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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.526.6037" ID-GBIF-Dataset="46cfb24b-675a-4bae-88a3-b55240ef8f8d" ID-PMC="PMC4607843" ID-Pensoft-Pub="1313-2970-526-19" ID-PubMed="26487823" ID-ZBK="648DED714CAD42E0A19DF2E7DB27EE05" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-526-19" ModsDocOrigin="ZooKeys 526" ModsDocTitle="Diagnostic survey of Malagasy Nesomyrmex species-groups and revision of hafahafa group species via morphology based cluster delimitation protocol" checkinTime="1451243915363" checkinUser="pensoft" docAuthor="Csosz, Sandor &amp; Fisher, Brian L." docDate="2015" docId="79B5FF699614CD89B02971B9F6C01EC7" docLanguage="en" docName="ZooKeys 526: 19-59" docOrigin="ZooKeys 526" docSource="http://dx.doi.org/10.3897/zookeys.526.6037" docTitle="Nesomyrmex spinosus Csosz &amp; Fisher, sp. n." docType="treatment" docUuid="D3643DB1-75EB-415A-9220-9F255A5FCB21" docUuidSource="ZooBank" docVersion="4" lastPageNumber="38" masterDocId="FF89FFBBD738BB54FFC1FF9EFFE7FF9C" masterDocTitle="Diagnostic survey of Malagasy Nesomyrmex species-groups and revision of hafahafa group species via morphology based cluster delimitation protocol" masterLastPageNumber="59" masterPageNumber="19" pageNumber="35" updateTime="1668160833139" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Diagnostic survey of Malagasy Nesomyrmex species-groups and revision of hafahafa group species via morphology based cluster delimitation protocol</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Csosz, Sandor</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>526</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>19</mods:start>
<mods:end>59</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.526.6037</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.526.6037</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-526-19</mods:identifier>
<mods:identifier type="ZBK">648DED714CAD42E0A19DF2E7DB27EE05</mods:identifier>
<mods:identifier type="ZooBank">648DED714CAD42E0A19DF2E7DB27EE05</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152062669" LSID="urn:lsid:zoobank.org:act:D3643DB1-75EB-415A-9220-9F255A5FCB21" httpUri="http://treatment.plazi.org/id/79B5FF699614CD89B02971B9F6C01EC7" lastPageId="19" lastPageNumber="38" pageId="16" pageNumber="35">
<subSubSection pageId="16" pageNumber="35" type="multiple">
<paragraph pageId="16" pageNumber="35">Taxon classification Animalia Hymenoptera Formicidae</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="35" type="nomenclature">
<paragraph pageId="16" pageNumber="35">
<taxonomicName LSID="http://zoobank.org/D3643DB1-75EB-415A-9220-9F255A5FCB21" authority="Csosz &amp; Fisher" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex spinosus" order="Hymenoptera" pageId="16" pageNumber="35" phylum="Arthropoda" rank="species" species="spinosus">
Nesomyrmex spinosus
<normalizedToken originalValue="Csősz">Csosz</normalizedToken>
&amp; Fisher
</taxonomicName>
<taxonomicNameLabel pageId="16" pageNumber="35">sp. n.</taxonomicNameLabel>
Figs 28-30, Table 4
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="36" type="etymology">
<paragraph pageId="17" pageNumber="36">
<pageBreakToken pageId="17" pageNumber="36" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="17" pageNumber="36">
Name
<normalizedToken originalValue="“spinosus”">&quot;spinosus&quot;</normalizedToken>
refers to the short, strong antero-dorsal spines on the petiolar node.
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36" type="type material">
<paragraph pageId="17" pageNumber="36">Type material investigated.</paragraph>
<paragraph pageId="17" pageNumber="36">
Holotype worker.CASENT0443515, BLF05489; MADGAGASCAR: Prov. Toliara,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Privé">Prive</normalizedToken>
Berenty,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAnjapolo">d'Anjapolo</normalizedToken>
, 21.4 km 325° NW Amboasary,
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-24.933332">24°56'S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="46.216667">46°13'E</geoCoordinate>
[-
<geoCoordinate direction="north" orientation="latitude" precision="1" value="24.92972">24.92972 N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="46.20967">46.20967 E</geoCoordinate>
], 65 m, 7.iii.2002 Fisher et al. (CASCASENT0443515);
</paragraph>
<paragraph lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36">
Paratypes. 24 workers and three males with the same label data with the holotype under CASENT codes: CASENT0443515, BLF05489, (2w, CAS);
<pageBreakToken pageId="18" pageNumber="37" start="start">CASENT</pageBreakToken>
0443516, BLF05489, (3w, CAS); CASENT0443517, BLF05489, (3w, CAS); CASENT0443518, BLF05489, (1w, CAS); CASENT0443519, BLF05489, (1w, CAS); CASENT0443520, BLF05489, (1w, CAS); CASENT0443521, BLF05489, (1w, CAS); CASENT0443522, BLF05489, (1w, CAS); CASENT0443523, BLF05489, (1w, CAS); CASENT0443524, BLF05489, (1w, CAS); CASENT0443525, BLF05489, (1w, CAS); CASENT0443526, BLF05489, (1w, CAS); CASENT0443527, BLF05489, (1w, CAS); CASENT0443530, BLF05489, (1w, CAS); CASENT0443531, BLF05489, (1w, CAS); CASENT0443532, BLF05489, (1w, CASCASENT0443532); CASENT0443533, BLF05489, (1w, CAS); CASENT0443534, BLF05489, (1w, CAS); CASENT0443535, BLF05489, (1w, CAS); CASENT0443536, BLF05489, (1m, CAS); CASENT0443537, BLF05489, (2m, CAS);
</paragraph>
<paragraph pageId="18" pageNumber="37">The list of 44 non-type individuals belonging to 26 nest samples of other material investigated is given in Table 1.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="37" type="diagnosis">
<paragraph pageId="18" pageNumber="37">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="37">In key.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="38" type="description of workers">
<paragraph pageId="19" pageNumber="38">
<pageBreakToken pageId="19" pageNumber="38" start="start">Description</pageBreakToken>
of workers.
