &amp; Wiesner, J. (2016) Tiger beetles (Coleoptera: Carabidae: Cicindelinae) of Mainit Hotspring and Mati Protected Landscape, Mindanao with notes on their ecology and threats. International Research Journal of Biological Sciences, 5 (9), 1 - 6." type="journal article" year="2016">Cabras &amp; Wiesner (2016)</bibRefCitation>
reported that
<taxonomicName id="CFDE4D63B82DFFCB4BC5D1E4FCD63895" authorityName="Wiesner" authorityYear="1980" box="[755,894,655,681]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lumawigi">
<emphasis id="3AAAEAF2B82DFFCB4BC5D1E4FCD63895" box="[755,894,655,681]" italics="true" pageId="19" pageNumber="20">H. lumawigi</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B82DFFCB4AB1D1E4FC583895" bold="true" box="[903,1008,655,681]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="21995789B82DFFCB4AB1D1E4FC583895" box="[903,1008,655,681]" pageId="19" pageNumber="20">syn. nov.</taxonomicNameLabel>
</emphasis>
occurs in secondary forest on shaded dark rocks near the river. Further,
<taxonomicName id="CFDE4D63B82DFFCB4B38D1D8FD1438F0" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[526,700,691,716]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B82DFFCB4B38D1D8FD1438F0" box="[526,700,691,716]" italics="true" pageId="19" pageNumber="20">H. nigrosericea</emphasis>
</taxonomicName>
seems to be sensitive to human disturbances, preferring areas with rich vegetation cover.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82DFFCB49F1D190FCD639D1" pageId="19" pageNumber="20" type="discussion">
<paragraph id="086136E0B82DFFCB49F1D190FCD639D1" blockId="19.[151,1437,151,1005]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D190FE933929" bold="true" box="[199,315,763,789]" pageId="19" pageNumber="20">Remarks.</emphasis>
Only recorded from Mindanao, with one uncertain exception (
<bibRefCitation id="6C4F4B11B82DFFCB4D2AD190FB353929" author="Horn, W." box="[1052,1181,763,789]" pageId="19" pageNumber="20" pagination="1 - 9" refId="ref39412" refString="Horn, W. (1930) Beitrage zur Kenntnis neuer und alter Cicindelinen des Indo-papuanischen Faunen-Gebietes. Wiener Entomologische Zeitung, 47 (1), 1 - 9." type="journal article" year="1930">Horn 1930</bibRefCitation>
;
<bibRefCitation id="6C4F4B11B82DFFCB4D9AD190FAE63929" author="Wiesner, J." box="[1196,1358,763,789]" pageId="19" pageNumber="20" pagination="119 - 127" refId="ref41195" refString="Wiesner, J. (1980) Beitrage zur Kenntnis der philippinischen Cicindelidae. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 119 - 127." type="journal article" year="1980">Wiesner 1980</bibRefCitation>
,
<bibRefCitation id="6C4F4B11B82DFFCB4C6AD190FA3F3929" author="Wiesner, J." box="[1372,1431,763,789]" pageId="19" pageNumber="20" refId="ref41340" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt, Checklist of the Tiger Beetles of the World, 27. Beitrag zur Kenntnis der Cicindelidae. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">1992</bibRefCitation>
;
<bibRefCitation id="6C4F4B11B82DFFCB49A1D074FE993905" author="Cassola, F." box="[151,305,799,825]" pageId="19" pageNumber="20" pagination="491 - 509" refId="ref37970" refString="Cassola, F. (2000) Studies on tiger beetles 102. The Cicindelidae collected by Roland A. Muller in the Philippine Islands, with description of three new species (Coleoptera: Cicindelidae). Zoologische Mededelingen, 73, 491 - 509." type="journal article" year="2000">Cassola 2000</bibRefCitation>
;
<bibRefCitation id="6C4F4B11B82DFFCB480AD074FDE43905" author="Cabras, A. A. &amp; Wiesner, J." box="[316,588,799,825]" pageId="19" pageNumber="20" pagination="1 - 6" refId="ref37814" refString="Cabras, A. A. &amp; Wiesner, J. (2016) Tiger beetles (Coleoptera: Carabidae: Cicindelinae) of Mainit Hotspring and Mati Protected Landscape, Mindanao with notes on their ecology and threats. International Research Journal of Biological Sciences, 5 (9), 1 - 6." type="journal article" year="2016">Cabras &amp; Wiesner 2016</bibRefCitation>
; Cabras
<emphasis id="3AAAEAF2B82DFFCB4B9DD074FD4B3905" box="[683,739,799,825]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2016). The record from Panay (
<bibRefCitation id="6C4F4B11B82DFFCB4D7FD074FB4F3905" author="Wiesner, J." box="[1097,1255,799,825]" pageId="19" pageNumber="20" pagination="119 - 127" refId="ref41195" refString="Wiesner, J. (1980) Beitrage zur Kenntnis der philippinischen Cicindelidae. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 119 - 127." type="journal article" year="1980">Wiesner 1980</bibRefCitation>
) is probably the result of a confusion: before his collection trip to Panay in 1978, Lumawig also collected tiger beetles at Mt. Apo on Mindanao in 1974. Since 24 further specimens of
<taxonomicName id="CFDE4D63B82DFFCB4BD9D00CFC3739BC" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[751,927,871,896]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B82DFFCB4BD9D00CFC3739BC" box="[751,927,871,896]" italics="true" pageId="19" pageNumber="20">H. nigrosericea</emphasis>
</taxonomicName>
were collected in 2010 and 2011 at Mt. Apo, this species seems to be quite common in that locality. As all additional material examined comes from Mindanao, and also the yet unclear record position of the
<typeStatus id="D7658842B82DFFCB4BA8D0C4FD6639F5" box="[670,718,943,969]" pageId="19" pageNumber="20">type</typeStatus>
specimens “Columbogan” is actually located on Mindanao (see
<bibRefCitation id="6C4F4B11B82DFFCB49A1D0B8FE9A39D1" author="Munroe, E." box="[151,306,978,1005]" pageId="19" pageNumber="20" pagination="234 - 235" refId="ref39854" refString="Munroe, E. (1958) Pycnarmon jaguaralis (Guenee) and an Allied Species (Lepidoptera: Pyralidae). The Canadian Entomologist, 90 (4), 234 - 235. https: // doi. org / 10.4039 / Ent 90234 - 4" type="journal article" year="1958">Munroe 1958</bibRefCitation>
),
<taxonomicName id="CFDE4D63B82DFFCB4870D0B8FE5D39D0" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[326,501,979,1004]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B82DFFCB4870D0B8FE5D39D0" box="[326,501,979,1004]" italics="true" pageId="19" pageNumber="20">H. nigrosericea</emphasis>
</taxonomicName>
is probably endemic to that island.
</paragraph>
</subSubSection>
</treatment>
</document>