</paragraph>
<paragraph pageId="19" pageNumber="38">
Body color: brown. Body color pattern: body concolorous, only clava darker. Absolute cephalic size: 1021 [935, 1121]
<normalizedToken originalValue="µm">µm</normalizedToken>
(n=46). Cephalic length vs. maximum width of head capsule (CL/CWb): 1.056 [0.980, 1.113]. Postocular distance vs. cephalic length (PoOc/CL): 0.374 [0.342, 0.393]. Postocular sides of cranium contour frontal view orientation: converging posteriorly. Postocular sides of cranium contour frontal view shape: broadly convex. Vertex contour line in frontal view shape: slightly concave. Vertex sculpture: main sculpture rugose, ground sculpture areolate. Gena contour line in frontal view shape: feebly convex. Genae contour from anterior view orientation: converging. Gena sculpture: rugo-reticulate with areolate ground sculpture. Concentric carinae laterally surrounding antennal foramen count: present. Eye length vs. absolute cephalic size (EL/CS): 0.239 [0.220, 0.265]. Frontal carina distance vs. absolute cephalic size (FRS/CS): 0.315 [0.291, 0.335]. Longitudinal carinae on median region of frons count: present. Longitudinal carinae on medial region of frons shape: forked. Smooth median region on frons count: absent. Antennomere count: 12. Scape length vs. absolute cephalic size (SL/CS): 0.880 [0.844, 0.919]. Facial area of the scape absolute setal angle: setae absent, pubescence only. Median clypeal notch count: present. Median clypeal notch depth vs. absolute cephalic size (Cdep/CS): 0.021 [0.015, 0.027]. Ground sculpture of submedian area of clypeus: smooth. Median carina of clypeus count: present. Lateral carinae of clypeus count: present. Median anatomical line of propodeal spine angle value to Weber length in lateral view: 65°. Spine length vs. absolute cephalic size (SPST/CS): 0.300 [0.258, 0.330]. Minimum spine distance vs. absolute cephalic size (SPBA/CS): 0.212 [0.184, 0.235]. Apical spine distance vs. absolute cephalic size (SPTI/CS): 0.307 [0.221, 0.361]. Propodeal spine shape: straight; slightly bent. Apical distance of pronotal spines vs. absolute cephalic size (PSTI/CS): 0.677 [0.624, 0.723]. Metanotal depression count: absent. Dorsal region of mesosoma sculpture: rugose with areolate ground sculpture. Lateral region of pronotum sculpture: areolate ground sculpture, superimposed by dispersed rugae. Mesopleuron sculpture: areolate ground sculpture, superimposed by dispersed rugae. Metapleuron sculpture: areolate ground sculpture, superimposed by dispersed rugae. Petiole width vs. absolute cephalic size (PEW/CS): 0.237 [0.206, 0.259]. Anterodorsal spines on petiolar node angle of deviation from each other: 60°. Apical distance of anterodorsal spines on petiolar node vs. absolute cephalic size (NSTI/CS): 0.216 [0.194, 0.276]. Frontal profile of petiolar node contour line in lateral view shape: straight. Dorso-caudal petiolar profile contour line in lateral view shape: convex. Dorsal region of petiole sculpture: ground sculpture areolate, main sculpture absent; ground sculpture areolate, main sculpture dispersed rugose. Postpetiole width vs. absolute cephalic size (PPW/CS): 0.491 [0.435, 0.529]. Dorsal region of postpetiole sculpture: ground sculpture areolate, main sculpture absent; ground sculpture areolate, main sculpture dispersed rugose.
</paragraph>
<caption pageId="19" pageNumber="38">
<paragraph pageId="19" pageNumber="38">
Figures 28-30.
<taxonomicName class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex spinosus" order="Hymenoptera" pageId="19" pageNumber="38" phylum="Arthropoda" rank="species" species="spinosus">Nesomyrmex spinosus</taxonomicName>
sp. n. paratype worker (CASENT0443532). Lateral view of the body (28), head of the holotype worker in full-face view (29), dorsal view of the body (30). Scale 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="19" pageNumber="38" type="distribution">
<paragraph pageId="19" pageNumber="38">Distribution.</paragraph>
<paragraph pageId="19" pageNumber="38">
This species is known to occur in small, highly isolated forests (
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Privé">Prive</normalizedToken>
Berenty,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAnjapolo">d'Anjapolo</normalizedToken>
and Parc National
<normalizedToken originalValue="dAndohahela">d'Andohahela</normalizedToken>
,
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAmbohibory">d'Ambohibory</normalizedToken>
) in the southern part of Madagascar (Fig. 13).
</paragraph>
</subSubSection>
</treatment>
</document>